132:
943:
1246:
1378:
111:
321:
were small, characterized by scattered tubercules and knobs on the outer surface of the exoskeleton. Their first two (or possibly three) pairs of walking legs had spines; the last two pairs were long and powerful, without spines. The prosoma (head) was subtrapezoidal, with arcuate compound eyes on
487:
455:
Dunlop, J. A., Penney, D. & Jekel, D. 2015. A summary list of fossil spiders and their relatives. In World Spider
Catalog. Natural History Museum Bern, online at
1324:
1419:
1311:
1458:
1022:
1012:
534:
488:"The eurypterid Adelophthalmus sievertsi (Chelicerata: Eurypterida) from the Lower Devonian (Emsian) Klerf Formation of Willwerath, Germany"
1448:
974:
989:
1233:
1017:
959:
1453:
1007:
997:
951:
1443:
1412:
969:
964:
1002:
979:
131:
1463:
635:
527:
1405:
942:
1438:
1264:
1350:
1249:
520:
333:
contains three valid species, with other named species now seen as invalid or part of other genera.
432:
126:
1355:
1337:
1329:
838:
771:
763:
1342:
1302:
1178:
892:
884:
874:
866:
846:
817:
733:
682:
499:
460:
1207:
1053:
910:
851:
807:
710:
697:
232:
1389:
1123:
1081:
1039:
902:
829:
789:
781:
705:
346:
303:
30:
1432:
1130:
1102:
1095:
1067:
1046:
799:
738:
728:
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674:
664:
656:
612:
576:
503:
318:
212:
199:
68:
1088:
920:
856:
812:
687:
241:
110:
1377:
1296:
1185:
1162:
1060:
754:
647:
585:
567:
173:
43:
1287:
1385:
1151:
1137:
1116:
1109:
1074:
915:
603:
594:
543:
322:
parallel axes. The male genital appendages were short with two distal spines.
295:
186:
88:
53:
1228:
1212:
1202:
561:
163:
143:
93:
37:
1281:
1171:
1144:
415:
396:
376:
361:
83:
78:
63:
58:
48:
1316:
474:
Treatise on
Invertebrate Paleontology, Part P Arthropoda 2, Chelicerata
307:
98:
73:
555:
153:
1258:
1262:
516:
512:
461:
http://www.wsc.nmbe.ch/resources/fossils/Fossils16.0.pdf
1393:
456:
1271:
1221:
1195:
1161:
1031:
988:
950:
901:
883:
865:
837:
826:
798:
780:
762:
751:
719:
696:
673:
655:
644:
628:
621:
383:Invalid or reassigned species are listed below:
1413:
528:
8:
1420:
1406:
1259:
834:
759:
652:
625:
535:
521:
513:
109:
20:
443:
302:have been recovered from deposits of
7:
1374:
1372:
451:
449:
447:
16:Extinct genus of Devonian organisms
14:
1376:
1245:
1244:
941:
504:10.1111/j.1475-4983.2005.00528.x
486:Poschmann, Markus (2006-01-19).
472:Størmer, L. (1955) Merostomata.
418:), reclassified as a species of
130:
1234:Timeline of eurypterid research
360:- Alken an der Mosel, Germany (
1:
1459:Fossil taxa described in 1936
354:Rhenopterus macrotuberculatus
1392:. You can help Knowledge by
1480:
1449:Prehistoric life of Europe
1371:
294:is an extinct prehistoric
1242:
939:
636:List of eurypterid genera
550:
265:
260:
247:
240:
127:Scientific classification
125:
117:
108:
23:
369:Rhenopterus tuberculatus
408:"Rhenopterus" sievertsi
1388:-related article is a
1454:Eurypterids of Europe
1351:Paleobiology Database
1444:Devonian eurypterids
269:R. macrotuberculatus
433:List of eurypterids
389:"Rhenopterus" latus
339:Rhenopterus diensti
250:Rhenopterus diensti
120:Rhenopterus diensti
764:Onychopterelloidea
457:http://wsc.nmbe.ch
118:Reconstruction of
1401:
1400:
1366:
1365:
1338:Open Tree of Life
1265:Taxon identifiers
1256:
1255:
937:
936:
933:
932:
929:
928:
885:Adelophthalmoidea
867:Waeringopteroidea
839:Carcinosomatoidea
772:Onychopterellidae
747:
746:
413:
394:
374:
359:
344:
287:
286:
281:
273:
236:
36:409.1–402.5
1471:
1464:Eurypterid stubs
1422:
1415:
1408:
1380:
1373:
1359:
1358:
1346:
1345:
1333:
1332:
1320:
1319:
1307:
1306:
1305:
1292:
1291:
1290:
1260:
1248:
1247:
1222:Related articles
1179:Merostomichnites
945:
893:Adelophthalmidae
875:Waeringopteridae
847:Carcinosomatidae
835:
818:Strobilopteridae
760:
734:Hibbertopteridae
683:Parastylonuridae
653:
626:
537:
530:
523:
514:
508:
507:
483:
477:
470:
464:
453:
411:
392:
372:
357:
342:
279:
271:
231:
224:
211:
198:
185:
135:
134:
113:
103:
40:
29:Temporal range:
21:
1479:
1478:
1474:
1473:
1472:
1470:
1469:
1468:
1429:
1428:
1427:
1426:
1369:
1367:
1362:
1354:
1349:
1341:
1336:
1328:
1323:
1315:
1310:
1301:
1300:
1295:
1286:
1285:
1280:
1267:
1257:
1252:
1238:
1217:
1208:Chasmataspidida
1191:
1157:
1054:Campylocephalus
1027:
984:
946:
925:
911:Hughmilleriidae
897:
879:
861:
852:Megalograptidae
828:
822:
808:Dolichopteridae
794:
782:Moselopteroidea
776:
753:
743:
729:Drepanopteridae
715:
711:Hardieopteridae
698:Kokomopteroidea
692:
669:
646:
640:
617:
546:
541:
511:
485:
484:
480:
471:
467:
459:, version 16.0
454:
445:
441:
429:
328:
316:
277:R. tuberculatus
256:
253:
230:
222:
209:
196:
183:
129:
104:
102:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
27:
17:
12:
11:
5:
1477:
1475:
1467:
1466:
1461:
1456:
1451:
1446:
1441:
1439:Rhenopteroidea
1431:
1430:
1425:
1424:
1417:
1410:
1402:
1399:
1398:
1381:
1364:
1363:
1361:
1360:
1347:
1334:
1321:
1308:
1293:
1277:
1275:
1269:
1268:
1263:
1254:
1253:
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1237:
1236:
1231:
1225:
1223:
1219:
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1216:
1215:
1210:
1205:
1199:
1197:
1196:Related groups
1193:
1192:
1190:
1189:
1182:
1175:
1167:
1165:
1159:
1158:
1156:
1155:
1148:
1141:
1134:
1127:
1124:Onychopterella
1120:
1113:
1106:
1099:
1092:
1085:
1082:Hibbertopterus
1078:
1071:
1064:
1057:
1050:
1043:
1040:Adelophthalmus
1035:
1033:
1032:Notable genera
1029:
1028:
1026:
1025:
1020:
1015:
1010:
1005:
1000:
994:
992:
986:
985:
983:
982:
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962:
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948:
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938:
935:
934:
931:
930:
927:
926:
924:
923:
918:
913:
907:
905:
903:Pterygotioidea
899:
898:
896:
895:
889:
887:
881:
880:
878:
877:
871:
869:
863:
862:
860:
859:
854:
849:
843:
841:
832:
830:Diploperculata
824:
823:
821:
820:
815:
810:
804:
802:
796:
795:
793:
792:
790:Moselopteridae
786:
784:
778:
777:
775:
774:
768:
766:
757:
749:
748:
745:
744:
742:
741:
736:
731:
725:
723:
717:
716:
714:
713:
708:
706:Kokomopteridae
702:
700:
694:
693:
691:
690:
685:
679:
677:
671:
670:
668:
667:
661:
659:
657:Rhenopteroidea
650:
642:
641:
639:
638:
632:
630:
623:
619:
618:
616:
615:
606:
597:
588:
579:
570:
564:
558:
551:
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542:
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539:
532:
525:
517:
510:
509:
478:
465:
442:
440:
437:
436:
435:
428:
425:
424:
423:
420:Adelophthalmus
404:
399:), synonym of
381:
380:
365:
350:
347:Lower Devonian
327:
324:
315:
312:
304:Lower Devonian
285:
284:
283:
282:
274:
263:
262:
261:Other species
258:
257:
254:
245:
244:
238:
237:
220:
216:
215:
207:
203:
202:
200:Rhenopteroidea
194:
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181:
177:
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171:
167:
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157:
156:
151:
147:
146:
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137:
136:
123:
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105:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
31:Lower Devonian
28:
15:
13:
10:
9:
6:
4:
3:
2:
1476:
1465:
1462:
1460:
1457:
1455:
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1442:
1440:
1437:
1436:
1434:
1423:
1418:
1416:
1411:
1409:
1404:
1403:
1397:
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1382:
1379:
1375:
1370:
1357:
1352:
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1335:
1331:
1326:
1322:
1318:
1313:
1309:
1304:
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1274:
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1251:
1241:
1235:
1232:
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1227:
1226:
1224:
1220:
1214:
1211:
1209:
1206:
1204:
1201:
1200:
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1194:
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1187:
1183:
1181:
1180:
1176:
1174:
1173:
1169:
1168:
1166:
1164:
1160:
1154:
1153:
1149:
1147:
1146:
1142:
1140:
1139:
1135:
1133:
1132:
1131:Pentecopterus
1128:
1126:
1125:
1121:
1119:
1118:
1114:
1112:
1111:
1107:
1105:
1104:
1103:Megalograptus
1100:
1098:
1097:
1096:Jaekelopterus
1093:
1091:
1090:
1086:
1084:
1083:
1079:
1077:
1076:
1072:
1070:
1069:
1068:Drepanopterus
1065:
1063:
1062:
1058:
1056:
1055:
1051:
1049:
1048:
1047:Brachyopterus
1044:
1042:
1041:
1037:
1036:
1034:
1030:
1024:
1023:South America
1021:
1019:
1016:
1014:
1013:North America
1011:
1009:
1006:
1004:
1001:
999:
996:
995:
993:
991:
987:
981:
978:
976:
975:Carboniferous
973:
971:
968:
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957:
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952:Geochronology
949:
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904:
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890:
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882:
876:
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868:
864:
858:
855:
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850:
848:
845:
844:
842:
840:
836:
833:
831:
825:
819:
816:
814:
811:
809:
806:
805:
803:
801:
800:Eurypteroidea
797:
791:
788:
787:
785:
783:
779:
773:
770:
769:
767:
765:
761:
758:
756:
750:
740:
739:Mycteroptidae
737:
735:
732:
730:
727:
726:
724:
722:
721:Mycteropoidea
718:
712:
709:
707:
704:
703:
701:
699:
695:
689:
686:
684:
681:
680:
678:
676:
675:Stylonuroidea
672:
666:
665:Rhenopteridae
663:
662:
660:
658:
654:
651:
649:
643:
637:
634:
633:
631:
627:
624:
620:
614:
613:Sclerophorata
610:
607:
605:
601:
598:
596:
592:
589:
587:
583:
580:
578:
577:Euchelicerata
574:
571:
569:
565:
563:
559:
557:
553:
552:
549:
545:
538:
533:
531:
526:
524:
519:
518:
515:
505:
501:
497:
493:
492:Palaeontology
489:
482:
479:
475:
469:
466:
462:
458:
452:
450:
448:
444:
438:
434:
431:
430:
426:
421:
417:
412:Størmer, 1969
410:
409:
405:
402:
398:
393:Størmer, 1936
391:
390:
386:
385:
384:
378:
373:Størmer, 1936
371:
370:
366:
363:
358:Størmer, 1974
356:
355:
351:
348:
343:Størmer, 1936
341:
340:
336:
335:
334:
332:
325:
323:
320:
313:
311:
309:
305:
301:
298:. Fossils of
297:
293:
292:
280:Størmer, 1936
278:
275:
272:Størmer, 1974
270:
267:
266:
264:
259:
255:Størmer, 1936
252:
251:
246:
243:
239:
234:
229:
228:
221:
218:
217:
214:
213:Rhenopteridae
208:
205:
204:
201:
195:
193:Superfamily:
192:
191:
188:
182:
179:
178:
175:
172:
169:
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162:
159:
158:
155:
152:
149:
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139:
138:
133:
128:
124:
121:
116:
112:
107:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
1394:expanding it
1383:
1368:
1272:
1184:
1177:
1170:
1150:
1143:
1136:
1129:
1122:
1115:
1108:
1101:
1094:
1089:Hughmilleria
1087:
1080:
1073:
1066:
1059:
1052:
1045:
1038:
921:Pterygotidae
857:Mixopteridae
813:Eurypteridae
688:Stylonuridae
608:
599:
590:
581:
572:
495:
491:
481:
473:
468:
419:
407:
406:
400:
388:
387:
382:
368:
367:
353:
352:
338:
337:
330:
329:
319:Rhenopterids
317:
299:
290:
289:
288:
276:
268:
249:
248:
242:Type species
226:
225:
119:
24:
18:
1303:Rhenopterus
1297:Wikispecies
1273:Rhenopterus
1186:Palmichnium
1163:Ichnogenera
1061:Carcinosoma
755:Eurypterina
648:Stylonurina
586:Prosomapoda
568:Chelicerata
566:Subphylum:
544:Eurypterids
414:- Germany (
395:- Germany (
375:- Germany (
345:- Germany (
331:Rhenopterus
314:Description
300:Rhenopterus
291:Rhenopterus
227:Rhenopterus
187:Eurypterida
174:Chelicerata
170:Subphylum:
25:Rhenopterus
1433:Categories
1386:eurypterid
1152:Stylonurus
1138:Pterygotus
1117:Mixopterus
1110:Megarachne
1075:Eurypterus
960:Ordovician
916:Slimonidae
827:Infraorder
604:Dekatriata
595:Planaterga
562:Arthropoda
439:References
401:R. diensti
296:eurypterid
164:Arthropoda
1229:Metastoma
1213:Xiphosura
1203:Arachnida
990:Geography
554:Kingdom:
498:: 67–82.
150:Kingdom:
144:Eukaryota
1288:Q7320403
1282:Wikidata
1250:Category
1172:Arcuites
1145:Slimonia
970:Devonian
965:Silurian
752:Suborder
645:Suborder
622:Taxonomy
560:Phylum:
556:Animalia
427:See also
416:Devonian
397:Devonian
377:Devonian
362:Devonian
206:Family:
160:Phylum:
154:Animalia
140:Domain:
1343:3581750
1330:1026727
1317:4650108
1018:Oceania
980:Permian
326:Species
308:Germany
306:age in
233:Størmer
219:Genus:
180:Order:
1008:Europe
998:Africa
629:Genera
476:, P39.
463:(PDF).
235:, 1936
1384:This
1356:18993
1325:IRMNG
609:Clade
600:Clade
591:Clade
582:Clade
573:Clade
1390:stub
1312:GBIF
1003:Asia
44:Preęž’
500:doi
1435::
1353::
1340::
1327::
1314::
1299::
1284::
611::
602::
593::
584::
575::
496:49
494:.
490:.
446:^
310:.
94:Pg
38:Ma
33:,
1421:e
1414:t
1407:v
1396:.
536:e
529:t
522:v
506:.
502::
422:.
403:.
379:)
364:)
349:)
223:â€
210:â€
197:â€
184:â€
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89:K
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79:T
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69:C
64:D
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54:O
49:ęž’
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