138:
106:
1020:"Observations concerning the evolution and the parasystematics of all the living and fossil Chlamydoselachiformes, Squatiniformes, Orectolobiformes, and Pristiophoriformes, based on both biological and odontological data. Suggestion of a possible origin of the order Pristiophoriformes, of the order Ganopristiformes and a global synthesis of the previous systematics proposals"
987:"Observations diverses et déductions concernant l'evolution et la systématique de quelques Euselachii, Neoselachii et Batoidei (Pisces - Elasmobranchii), actuels et fossiles. Données complémentaires"
1302:
Smith, Roy E.; Martill, David M.; Longrich, N.; Unwin, David M.; Ibrahim, N.; Zouhri, S. (2023). "Comparative taphonomy of Kem Kem Group (Cretaceous) pterosaurs of southeast
Morocco".
1275:
Mulcahy, Kevin D. (2023). "Relative abundance of vertebrate fossil taxa in the Upper
Cretaceous exposures of Monmouth County brooks & a test of species richness extrapolators".
954:"Observations diverses et déductions concernant l'evolution et la systématique de quelques Euselachii, Neoselachii et Batoidei (Pisces - Elasmobranchii), actuels et fossiles"
986:
927:
Kriwet, J. (2004). "The systematic position of the
Cretaceous sclerorhynchid sawfishes (Elasmobranchii, Pristiorajea)". In Arratia, G.; Tintori, A. (eds.).
396:
1019:
953:
1410:
1051:
Greenfield, T. (2024). "Pristification: Defining the convergent evolution of saws in sharks and rays (Chondrichthyes, Neoselachii)".
936:
137:
929:
Mesozoic Fishes 3 – Systematics, Paleoenvironments and
Biodiversity. Proceedings of the International Meeting Serpiano, 2001
1001:
391:. Several genera (see below) are not currently placed in any of these families. Sclerorhynchoids first appeared in the
1530:
1339:"The famous fish beds of Lebanon: the Upper Cretaceous Lagerstätten of Haqel, Hjoula, Nammoura, and Sahel Aalma"
869:
1535:
1084:
gen. et sp. nov., and implications for the phylogenetic relations of the
Sclerorhynchoidei (Chondrichthyes)"
1455:
1034:
968:
1493:
1190:
1502:
1379:
1240:
Greenfield, T. (2021). "Corrections to the nomenclature of sawskates (Rajiformes, Sclerorhynchoidei)".
1311:
1152:
1098:
774:
353:
1426:
Capasso, L. (2003). "Embryos and trophonems in
Cenomanian rays from Lebanon (Batomorphii, Pisces)".
1403:
Handbook of
Paleoichthyology. Volume 3E. Chondrichthyes. Mesozoic and Cenozoic Elasmobranchii: Teeth
846:
908:
Case, G.R. (1978). "A new selachian fauna from the Judith River
Formation (Campanian) of Montana".
782:
1257:
1168:
1114:
830:
277:
132:
124:
1507:
790:
1189:
Villalobos-Segura, E.; Kriwet, J.; Vullo, R.; Stumpf, S.; Ward, D.J.; Underwood, C.J. (2021b).
1435:
1406:
1380:"Paleobiology and paleobiogeography of sclerorhynchid sawfishes (Chondrichthyes, Batomorphii)"
1360:
1217:
932:
862:
368:
proposed in 2021 for sclerorhynchoids, which has been subsequently used by other researchers.
337:
114:
822:
1350:
1319:
1284:
1249:
1209:
1160:
1106:
1060:
742:
734:
559:
384:
262:
105:
750:
356:, recently proposed as 'pristification', and their closest living relatives are actually
1337:
George, H.; Bazzi, M.; El Hossny, T.; Ashraf, N.; Saad, Pierre A.; Clements, T. (2024).
1315:
1156:
1133:
1102:
1079:
429:
Below is a cladogram of
Sclerorhynchoidei, with the topology based on Villalobos-Segura
726:
658:
473:
388:
268:
256:
227:
189:
179:
119:
1524:
1261:
1172:
1118:
758:
624:
610:
418:
409:
372:
244:
63:
32:
1195:(Elasmobranchii) from the 'Mid'-Cretaceous and their palaeontological implications"
798:
641:
588:
530:
380:
357:
1110:
889:
Cappetta, H. (1980). "Les Sélaciens du Crétacé supérieur du Liban. II. Batoïdes".
1478:
1213:
1487:
854:
838:
814:
766:
480:
365:
38:
1323:
1253:
512:
440:
376:
341:
330:
250:
209:
83:
48:
1439:
1364:
1221:
1064:
806:
400:
392:
149:
88:
28:
1288:
1080:"The first three-dimensional fossils of Cretaceous sclerorhynchid sawfish:
1078:
Villalobos-Segura, E.; Underwood, C.J.; Ward, D.J.; Claeson, K.M. (2019).
1472:
349:
333:
199:
169:
78:
73:
58:
53:
43:
1355:
1338:
453:
345:
93:
68:
1164:
1134:"The first skeletal record of the enigmatic Cretaceous sawfish genus
404:
159:
1449:
1184:
1182:
1453:
1132:
Villalobos-Segura, E.; Underwood, C.J.; Ward, D.J. (2021a).
433:(2021b) and the family taxonomy based on Greenfield (2021).
1191:"The skeletal remains of the euryhaline sclerorhynchoid â€
1138:(Chondrichthyes, Batoidea) from the Turonian of Morocco"
417:
represents one of the first fossil evidence of batoid
1462:
1428:
Atti del Museo Civico di Storia
Naturale di Trieste
1235:
1233:
1231:
931:. Verlag Dr. Friedrich Pfeil. pp. 57–73.
8:
371:The suborder contains five named families:
360:. While they are often called "sawfishes",
1450:
104:
20:
1354:
1202:Zoological Journal of the Linnean Society
403:occurrences being misidentifications or
16:Extinct suborder of cartilaginous fishes
881:
1405:. Munich: Verlag Dr. Friedrich Pfeil.
397:Cretaceous–Paleogene extinction event
7:
1091:Journal of Systematic Palaeontology
616:
565:
555:
503:
496:
469:
446:
436:
985:Herman, J.; Van Waes, H. (2012b).
952:Herman, J.; Van Waes, H. (2012a).
407:specimens. One female specimen of
14:
1384:Revista Española de PaleontologĂa
1343:Journal of the Geological Society
1018:Herman, J.; Van Waes, H. (2014).
1378:Kriwet, J.; Kussius, K. (2001).
136:
910:Palaeontographica, Abteilung A
891:Palaeontographica, Abteilung A
1:
1111:10.1080/14772019.2019.1578832
413:with nine embryos preserved
395:and went extinct during the
310:Herman & Van Waes, 2012b
303:Herman & Van Waes, 2012a
317:Herman & Van Waes, 2014
1552:
1214:10.1093/zoolinnean/zlaa166
1033:(1): 1–347. Archived from
329:is an extinct suborder of
1324:10.1016/j.eve.2023.100006
1254:10.11646/bionomina.22.1.3
1082:Asflapristis cristadentis
1000:(2): 1–12. Archived from
967:(1): 1–89. Archived from
655:
638:
621:
614:
585:
570:
563:
553:
527:
509:
501:
494:
477:
467:
451:
444:
283:
276:
240:
235:
133:Scientific classification
131:
112:
103:
23:
870:Sao Khua sclerorhynchoid
1145:Papers in Palaeontology
1065:10.11646/MESOZOIC.1.2.3
1289:10.5281/zenodo.7905373
1503:Paleobiology Database
1401:Cappetta, H. (2012).
113:Comparison between a
354:convergently evolved
307:Ptychotrygoniformes
293:Sclerorhynchiformes
1356:10.1144/jgs2023-210
1316:2023EvEar...100006S
1157:2021PPal....7..353V
1103:2019JSPal..17.1847V
364:is a more accurate
352:. This feature was
127:(bottom and right)
465:Sclerorhynchoidei
314:Ganopristomorphii
125:largetooth sawfish
1531:Sclerorhynchoidei
1518:
1517:
1494:Sclerorhynchoidei
1464:Sclerorhynchoidei
1456:Taxon identifiers
1412:978-3-89937-148-2
1165:10.1002/spp2.1287
1097:(21): 1847–1870.
1027:GĂ©ominpal Belgica
994:GĂ©ominpal Belgica
961:GĂ©ominpal Belgica
715:
714:
706:
705:
697:
696:
688:
687:
679:
678:
670:
669:
600:
599:
542:
541:
327:Sclerorhynchoidei
324:
323:
318:
311:
304:
300:Ganopristiformes
297:
290:
231:
223:Sclerorhynchoidei
115:longnose sawshark
24:Sclerorhynchoidei
1543:
1511:
1510:
1498:
1497:
1496:
1483:
1482:
1481:
1451:
1444:
1443:
1423:
1417:
1416:
1398:
1392:
1391:
1375:
1369:
1368:
1358:
1334:
1328:
1327:
1299:
1293:
1292:
1272:
1266:
1265:
1237:
1226:
1225:
1199:
1186:
1177:
1176:
1142:
1129:
1123:
1122:
1088:
1075:
1069:
1068:
1048:
1042:
1041:
1039:
1024:
1015:
1009:
1008:
1006:
991:
982:
976:
975:
973:
958:
949:
943:
942:
924:
918:
917:
905:
899:
898:
886:
743:Atlanticopristis
735:Ankistrorhynchus
617:
566:
560:Ptychotrygonidae
556:
504:
497:
470:
447:
437:
385:Ptychotrygonidae
316:
309:
302:
295:
288:
286:Ganopristinidea
263:Ptychotrygonidae
226:
221:
141:
140:
123:(middle), and a
117:(top and left),
108:
98:
27:Temporal range:
21:
1551:
1550:
1546:
1545:
1544:
1542:
1541:
1540:
1521:
1520:
1519:
1514:
1506:
1501:
1492:
1491:
1486:
1477:
1476:
1471:
1458:
1448:
1447:
1425:
1424:
1420:
1413:
1400:
1399:
1395:
1377:
1376:
1372:
1336:
1335:
1331:
1301:
1300:
1296:
1274:
1273:
1269:
1239:
1238:
1229:
1197:
1188:
1187:
1180:
1140:
1131:
1130:
1126:
1086:
1077:
1076:
1072:
1050:
1049:
1045:
1037:
1022:
1017:
1016:
1012:
1004:
989:
984:
983:
979:
971:
956:
951:
950:
946:
939:
926:
925:
921:
916:(1–6): 176–205.
907:
906:
902:
897:(5–6): 149–229.
888:
887:
883:
878:
775:Borodinopristis
751:Australopristis
721:
716:
707:
698:
689:
680:
671:
601:
543:
525:Onchopristidae
427:
315:
308:
301:
294:
287:
225:
219:
135:
99:
97:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
35:
25:
17:
12:
11:
5:
1549:
1547:
1539:
1538:
1536:Fish suborders
1533:
1523:
1522:
1516:
1515:
1513:
1512:
1499:
1484:
1468:
1466:
1460:
1459:
1454:
1446:
1445:
1418:
1411:
1393:
1370:
1329:
1304:Evolving Earth
1294:
1267:
1227:
1208:(2): 746–771.
1178:
1151:(1): 353–376.
1124:
1070:
1059:(2): 121–124.
1043:
1040:on 2015-02-17.
1010:
1007:on 2015-02-17.
977:
974:on 2015-02-17.
944:
937:
919:
900:
880:
879:
877:
874:
873:
872:
866:
858:
850:
847:Plicatopristis
842:
834:
826:
818:
810:
802:
794:
786:
778:
770:
762:
754:
746:
738:
730:
727:Agaleorhynchus
720:
717:
713:
712:
709:
708:
704:
703:
700:
699:
695:
694:
691:
690:
686:
685:
682:
681:
677:
676:
673:
672:
668:
667:
664:
663:
659:Sclerorhynchus
654:
651:
650:
647:
646:
637:
634:
633:
630:
629:
620:
615:
613:
607:
606:
603:
602:
598:
597:
594:
593:
584:
581:
580:
577:
576:
569:
564:
562:
554:
552:
549:
548:
545:
544:
540:
539:
536:
535:
526:
522:
521:
518:
517:
508:
507:Ischyrhizidae
502:
500:
495:
493:
490:
489:
486:
485:
476:
474:Schizorhizidae
468:
466:
462:
461:
458:
457:
450:
445:
443:
435:
426:
423:
399:, with former
389:Schizorhizidae
381:Onchopristidae
336:that had long
322:
321:
320:
319:
312:
305:
298:
291:
281:
280:
274:
273:
272:
271:
269:Schizorhizidae
265:
259:
257:Onchopristidae
253:
247:
238:
237:
233:
232:
217:
213:
212:
207:
203:
202:
197:
193:
192:
190:Elasmobranchii
187:
183:
182:
180:Chondrichthyes
177:
173:
172:
167:
163:
162:
157:
153:
152:
147:
143:
142:
129:
128:
120:Sclerorhynchus
110:
109:
101:
100:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
36:
26:
15:
13:
10:
9:
6:
4:
3:
2:
1548:
1537:
1534:
1532:
1529:
1528:
1526:
1509:
1504:
1500:
1495:
1489:
1485:
1480:
1474:
1470:
1469:
1467:
1465:
1461:
1457:
1452:
1441:
1437:
1433:
1429:
1422:
1419:
1414:
1408:
1404:
1397:
1394:
1389:
1385:
1381:
1374:
1371:
1366:
1362:
1357:
1352:
1348:
1344:
1340:
1333:
1330:
1325:
1321:
1317:
1313:
1309:
1305:
1298:
1295:
1290:
1286:
1282:
1278:
1271:
1268:
1263:
1259:
1255:
1251:
1247:
1243:
1236:
1234:
1232:
1228:
1223:
1219:
1215:
1211:
1207:
1203:
1196:
1194:
1185:
1183:
1179:
1174:
1170:
1166:
1162:
1158:
1154:
1150:
1146:
1139:
1137:
1128:
1125:
1120:
1116:
1112:
1108:
1104:
1100:
1096:
1092:
1085:
1083:
1074:
1071:
1066:
1062:
1058:
1054:
1047:
1044:
1036:
1032:
1028:
1021:
1014:
1011:
1003:
999:
995:
988:
981:
978:
970:
966:
962:
955:
948:
945:
940:
938:3-89937-053-8
934:
930:
923:
920:
915:
911:
904:
901:
896:
892:
885:
882:
875:
871:
867:
865:
864:
859:
857:
856:
851:
849:
848:
843:
841:
840:
835:
833:
832:
827:
825:
824:
819:
817:
816:
811:
809:
808:
803:
801:
800:
795:
793:
792:
787:
785:
784:
779:
777:
776:
771:
769:
768:
763:
761:
760:
759:Baharipristis
755:
753:
752:
747:
745:
744:
739:
737:
736:
731:
729:
728:
723:
722:
718:
711:
710:
702:
701:
693:
692:
684:
683:
675:
674:
666:
665:
662:
661:
660:
653:
652:
649:
648:
645:
644:
643:
636:
635:
632:
631:
628:
627:
626:
625:Libanopristis
619:
618:
612:
611:Ganopristidae
609:
608:
605:
604:
596:
595:
592:
591:
590:
583:
582:
579:
578:
575:
574:
568:
567:
561:
558:
557:
551:
550:
547:
546:
538:
537:
534:
533:
532:
524:
523:
520:
519:
516:
515:
514:
506:
505:
499:
498:
492:
491:
488:
487:
484:
483:
482:
475:
472:
471:
464:
463:
460:
459:
456:
455:
449:
448:
442:
439:
438:
434:
432:
424:
422:
420:
419:ovoviviparity
416:
412:
411:
410:Libanopristis
406:
402:
398:
394:
390:
386:
382:
378:
377:Ischyrhizidae
374:
373:Ganopristidae
369:
367:
363:
359:
355:
351:
347:
343:
339:
335:
332:
328:
313:
306:
299:
292:
285:
284:
282:
279:
275:
270:
266:
264:
260:
258:
254:
252:
251:Ischyrhizidae
248:
246:
245:Ganopristidae
242:
241:
239:
234:
229:
224:
218:
215:
214:
211:
208:
205:
204:
201:
198:
195:
194:
191:
188:
185:
184:
181:
178:
175:
174:
171:
168:
165:
164:
161:
158:
155:
154:
151:
148:
145:
144:
139:
134:
130:
126:
122:
121:
116:
111:
107:
102:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
34:
33:Maastrichtian
30:
22:
19:
1463:
1431:
1427:
1421:
1402:
1396:
1387:
1383:
1373:
1346:
1342:
1332:
1307:
1303:
1297:
1280:
1277:The Mosasaur
1276:
1270:
1248:(1): 39–41.
1245:
1241:
1205:
1201:
1193:Onchopristis
1192:
1148:
1144:
1136:Ptychotrygon
1135:
1127:
1094:
1090:
1081:
1073:
1056:
1052:
1046:
1035:the original
1030:
1026:
1013:
1002:the original
997:
993:
980:
969:the original
964:
960:
947:
928:
922:
913:
909:
903:
894:
890:
884:
861:
853:
845:
837:
829:
821:
813:
805:
799:Ctenopristis
797:
789:
783:Celtipristis
781:
773:
765:
757:
749:
741:
733:
725:
719:Other genera
657:
656:
642:Micropristis
640:
639:
623:
622:
589:Ptychotrygon
587:
586:
573:Asflapristis
572:
571:
531:Onchopristis
529:
528:
511:
510:
479:
478:
452:
430:
428:
414:
408:
370:
361:
326:
325:
296:Kriwet, 2004
222:
196:Superorder:
118:
18:
1488:Wikispecies
1390:(3): 35–46.
855:Pucapristis
839:Onchosaurus
831:Marckgrafia
815:Iberotrygon
767:Biropristis
481:Schizorhiza
366:common name
344:similar to
340:with large
1525:Categories
1310:. 100006.
876:References
791:Columbusia
513:Ischyrhiza
441:Rajiformes
289:Case, 1978
216:Suborder:
210:Rajiformes
186:Subclass:
1479:Q25095431
1440:0365-1576
1434:: 23–39.
1365:0016-7649
1262:239067365
1242:Bionomina
1222:0024-4082
1173:210302939
1119:145940997
807:Dalpiazia
425:Phylogeny
401:Paleocene
393:Barremian
362:sawskates
350:sawsharks
346:sawfishes
342:denticles
236:Families
156:Kingdom:
150:Eukaryota
29:Barremian
1473:Wikidata
1283:: 1–14.
1053:Mesozoic
863:Renpetia
405:reworked
331:rajiform
278:Synonyms
228:Cappetta
200:Batoidea
170:Chordata
166:Phylum:
160:Animalia
146:Domain:
1312:Bibcode
1153:Bibcode
1099:Bibcode
823:Kiestus
454:Rajidae
415:in situ
206:Order:
176:Class:
1508:131691
1438:
1409:
1363:
1260:
1220:
1171:
1117:
935:
431:et al.
387:, and
358:skates
338:rostra
230:, 1980
1349:(5).
1258:S2CID
1198:(PDF)
1169:S2CID
1141:(PDF)
1115:S2CID
1087:(PDF)
1038:(PDF)
1023:(PDF)
1005:(PDF)
990:(PDF)
972:(PDF)
957:(PDF)
1436:ISSN
1407:ISBN
1361:ISSN
1218:ISSN
933:ISBN
348:and
334:rays
39:Preęž’
1351:doi
1347:181
1320:doi
1285:doi
1250:doi
1210:doi
1206:193
1161:doi
1107:doi
1061:doi
914:160
895:168
1527::
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220:â€
94:N
84:K
79:J
74:T
69:P
64:C
59:D
54:S
49:O
44:ęž’
31:-
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