187:
66:
41:
3488:
259:, derived from Latin, translates as 'little hammer' and the diminutive form of Marc Argeloo's name, a dedication to his collections of bats for the Amsterdam Zoological Museum. Two subspecies were recognised by
3548:
299:, with recorded forearm measurements of 51.0 to 54.9 millimetres, and the greatest length of the skull in a range of 25.2–27.2 mm for both subspecies.
3529:
3369:
3027:
3395:
3563:
3138:
3003:
2994:
468:
2920:
416:
3558:
1332:
1323:
2852:
1377:
3568:
2938:
2888:
1422:
930:
566:
2870:
1341:
373:"Notes on distribution and taxonomy of australasian bats. I. Pteropodinae and Nyctimeninae (Mammalia, Megachiroptera, Pteropodidae)"
3129:
3421:
2929:
2834:
2825:
2358:
1350:
948:
602:
439:
3578:
2535:
1732:
1431:
1395:
1368:
665:
620:
3439:
2861:
2526:
1413:
1404:
3400:
2081:
1932:
1896:
1783:
1386:
1088:
939:
593:
3522:
2843:
2816:
1905:
1022:
894:
647:
638:
65:
3184:
3583:
3434:
3230:
1492:
1359:
707:
629:
611:
575:
260:
989:
398:
2879:
2430:
2162:
2027:
1555:
980:
683:
584:
461:
3147:
2695:
1614:
3588:
3573:
2484:
2234:
1923:
1914:
1824:
1455:
1097:
1079:
831:
807:
2439:
2144:
2108:
1046:
903:
3334:
3515:
3221:
2331:
2063:
1991:
1872:
1848:
1774:
1664:
1573:
1564:
1522:
1210:
1177:
3553:
3296:
2668:
2663:
2340:
2243:
2126:
2018:
1531:
1186:
862:
841:
674:
2412:
2394:
2054:
1655:
1605:
1244:
656:
454:
186:
160:
3263:
1291:
912:
3356:
3239:
2740:
2722:
2421:
2385:
2270:
2198:
2189:
1964:
1815:
372:
277:
340:
2749:
2611:
2457:
2367:
2252:
2036:
1714:
252:
215:
30:
2686:
2593:
2448:
2376:
2307:
2180:
2090:
2000:
1973:
1646:
1070:
60:
3460:
3343:
3097:
3088:
2962:
2677:
2466:
2322:
2297:
2279:
2225:
2216:
2207:
2171:
2072:
2045:
1741:
1723:
1269:
549:
422:
412:
255:, whose retirement was announced when the new species was discovered. The subspecific epithet
231:
3499:
3465:
3413:
3064:
2704:
2650:
2502:
2493:
2475:
2403:
2349:
2313:
2261:
2135:
2099:
1982:
1750:
921:
774:
406:
335:
251:, derived from the Latin for 'steadfast', was intended to describe the conservation work of
223:
3447:
2713:
2602:
2288:
2117:
1798:
1464:
1446:
1234:
1145:
1121:
998:
783:
446:
3202:
2153:
2009:
514:
317:
3542:
3408:
3254:
3079:
3055:
2977:
2777:
2735:
1013:
822:
698:
402:
394:
326:
137:
50:
45:
3348:
3212:
3045:
3018:
2953:
2789:
2517:
1692:
1629:
1546:
1505:
798:
744:
536:
3426:
3382:
3328:
3112:
2799:
2563:
1160:
1112:
963:
3319:
2576:
1887:
1765:
1588:
1225:
877:
520:
3495:
3165:
2626:
1863:
1282:
1201:
1136:
1061:
757:
426:
219:
77:
3452:
3313:
3175:
1947:
1839:
1705:
1306:
1037:
853:
508:
502:
496:
490:
176:
97:
3374:
2903:
478:
211:
127:
3487:
3387:
2645:
107:
87:
3290:
438:"Mammals described from the Papuan region between 1995 and 2010."
411:(3rd ed.). Johns Hopkins University Press. pp. 312–529.
408:
Mammal
Species of the World: A Taxonomic and Geographic Reference
3361:
281:(Lesser tube-nosed fruit bat), the type, first proposed in 1942.
3294:
2774:
2560:
1689:
1489:
1266:
741:
533:
450:
229:
The species was first proposed as a subgeneric arrangement of
117:
3503:
3303:
3252:
3210:
3201:
3173:
3164:
3110:
3077:
3053:
3044:
3016:
2975:
2951:
2901:
2797:
2788:
2624:
2574:
2515:
1945:
1885:
1861:
1837:
1796:
1763:
1703:
1627:
1586:
1544:
1503:
1444:
1304:
1280:
1223:
1199:
1158:
1134:
1110:
1059:
1035:
1011:
961:
875:
851:
820:
796:
755:
696:
547:
341:10.2305/IUCN.UK.2021-3.RLTS.T136836A22041117.en
289:Bergmans, 2001 (Steadfast tube-nosed fruit bat)
366:
364:
362:
360:
358:
3523:
462:
8:
3530:
3516:
3291:
3207:
3170:
3050:
2794:
2785:
2771:
2571:
2557:
1700:
1686:
1500:
1486:
1277:
1263:
752:
738:
544:
530:
469:
455:
447:
185:
39:
20:
339:
3139:East African little collared fruit bat (
3028:Veldkamp's dwarf epauletted fruit bat (
308:
2995:Hayman's dwarf epauletted fruit bat (
2853:Lesser Angolan epauletted fruit bat (
7:
3484:
3482:
3004:Peters' dwarf epauletted fruit bat (
2921:Buettikofer's epauletted fruit bat (
1423:New Britain naked-backed fruit bat (
931:Nusatenggara short-nosed fruit bat (
567:Broad-striped tube-nosed fruit bat (
214:in the family Pteropodidae found in
3549:IUCN Red List least concern species
2871:East African epauletted fruit bat (
1342:Philippine naked-backed fruit bat (
1333:Beaufort's naked-backed fruit bat (
1324:Andersen's naked-backed fruit bat (
327:IUCN Red List of Threatened Species
3502:. You can help Knowledge (XXG) by
2359:Little golden-mantled flying fox (
1378:Solomon's naked-backed fruit bat (
1351:Halmahera naked-backed fruit bat (
949:Indonesian short-nosed fruit bat (
603:Round-eared tube-nosed fruit bat (
14:
2939:Franquet's epauletted fruit bat (
2889:Wahlberg's epauletted fruit bat (
2536:Sulawesi stripe-faced fruit bat (
1733:Giant golden-crowned flying fox (
1432:Greenish naked-backed fruit bat (
1396:Moluccan naked-backed fruit bat (
1369:Sulawesi naked-backed fruit bat (
666:Philippine tube-nosed fruit bat (
3486:
2862:Ethiopian epauletted fruit bat (
2527:Mindoro stripe-faced fruit bat (
1414:Western naked-backed fruit bat (
1405:Panniet naked-backed fruit bat (
717:Steadfast tube-nosed fruit bat (
64:
2835:Peters's epauletted fruit bat (
2826:Ansell's epauletted fruit bat (
2082:New Ireland masked flying fox (
1933:New Georgian monkey-faced Bat (
1897:Bougainville monkey-faced Bat (
1784:Small white-winged flying fox (
1387:Lesser naked-backed fruit bat (
940:Greater short-nosed fruit bat (
594:Mountain tube-nosed fruit bat (
24:Steadfast tube-nosed fruit bat
2844:Gambian epauletted fruit bat (
2817:Angolan epauletted fruit bat (
1906:Guadalcanal monkey-faced Bat (
895:Lesser short-nosed fruit bat (
648:Demonic tube-nosed fruit bat (
639:Malaita tube-nosed fruit bat (
621:Keast's tube-nosed fruit bat (
202:steadfast tube-nosed fruit bat
1:
3564:Mammals of Western New Guinea
3130:São Tomé collared fruit bat (
1360:Biak naked-backed fruit bat (
708:Lesser tube-nosed fruit bat (
630:Island tube-nosed fruit bat (
612:Dragon tube-nosed fruit bat (
576:Common tube-nosed fruit bat (
3120:(Little collared fruit bats)
2880:Minor epauletted fruit bat (
2431:Philippine gray flying fox (
2028:Bismarck masked flying fox (
1089:Ratanaworabhan's fruit bat (
684:Umboi tube-nosed fruit bat (
3559:Mammals of Papua New Guinea
3148:Little collared fruit bat (
1806:(Straw-coloured fruit bats)
1615:New Caledonia blossom bat (
1023:Fischer's pygmy fruit bat (
295:The species is larger than
261:Mammal Species of the World
3605:
3481:
2485:New Caledonia flying fox (
2235:Black-bearded flying fox (
1924:Montane monkey-faced Bat (
1915:Greater monkey-faced Bat (
1825:Straw-coloured fruit bat (
1456:Sulawesi harpy fruit bat (
1098:White-collared fruit bat (
1080:Javan tailless fruit bat (
990:Rickart's dyak fruit bat (
832:Spotted-winged fruit bat (
808:Mindanao pygmy fruit bat (
3569:Mammals described in 2001
3222:Short-palated fruit bat (
2784:
2770:
2570:
2556:
2332:Geelvink Bay flying fox (
2163:Livingstone's fruit bat (
2064:Large Samoan flying fox (
1992:Small Samoan flying fox (
1873:Small-toothed fruit bat (
1849:Fijian monkey-faced Bat (
1775:White-winged flying fox (
1699:
1685:
1665:Moss-forest blossom bat (
1565:Black-bellied fruit bat (
1556:Fardoulis's blossom bat (
1523:Long-tongued nectar bat (
1513:(Long-tongued fruit bats)
1499:
1485:
1276:
1262:
1178:Greater musky fruit bat (
981:Brooks's dyak fruit bat (
751:
737:
585:Pallas's tube-nosed bat (
543:
529:
485:
477:Extant species of family
193:
184:
166:
159:
61:Scientific classification
59:
37:
28:
23:
3494:This article related to
3185:D'Anchieta's fruit bat (
2696:Leschenault's rousette (
2341:Grey-headed flying fox (
2244:Black-eared flying fox (
2127:Ontong Java flying fox (
2019:Ashy-headed flying fox (
1596:(Long-tailed fruit bats)
1532:Long-tongued fruit bat (
1314:(Bare-backed fruit bats)
1187:Lesser musky fruit bat (
863:Black-capped fruit bat (
675:Eastern tube-nosed Bat (
316:Armstrong, K.N. (2021).
3231:Campo-Ma’an fruit bat (
2440:Temminck's flying fox (
2413:Little red flying fox (
2395:Madagascan flying fox (
2145:Andersen's flying fox (
2109:Gilliard's flying fox (
2055:Spectacled flying fox (
1656:Halmahera blossom bat (
1606:Long-tailed fruit bat (
1047:Salim Ali's fruit bat (
904:Horsfield's fruit bat (
657:Lesser tube-nosed bat (
557:(tube-nosed fruit bats)
3579:Bats of Southeast Asia
2985:(Dwarf epauleted bats)
2807:(Epauleted fruit bats)
2741:Long-haired rousette (
2723:Bare-backed rousette (
2422:Seychelles fruit bat (
2386:Rodrigues flying fox (
2271:Mauritian flying fox (
2199:Big-eared flying fox (
1965:Admiralty flying fox (
1816:Madagascan fruit bat (
1574:Woodford's fruit bat (
1211:Blanford's fruit bat (
885:(Dog-faced fruit bats)
842:Balionycteris seimundi
405:; Reeder, D.M (eds.).
334:: e.T136836A22041117.
2750:Madagascan rousette (
2669:Geoffroy's rousette (
2634:(rousette fruit bats)
2612:Philippine dawn bat (
2458:Vanikoro flying fox (
2368:Solomons flying fox (
2253:Caroline flying fox (
2037:Moluccan flying fox (
1715:Sulawesi flying fox (
371:Bergmans, W. (2001).
3264:Zenker's fruit bat (
2930:Dobson's fruit bat (
2594:Greater nectar bat (
2449:Insular flying fox (
2377:Rennell flying fox (
2181:Okinawa flying fox (
2091:Nicobar flying fox (
2001:Vanuatu flying fox (
1974:Aldabra flying fox (
1647:Common blossom bat (
1292:Bulmer's fruit bat (
1071:Tailless fruit bat (
913:Peters's fruit bat (
297:Paranyctimene raptor
286:Paranyctimene tenax
278:Paranyctimene raptor
3335:Paranyctimene tenax
3305:Paranyctimene tenax
3240:Pohle's fruit bat (
3089:Azagnyi fruit bat (
2963:Hammer-headed Bat (
2752:R. madagascariensis
2687:Egyptian rousette (
2678:Sulawesi rousette (
2467:Kosrae flying fox (
2323:Masked flying fox (
2298:Ornate flying fox (
2280:Temotu flying fox (
2226:Indian flying fox (
2217:Mariana fruit bat (
2208:Lesser flying fox (
2190:Lyle's flying fox (
2172:Lombok flying fox (
2073:Ryukyu flying fox (
2046:Makira flying fox (
1786:D. microleucopterus
1742:Palawan fruit bat (
1724:Talaud flying fox (
320:Paranyctimene tenax
207:Paranyctimene tenax
170:Paranyctimene tenax
31:Conservation status
3584:Bats of New Guinea
3065:Angolan rousette (
2705:Linduan Rousette (
2671:R. amplexicaudatus
2651:Manado fruit bat (
2503:Dwarf flying fox (
2494:Pemba flying fox (
2476:Large flying fox (
2404:Samoa flying fox (
2350:Bonin flying fox (
2314:Pelew flying fox (
2262:Great flying fox (
2136:Small flying fox (
2100:Banks flying fox (
1983:Black flying fox (
1751:Sunda flying fox (
1168:(Musky fruit bats)
971:(Dayak fruit bats)
922:Minute fruit bat (
775:Borneo fruit bat (
765:(Pygmy fruit bats)
399:"Order Chiroptera"
210:) is a species of
3511:
3510:
3476:
3475:
3461:Open Tree of Life
3297:Taxon identifiers
3288:
3287:
3284:
3283:
3280:
3279:
3276:
3275:
3233:C. campomaanensis
3197:
3196:
3160:
3159:
3123:
3040:
3039:
2988:
2914:
2810:
2766:
2765:
2762:
2761:
2714:Comoro rousette (
2637:
2603:Cave nectar bat (
2587:
2584:(Dawn fruit bats)
2552:
2551:
2548:
2547:
2289:Ceram fruit bat (
2118:Gray flying fox (
2057:P. conspicillatus
1958:
1809:
1681:
1680:
1677:
1676:
1640:
1599:
1516:
1481:
1480:
1477:
1476:
1465:Harpy fruit bat (
1317:
1270:Harpyionycterinae
1258:
1257:
1254:
1253:
1235:Swift fruit bat (
1171:
1146:Dusky fruit bat (
1124:O. cartilagonodus
1122:Luzon fruit bat (
999:Dayak fruit bat (
974:
951:C. titthaecheilus
888:
865:C. melanocephalus
784:Pygmy fruit bat (
768:
733:
732:
729:
728:
560:
418:978-0-8018-8221-0
198:
197:
54:
3596:
3532:
3525:
3518:
3490:
3483:
3469:
3468:
3456:
3455:
3443:
3442:
3430:
3429:
3417:
3416:
3404:
3403:
3391:
3390:
3378:
3377:
3365:
3364:
3352:
3351:
3339:
3338:
3337:
3324:
3323:
3322:
3292:
3208:
3171:
3132:M. brachycephala
3122:
3121:
3117:
3098:Woermann's bat (
3051:
2987:
2986:
2982:
2913:
2912:
2911:(Epauleted bats)
2908:
2809:
2808:
2804:
2795:
2786:
2772:
2636:
2635:
2631:
2586:
2585:
2581:
2572:
2558:
2424:P. seychellensis
2343:P. poliocephalus
2264:P. neohibernicus
2154:Kei flying fox (
2039:P. chrysoproctus
2010:Aru flying fox (
1967:P. admiralitatum
1957:
1956:
1952:
1808:
1807:
1803:
1701:
1687:
1639:
1638:
1634:
1617:N. neocaledonica
1598:
1597:
1593:
1515:
1514:
1510:
1501:
1487:
1316:
1315:
1311:
1278:
1264:
1170:
1169:
1165:
973:
972:
968:
887:
886:
882:
767:
766:
762:
753:
739:
559:
558:
554:
545:
531:
471:
464:
457:
448:
431:
430:
391:
385:
384:
368:
353:
352:
350:
348:
343:
313:
224:Papua New Guinea
189:
172:
69:
68:
48:
43:
42:
21:
3604:
3603:
3599:
3598:
3597:
3595:
3594:
3593:
3589:Fruit bat stubs
3574:Bats of Oceania
3539:
3538:
3537:
3536:
3479:
3477:
3472:
3464:
3459:
3451:
3448:Observation.org
3446:
3438:
3433:
3425:
3420:
3412:
3407:
3399:
3394:
3386:
3381:
3373:
3368:
3360:
3355:
3347:
3342:
3333:
3332:
3327:
3318:
3317:
3312:
3299:
3289:
3272:
3248:
3193:
3156:
3119:
3118:
3116:
3106:
3073:
3036:
3012:
2984:
2983:
2981:
2971:
2947:
2923:E. buettikoferi
2910:
2909:
2907:
2897:
2806:
2805:
2803:
2780:
2758:
2698:R. leschenaulti
2633:
2632:
2630:
2620:
2583:
2582:
2580:
2566:
2544:
2511:
2460:P. tuberculatus
2282:P. nitendiensis
2165:P. livingstonii
2111:P. gilliardorum
1954:
1953:
1951:
1941:
1881:
1857:
1833:
1805:
1804:
1802:
1792:
1759:
1695:
1673:
1636:
1635:
1633:
1623:
1595:
1594:
1592:
1582:
1540:
1512:
1511:
1509:
1495:
1473:
1447:Harpyionycteris
1440:
1407:D. pannietensis
1313:
1312:
1310:
1300:
1272:
1250:
1219:
1195:
1167:
1166:
1164:
1154:
1130:
1106:
1055:
1031:
1007:
970:
969:
967:
957:
933:C. nusatenggara
884:
883:
881:
871:
847:
816:
810:A. paucidentata
792:
764:
763:
761:
747:
725:
692:
556:
555:
553:
539:
525:
481:
475:
444:
435:
434:
419:
393:
392:
388:
370:
369:
356:
346:
344:
315:
314:
310:
305:
235:, published as
180:
174:
168:
155:
63:
55:
44:
40:
33:
17:
12:
11:
5:
3602:
3600:
3592:
3591:
3586:
3581:
3576:
3571:
3566:
3561:
3556:
3551:
3541:
3540:
3535:
3534:
3527:
3520:
3512:
3509:
3508:
3491:
3474:
3473:
3471:
3470:
3457:
3444:
3431:
3418:
3405:
3392:
3379:
3366:
3353:
3340:
3325:
3309:
3307:
3301:
3300:
3295:
3286:
3285:
3282:
3281:
3278:
3277:
3274:
3273:
3271:
3270:
3260:
3258:
3250:
3249:
3247:
3246:
3237:
3228:
3218:
3216:
3205:
3203:Scotonycterini
3199:
3198:
3195:
3194:
3192:
3191:
3181:
3179:
3168:
3162:
3161:
3158:
3157:
3155:
3154:
3145:
3136:
3126:
3124:
3108:
3107:
3105:
3104:
3095:
3085:
3083:
3075:
3074:
3072:
3071:
3061:
3059:
3048:
3042:
3041:
3038:
3037:
3035:
3034:
3024:
3022:
3014:
3013:
3011:
3010:
3001:
2997:M. intermedius
2991:
2989:
2973:
2972:
2970:
2969:
2959:
2957:
2949:
2948:
2946:
2945:
2936:
2927:
2917:
2915:
2899:
2898:
2896:
2895:
2886:
2877:
2868:
2859:
2850:
2841:
2832:
2823:
2813:
2811:
2792:
2782:
2781:
2775:
2768:
2767:
2764:
2763:
2760:
2759:
2757:
2756:
2747:
2730:
2729:
2720:
2711:
2702:
2693:
2689:R. aegyptiacus
2684:
2675:
2658:
2657:
2640:
2638:
2622:
2621:
2619:
2618:
2609:
2600:
2590:
2588:
2568:
2567:
2561:
2554:
2553:
2550:
2549:
2546:
2545:
2543:
2542:
2533:
2529:S. mindorensis
2523:
2521:
2513:
2512:
2510:
2509:
2500:
2491:
2482:
2473:
2464:
2455:
2446:
2437:
2428:
2419:
2410:
2401:
2392:
2388:P. rodricensis
2383:
2374:
2365:
2356:
2347:
2338:
2329:
2320:
2311:
2304:
2295:
2286:
2277:
2268:
2259:
2250:
2241:
2237:P. melanopogon
2232:
2223:
2214:
2205:
2196:
2187:
2183:P. loochoensis
2178:
2174:P. lombocensis
2169:
2160:
2151:
2147:P. intermedius
2142:
2138:P. hypomelanus
2133:
2124:
2115:
2106:
2097:
2088:
2079:
2070:
2061:
2052:
2043:
2034:
2030:P. capistratus
2025:
2016:
2007:
1998:
1989:
1980:
1976:P. aldabrensis
1971:
1961:
1959:
1955:(flying foxes)
1943:
1942:
1940:
1939:
1930:
1921:
1912:
1903:
1893:
1891:
1883:
1882:
1880:
1879:
1869:
1867:
1859:
1858:
1856:
1855:
1845:
1843:
1835:
1834:
1832:
1831:
1822:
1812:
1810:
1794:
1793:
1791:
1790:
1781:
1777:D. leucopterus
1771:
1769:
1761:
1760:
1758:
1757:
1748:
1739:
1730:
1721:
1711:
1709:
1697:
1696:
1690:
1683:
1682:
1679:
1678:
1675:
1674:
1672:
1671:
1662:
1653:
1643:
1641:
1637:(blossom bats)
1625:
1624:
1622:
1621:
1612:
1602:
1600:
1584:
1583:
1581:
1580:
1571:
1562:
1552:
1550:
1542:
1541:
1539:
1538:
1529:
1519:
1517:
1497:
1496:
1493:Macroglossinae
1490:
1483:
1482:
1479:
1478:
1475:
1474:
1472:
1471:
1462:
1452:
1450:
1442:
1441:
1439:
1438:
1429:
1420:
1411:
1402:
1398:D. moluccensis
1393:
1384:
1375:
1366:
1357:
1348:
1339:
1330:
1320:
1318:
1302:
1301:
1299:
1298:
1288:
1286:
1274:
1273:
1267:
1260:
1259:
1256:
1255:
1252:
1251:
1249:
1248:
1241:
1231:
1229:
1221:
1220:
1218:
1217:
1207:
1205:
1197:
1196:
1194:
1193:
1184:
1174:
1172:
1156:
1155:
1153:
1152:
1142:
1140:
1132:
1131:
1129:
1128:
1118:
1116:
1108:
1107:
1105:
1104:
1095:
1086:
1077:
1067:
1065:
1057:
1056:
1054:
1053:
1043:
1041:
1033:
1032:
1030:
1029:
1019:
1017:
1009:
1008:
1006:
1005:
996:
987:
977:
975:
959:
958:
956:
955:
946:
937:
928:
919:
915:C. luzoniensis
910:
906:C. horsfieldii
901:
891:
889:
873:
872:
870:
869:
859:
857:
849:
848:
846:
845:
838:
828:
826:
818:
817:
815:
814:
804:
802:
794:
793:
791:
790:
781:
771:
769:
749:
748:
742:
735:
734:
731:
730:
727:
726:
724:
723:
714:
704:
702:
694:
693:
691:
690:
681:
672:
663:
654:
645:
641:N. malaitensis
636:
627:
618:
609:
600:
591:
582:
573:
563:
561:
541:
540:
534:
527:
526:
524:
523:
517:
515:Laurasiatheria
511:
505:
499:
493:
486:
483:
482:
476:
474:
473:
466:
459:
451:
442:
441:
433:
432:
417:
386:
354:
307:
306:
304:
301:
293:
292:
291:
290:
282:
271:
270:
247:. The epithet
196:
195:
191:
190:
182:
181:
175:
164:
163:
157:
156:
149:
147:
143:
142:
135:
131:
130:
125:
121:
120:
115:
111:
110:
105:
101:
100:
95:
91:
90:
85:
81:
80:
75:
71:
70:
57:
56:
38:
35:
34:
29:
26:
25:
16:Species of bat
15:
13:
10:
9:
6:
4:
3:
2:
3601:
3590:
3587:
3585:
3582:
3580:
3577:
3575:
3572:
3570:
3567:
3565:
3562:
3560:
3557:
3555:
3554:Paranyctimene
3552:
3550:
3547:
3546:
3544:
3533:
3528:
3526:
3521:
3519:
3514:
3513:
3507:
3505:
3501:
3497:
3492:
3489:
3485:
3480:
3467:
3462:
3458:
3454:
3449:
3445:
3441:
3436:
3432:
3428:
3423:
3419:
3415:
3410:
3406:
3402:
3397:
3393:
3389:
3384:
3380:
3376:
3371:
3367:
3363:
3358:
3354:
3350:
3345:
3341:
3336:
3330:
3326:
3321:
3315:
3311:
3310:
3308:
3306:
3302:
3298:
3293:
3269:
3267:
3262:
3261:
3259:
3257:
3256:
3255:Scotonycteris
3251:
3245:
3243:
3238:
3236:
3234:
3229:
3227:
3225:
3220:
3219:
3217:
3215:
3214:
3209:
3206:
3204:
3200:
3190:
3188:
3183:
3182:
3180:
3178:
3177:
3172:
3169:
3167:
3163:
3153:
3151:
3146:
3144:
3142:
3137:
3135:
3133:
3128:
3127:
3125:
3115:
3114:
3109:
3103:
3101:
3096:
3094:
3092:
3087:
3086:
3084:
3082:
3081:
3080:Megaloglossus
3076:
3070:
3068:
3067:L. angolensis
3063:
3062:
3060:
3058:
3057:
3056:Lissonycteris
3052:
3049:
3047:
3043:
3033:
3031:
3030:N. veldkampii
3026:
3025:
3023:
3021:
3020:
3015:
3009:
3007:
3002:
3000:
2998:
2993:
2992:
2990:
2980:
2979:
2978:Micropteropus
2974:
2968:
2966:
2965:H. monstrosus
2961:
2960:
2958:
2956:
2955:
2950:
2944:
2942:
2937:
2935:
2933:
2928:
2926:
2924:
2919:
2918:
2916:
2906:
2905:
2900:
2894:
2892:
2887:
2885:
2883:
2878:
2876:
2874:
2869:
2867:
2865:
2860:
2858:
2856:
2851:
2849:
2847:
2842:
2840:
2838:
2833:
2831:
2829:
2824:
2822:
2820:
2819:E. angolensis
2815:
2814:
2812:
2802:
2801:
2796:
2793:
2791:
2787:
2783:
2779:
2778:Epomophorinae
2773:
2769:
2755:
2753:
2748:
2746:
2744:
2738:
2737:
2736:Stenonycteris
2732:
2731:
2728:
2726:
2725:R. spinalatus
2721:
2719:
2717:
2716:R. obliviosus
2712:
2710:
2708:
2707:R. linduensis
2703:
2701:
2699:
2694:
2692:
2690:
2685:
2683:
2681:
2680:R. celebensis
2676:
2674:
2672:
2666:
2665:
2660:
2659:
2656:
2654:
2648:
2647:
2642:
2641:
2639:
2629:
2628:
2623:
2617:
2615:
2610:
2608:
2606:
2601:
2599:
2597:
2592:
2591:
2589:
2579:
2578:
2573:
2569:
2565:
2559:
2555:
2541:
2539:
2534:
2532:
2530:
2525:
2524:
2522:
2520:
2519:
2514:
2508:
2506:
2501:
2499:
2497:
2496:P. voeltzkowi
2492:
2490:
2488:
2483:
2481:
2479:
2474:
2472:
2470:
2465:
2463:
2461:
2456:
2454:
2452:
2447:
2445:
2443:
2442:P. temminckii
2438:
2436:
2434:
2429:
2427:
2425:
2420:
2418:
2416:
2415:P. scapulatus
2411:
2409:
2407:
2402:
2400:
2398:
2393:
2391:
2389:
2384:
2382:
2380:
2375:
2373:
2371:
2366:
2364:
2362:
2357:
2355:
2353:
2348:
2346:
2344:
2339:
2337:
2335:
2330:
2328:
2326:
2325:P. personatus
2321:
2319:
2317:
2316:P. pelewensis
2312:
2310:
2309:
2305:
2303:
2301:
2296:
2294:
2292:
2287:
2285:
2283:
2278:
2276:
2274:
2269:
2267:
2265:
2260:
2258:
2256:
2255:P. molossinus
2251:
2249:
2247:
2242:
2240:
2238:
2233:
2231:
2229:
2224:
2222:
2220:
2215:
2213:
2211:
2206:
2204:
2202:
2197:
2195:
2193:
2188:
2186:
2184:
2179:
2177:
2175:
2170:
2168:
2166:
2161:
2159:
2157:
2152:
2150:
2148:
2143:
2141:
2139:
2134:
2132:
2130:
2125:
2123:
2121:
2116:
2114:
2112:
2107:
2105:
2103:
2098:
2096:
2094:
2089:
2087:
2085:
2080:
2078:
2076:
2075:P. dasymallus
2071:
2069:
2067:
2062:
2060:
2058:
2053:
2051:
2049:
2044:
2042:
2040:
2035:
2033:
2031:
2026:
2024:
2022:
2017:
2015:
2013:
2008:
2006:
2004:
1999:
1997:
1995:
1990:
1988:
1986:
1981:
1979:
1977:
1972:
1970:
1968:
1963:
1962:
1960:
1950:
1949:
1944:
1938:
1936:
1931:
1929:
1927:
1922:
1920:
1918:
1913:
1911:
1909:
1904:
1902:
1900:
1895:
1894:
1892:
1890:
1889:
1884:
1878:
1876:
1871:
1870:
1868:
1866:
1865:
1860:
1854:
1852:
1847:
1846:
1844:
1842:
1841:
1836:
1830:
1828:
1823:
1821:
1819:
1814:
1813:
1811:
1801:
1800:
1795:
1789:
1787:
1782:
1780:
1778:
1773:
1772:
1770:
1768:
1767:
1762:
1756:
1754:
1749:
1747:
1745:
1740:
1738:
1736:
1731:
1729:
1727:
1722:
1720:
1718:
1717:A. celebensis
1713:
1712:
1710:
1708:
1707:
1702:
1698:
1694:
1688:
1684:
1670:
1668:
1663:
1661:
1659:
1654:
1652:
1650:
1645:
1644:
1642:
1632:
1631:
1626:
1620:
1618:
1613:
1611:
1609:
1608:N. macdonaldi
1604:
1603:
1601:
1591:
1590:
1585:
1579:
1577:
1572:
1570:
1568:
1563:
1561:
1559:
1558:M. fardoulisi
1554:
1553:
1551:
1549:
1548:
1543:
1537:
1535:
1530:
1528:
1526:
1521:
1520:
1518:
1508:
1507:
1502:
1498:
1494:
1488:
1484:
1470:
1468:
1467:H. whiteheadi
1463:
1461:
1459:
1458:H. celebensis
1454:
1453:
1451:
1449:
1448:
1443:
1437:
1435:
1430:
1428:
1426:
1425:D. praedatrix
1421:
1419:
1417:
1412:
1410:
1408:
1403:
1401:
1399:
1394:
1392:
1390:
1385:
1383:
1381:
1376:
1374:
1372:
1367:
1365:
1363:
1358:
1356:
1354:
1349:
1347:
1345:
1340:
1338:
1336:
1331:
1329:
1327:
1322:
1321:
1319:
1309:
1308:
1303:
1297:
1295:
1290:
1289:
1287:
1285:
1284:
1279:
1275:
1271:
1265:
1261:
1247:
1246:
1245:T. suhaniahae
1242:
1240:
1238:
1237:T. nigrescens
1233:
1232:
1230:
1228:
1227:
1222:
1216:
1214:
1209:
1208:
1206:
1204:
1203:
1198:
1192:
1190:
1185:
1183:
1181:
1176:
1175:
1173:
1163:
1162:
1157:
1151:
1149:
1144:
1143:
1141:
1139:
1138:
1133:
1127:
1125:
1120:
1119:
1117:
1115:
1114:
1109:
1103:
1101:
1096:
1094:
1092:
1087:
1085:
1083:
1078:
1076:
1074:
1069:
1068:
1066:
1064:
1063:
1058:
1052:
1050:
1045:
1044:
1042:
1040:
1039:
1034:
1028:
1026:
1021:
1020:
1018:
1016:
1015:
1014:Haplonycteris
1010:
1004:
1002:
997:
995:
993:
988:
986:
984:
979:
978:
976:
966:
965:
960:
954:
952:
947:
945:
943:
938:
936:
934:
929:
927:
925:
920:
918:
916:
911:
909:
907:
902:
900:
898:
897:C. brachyotis
893:
892:
890:
880:
879:
874:
868:
866:
861:
860:
858:
856:
855:
850:
844:
843:
839:
837:
835:
830:
829:
827:
825:
824:
823:Balionycteris
819:
813:
811:
806:
805:
803:
801:
800:
795:
789:
787:
782:
780:
778:
773:
772:
770:
760:
759:
754:
750:
746:
740:
736:
722:
720:
715:
713:
711:
706:
705:
703:
701:
700:
699:Paranyctimene
695:
689:
687:
682:
680:
678:
673:
671:
669:
664:
662:
660:
655:
653:
651:
646:
644:
642:
637:
635:
633:
628:
626:
624:
619:
617:
615:
614:N. draconilla
610:
608:
606:
601:
599:
597:
592:
590:
588:
587:N. cephalotes
583:
581:
579:
578:N. albiventer
574:
572:
570:
565:
564:
562:
552:
551:
546:
542:
538:
532:
528:
522:
518:
516:
512:
510:
506:
504:
500:
498:
494:
492:
488:
487:
484:
480:
472:
467:
465:
460:
458:
453:
452:
449:
445:
440:
437:
436:
428:
424:
420:
414:
410:
409:
404:
400:
396:
395:Simmons, N.B.
390:
387:
383:(8): 119–152.
382:
378:
374:
367:
365:
363:
361:
359:
355:
342:
337:
333:
329:
328:
323:
321:
312:
309:
302:
300:
298:
288:
287:
283:
280:
279:
275:
274:
273:
272:
269:
268:Paranyctimene
266:
265:
264:
262:
258:
254:
253:Peter Nijhoff
250:
246:
242:
241:Paranyctimene
238:
234:
233:
227:
225:
221:
217:
213:
209:
208:
203:
192:
188:
183:
178:
173:
171:
165:
162:
161:Binomial name
158:
154:
153:
152:P. tenax
148:
145:
144:
141:
140:
139:Paranyctimene
136:
133:
132:
129:
126:
123:
122:
119:
116:
113:
112:
109:
106:
103:
102:
99:
96:
93:
92:
89:
86:
83:
82:
79:
76:
73:
72:
67:
62:
58:
52:
47:
46:Least Concern
36:
32:
27:
22:
19:
3504:expanding it
3493:
3478:
3304:
3265:
3253:
3241:
3232:
3223:
3213:Casinycteris
3211:
3187:P. anchietae
3186:
3174:
3149:
3140:
3131:
3111:
3100:M. woermanni
3099:
3090:
3078:
3066:
3054:
3046:Myonycterini
3029:
3019:Nanonycteris
3017:
3005:
2996:
2976:
2964:
2954:Hypsignathus
2952:
2941:E. franqueti
2940:
2931:
2922:
2902:
2891:E. wahlbergi
2890:
2881:
2872:
2863:
2854:
2846:E. gambianus
2845:
2837:E. crypturus
2836:
2827:
2818:
2798:
2790:Epomophorini
2751:
2742:
2733:
2724:
2715:
2706:
2697:
2688:
2679:
2670:
2661:
2652:
2643:
2625:
2613:
2604:
2595:
2575:
2537:
2528:
2518:Styloctenium
2516:
2505:P. woodfordi
2504:
2495:
2486:
2477:
2468:
2459:
2450:
2441:
2433:P. speciosus
2432:
2423:
2414:
2406:P. samoensis
2405:
2396:
2387:
2378:
2369:
2360:
2352:P. pselaphon
2351:
2342:
2333:
2324:
2315:
2308:P. pelagicus
2306:
2299:
2290:
2281:
2272:
2263:
2254:
2246:P. melanotus
2245:
2236:
2227:
2219:P. mariannus
2218:
2210:P. mahaganus
2209:
2200:
2191:
2182:
2173:
2164:
2155:
2146:
2137:
2128:
2119:
2110:
2101:
2092:
2083:
2074:
2065:
2056:
2047:
2038:
2029:
2020:
2011:
2003:P. anetianus
2002:
1994:P. allenorum
1993:
1984:
1975:
1966:
1946:
1934:
1925:
1917:P. flanneryi
1916:
1907:
1898:
1886:
1874:
1862:
1851:M. acrodonta
1850:
1838:
1826:
1818:E. dupreanum
1817:
1797:
1785:
1776:
1764:
1752:
1743:
1734:
1725:
1716:
1704:
1693:Pteropodinae
1666:
1658:S. carolinae
1657:
1649:S. australis
1648:
1630:Syconycteris
1628:
1616:
1607:
1587:
1576:M. woodfordi
1575:
1566:
1557:
1547:Melonycteris
1545:
1533:
1524:
1506:Macroglossus
1504:
1466:
1457:
1445:
1433:
1424:
1415:
1406:
1397:
1388:
1379:
1370:
1361:
1353:D. crenulata
1352:
1343:
1335:D. beauforti
1334:
1326:D. anderseni
1325:
1305:
1293:
1281:
1243:
1236:
1224:
1213:S. blanfordi
1212:
1200:
1188:
1179:
1159:
1147:
1135:
1123:
1111:
1099:
1090:
1081:
1073:M. ecaudatus
1072:
1060:
1049:L. salimalii
1048:
1036:
1024:
1012:
1001:D. spadiceus
1000:
991:
982:
962:
950:
941:
932:
923:
914:
905:
896:
876:
864:
852:
840:
833:
821:
809:
799:Alionycteris
797:
785:
776:
756:
745:Cynopterinae
718:
716:
709:
697:
686:N. vizcaccia
685:
677:N. robinsoni
676:
667:
658:
649:
640:
631:
622:
613:
604:
595:
586:
577:
568:
548:
537:Nyctimeninae
513:Superorder:
507:Infraclass:
479:Pteropodidae
443:
407:
403:Wilson, D.E.
389:
380:
376:
345:. Retrieved
331:
325:
319:
311:
296:
294:
285:
284:
276:
267:
256:
248:
244:
240:
236:
230:
228:
206:
205:
201:
199:
169:
167:
151:
150:
138:
128:Pteropodidae
18:
3383:iNaturalist
3329:Wikispecies
3242:C. ophiodon
3224:C. argynnis
3150:M. torquata
3113:Myonycteris
3006:M. pusillus
2864:E. labiatus
2800:Epomophorus
2564:Rousettinae
2538:S. wallacei
2478:P. vampyrus
2451:P. tonganus
2379:P. rennelli
2291:P. ocularis
2201:P. macrotis
2156:P. keyensis
2129:P. howensis
2102:P. fundatus
2093:P. faunulus
2048:P. cognatus
2021:P. caniceps
2012:P. aruensis
1766:Desmalopex
1753:A. mackloti
1744:A. leucotis
1567:M. melanops
1534:M. sobrinus
1344:D. chapmani
1294:A. bulmerae
1161:Ptenochirus
1113:Otopteropus
1100:M. wetmorei
1091:M. niphanae
1082:M. kusnotoi
1025:H. fischeri
992:D. rickarti
964:Dyacopterus
834:B. maculata
777:A. aequalis
605:N. cyclotis
347:16 December
3543:Categories
3496:fruit bats
3266:S. zenkeri
3141:M. relicta
3091:M. azagnyi
2932:E. dobsoni
2873:E. minimus
2855:E. grandis
2828:E. anselli
2776:Subfamily
2743:R. lanosus
2614:E. robusta
2605:E. spelaea
2577:Eonycteris
2562:Subfamily
2487:P. vetulus
2469:P. ualanus
2370:P. rayneri
2361:P. pumilus
2300:P. ornatus
2120:P. griseus
2084:P. ennisae
1926:P. pulchra
1888:Pteralopex
1735:A. jubatus
1726:A. humilis
1691:Subfamily
1589:Notopteris
1525:M. minimus
1491:Subfamily
1434:D. viridis
1416:D. peronii
1380:D. inermis
1371:D. exoleta
1268:Subfamily
1226:Thoopterus
983:D. brooksi
924:C. minutus
878:Cynopterus
743:Subfamily
659:N. minutus
650:N. masalai
596:N. certans
535:Subfamily
521:Chiroptera
377:Beaufortia
303:References
216:West Papua
118:Chiroptera
3166:Plerotini
2734:Subgenus
2664:Rousettus
2662:Subgenus
2653:R. bidens
2644:Subgenus
2627:Rousettus
2334:P. pohlei
2228:P. medius
1985:P. alecto
1908:P. atrata
1899:P. anceps
1875:N. frosti
1864:Neopteryx
1827:E. helvum
1667:S. hobbit
1362:D. emersa
1283:Aproteles
1202:Sphaerias
1180:P. jagori
1148:P. lucasi
1137:Penthetor
1062:Megaerops
942:C. sphinx
786:A. alecto
758:Aethalops
710:P. raptor
668:N. rabori
623:N. keasti
550:Nyctimene
489:Kingdom:
237:Nyctimene
232:Nyctimene
220:Indonesia
146:Species:
84:Kingdom:
78:Eukaryota
3440:13800223
3320:Q1770246
3314:Wikidata
3176:Plerotes
2882:E. minor
2596:E. major
2397:P. rufus
2273:P. niger
2192:P. lylei
1948:Pteropus
1840:Mirimiri
1706:Acerodon
1389:D. minor
1307:Dobsonia
1189:P. minor
1038:Latidens
854:Chironax
719:P. tenax
632:N. major
569:N. aello
509:Eutheria
503:Mammalia
497:Chordata
495:Phylum:
491:Animalia
427:62265494
397:(2005).
263:(2005).
257:marculus
177:Bergmans
124:Family:
108:Mammalia
98:Chordata
94:Phylum:
88:Animalia
74:Domain:
51:IUCN 3.1
3466:3613528
3427:1004430
3375:4265736
3362:7245151
2904:Epomops
2066:P. coxi
1935:P. taki
1799:Eidolon
519:Order:
501:Class:
212:megabat
134:Genus:
114:Order:
104:Class:
49: (
3453:201010
3414:136836
3401:945978
2646:Boneia
425:
415:
222:, and
194:Range
179:, 2001
3498:is a
3388:74844
3349:4D6W4
401:. In
249:tenax
245:tenax
3500:stub
3409:IUCN
3396:ITIS
3370:GBIF
423:OCLC
413:ISBN
349:2023
332:2021
200:The
3435:MSW
3422:MDD
3357:EoL
3344:CoL
336:doi
3545::
3463::
3450::
3437::
3424::
3411::
3398::
3385::
3372::
3359::
3346::
3331::
3316::
2739::
2667::
2649::
421:.
381:51
379:.
375:.
357:^
330:.
324:.
243:)
226:.
218:,
3531:e
3524:t
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470:e
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