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Strudiella

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104: 877: 378:(2013). They supported mandibular teeth structure again. They argued that visible maxillary palps are abnormal under the crustacean hypothesis. They considered that scape, pedicel and flagellum can be confirmed from width of antennae, and antennae being wider than legs would not deny insect affinity. For the extra legs that Hörnschemeyer 619: 465:
Romain Garrouste, Gaël Clément, Patricia Nel, Michael S. Engel, Philippe Grandcolas, Cyrille D’Haese, Linda Lagebro, Julien Denayer, Pierre Gueriau, Patrick Lafaite, Sébastien Olive, Cyrille Prestianni and André Nel (2012). "A complete insect from the Late Devonian period". Nature. 488 (7409): 82–85.
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is considered as arbitrary. The scape, pedicel and flagellum on its antennae cannot be confirmed, and the whole antennae is wider than the legs which is highly unusual for insects. The alleged subdivision of the trunk into thorax and abdomen is also questionable. Importantly, over 3 pairs of legs can
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Ward, P.; Labandeira, C.; Laurin, M.; Berner, R. A. (2006). "Confirmation of Romer's Gap as a low oxygen interval constraining the timing of initial arthropod and vertebrate terrestrialization". Proceedings of the National Academy of Sciences. 103 (45): 16818–22. Bibcode:2006PNAS..10316818W. PMC
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is known from a single specimen. It is a small arthropod with length about 8 mm (0.31 in). Structures like antennae and a number of pairs of legs can be seen. Due to the poorly preserved nature of this fossil, its interpretation and classification depend on the authors.
674: 351:(2013) questioned its interpretation as an insect. The mandible and its teeth cannot be confirmed, the mandibular teeth would be caused by the idiosyncratic way the rock parted. The eyes are vague, the indication of eye rims by Garrouste 562:
Garrouste, Romain; Clément, Gaël; Nel, Patricia; Engel, Michael S.; Grandcolas, Philippe; D’Haese, Cyrille A.; Lagebro, Linda; Denayer, Julien; Gueriau, Pierre; Lafaite, Patrick; Olive, Sébastien; Prestianni, Cyrille; Nel, André (2013).
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have a continuous series of sharp, small irregular molar and incisor cusps. Its small head had large eyes. The large thorax had a rounded structure covering its head, corresponding to an expanded pronotum. The
410:(the 'gap' still occurs in the early Carboniferous, coinciding and extending past the Romer's gap for tetrapods, which may have been caused by low oxygen levels in the atmosphere). However, Hörnschemeyer 498:
Hörnschemeyer, Thomas; Haug, Joachim T.; Bethoux, Olivier; Beutel, Rolf G.; Charbonnier, Sylvain; Hegna, Thomas A.; Koch, Markus; Rust, Jes; Wedmann, Sonja; Bradler, Sven; Willmann, Rainer (2013-02-20).
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as "putative insect". Haug and Haug (2017) listed presumed Devonian insect fossils, and commented that "Its very incomplete preservation makes its interpretation problematic".
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Gueriau, Pierre; Rabet, Nicolas; Clément, Gaël; Lagebro, Linda; Vannier, Jean; Briggs, Derek E. G.; Charbonnier, Sylvain; Olive, Sébastien; Béthoux, Olivier (2016-02-08).
1066: 294:(2012), it was described as the first complete Devonian insect. The median abdominal structures are filled with guts, which excludes the possibility that is a 323:
or other appendicular structures. There is a rounded apical structure on the tip of the abdomen. The lack of wings and small size would indicate that it is a
270:. It was originally described as the first complete Late Devonian terrestrial insect, but due to its poor state of preservation, its affinity is discussed. 1053: 961: 1145: 1155: 1120: 675:"Stylonurine eurypterids from the Strud locality (Upper Devonian, Belgium): new insights into the ecology of freshwater sea scorpions" 319:
had 10 segments (while a drawing in the paper shows 11 segments), and the authors considered that it lacked any lateral leglets,
1125: 212: 1140: 881: 620:"The Strud crustacean fauna (Late Devonian, Belgium): updated review and palaeoecology of an early continental ecosystem" 1150: 1135: 311: 730:"A 365-Million-Year-Old Freshwater Community Reveals Morphological and Ecological Stasis in Branchiopod Crustaceans" 406:
as a Devonian insect reduces a previous gap of 45 million years in the evolutionary history of insects, part of the
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be observed. This poor state of preservation allows numbers of alternate interpretations like a decayed crustacean.
414:(2013) which denied its interpretation as insect commented that it is crucial to prevent this fossil from entering 299: 263: 942: 1130: 904: 1092: 987: 892: 741: 686: 631: 576: 512: 439: 673:
Lamsdell, James C.; Lagebro, Linda; Edgecombe, Gregory D.; Budd, Graham E.; Gueriau, Pierre (2019).
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are questionable, and the early fossil records of insects still remains scarce and problematic.
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confirmed, they considered as internal organs extruded during compression and decay.
307: 61: 1019: 604: 420: 220: 310:, the scape and pedicel are wider than the 10-segmented flagellum. The triangular 927: 1028: 936: 426: 36: 876: 754: 729: 698: 643: 624:
Earth and Environmental Science Transactions of the Royal Society of Edinburgh
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insect, and the mandible morphology suggests its diet would be
790:"The presumed oldest flying insect: more likely a myriapod?" 418:
textbooks. Most of fossil records of Devonian insects like
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Gueriau, Pierre; Rabet, Nicolas; Hat, Eva Du Tien (2016).
298:. It is interpreted to have 3 pairs of legs from the 1003: 911: 390:Multiple later studies about Strud fossils call 302:, the tibiae and femora are long and thin. The 788:Haug, Carolin; Haug, Joachim T. (2017-05-30). 8: 250:is a species of extinct arthropod from the 899: 102: 20: 823: 805: 753: 451: 290:In the first description by Garrouste 7: 783: 781: 493: 491: 489: 487: 461: 459: 457: 455: 501:"Is Strudiella a Devonian insect?" 14: 875: 262:, Belgium) environment from the 111:Interpretation after Garrouste 1146:Taxa named by Michael S. Engel 1: 1156:Devonian arthropods of Europe 1121:Fossil taxa described in 2012 327:. It is interpreted to be a 16:Extinct species of arthropod 847:1636538 Freely accessible. 343:Hörnschemeyer et al. (2013) 1172: 264:Bois des Mouches Formation 254:. It was recovered in the 755:10.1016/j.cub.2015.12.039 699:10.1017/S0016756818000936 644:10.1017/S1755691017000275 374:replied to Hörnschemeyer 226: 219: 125:Scientific classification 123: 115:(2012) and Hörnschemeyer 110: 101: 23: 565:"Garrouste et al. reply" 861:10.1073/pnas.0607824103 398:Importance of discovery 360:Garrouste et al. (2013) 347:However, Hörnschemeyer 286:Garrouste et al. (2012) 364:In the same volume of 1126:Late Devonian animals 1093:Paleobiology Database 988:Paleobiology Database 1141:Evolution of insects 893:Encyclopedia of Life 440:Evolution of insects 943:Strudiella devonica 913:Strudiella devonica 746:2016CBio...26..383G 691:2019GeoM..156.1708L 679:Geological Magazine 636:2016EESTR.107...79G 589:10.1038/nature11888 581:2013Natur.494E...4G 525:10.1038/nature11887 517:2013Natur.494E...3H 480:10.1038/nature11281 247:Strudiella devonica 229:Strudiella devonica 1151:Controversial taxa 1136:Fossils of Belgium 807:10.7717/peerj.3402 1108: 1107: 1080:Open Tree of Life 975:Open Tree of Life 905:Taxon identifiers 685:(10): 1708–1714. 402:The discovery of 243: 242: 1163: 1101: 1100: 1088: 1087: 1075: 1074: 1062: 1061: 1049: 1048: 1039: 1038: 1037: 1024: 1023: 1022: 996: 995: 983: 982: 970: 969: 957: 956: 947: 946: 945: 932: 931: 930: 900: 880:Data related to 879: 863: 844: 838: 837: 827: 809: 785: 776: 775: 757: 725: 719: 718: 670: 664: 663: 615: 609: 608: 559: 553: 552: 495: 482: 463: 106: 96: 33: 29:Temporal range: 21: 1171: 1170: 1166: 1165: 1164: 1162: 1161: 1160: 1111: 1110: 1109: 1104: 1096: 1091: 1083: 1078: 1070: 1065: 1057: 1052: 1044: 1042: 1033: 1032: 1027: 1018: 1017: 1012: 999: 991: 986: 978: 973: 965: 960: 952: 950: 941: 940: 935: 926: 925: 920: 907: 872: 867: 866: 845: 841: 787: 786: 779: 734:Current Biology 727: 726: 722: 672: 671: 667: 617: 616: 612: 575:(7437): E4–E5. 561: 560: 556: 511:(7437): E3–E4. 497: 496: 485: 464: 453: 448: 436: 400: 388: 362: 345: 288: 276: 239: 232: 215: 205: 204: 189: 176: 163: 150: 138: 97: 95: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 39: 31: 30: 27: 17: 12: 11: 5: 1169: 1167: 1159: 1158: 1153: 1148: 1143: 1138: 1133: 1131:Famennian life 1128: 1123: 1113: 1112: 1106: 1105: 1103: 1102: 1089: 1076: 1063: 1050: 1040: 1025: 1009: 1007: 1001: 1000: 998: 997: 984: 971: 958: 948: 933: 917: 915: 909: 908: 903: 897: 896: 885: 884:at Wikispecies 871: 870:External links 868: 865: 864: 839: 777: 740:(3): 383–390. 720: 665: 630:(2–3): 79–90. 610: 554: 483: 450: 449: 447: 444: 443: 442: 435: 432: 399: 396: 387: 384: 361: 358: 344: 341: 287: 284: 275: 272: 241: 240: 233: 224: 223: 217: 216: 206: 197: 195: 191: 190: 184: 182: 178: 177: 171: 169: 165: 164: 158: 156: 152: 151: 146: 144: 140: 139: 134: 132: 128: 127: 121: 120: 108: 107: 99: 98: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 40: 35: 34: 28: 15: 13: 10: 9: 6: 4: 3: 2: 1168: 1157: 1154: 1152: 1149: 1147: 1144: 1142: 1139: 1137: 1134: 1132: 1129: 1127: 1124: 1122: 1119: 1118: 1116: 1099: 1094: 1090: 1086: 1081: 1077: 1073: 1068: 1064: 1060: 1055: 1051: 1047: 1041: 1036: 1030: 1026: 1021: 1015: 1011: 1010: 1008: 1006: 1002: 994: 989: 985: 981: 976: 972: 968: 963: 959: 955: 949: 944: 938: 934: 929: 923: 919: 918: 916: 914: 910: 906: 901: 895: 894: 889: 886: 883: 878: 874: 873: 869: 862: 858: 854: 850: 843: 840: 835: 831: 826: 821: 817: 813: 808: 803: 799: 795: 791: 784: 782: 778: 773: 769: 765: 761: 756: 751: 747: 743: 739: 735: 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678: 668: 627: 623: 613: 572: 568: 557: 508: 504: 425: 421:Rhyniognatha 419: 411: 403: 401: 391: 389: 379: 375: 371: 370:, Garrouste 365: 363: 352: 348: 346: 291: 289: 278: 277: 246: 245: 244: 235: 228: 227: 221:Type species 208: 200: 199: 181:(unranked): 116: 112: 24: 18: 1029:Wikispecies 937:Wikispecies 427:Leverhulmia 329:terrestrial 274:Description 155:Subphylum: 1115:Categories 1035:Strudiella 1020:Q119983774 1005:Strudiella 882:Strudiella 446:References 416:entomology 404:Strudiella 392:Strudiella 337:saprophagy 279:Strudiella 234:Garrouste 207:Garrouste 201:Strudiella 187:Dicondylia 148:Arthropoda 25:Strudiella 928:Q12072575 816:2167-8359 800:: e3402. 764:0960-9822 715:135411044 707:0016-7568 660:133838663 652:1755-6910 597:1476-4687 549:205232661 533:1476-4687 333:fungivore 312:mandibles 308:uniramous 268:Famennian 131:Kingdom: 32:Famennian 1043:BioLib: 1014:Wikidata 951:BioLib: 922:Wikidata 853:17065318 834:28584727 772:26776738 541:23426326 472:22859205 434:See also 304:antennae 266:, Upper 252:Devonian 161:Hexapoda 143:Phylum: 136:Animalia 1085:6324396 1072:1484519 1059:8532529 1046:1571825 980:6324397 967:8668604 954:1571826 890:at the 825:5452959 742:Bibcode 687:Bibcode 632:Bibcode 605:4321678 577:Bibcode 513:Bibcode 335:and/or 317:abdomen 194:Genus: 174:Insecta 168:Class: 119:(2013) 1098:243457 993:243459 851:  832:  822:  814:  770:  762:  713:  705:  658:  650:  603:  595:  569:Nature 547:  539:  531:  505:Nature 470:  412:et al. 380:et al. 376:et al. 372:et al. 367:Nature 353:et al. 349:et al. 300:thorax 292:et al. 260:Gesves 238:, 2012 236:et al. 209:et al. 117:et al. 113:et al. 1067:IRMNG 794:PeerJ 711:S2CID 656:S2CID 601:S2CID 545:S2CID 325:nymph 321:gills 256:Strud 1054:GBIF 962:GBIF 849:PMID 830:PMID 812:ISSN 768:PMID 760:ISSN 703:ISSN 648:ISSN 593:ISSN 537:PMID 529:ISSN 468:PMID 306:are 213:2012 37:Preęž’ 857:doi 820:PMC 802:doi 750:doi 695:doi 683:156 640:doi 628:107 585:doi 573:494 521:doi 509:494 476:doi 424:or 1117:: 1095:: 1082:: 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Index

Preęž’
ęž’
O
S
D
C
P
T
J
K
Pg
N

Scientific classification
Animalia
Arthropoda
Hexapoda
Insecta
Dicondylia
2012
Type species
Devonian
Strud
Gesves
Bois des Mouches Formation
Famennian
molting shell
thorax
antennae
uniramous

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