252:
513:
664:
652:
56:
105:
42:
640:
679:
80:
229:
577:
fighting actual combat. They mainly attack through biting each other and aim at the heads or other vulnerable spots. Occasionally, even extremely deep cuts into jaw muscles and tissues were observed. Generally, equally sized males will have longer fights than two males with differing sizes; however, males fighting intruders in their own areas of residence did not win significantly more than intruders.
703:
694:
624:, lack of habitat management, and inappropriate habitat management. Although the sand lizard is under strict protection in the UK, there are still actions needed to be taken, including habitat protection, habitat management, species protection, species management, distribution surveys, population and conservation status monitoring, scientific research, and public awareness.
474:
laying timing from year-to-year, with the first ones to do so generally being larger, more physically fit females. These early clutches ended up producing offspring that were larger, more successful, and had higher survival rates. Generally offspring from earlier clutches were higher quality offspring.
503:
Natural malformed offspring have been observed in sand lizards most likely due to close kin inbreeding defects. Because of their relatively long lifespan and male polygynous mating, there is a decent chance of inbreeding occurring during their lifespan. Inbreeding has been seen to cause malformations
473:
Female sand lizards usually only lay a single clutch each year and clutch sizes that range between six and 15 eggs in a single one. However, there is a lot of variation in the time of year when females end up laying their eggs. But generally a single female sand lizard has a relatively consistent egg
627:
Efforts in the UK have been made to protect and conserve their habitats because the sand lizard is one of only six reptile species found in
Britain. Lizard populations were analyzed and their habitats were studied and specific recommendations were made for maintaining the quality of the environments
588:
Because of this, a several day time commitment means the investment male sand lizards put into mating is incredibly high and as a result males can begin to develop selective preferences for certain females. Because female sand lizards are often found close together in groups, males have more choices
580:
If male sand lizards engage in a fight with another multiple times, they will consider them a rival. However, subsequent aggressive interactions beyond the first one tend to be much shorter on average than the first encounter between the two. This is most likely because they are able to individually
572:
Male sand lizards are polygynous and compete with each other for female mates through incredibly aggressive manners. Sand lizard copulation is a process that lasts 2–4 minutes and after it is completed, the males will guard the individual female for hours or up to several days to protect them. After
559:
Sand lizard locomotor performance, agility, and speed is related to their ability to avoid predators (as well as foraging efficiency). Like many lizards, sand lizards are able to separate their tail from the rest of their body in order to escape predators. Sand lizards with autotomy were able to run
486:
Sand lizards have demonstrated that as their age increases, so does their reproductive success. However, a study showed that this relationship is most likely due to the increase in body size as sand lizards mature rather than increased experience they acquire as they live longer. This may be because
462:
Generally, males reach sexual maturity at a smaller size compared with females. Vitellogenesis happens when females are 45 days for the whole population. Both sexes tend to lose body fat during mating period, since their main energy resources come from body fat and from the liver and proximal at the
449:
Generally females will simply reject male attempts at mating outside of their receptive mating period, doing so through a unique head bobbing behavior. However, during their period of receptivity, females have been known to not discriminate against different males and do not reject them besides when
343:
Sand lizards spend most of their time basking, foraging, or under vegetation. They prefer to live in diverse habitats. They are largely solitary outside of mating season. Male sand lizards typically have larger territories than females, and they will compete with other males when territory overlaps.
453:
Male sand lizards have been shown to highly prefer larger females when selecting mates, but they are still willing to mate with smaller females if the opportunity presents itself. It was observed that some males that were too small for the large females they attempted to court had troubles gripping
432:
on logs. This is most likely due to its absorption and slow release of heat radiation as well as providing a vantage point to spot potential predators and prey. They very rarely share basking sites with other lizards. Male sand lizards especially avoid sharing basking sites with each other, and are
413:
Male sand lizards have home ranges with an extremely high amount of overlap; however, female sand lizards have much smaller home ranges (generally less than 100 meters squared) that neighbor each other or they will even share burrows or basking areas with each other. They highly prefer and are much
363:
legged lizard. In northwest Europe, both sexes are characterised by lateral and dorsal strips of ocellated (eye-shaped) markings, which appear as dark patches with pale centres. Colouration varies across their
European and Russian range. Males have finer markings than females, and their flanks turn
427:
Sand lizards spend extended periods of the day after emerging on long basking sessions. They occasionally take breaks to forage for resources. They also spend a considerable amount of time simply doing normal activity in areas under concealed vegetation. Under poor weather conditions, sand lizards
477:
When female sand lizards produce larger clutch sizes, the size of each individual offspring in it has been observed to decrease. On the opposite side, smaller clutch sizes had fewer, but larger, offspring. When food resources were increased so it was not a limiting factor, clutch size increased
576:
When male sand lizards compete over females, they perform ritualistic displays that often escalate into aggressive behaviors and actual combat. Males raise themselves up on all four limbs and turn over to intimidate their opponents, one may back down at this stage but if not, the two can begin
495:
When a female sand lizard mates with two or more males, sperm competition within the female's reproductive tract may occur. Active selection of sperm by females appears to occur in a manner that enhances female fitness. On the basis of this selective process, the sperm of males that are more
563:
Sand lizards are a frequent carrier of common ticks which are a great risk to the health of the lizards, especially males during breeding seasons when they have great mobility. Males with larger home territory ranges were also subject to carrying even greater loads of parasites. Trematodes,
584:
Sand lizard mating seasons are very short, lasting only 17 days of the entire year. Generally males are only able to mate with an absolute maximum of six different females during this time period and because a single mating guarding session can take up to 18% of the entire mating season.
394:
In males, the bright green genital coloration has been shown to be brighter depending on body mass and fighting ability. Males with brighter colors were more likely to initiate aggressive behaviors and win fights, which generally leads to them having a higher mating success.
500:. Multiple inseminations of multiple fully fertile males can result in multiple paternity of the offspring. Mating order and time between copulations has been found to have no effect on the reproductive success of the first or last male to mate.
478:
significantly, so it is likely that sand lizards produce clutch sizes with different offspring sizes based on resource availability. The other factor that contributes is the physical constraints of the space available to lay their clutches in.
445:
in organisms is more selective than male mate choice. This is due to the fact that females generally have to invest more time and resources into offspring than males who are under less selective pressures due to their low parental investment.
251:
410:, sand dunes, and rock gardens. While in continental Europe the sand lizard has a wide variety of habitat, in colder places like Great Britain they depend on sand to incubate their eggs and such are restricted to coastal habitats.
433:
only found occasionally sharing with females. They avoid basking on open sand or soil and prefer to use vegetation if they bask on the ground. They thermoregulate themselves by moving between areas of high and low insolation.
379:
also includes a plain red or brown-backed phase without any dorsal markings. In these two subspecies, only the flanks of the males turn green in the mating season, but in the eastern subspecies (predominantly
340:, the sand lizard's mating season is very short. Males choose mates selectively, whereas females mate more indiscriminately. Females usually only lay a single clutch of eggs per year.
414:
more active in areas with a wide variety of structural characteristics. This can create different temperature micro areas that they are able to move between depending on their needs.
1693:
496:
distantly related to the female are preferentially used for fertilization, rather than the sperm of close relatives. This preference may enhance the fitness of progeny by reducing
1974:
581:
recognize others and the result of the fight between two rivals will most likely be similar to the first, so they are able to predict the outcome and end their battle early.
364:
bright green during the spring mating season, fading again in the late summer. Male adults may reach a total body length of 19.3 cm, where female adults may reach 18.5 cm.
1865:
512:
1917:
2010:
463:
tail. After a few weeks from the hibernation, male adults become extremely aggressive towards each other, trying to mate as many females as they can.
560:
faster and as a result were better at avoiding predators that were chasing them. These tails do not regrow past 80% the original length after autotomy.
564:
metacercariae, cestode larva, nematodes, nematode larvae and acarines are all examples of parasites that inhabit the alimentary tract of this lizard.
1839:
610:
1878:
1246:"Does reproductive success increase with age or with size in species with indeterminate growth? A case study using sand lizards (Lacerta agilis)"
840:"Systematics of the Palaearctic and Oriental lizard tribe Lacertini (Squamata: Lacertidae: Lacertinae), with descriptions of eight new genera"
1943:
866:
336:
Males are known for their bright coloration and aggressive, possessive behaviors when seasonally competing for females. In contrast to other
2035:
678:
651:
1549:
1123:
Corbett, K.F. and D.L. Tamarind. 1979. Conservation of the sand lizard, Lacerta agilis, by habitat management. Brit. J. Herp. 5: 799-823.
2050:
2030:
723:
685:
1800:
729:
663:
317:. Its distribution is often patchy. In the northern extremes of the sand lizard's distribution, it survives by inhabiting seaside
597:
This lizard is regarded as threatened and is strictly protected under UK law – as it is throughout most of Europe (it is a
454:
females with their jaws during copulation due to their small size. This may be one constricting factor in their choice of mate.
219:
1883:
613:
coordinates conservation action for the sand lizard, including a successful captive-breeding and reintroduction programme.
1904:
1132:
Olsson, Mats (1988-01-01). "Ecology of a
Swedish population of the sand lizard (Lacerta agilis) - a preliminary report".
406:
The sand lizard can be found in a variety of habitat. They prefer dry habitats such as outcrops, fields, hills, beaches,
1114:
Strasbourg (26 October 2006). "Action Plan for the
Conservation of the Sand Lizard (Lacerta agilis) in Northwest Europe"
487:
female sand lizards that are larger are simply able to produce more clutches with the increased space in their bodies.
2045:
1818:
2020:
598:
1922:
1389:
Ekner-Grzyb, Anna; Sajkowska, Zofia; Dudek, Krzysztof; Gawałek, Monika; Skórka, Piotr; Tryjanowski, Piotr (2013).
1831:
734:
104:
2040:
2025:
2015:
1151:
1987:
943:
1004:"Male preference for large females and assortative mating for body size in the sand lizard (Lacerta agilis)"
944:"Malformed offspring, sibling matings, and selection against inbreeding in the sand lizard (Lacerta agilis)"
702:
504:
in sand lizards with studies in laboratory settings mirroring the deformations seen naturally in the wild.
1704:
1625:
Russell, Liam (December 2012). "THE CONSERVATION AND LANDSCAPE GENETICS OF THE SAND LIZARD Lacerta agilis"
639:
1391:"Locomotor performance of sand lizards (Lacerta agilis): effects of predatory pressure and parasite load"
428:
emerge from resting locations later and bask less often throughout the day. Sand lizards most frequently
621:
497:
199:
1343:
1787:
1751:
1648:
1459:
1257:
1199:"The Limits to Reproductive Output: Offspring Size Versus Number in the Sand Lizard (Lacerta agilis)"
1065:
1152:"The seasonal timing of oviposition in sand lizards (Lacerta agilis): why early clutches are better"
778:
1742:
617:
429:
69:
1054:"Patterns of daily behaviour in two lizard species Lacerta agilis L. and Lacerta vivipara Jacquin"
1672:
1608:
1530:
1483:
1371:
1289:
1226:
1179:
1097:
1031:
971:
921:
719:
398:
Sand lizards can live for over ten years, with their average lifespan lasting between 5–6 years.
99:
55:
1870:
1502:
1956:
1774:
1664:
1600:
1561:
1522:
1475:
1428:
1410:
1363:
1324:
1281:
1273:
1218:
1171:
1089:
1081:
1023:
963:
913:
862:
839:
360:
41:
1637:"Ecology and Conservation of the Sand Lizard (Lacerta agilis L.) Habitat in Southern England"
470:
in loose sand in a sunny location, leaving them to be incubated by the warmth of the ground.
1969:
1961:
1896:
1656:
1592:
1514:
1467:
1418:
1402:
1355:
1316:
1265:
1210:
1163:
1073:
1015:
955:
905:
854:
773:
606:
326:
1198:
347:
Sand lizards may live up to ten years. Due to their longevity, they are sometimes prone to
1930:
1501:
Wieczorek, Magdalena; Rektor, Robert; Najbar, Bartłomiej; Morelli, Federico (2020-05-26).
813:
387:
Sand lizards can self-amputate their tails as a defence mechanism. This ability is called
176:
1307:
Olsson M, Shine R, Madsen T, Gullberg A, Tegelström H (1997). "Sperm choice by females".
1652:
1581:"Contest success in relation to size and residency in male sand lizards, Lacerta agilis"
1463:
1261:
1167:
1069:
1826:
1813:
1756:
1503:"Tick parasitism is associated with home range area in the sand lizard, Lacerta agilis"
1423:
1390:
959:
517:
330:
302:
1580:
1320:
755:
2004:
1891:
1596:
1534:
764:
215:
89:
84:
1612:
1487:
1375:
1293:
1230:
1183:
1101:
1035:
975:
925:
1779:
669:
537:
894:"Rival recognition affects male contest behavior in sand lizards (Lacerta agilis)"
589:
and opportunities to choose a mate depending on their individual characteristics.
1448:"Some specific features of tail regeneration in the sand lizard (Lacerta agilis)"
1909:
1852:
1736:
693:
442:
306:
228:
1518:
1406:
858:
525:
348:
280:
273:
166:
1727:
1668:
1604:
1565:
1526:
1479:
1414:
1367:
1277:
1222:
1175:
1085:
1027:
967:
917:
344:
Females neighbour each other more amicably, occasionally sharing habitats.
116:
17:
1432:
1359:
1328:
1285:
1093:
616:
The sand lizard is facing multiple threats throughout its range, including
1805:
1721:
545:
388:
337:
156:
136:
1982:
1935:
1844:
1766:
1676:
1636:
1471:
1269:
1077:
1019:
909:
549:
146:
1857:
1447:
1245:
1053:
1003:
893:
533:
375:), the dorsal stripe is thin and interrupted, or not present at all.
314:
310:
298:
276:
126:
1698:
1660:
1550:"Parasites of the sand lizard [Lacerta agilis L.] in Poland"
1214:
541:
407:
318:
250:
1948:
573:
this time period, they will continue their search for new mates.
524:
Sand lizards are preyed upon by a variety of predators including
838:
Arnold, E. Nicholas; Arribas, Oscar; Carranza, Salvador (2007).
384:), males can be wholly green, even outside the breeding season.
256:
1792:
1702:
1342:
Olsson, Mats; Gullberg, Annica; Tegelströ, Håkan (1994-07-01).
1052:
House, S. M.; Taylor, P. J.; Spellerberg, I. F. (1979-01-01).
553:
529:
467:
322:
544:. In addition to wild predators, domestic species, such as
726:(UK protected land), a UK habitat where sand lizards exist
754:
Aghasyan, A.; Avci, A.; Tuniyev, B.; et al. (2021).
628:
so sand lizards can continue to inhabit those regions.
1344:"Sperm competition in the sand lizard, Lacerta agilis"
1711:
321:, where the ground temperature is elevated by the
942:Olsson, M.; Gullberg, A.; Tegelström, H. (1996).
853:. Auckland, New Zealand: Magnolia Press: 1–86.
779:10.2305/IUCN.UK.2021-2.RLTS.T157288A49644624.en
609:name translates as the "common lizard". The UK
297:The sand lizard is distributed across most of
1635:House, Susan M.; Spellerberg, Ian F. (1983).
8:
1244:Olsson, Mats; Shine, Richard (1996-01-01).
1699:
227:
78:
54:
40:
31:
1422:
777:
611:Amphibian and Reptile Conservation Trust
511:
746:
635:
1197:Olsson, M.; Shine, R. (January 1997).
7:
1988:E58D3BE4-3162-4D60-9387-36907AA5DD6A
1832:caeb7132-93b4-47e5-a542-83f46c003f06
1047:
1045:
997:
995:
993:
991:
989:
987:
985:
937:
935:
887:
885:
657:Sand lizard, mating season colouring
367:In the two main western subspecies (
325:. The sand lizard uses warm sand to
2011:IUCN Red List least concern species
1168:10.1046/j.1420-9101.1997.10030369.x
1008:Behavioral Ecology and Sociobiology
898:Behavioral Ecology and Sociobiology
801:The British Amphibians and Reptiles
765:IUCN Red List of Threatened Species
724:Site of Special Scientific Interest
1694:List of European Protected Species
960:10.1046/j.1420-9101.1996.9020229.x
25:
1446:Khabibullin, V. F. (2000-01-01).
730:List of reptiles of Great Britain
450:they are outside of this period.
701:
692:
677:
662:
650:
638:
313:. It does not occur in European
103:
1156:Journal of Evolutionary Biology
948:Journal of Evolutionary Biology
814:"Lacerta Agilis LINNAEUS, 1758"
1150:Olsson, M.; Shine, R. (1997).
1:
1321:10.1016/s0169-5347(97)85751-5
686:Põhja-Kõrvemaa Nature Reserve
684:Sand lizard at Jussi heat in
482:Reproductive success with age
1597:10.1016/0003-3472(92)90046-C
466:The female sand lizard lays
305:and across the continent to
2036:Taxa named by Carl Linnaeus
1579:Olsson, Mats (1992-08-01).
1002:Olsson, Mats (1993-05-01).
892:Olsson, Mats (1994-10-01).
301:from the southern coast of
2067:
2051:Habitats Directive species
2031:Reptiles described in 1758
1641:Journal of Applied Ecology
1452:Russian Journal of Ecology
601:). This is in contrast to
599:European Protected Species
520:with a caught sand lizard
1519:10.1163/15685381-bja10018
1407:10.1007/s10211-013-0148-2
859:10.11646/zootaxa.1430.1.1
735:List of reptiles of Italy
708:Male and female in Poland
508:Predators and competitors
235:
226:
205:
198:
100:Scientific classification
98:
76:
67:
62:
53:
48:
39:
34:
402:Habitat and distribution
1203:The American Naturalist
620:, habitat degradation,
1360:10.1006/anbe.1994.1226
803:. Collins, London, UK.
772:: e.T157288A49644624.
521:
259:
255:Puscha-Vodytsia Park,
244: Extant, resident
622:habitat fragmentation
568:Competitive behaviour
515:
498:inbreeding depression
359:The sand lizard is a
254:
63:A female sand lizard
1827:Fauna Europaea (new)
491:Inbreeding avoidance
279:. There are several
1653:1983JApEc..20..417H
1554:Acta Parasitologica
1464:2000RuJEc..31...66K
1262:1996Oecol.105..175O
1070:1979Oecol..44..396H
618:habitat destruction
593:Conservation status
441:In general, female
70:Conservation status
49:A male sand lizard
2046:Reptiles of Russia
1548:Lewin, J. (1992).
1472:10.1007/BF02799730
1270:10.1007/BF00328543
1078:10.1007/BF00545244
1020:10.1007/BF00183789
910:10.1007/BF00170705
799:M., Smith (1969).
720:Hesketh Golf Links
522:
361:sexually dimorphic
260:
2021:Lizards of Europe
1998:
1997:
1957:Open Tree of Life
1705:Taxon identifiers
1507:Amphibia-Reptilia
1309:Trends Ecol. Evol
868:978-1-86977-097-6
249:
248:
93:
27:Species of lizard
16:(Redirected from
2058:
1991:
1990:
1978:
1977:
1965:
1964:
1952:
1951:
1939:
1938:
1926:
1925:
1913:
1912:
1910:NBNSYS0000005070
1900:
1899:
1887:
1886:
1874:
1873:
1861:
1860:
1848:
1847:
1835:
1834:
1822:
1821:
1809:
1808:
1796:
1795:
1783:
1782:
1770:
1769:
1760:
1759:
1747:
1746:
1745:
1732:
1731:
1730:
1700:
1681:
1680:
1632:
1626:
1623:
1617:
1616:
1585:Animal Behaviour
1576:
1570:
1569:
1545:
1539:
1538:
1498:
1492:
1491:
1443:
1437:
1436:
1426:
1386:
1380:
1379:
1348:Animal Behaviour
1339:
1333:
1332:
1304:
1298:
1297:
1241:
1235:
1234:
1194:
1188:
1187:
1147:
1141:
1130:
1124:
1121:
1115:
1112:
1106:
1105:
1049:
1040:
1039:
999:
980:
979:
939:
930:
929:
889:
880:
879:
877:
875:
844:
835:
829:
828:
826:
824:
810:
804:
797:
791:
790:
788:
786:
781:
751:
705:
696:
681:
666:
654:
642:
423:General activity
243:
231:
211:
108:
107:
87:
82:
81:
58:
44:
32:
21:
2066:
2065:
2061:
2060:
2059:
2057:
2056:
2055:
2041:Fauna of Poland
2026:Lizards of Asia
2016:Lacerta (genus)
2001:
2000:
1999:
1994:
1986:
1981:
1973:
1968:
1960:
1955:
1947:
1942:
1934:
1931:Observation.org
1929:
1921:
1916:
1908:
1903:
1895:
1890:
1882:
1877:
1869:
1864:
1856:
1851:
1843:
1838:
1830:
1825:
1817:
1812:
1804:
1799:
1791:
1786:
1778:
1773:
1765:
1763:
1755:
1750:
1741:
1740:
1735:
1726:
1725:
1720:
1707:
1690:
1685:
1684:
1661:10.2307/2403517
1634:
1633:
1629:
1624:
1620:
1578:
1577:
1573:
1547:
1546:
1542:
1500:
1499:
1495:
1445:
1444:
1440:
1395:Acta Ethologica
1388:
1387:
1383:
1341:
1340:
1336:
1306:
1305:
1301:
1243:
1242:
1238:
1196:
1195:
1191:
1149:
1148:
1144:
1131:
1127:
1122:
1118:
1113:
1109:
1051:
1050:
1043:
1001:
1000:
983:
941:
940:
933:
891:
890:
883:
873:
871:
869:
842:
837:
836:
832:
822:
820:
812:
811:
807:
798:
794:
784:
782:
753:
752:
748:
743:
716:
709:
706:
697:
688:
682:
673:
667:
658:
655:
646:
643:
634:
595:
570:
510:
493:
484:
460:
439:
425:
420:
404:
357:
245:
241:
222:
213:
207:
194:
102:
94:
83:
79:
72:
28:
23:
22:
15:
12:
11:
5:
2064:
2062:
2054:
2053:
2048:
2043:
2038:
2033:
2028:
2023:
2018:
2013:
2003:
2002:
1996:
1995:
1993:
1992:
1979:
1966:
1953:
1940:
1927:
1914:
1901:
1888:
1875:
1862:
1849:
1836:
1823:
1814:Fauna Europaea
1810:
1797:
1784:
1771:
1761:
1757:Lacerta_agilis
1748:
1743:Lacerta agilis
1733:
1717:
1715:
1713:Lacerta agilis
1709:
1708:
1703:
1697:
1696:
1689:
1688:External links
1686:
1683:
1682:
1647:(2): 417–437.
1627:
1618:
1571:
1540:
1513:(4): 479–488.
1493:
1438:
1401:(3): 173–179.
1381:
1354:(1): 193–200.
1334:
1299:
1256:(2): 175–178.
1236:
1215:10.1086/285985
1209:(1): 179–188.
1189:
1162:(3): 369–381.
1142:
1125:
1116:
1107:
1064:(3): 396–402.
1041:
1014:(5): 337–341.
981:
954:(2): 229–242.
931:
904:(4): 249–252.
881:
867:
830:
818:www.lacerta.de
805:
792:
758:Lacerta agilis
745:
744:
742:
739:
738:
737:
732:
727:
715:
712:
711:
710:
707:
700:
698:
691:
689:
683:
676:
674:
668:
661:
659:
656:
649:
647:
644:
637:
633:
630:
594:
591:
569:
566:
518:common kestrel
509:
506:
492:
489:
483:
480:
459:
456:
438:
437:Mate selection
435:
424:
421:
419:
416:
403:
400:
356:
353:
329:itself and to
327:thermoregulate
269:Lacerta agilis
247:
246:
240:
233:
232:
224:
223:
214:
209:Lacerta agilis
203:
202:
196:
195:
191:L. agilis
188:
186:
182:
181:
174:
170:
169:
164:
160:
159:
154:
150:
149:
144:
140:
139:
134:
130:
129:
124:
120:
119:
114:
110:
109:
96:
95:
77:
74:
73:
68:
65:
64:
60:
59:
51:
50:
46:
45:
37:
36:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
2063:
2052:
2049:
2047:
2044:
2042:
2039:
2037:
2034:
2032:
2029:
2027:
2024:
2022:
2019:
2017:
2014:
2012:
2009:
2008:
2006:
1989:
1984:
1980:
1976:
1971:
1967:
1963:
1958:
1954:
1950:
1945:
1941:
1937:
1932:
1928:
1924:
1919:
1915:
1911:
1906:
1902:
1898:
1893:
1889:
1885:
1880:
1876:
1872:
1867:
1863:
1859:
1854:
1850:
1846:
1841:
1837:
1833:
1828:
1824:
1820:
1815:
1811:
1807:
1802:
1798:
1794:
1789:
1785:
1781:
1776:
1772:
1768:
1762:
1758:
1753:
1749:
1744:
1738:
1734:
1729:
1723:
1719:
1718:
1716:
1714:
1710:
1706:
1701:
1695:
1692:
1691:
1687:
1678:
1674:
1670:
1666:
1662:
1658:
1654:
1650:
1646:
1642:
1638:
1631:
1628:
1622:
1619:
1614:
1610:
1606:
1602:
1598:
1594:
1590:
1586:
1582:
1575:
1572:
1567:
1563:
1559:
1555:
1551:
1544:
1541:
1536:
1532:
1528:
1524:
1520:
1516:
1512:
1508:
1504:
1497:
1494:
1489:
1485:
1481:
1477:
1473:
1469:
1465:
1461:
1457:
1453:
1449:
1442:
1439:
1434:
1430:
1425:
1420:
1416:
1412:
1408:
1404:
1400:
1396:
1392:
1385:
1382:
1377:
1373:
1369:
1365:
1361:
1357:
1353:
1349:
1345:
1338:
1335:
1330:
1326:
1322:
1318:
1315:(11): 445–6.
1314:
1310:
1303:
1300:
1295:
1291:
1287:
1283:
1279:
1275:
1271:
1267:
1263:
1259:
1255:
1251:
1247:
1240:
1237:
1232:
1228:
1224:
1220:
1216:
1212:
1208:
1204:
1200:
1193:
1190:
1185:
1181:
1177:
1173:
1169:
1165:
1161:
1157:
1153:
1146:
1143:
1139:
1135:
1129:
1126:
1120:
1117:
1111:
1108:
1103:
1099:
1095:
1091:
1087:
1083:
1079:
1075:
1071:
1067:
1063:
1059:
1055:
1048:
1046:
1042:
1037:
1033:
1029:
1025:
1021:
1017:
1013:
1009:
1005:
998:
996:
994:
992:
990:
988:
986:
982:
977:
973:
969:
965:
961:
957:
953:
949:
945:
938:
936:
932:
927:
923:
919:
915:
911:
907:
903:
899:
895:
888:
886:
882:
870:
864:
860:
856:
852:
848:
841:
834:
831:
819:
815:
809:
806:
802:
796:
793:
780:
775:
771:
767:
766:
761:
759:
750:
747:
740:
736:
733:
731:
728:
725:
721:
718:
717:
713:
704:
699:
695:
690:
687:
680:
675:
671:
665:
660:
653:
648:
641:
636:
631:
629:
625:
623:
619:
614:
612:
608:
604:
600:
592:
590:
586:
582:
578:
574:
567:
565:
561:
557:
555:
551:
547:
543:
539:
538:birds of prey
535:
531:
527:
519:
514:
507:
505:
501:
499:
490:
488:
481:
479:
475:
471:
469:
464:
457:
455:
451:
447:
444:
436:
434:
431:
422:
417:
415:
411:
409:
401:
399:
396:
392:
390:
385:
383:
378:
374:
370:
365:
362:
354:
352:
350:
345:
341:
339:
334:
332:
328:
324:
320:
316:
312:
308:
304:
300:
295:
294:
293:L. a. exigua.
290:
286:
282:
278:
275:
271:
270:
265:
258:
253:
239:
234:
230:
225:
221:
217:
212:
210:
204:
201:
200:Binomial name
197:
193:
192:
187:
184:
183:
180:
179:
175:
172:
171:
168:
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
106:
101:
97:
91:
86:
85:Least Concern
75:
71:
66:
61:
57:
52:
47:
43:
38:
33:
30:
19:
1712:
1644:
1640:
1630:
1621:
1588:
1584:
1574:
1557:
1553:
1543:
1510:
1506:
1496:
1458:(1): 66–68.
1455:
1451:
1441:
1398:
1394:
1384:
1351:
1347:
1337:
1312:
1308:
1302:
1253:
1249:
1239:
1206:
1202:
1192:
1159:
1155:
1145:
1137:
1134:Mertensiella
1133:
1128:
1119:
1110:
1061:
1057:
1011:
1007:
951:
947:
901:
897:
872:. Retrieved
850:
846:
833:
821:. Retrieved
817:
808:
800:
795:
783:. Retrieved
769:
763:
757:
749:
670:Penza Oblast
626:
615:
603:L. a. exigua
602:
596:
587:
583:
579:
575:
571:
562:
558:
523:
502:
494:
485:
476:
472:
465:
461:
458:Reproduction
452:
448:
440:
426:
412:
405:
397:
393:
386:
382:L. a. exigua
381:
376:
372:
369:L. a. agilis
368:
366:
358:
346:
342:
335:
296:
292:
288:
285:L. a. agilis
284:
283:, including
268:
267:
263:
261:
237:
208:
206:
190:
189:
177:
35:Sand lizard
29:
18:Sand lizards
1853:iNaturalist
1737:Wikispecies
1591:: 386–388.
785:12 November
645:Sand lizard
443:mate choice
377:L. a. argus
373:L. a. argus
355:Description
307:Lake Baikal
289:L. a. argus
264:sand lizard
2005:Categories
741:References
349:inbreeding
333:its eggs.
319:heathlands
281:subspecies
167:Lacertidae
1669:0021-8901
1605:0003-3472
1566:1230-2821
1535:219744018
1527:1568-5381
1480:1608-3334
1415:0873-9749
1368:0003-3472
1278:1432-1939
1250:Oecologia
1223:0003-0147
1176:1420-9101
1086:1432-1939
1058:Oecologia
1028:1432-0762
968:1420-9101
918:1432-0762
823:30 August
546:pheasants
526:mustelids
516:A female
408:heathland
338:squamates
238:L. agilis
236:Range of
185:Species:
123:Kingdom:
117:Eukaryota
1871:10365851
1764:BioLib:
1722:Wikidata
1613:53146160
1488:44869955
1433:24052686
1376:53190531
1329:21238151
1294:13366940
1286:28307079
1231:84986874
1184:84691498
1140:: 86–91.
1102:22058312
1094:28310296
1036:37769992
976:84455614
926:24228867
714:See also
672:, Russia
605:, whose
550:chickens
418:Behavior
389:autotomy
331:incubate
274:lacertid
216:Linnaeus
163:Family:
157:Squamata
147:Reptilia
137:Chordata
133:Phylum:
127:Animalia
113:Domain:
90:IUCN 3.1
1983:ZooBank
1949:1506598
1884:1155305
1845:6159223
1728:Q148595
1677:2403517
1649:Bibcode
1460:Bibcode
1424:3775096
1258:Bibcode
1066:Bibcode
874:12 July
847:Zootaxa
632:Gallery
607:Russian
534:badgers
303:Britain
272:) is a
178:Lacerta
173:Genus:
153:Order:
143:Class:
88: (
1975:agilis
1962:862848
1897:157288
1819:214645
1793:791768
1675:
1667:
1611:
1603:
1564:
1533:
1525:
1486:
1478:
1431:
1421:
1413:
1374:
1366:
1327:
1292:
1284:
1276:
1229:
1221:
1182:
1174:
1100:
1092:
1084:
1034:
1026:
974:
966:
924:
916:
865:
552:, and
542:snakes
540:, and
315:Turkey
311:Russia
299:Europe
291:, and
277:lizard
242:
1923:80427
1866:IRMNG
1858:35912
1801:EUNIS
1780:6NRCH
1673:JSTOR
1609:S2CID
1560:(1).
1531:S2CID
1484:S2CID
1372:S2CID
1290:S2CID
1227:S2CID
1180:S2CID
1098:S2CID
1032:S2CID
972:S2CID
922:S2CID
843:(PDF)
530:foxes
1944:OBIS
1918:NCBI
1892:IUCN
1879:ITIS
1840:GBIF
1665:ISSN
1601:ISSN
1562:ISSN
1523:ISSN
1476:ISSN
1429:PMID
1411:ISSN
1364:ISSN
1325:PMID
1282:PMID
1274:ISSN
1219:ISSN
1172:ISSN
1090:PMID
1082:ISSN
1024:ISSN
964:ISSN
914:ISSN
876:2017
863:ISBN
851:1430
825:2021
787:2021
770:2021
722:, a
554:cats
468:eggs
430:bask
371:and
262:The
257:Kyiv
220:1758
1936:459
1905:NBN
1806:713
1788:EoL
1775:CoL
1767:356
1752:ADW
1657:doi
1593:doi
1515:doi
1468:doi
1419:PMC
1403:doi
1356:doi
1317:doi
1266:doi
1254:105
1211:doi
1207:149
1164:doi
1074:doi
1016:doi
956:doi
906:doi
855:doi
774:doi
323:sun
309:in
2007::
1985::
1972::
1970:RD
1959::
1946::
1933::
1920::
1907::
1894::
1881::
1868::
1855::
1842::
1829::
1816::
1803::
1790::
1777::
1754::
1739::
1724::
1671:.
1663:.
1655:.
1645:20
1643:.
1639:.
1607:.
1599:.
1589:44
1587:.
1583:.
1558:37
1556:.
1552:.
1529:.
1521:.
1511:41
1509:.
1505:.
1482:.
1474:.
1466:.
1456:31
1454:.
1450:.
1427:.
1417:.
1409:.
1399:16
1397:.
1393:.
1370:.
1362:.
1352:48
1350:.
1346:.
1323:.
1313:12
1311:.
1288:.
1280:.
1272:.
1264:.
1252:.
1248:.
1225:.
1217:.
1205:.
1201:.
1178:.
1170:.
1160:10
1158:.
1154:.
1136:.
1096:.
1088:.
1080:.
1072:.
1062:44
1060:.
1056:.
1044:^
1030:.
1022:.
1012:32
1010:.
1006:.
984:^
970:.
962:.
950:.
946:.
934:^
920:.
912:.
902:35
900:.
896:.
884:^
861:.
849:.
845:.
816:.
768:.
762:.
556:.
548:,
536:,
532:,
528:,
391:.
351:.
287:,
218:,
1679:.
1659::
1651::
1615:.
1595::
1568:.
1537:.
1517::
1490:.
1470::
1462::
1435:.
1405::
1378:.
1358::
1331:.
1319::
1296:.
1268::
1260::
1233:.
1213::
1186:.
1166::
1138:1
1104:.
1076::
1068::
1038:.
1018::
978:.
958::
952:9
928:.
908::
878:.
857::
827:.
789:.
776::
760:"
756:"
266:(
92:)
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
