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408:
is known from a partial skull that has bony characters placing it in the booby family
Sulidae. Unlike living boobies, it has a large bill with a convex upper mandible, reminiscent of a toucan. The remains were recovered from the west side of Sud Sacaco Level (4β5 Million year-old) of the Pisco
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1955:
1960:
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Stucchi, Marcelo; Urbina, Mario (2004). ""Ramphastosula" (Aves, Sulidae): A New Genus from the Early
Pliocene of the Pisco Formation, Peru".
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1323:
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289:
183:
148:
392:. The genus name is a combination of "Rhamphastos", "toucan" and "sula", an
1720:
1642:
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871:
782:
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653:
628:
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word for "fool" that has been used to describe boobies (Sulidae,
409:
Formation. This area was a sheltered beach with rock platforms.
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852:
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10.1671/0272-4634(2004)024[0974:RASANG]2.0.CO;2
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1602:
1587:
1570:
1553:
1536:
1519:
1502:
1478:
1455:
1423:
1413:
1403:
1393:
1383:
1353:
1343:
1037:
1707:
1652:
1601:
1469:
1439:
1367:
1334:
1119:
1099:
1079:
1049:
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894:
870:
805:
781:
714:
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388:, named for Peruvian vertebrate paleontologist
1915:
1243:
476:
8:
51:introducing citations to additional sources
424:
422:
1922:
1908:
1649:
1331:
1318:
1250:
1236:
1228:
876:
867:
849:
787:
778:
753:
633:
624:
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530:
504:
483:
469:
461:
211:
115:
41:Relevant discussion may be found on the
418:
7:
1876:
1874:
1180:(imperial cormorant, blue-eyed shag)
1894:. You can help Knowledge (XXG) by
431:Journal of Vertebrate Paleontology
14:
1878:
1855:
236:
34:relies largely or entirely on a
23:
1956:Pliocene birds of South America
1:
1961:Fossil taxa described in 2004
339:Stucchi & Urbina, 2004 (
579:Christmas Island frigatebird
1982:
1873:
384:and only named species is
1853:
1330:
1317:
1279:
879:
866:
848:
790:
777:
752:
636:
623:
605:
542:
529:
503:
332:
327:
233:Scientific classification
231:
219:
210:
118:
1274:and their extinct allies
1061:Double-crested cormorant
956:White-breasted cormorant
1004:(long-tailed cormorant)
569:Magnificent frigatebird
1966:Prehistoric bird stubs
1133:(Magellanic cormorant)
963:(Temminck's cormorant)
936:Little black cormorant
498:(Phalacrocoraciformes)
319:Stucchi & Urbina,
222:Ramphastosula aguirrei
124:Temporal range: early
1142:New Zealand king shag
997:Little pied cormorant
931:Black-faced cormorant
574:Ascension frigatebird
16:Extinct genus of bird
1111:Red-legged cormorant
1071:Flightless cormorant
1041:Spectacled cormorant
861:: Phalacrocoracidae)
765:: Anhingidae ·
516:: Fregatidae ·
376:discovered in early
47:improve this article
1066:Neotropic cormorant
1033:Red-faced cormorant
832:Australasian darter
736:Australasian gannet
612:Gannets and boobies
380:rocks of Peru. The
1209:South Georgia shag
1144:(rough-faced shag)
1023:Brandt's cormorant
961:Japanese cormorant
589:Lesser frigatebird
1903:
1902:
1868:
1867:
1864:
1863:
1851:
1850:
1703:
1702:
1471:Phalacrocoracidae
1225:
1224:
1221:
1220:
1217:
1216:
1184:Heard Island shag
1028:Pelagic cormorant
982:Crowned cormorant
906:Socotra cormorant
890:
889:
844:
843:
840:
839:
801:
800:
748:
747:
744:
743:
663:Blue-footed booby
647:
646:
601:
600:
597:
596:
584:Great frigatebird
553:
552:
350:
349:
344:
323:
112:
111:
97:
1973:
1924:
1917:
1910:
1888:prehistoric bird
1882:
1875:
1859:
1832:
1822:
1815:Prophalacrocorax
1812:
1795:
1778:
1768:
1758:
1748:
1738:
1728:
1718:
1655:
1650:
1604:
1589:
1572:
1555:
1538:
1521:
1504:
1480:
1457:
1425:
1415:
1405:
1395:
1385:
1355:
1345:
1332:
1319:
1252:
1245:
1238:
1229:
1137:Guanay cormorant
1039:
987:Little cormorant
941:Indian cormorant
877:
868:
850:
788:
779:
754:
683:Red-footed booby
634:
625:
607:
540:
531:
505:
485:
478:
471:
462:
455:
454:
426:
390:Gregorio Ramirez
372:bird known from
338:
318:
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241:
240:
215:
205:
135:
116:
107:
104:
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96:
62:"Rhamphastosula"
55:
27:
19:
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1326:
1313:
1275:
1256:
1226:
1213:
1115:
1095:
1075:
1045:
1007:
992:Pygmy cormorant
966:
951:Great cormorant
886:
862:
836:
822:Oriental darter
797:
773:
740:
726:Northern gannet
710:
692:
643:
619:
593:
549:
525:
499:
489:
459:
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428:
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420:
415:
402:) in general.
317:
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226:Pisco Formation
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40:
28:
17:
12:
11:
5:
1979:
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1968:
1963:
1958:
1953:
1948:
1946:Pliocene birds
1943:
1933:
1932:
1927:
1926:
1919:
1912:
1904:
1901:
1900:
1883:
1866:
1865:
1862:
1861:
1854:
1852:
1849:
1848:
1846:
1845:
1838:
1828:
1825:Rhamphastosula
1818:
1808:
1801:
1791:
1784:
1774:
1764:
1754:
1744:
1734:
1724:
1713:
1711:
1705:
1704:
1701:
1700:
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1690:
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1676:
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1659:
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1646:
1639:
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1610:
1608:
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1585:
1578:
1568:
1561:
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1544:
1534:
1527:
1517:
1510:
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1493:
1486:
1475:
1473:
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1466:
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1463:
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1443:
1437:
1436:
1434:
1433:
1421:
1411:
1401:
1391:
1381:
1373:
1371:
1365:
1364:
1362:
1361:
1351:
1340:
1338:
1336:incertae sedis
1328:
1327:
1322:
1315:
1314:
1312:
1311:
1305:
1299:
1293:
1287:
1280:
1277:
1276:
1257:
1255:
1254:
1247:
1240:
1232:
1223:
1222:
1219:
1218:
1215:
1214:
1212:
1211:
1206:
1204:Macquarie shag
1201:
1199:Kerguelen shag
1196:
1194:Antarctic shag
1191:
1186:
1181:
1175:
1170:
1165:
1160:
1155:
1150:
1145:
1139:
1134:
1127:
1125:
1117:
1116:
1114:
1113:
1107:
1105:
1097:
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1093:
1087:
1085:
1077:
1076:
1074:
1073:
1068:
1063:
1057:
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1046:
1044:
1043:
1035:
1030:
1025:
1019:
1017:
1009:
1008:
1006:
1005:
1002:Reed cormorant
999:
994:
989:
984:
978:
976:
968:
967:
965:
964:
958:
953:
948:
946:Cape cormorant
943:
938:
933:
928:
926:Pied cormorant
923:
918:
913:
911:Bank cormorant
908:
902:
900:
892:
891:
888:
887:
880:
874:
864:
863:
853:
846:
845:
842:
841:
838:
837:
835:
834:
829:
827:African darter
824:
819:
813:
811:
803:
802:
799:
798:
791:
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775:
774:
757:
750:
749:
746:
745:
742:
741:
739:
738:
733:
728:
722:
720:
712:
711:
709:
708:
706:Abbott's booby
702:
700:
694:
693:
691:
690:
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680:
675:
670:
668:Peruvian booby
665:
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631:
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473:
465:
457:
456:
437:(4): 974β978.
417:
416:
414:
411:
406:Rhamphastosula
354:Rhamphastosula
348:
347:
346:
345:
330:
329:
325:
324:
314:Rhamphastosula
307:
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137:
136:
123:
120:Rhamphastosula
110:
109:
45:. Please help
31:
29:
22:
15:
13:
10:
9:
6:
4:
3:
2:
1978:
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1951:Pliocene Peru
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1630:
1626:
1624:
1623:
1619:
1617:
1616:
1612:
1611:
1609:
1607:
1606:Plotopteridae
1600:
1594:
1593:
1586:
1584:
1583:
1579:
1577:
1576:
1569:
1567:
1566:
1562:
1560:
1559:
1552:
1550:
1549:
1548:Phalacrocorax
1545:
1543:
1542:
1535:
1533:
1532:
1528:
1526:
1525:
1518:
1516:
1515:
1511:
1509:
1508:
1501:
1499:
1498:
1494:
1492:
1491:
1487:
1485:
1484:
1477:
1476:
1474:
1472:
1468:
1462:
1461:
1454:
1452:
1451:
1447:
1446:
1444:
1442:
1438:
1431:
1430:
1422:
1420:
1419:
1412:
1410:
1409:
1402:
1400:
1399:
1392:
1390:
1389:
1382:
1380:
1379:
1375:
1374:
1372:
1370:
1366:
1360:
1359:
1352:
1350:
1349:
1342:
1341:
1339:
1337:
1333:
1329:
1325:
1320:
1316:
1310:
1306:
1304:
1300:
1298:
1294:
1292:
1288:
1286:
1282:
1281:
1278:
1273:
1269:
1265:
1261:
1253:
1248:
1246:
1241:
1239:
1234:
1233:
1230:
1210:
1207:
1205:
1202:
1200:
1197:
1195:
1192:
1190:
1187:
1185:
1182:
1179:
1178:Imperial shag
1176:
1174:
1171:
1169:
1168:Campbell shag
1166:
1164:
1163:Auckland shag
1161:
1159:
1156:
1154:
1151:
1149:
1146:
1143:
1140:
1138:
1135:
1132:
1129:
1128:
1126:
1124:
1123:
1118:
1112:
1109:
1108:
1106:
1104:
1103:
1098:
1092:
1091:European shag
1089:
1088:
1086:
1084:
1083:
1078:
1072:
1069:
1067:
1064:
1062:
1059:
1058:
1056:
1054:
1053:
1048:
1042:
1036:
1034:
1031:
1029:
1026:
1024:
1021:
1020:
1018:
1016:
1015:
1010:
1003:
1000:
998:
995:
993:
990:
988:
985:
983:
980:
979:
977:
975:
974:
969:
962:
959:
957:
954:
952:
949:
947:
944:
942:
939:
937:
934:
932:
929:
927:
924:
922:
919:
917:
914:
912:
909:
907:
904:
903:
901:
899:
898:
897:Phalacrocorax
893:
885:
884:
878:
875:
873:
869:
865:
860:
856:
851:
847:
833:
830:
828:
825:
823:
820:
818:
815:
814:
812:
810:
809:
804:
796:
795:
789:
786:
784:
780:
776:
771:
768:
764:
760:
755:
751:
737:
734:
732:
729:
727:
724:
723:
721:
719:
718:
713:
707:
704:
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689:
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64: β
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58:Find sources:
52:
48:
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38:
37:
36:single source
32:This article
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1680:Klallamornis
1678:
1673:Hokkaidornis
1671:
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1615:Empeirodytes
1613:
1592:Valenticarbo
1590:
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1565:Poikilocarbo
1563:
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1539:
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1507:Limicorallus
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1460:Limnofregata
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1301:Infraclass:
1260:frigatebirds
1158:Chatham shag
1148:Foveaux shag
1120:
1102:Poikilocarbo
1100:
1080:
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1012:
971:
921:Spotted shag
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673:Masked booby
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510:Frigatebirds
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220:Holotype of
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103:October 2021
100:
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1835:Sarmatosula
1731:Empheresula
1687:Olympidytes
1629:Plotopterum
1531:Nannopterum
1408:Macranhinga
1358:Protoplotus
1348:Mangystania
1189:Crozet shag
1173:Bounty shag
1052:Nannopterum
731:Cape gannet
688:Brown booby
678:Nazca booby
386:R. ramirezi
336:R. ramirezi
224:, from the
1935:Categories
1741:Enkurosula
1666:Copepteryx
1657:Tonsalinae
1514:Microcarbo
1497:Leucocarbo
1483:Borvocarbo
1441:Fregatidae
1418:Meganhinga
1388:Giganhinga
1369:Anhingidae
1324:Suliformes
1309:Suliformes
1303:Neognathae
1268:cormorants
1258:Genera of
1153:Otago shag
1122:Leucocarbo
973:Microcarbo
855:Cormorants
618:: Sulidae)
496:Suliformes
413:References
290:Suliformes
73:newspapers
1798:Paleosula
1771:Microsula
1721:Bimbisula
1643:Stenornis
1575:Praecarbo
1541:Nectornis
1524:Nambashag
1429:Paranavis
1398:Liptornis
1283:Kingdom:
1131:Rock shag
916:Pitt shag
394:Icelandic
361:") is an
357:("toucan
256:Kingdom:
250:Eukaryota
131:5β4
43:talk page
1805:Papasula
1622:Phocavis
1558:Piscator
1291:Chordata
1289:Phylum:
1285:Animalia
1272:anhingas
698:Papasula
378:Pliocene
328:Species
296:Family:
270:Chordata
266:Phylum:
260:Animalia
246:Domain:
126:Pliocene
1941:Sulidae
1781:Miosula
1751:Eostega
1709:Sulidae
1694:Tonsala
1490:Gulosus
1450:Fregata
1378:Anhinga
1307:Clade:
1295:Class:
1264:boobies
1082:Gulosus
883:Species
817:Anhinga
808:Anhinga
794:Species
770:Anhinga
759:Darters
640:Species
560:Fregata
546:Species
522:Fregata
451:4524793
374:fossils
363:extinct
306:Genus:
300:Sulidae
286:Order:
276:Class:
200:↓
87:scholar
1636:Stemec
859:family
763:family
616:family
514:family
449:
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1886:This
1788:Morus
1582:Urile
1014:Urile
872:Genus
783:Genus
767:genus
717:Morus
629:Genus
535:Genus
518:genus
492:Order
447:JSTOR
370:sulid
366:genus
359:booby
94:JSTOR
80:books
1892:stub
1842:Sula
1297:Aves
654:Sula
399:Sula
382:type
341:type
321:2004
280:Aves
139:Preκ
66:news
439:doi
368:of
49:by
1937::
1270:,
1266:,
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520::
494::
445:.
435:24
433:.
421:^
189:Pg
133:Ma
1923:e
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1432:"
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310:β
194:N
184:K
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169:P
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159:D
154:S
149:O
144:κ
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91:Β·
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