49:
553:
404:
1047:
500:. Coming in second to Jonathan is Esmeralda, at 180 years old. Esmeralda is an Aldabra giant tortoise living in the Galapagos Islands. There is a report that a tortoise was kept in the garrison by French explorer Chevalier Marc-Joseph Marion du Fresne. The creature lived in the fort for 118 years. It died in 1918 when it became blind and accidentally fell to its death from atop a gun turret.
31:
544:
papers on the genetics of the
Seychelles and Indian Ocean tortoises provide conflicting results. Some studies suggest only one species (with multiple variants) was ever present in the islands, whilst others suggest three distinct, but closely related species. These different views derive from studies of different genes.
512:, but is considered by many researchers to be either synonymous with or only subspecifically distinct from that taxon. This identification is based primarily on morphological characters. Published molecular identifications are unclear with several different indications provided by different data sources.
564:
testing, tortoises of the "extinct" subspecies were identified and acquired by the Nature
Protection Trust of Seychelles for conservation. They were brought to Silhouette Island and the captive-breeding program was initiated in 1997. For several years the female tortoises produced infertile eggs. In
519:
It was apparently extirpated from the wild but is now known only from 37 adults, including 28 captive, and 8 on
Cousine Island, 6 of which were released in 2011 along with 40 captive-bred juveniles. Captive-reared juveniles show that there is a presumed genetic basis to the morphotype and further
581:
island. The Nature
Protection Trust of Seychelles takes consolation from producing a new generation of many young tortoises which will live for at least 100 years. In that time there may be an opportunity to establish pure populations of these tortoises. These animals live longer than short-term
576:
agreed to provide a new home for the
Seychelles tortoises. In April 2011, the Seychelles giant tortoises went to Cousine. If they had been able to release them on Silhouette island, they would have established a separate wild population, now they will be mixed with the Aldabra tortoises that are
543:
to examine the identity of the living tortoises. Examination of museum specimens of the 'extinct' Seychelles subspecies by Dr. Justin
Gerlach and Laura Canning confirmed that some living tortoises exhibited characteristics of the supposedly extinct subspecies. Some recently published scientific
515:
It is a domed grazing subspecies, differing from the
Aldabra tortoise in its broader shape and reduced ossification of the skeleton; it differs also from the other controversial giant tortoise in the Seychelles, the saddle-backed morphotype (Arnold's giant tortoise).
568:
The decision of the
Islands Development Company to evict the Nature Protection Trust of Seychelles from Silhouette island by March 2011, and their refusal to permit wild tortoises to live on the island forced them to remove and find new homes for the tortoises.
394:
It was recently rediscovered. Currently, a little over a hundred individuals exist. Many had been reestablished in the wild on forested islands such as
Silhouette, but were evicted in 2011 by the Seychelles Islands Development Company.
463:
Giant tortoises are among the longest-lived animals on the planet. Some individual
Aldabra giant tortoises are thought to be over 200 years of age, but this is difficult to verify because they tend to outlive their human observers.
446:
Like other species of tortoise, the Seychelles giant tortoise is not strictly herbivorous. There have been anecdotal reports of tortoises consuming birds, crabs, and bones. In 2021, a female tortoise on
539:
in the Seychelles, it is occasionally, most recently in 1995, suggested that some Seychelles granitic island tortoises survived in captivity. The report of oddly shaped captive tortoises prompted the
532:
inhabited the Seychelles. Large and slow, the tortoises were reportedly friendly, with little fear of man. Sailors and settlers slaughtered thousands and swiftly drove most populations to extinction.
565:
November 2002 eggs laid by a young female started to hatch. By the time the Nature Protection Trust of Seychelles was evicted, they had produced 40 juveniles of the Seychelles giant tortoise.
4357:
4298:
4324:
480:
in 1875. At his death in March 2006 at the Kolkata (formerly Calcutta) Zoo in India, Adwaita is reputed to have reached the longest ever measured
1292:
1481:
1475:
948:
540:
2052:
4367:
4276:
508:
This is a controversial subspecies possibly distinct from the Aldabra giant tortoise. The species is a morphologically distinctive
1103:
4362:
1334:
4085:
3168:
1463:
1457:
1329:
1088:
388:
4329:
4212:
Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status
2021:
turtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status
1717:
1532:
598:
492:-dwelling Seychelles giant tortoise, is thought to be the oldest living terrestrial animal on Earth since the 2006 death of
1702:
1692:
1354:
4062:
48:
4210:
2020:
757:
4221:
2744:
2128:
1842:
1707:
1359:
1096:
798:
2477:
1697:
1303:
941:
473:
1712:
1469:
1120:
3123:
2045:
1623:
1344:
1313:
1282:
1170:
1379:
1252:
772:"Release of Arnold's giant tortoises Dipsochelys arnoldi on Silhouette island, Seychelles; By Justin Gerlach"
3979:
1992:
1542:
1391:
1385:
1939:
1933:
1891:
4352:
4238:
3951:
2751:
2709:
2349:
1951:
1897:
1591:
1573:
1349:
1287:
1246:
1222:
1216:
1188:
1182:
1176:
1151:
1081:
431:
177:
154:
1927:
1597:
1579:
2695:
2197:
2008:
2002:
1963:
1945:
1909:
1796:
1784:
1753:
1585:
1499:
1432:
1339:
1319:
1234:
1112:
934:
1987:
1969:
1669:
1603:
1567:
1267:
1262:
1921:
3937:
2603:
2227:
1957:
1504:
1240:
1228:
679:
1915:
1903:
1885:
3922:
2281:
2038:
1997:
1790:
1674:
1437:
1211:
485:
1370:
383:
islands, but was hunted in vast numbers by European sailors. By around 1840 it was presumed to be
3329:
3294:
3161:
2959:
2900:
2870:
1814:
1778:
1758:
1664:
1558:
1277:
493:
416:
203:
43:
2878:
2632:
4285:
3872:
2855:
2807:
2798:
2681:
2258:
1819:
1646:
1641:
1409:
1298:
1144:
1139:
697:
552:
536:
481:
448:
194:
3664:
3052:
3027:
2988:
2936:
2667:
2212:
1865:
1735:
1427:
1307:
1063:
687:
468:
was reputedly one of four brought by British seamen from the Seychelles Islands as gifts to
535:
Though generally assumed that the Aldabra giant tortoise was the only one to have survived
254:
4112:
3760:
3489:
3457:
3094:
3003:
2779:
2365:
2110:
1979:
1847:
1537:
1527:
1522:
1323:
846:
683:
4007:
3997:
3894:
3744:
3584:
3286:
3231:
2973:
2908:
2448:
2439:
2408:
2316:
2205:
2119:
1834:
1614:
1547:
1206:
578:
573:
403:
4346:
4187:
4070:
3970:
3959:
3930:
3844:
3728:
3712:
3672:
3606:
3516:
3481:
3425:
3409:
3393:
3367:
3355:
3339:
3317:
3309:
3116:
2827:
2660:
2521:
2468:
2416:
1824:
1727:
1073:
1006:
529:
497:
411:
The three Aldabra-Seychelles giant tortoise subspecies can be distinguished based on
372:
140:
120:
716:
4290:
4122:
4047:
4023:
3540:
3417:
3401:
3377:
3278:
3223:
3147:
3063:
3037:
2951:
2862:
2841:
2765:
2730:
2540:
2506:
2373:
2289:
2235:
1857:
1769:
1448:
1162:
1012:
489:
477:
469:
771:
4261:
4311:
4270:
4100:
4055:
3901:
3886:
3808:
3800:
3792:
3768:
3648:
3632:
3616:
3532:
3524:
3473:
3449:
3441:
3433:
3301:
3253:
3176:
3101:
2966:
2922:
2848:
2834:
2786:
2643:
2561:
2499:
2492:
2485:
2400:
2342:
2334:
2251:
2243:
2190:
2180:
1806:
1656:
1198:
1131:
226:
909:
30:
4179:
4092:
4078:
4039:
3816:
3736:
3704:
3688:
3656:
3594:
3567:
3548:
3465:
3270:
3190:
3183:
3131:
3108:
3018:
2885:
2617:
2424:
2357:
2274:
2266:
2169:
2160:
1491:
1419:
1055:
1000:
994:
726:
692:
667:
509:
380:
367:
110:
867:
747:
4171:
4031:
4015:
3944:
3908:
3858:
3851:
3776:
3752:
3696:
3640:
3347:
3139:
3087:
3045:
2758:
2716:
2624:
2610:
2596:
2589:
2582:
2575:
2547:
2392:
2220:
822:
282:
268:
240:
60:
701:
645:
624:
888:
419:
animals may have distorted carapaces and so may be difficult to identify.
4255:
3986:
3915:
3835:
3824:
3720:
3385:
3262:
3154:
2996:
2981:
2943:
2892:
2818:
2723:
2702:
2688:
2674:
2530:
2457:
2326:
2305:
2297:
2079:
1876:
1745:
1514:
1401:
1032:
976:
957:
752:
721:
412:
364:
130:
80:
604:
4303:
4130:
3879:
3784:
3680:
3624:
3204:
3197:
3079:
2929:
2915:
2772:
2737:
2652:
2554:
2094:
2085:
1684:
1633:
988:
982:
465:
384:
90:
4316:
3865:
3508:
2100:
2073:
2061:
970:
100:
70:
4232:
556:
A baby and a possibility for the reestablishment of this subspecies
3072:
2139:
926:
551:
452:
427:
402:
320:
2568:
4236:
2034:
930:
430:; it is usually a brownish-grey color. In comparison, the true
561:
2030:
451:
was recorded deliberately hunting, killing, and eating a
748:"Meet Jonathan, St Helena's 182-year-old giant tortoise"
496:; a 176 year old Galapagos giant tortoise who lived at
4245:
4162:
4155:
4148:
4110:
3996:
3969:
3834:
3604:
3582:
3575:
3566:
3499:
3365:
3327:
3252:
3245:
3214:
3062:
3026:
3017:
2817:
2796:
2642:
2529:
2520:
2467:
2447:
2438:
2383:
2315:
2179:
2168:
2159:
2127:
2118:
2109:
1978:
1874:
1856:
1833:
1805:
1767:
1744:
1726:
1683:
1655:
1632:
1612:
1556:
1513:
1490:
1446:
1418:
1400:
1368:
1261:
1197:
1161:
1130:
1072:
1054:
1021:
646:"The identification of Seychelles giant tortoises"
426:) is broad, flattened on the back and with raised
817:
815:
438:) is a roundly-domed, black-colored subspecies.
391:, a subspecies which shared the same islands.
4209:Phylogenetic arrangement of turtles based on
2046:
2019:Phylogenetic arrangement of turtles based on
942:
8:
793:
791:
528:Originally, several different subspecies of
484:of 255 years (birth year 1750). As of 2024,
666:Zora, Anna; Gerlach, Justin (August 2021).
4233:
4159:
4152:
3579:
3572:
3249:
3023:
2526:
2444:
2176:
2165:
2133:
2124:
2115:
2053:
2039:
2031:
1025:
949:
935:
927:
520:genetic work is needed to elucidate this.
294:DUMÉRIL & BIBRON, 1835 — BOUR 1994: 85
29:
20:
691:
619:
617:
615:
613:
582:management and development perspectives.
668:"Giant tortoises hunt and consume birds"
379:It inhabited the large central granitic
841:
839:
591:
1482:Saddle-backed Rodrigues giant tortoise
1476:Saddle-backed Mauritius giant tortoise
541:Nature Protection Trust of Seychelles
7:
715:BBC News - South Asia (2006-03-23).
717:"'Clive of India's' tortoise dies"
14:
476:in the 18th century, and came to
238:DUMÉRIL & BIBRON, 1835: 221 (
4277:Aldabrachelys gigantea hololissa
4247:Aldabrachelys gigantea hololissa
1045:
760:from the original on 2014-04-09.
572:The privately managed island of
356:Aldabrachelys gigantea hololissa
337:Aldabrachelys gigantea hololissa
329:Dipsochelys dussumieri hololissa
213:Aldabrachelys gigantea hololissa
187:Aldabrachelys gigantea hololissa
47:
746:Kettle, Sally (13 March 2014).
422:The Seychelles giant tortoise (
361:Seychelles domed giant tortoise
1464:Domed Rodrigues giant tortoise
1458:Domed Mauritius giant tortoise
1330:Santiago Island giant tortoise
847:"Aldabrachelys hololissa, 061"
1:
1283:Chatham Island giant tortoise
605:Reptarium.cz Reptile Database
4358:Taxa named by Albert Günther
1350:Volcán Darwin giant tortoise
1345:Volcán Alcedo giant tortoise
1335:Pinzón Island giant tortoise
280:DUMÉRIL & BIBRON, 1835 (
4222:List of Testudines families
1843:Flat-backed spider tortoise
1718:Speke's hinge-back tortoise
1708:Lobatse hinge-back tortoise
1533:Goode's thornscrub tortoise
1360:Western Santa Cruz tortoise
1340:Sierra Negra giant tortoise
1288:Eastern Santa Cruz tortoise
4384:
4368:Reptiles described in 1877
1703:Home's hinge-back tortoise
1698:Forest hinge-back tortoise
1693:Bell's hinge-back tortoise
1355:Volcán Wolf giant tortoise
1304:Hood Island giant tortoise
1293:Fernandina Island tortoise
1043:
524:Extinction and rediscovery
474:British East India Company
24:Seychelles giant tortoise
4205:
2136:
2068:
2017:
1713:Natal hinge-back tortoise
1278:Cerro Azul giant tortoise
1121:Aldabrachelys grandidieri
1028:
965:
823:"Tortoise reintroduction"
693:10.1016/j.cub.2021.06.088
351:Seychelles giant tortoise
209:
202:
183:
176:
44:Scientific classification
42:
37:
28:
23:
1624:Hesperotestudo percrassa
1380:Cheirogaster bacharidisi
1314:Santa Fe Island tortoise
1299:Floreana Island tortoise
1253:Chelonoidis sombrerensis
1171:African spurred tortoise
625:"Moving giant tortoises"
607:. Accessed 05 June 2015.
1543:Sonoran Desert tortoise
389:Arnold's giant tortoise
310:FRITZ & HAVAS, 2006
291:Dipsochelys elephantina
4363:Reptiles of Seychelles
1470:Réunion giant tortoise
1392:Cheirogaster schafferi
1386:Cheirogaster gymnesica
1223:Chelonoidis alburyorum
1217:Yellow-footed tortoise
1189:Centrochelys vulcanica
1183:Centrochelys burchardi
1177:Centrochelys atlantica
1152:Astrochelys rogerbouri
1082:Aldabra giant tortoise
600:Aldabrachelys gigantea
557:
432:Aldabra giant tortoise
408:
407:Living adult specimens
286:) ROTHSCHILD 1915: 433
272:) ROTHSCHILD 1897: 407
16:Subspecies of tortoise
2003:Spur-thighed tortoise
1940:Stylemys nebrascensis
1934:Stylemys karakolensis
1892:Stylemys calaverensis
1797:Megalochelys sondaari
1785:Megalochelys cautleyi
1754:Asian forest tortoise
1500:Burmese star tortoise
1320:Pinta Island tortoise
1113:Aldabrachelys abrupta
555:
415:shape, however, many
406:
359:), also known as the
315:Dipsochelys hololissa
299:Dipsochelys hololissa
1993:Kleinmann's tortoise
1952:Stylemys oregonensis
1898:Stylemys canetotiana
1592:Hadrianus schucherti
1574:Hadrianus majusculus
1505:Indian star tortoise
1247:Chelonoidis sellovii
1241:Chelonoidis monensis
1229:Chelonoidis cubensis
307:Geochelone hololissa
244:) — GÜNTHER 1877: 21
224:GÜNTHER, 1875: 296 (
169:A. g. hololissa
1998:Marginated tortoise
1928:Stylemys frizaciana
1791:Megalochelys margae
1675:Travancore tortoise
1598:Hadrianus utahensis
1580:Hadrianus octonaria
1268:Chelonoidis complex
1212:Red-footed tortoise
684:2021CBio...31.R989Z
252:GÜNTHER, 1877: 39 (
235:Testudo elephantina
3330:Nanhsiungchelyidae
3295:Macrocephalochelys
2009:Testudo brevitesta
1988:Hermann's tortoise
1964:Stylemys uintensis
1946:Stylemys neglectus
1910:Stylemys conspecta
1815:Geometric tortoise
1779:Megalochelys atlas
1759:Impressed tortoise
1670:Forsten's tortoise
1665:Elongated tortoise
1586:Hadrianus robustus
1263:Galápagos tortoise
1235:Chelonoidis lutzae
558:
488:, a 192-year-old,
409:
266:SCHWEIGGER, 1812 (
38:A living specimen
4340:
4339:
4239:Taxon identifiers
4230:
4229:
4201:
4200:
4197:
4196:
4144:
4143:
4140:
4139:
3562:
3561:
3558:
3557:
3241:
3240:
3013:
3012:
2516:
2515:
2434:
2433:
2155:
2154:
2151:
2150:
2147:
2146:
2028:
2027:
2023:. Key: †=extinct.
1970:Stylemys undabuna
1820:Serrated tortoise
1604:Hadrianus vialovi
1568:Hadrianus corsoni
1438:Speckled padloper
1410:Angulate tortoise
1145:Radiated tortoise
1140:Angonoka tortoise
1041:
1040:
868:"Giant tortoises"
678:(16): R989–R990.
537:over-exploitation
387:, along with the
347:
346:
341:
333:
325:
311:
303:
295:
287:
273:
259:
249:Testudo hololissa
245:
231:
221:Testudo hololissa
217:
4375:
4333:
4332:
4320:
4319:
4307:
4306:
4294:
4293:
4281:
4280:
4279:
4266:
4265:
4264:
4234:
4160:
4153:
3665:Chupacabrachelys
3580:
3573:
3250:
3028:Carettochelyidae
3024:
2527:
2445:
2177:
2166:
2134:
2125:
2116:
2055:
2048:
2041:
2032:
1922:Stylemys emiliae
1866:Leopard tortoise
1736:Pancake tortoise
1647:Greater padloper
1064:Russian tortoise
1049:
1048:
1026:
951:
944:
937:
928:
921:
920:
918:
916:
906:
900:
899:
897:
895:
885:
879:
878:
876:
874:
864:
858:
857:
855:
853:
843:
834:
833:
831:
829:
819:
810:
809:
807:
805:
795:
786:
785:
783:
781:
776:
768:
762:
761:
743:
737:
736:
734:
733:
712:
706:
705:
695:
663:
657:
656:
654:
652:
642:
636:
635:
633:
631:
621:
608:
596:
339:
331:
317:
309:
301:
293:
279:
277:Testudo daudinii
265:
263:Testudo gigantea
251:
237:
223:
215:
189:
155:A. gigantea
52:
51:
33:
21:
4383:
4382:
4378:
4377:
4376:
4374:
4373:
4372:
4343:
4342:
4341:
4336:
4328:
4323:
4315:
4310:
4302:
4297:
4289:
4284:
4275:
4274:
4269:
4260:
4259:
4254:
4241:
4231:
4226:
4193:
4136:
4113:Sahonachelyidae
4106:
3992:
3965:
3830:
3761:Phosphatochelys
3600:
3554:
3495:
3490:Terlinguachelys
3458:Pneumatoarthrus
3361:
3323:
3237:
3210:
3058:
3009:
2813:
2792:
2780:Siebenrockiella
2638:
2512:
2463:
2430:
2379:
2366:Mesodermochelys
2311:
2172:
2143:
2105:
2064:
2059:
2029:
2024:
2013:
1974:
1958:Stylemys pygmea
1870:
1852:
1848:Spider tortoise
1829:
1801:
1763:
1740:
1722:
1679:
1651:
1642:Common padloper
1628:
1608:
1552:
1538:Gopher tortoise
1528:Desert tortoise
1523:Bolson tortoise
1509:
1486:
1442:
1414:
1396:
1364:
1324:Lonesome George
1257:
1193:
1157:
1126:
1104:A. g. hololissa
1068:
1050:
1046:
1037:
1030:Species of the
1017:
961:
955:
925:
924:
914:
912:
910:"Tortoise news"
908:
907:
903:
893:
891:
887:
886:
882:
872:
870:
866:
865:
861:
851:
849:
845:
844:
837:
827:
825:
821:
820:
813:
803:
801:
797:
796:
789:
779:
777:
774:
770:
769:
765:
745:
744:
740:
731:
729:
714:
713:
709:
672:Current Biology
665:
664:
660:
650:
648:
644:
643:
639:
629:
627:
623:
622:
611:
597:
593:
588:
550:
526:
506:
461:
444:
424:A. g. hololissa
401:
198:
191:
185:
172:
158:
46:
17:
12:
11:
5:
4381:
4379:
4371:
4370:
4365:
4360:
4355:
4345:
4344:
4338:
4337:
4335:
4334:
4321:
4308:
4295:
4282:
4267:
4251:
4249:
4243:
4242:
4237:
4228:
4227:
4225:
4224:
4217:
4216:
4215:. † = extinct.
4206:
4203:
4202:
4199:
4198:
4195:
4194:
4192:
4191:
4183:
4175:
4166:
4164:
4157:
4150:
4146:
4145:
4142:
4141:
4138:
4137:
4135:
4134:
4126:
4117:
4115:
4108:
4107:
4105:
4104:
4096:
4089:
4082:
4074:
4066:
4059:
4051:
4043:
4035:
4027:
4019:
4011:
4008:Albertwoodemys
4002:
4000:
3998:Podocnemididae
3994:
3993:
3991:
3990:
3983:
3975:
3973:
3967:
3966:
3964:
3963:
3955:
3948:
3941:
3934:
3926:
3919:
3912:
3905:
3898:
3895:Lomalatachelys
3890:
3883:
3876:
3869:
3862:
3855:
3848:
3840:
3838:
3832:
3831:
3829:
3828:
3820:
3812:
3804:
3796:
3788:
3780:
3772:
3764:
3756:
3748:
3745:Labrostochelys
3740:
3732:
3724:
3716:
3708:
3700:
3692:
3684:
3676:
3668:
3660:
3652:
3644:
3636:
3628:
3620:
3611:
3609:
3602:
3601:
3599:
3598:
3589:
3587:
3585:Araripemydidae
3577:
3570:
3564:
3563:
3560:
3559:
3556:
3555:
3553:
3552:
3544:
3536:
3528:
3520:
3512:
3503:
3501:
3497:
3496:
3494:
3493:
3485:
3477:
3469:
3461:
3453:
3445:
3437:
3429:
3421:
3413:
3405:
3397:
3389:
3381:
3372:
3370:
3363:
3362:
3360:
3359:
3351:
3343:
3334:
3332:
3325:
3324:
3322:
3321:
3313:
3305:
3298:
3290:
3287:Emarginachelys
3282:
3274:
3266:
3258:
3256:
3247:
3243:
3242:
3239:
3238:
3236:
3235:
3232:Sinaspideretes
3227:
3218:
3216:
3212:
3211:
3209:
3208:
3201:
3194:
3187:
3180:
3172:
3165:
3158:
3151:
3143:
3135:
3127:
3120:
3112:
3105:
3098:
3091:
3083:
3076:
3068:
3066:
3060:
3059:
3057:
3056:
3049:
3041:
3032:
3030:
3021:
3015:
3014:
3011:
3010:
3008:
3007:
3000:
2992:
2985:
2977:
2970:
2963:
2955:
2947:
2940:
2933:
2926:
2919:
2912:
2909:Hesperotestudo
2904:
2896:
2889:
2882:
2874:
2866:
2859:
2852:
2845:
2838:
2831:
2823:
2821:
2815:
2814:
2812:
2811:
2803:
2801:
2799:Platysternidae
2794:
2793:
2791:
2790:
2783:
2776:
2769:
2762:
2755:
2748:
2741:
2734:
2727:
2720:
2713:
2706:
2699:
2692:
2685:
2678:
2671:
2664:
2656:
2648:
2646:
2640:
2639:
2637:
2636:
2628:
2621:
2614:
2607:
2600:
2593:
2586:
2579:
2572:
2565:
2558:
2551:
2544:
2535:
2533:
2524:
2518:
2517:
2514:
2513:
2511:
2510:
2503:
2496:
2489:
2481:
2473:
2471:
2465:
2464:
2462:
2461:
2453:
2451:
2449:Dermatemydidae
2442:
2440:Kinosternoidea
2436:
2435:
2432:
2431:
2429:
2428:
2420:
2412:
2409:Procolpochelys
2404:
2396:
2387:
2385:
2381:
2380:
2378:
2377:
2369:
2361:
2353:
2346:
2338:
2330:
2321:
2319:
2317:Dermochelyidae
2313:
2312:
2310:
2309:
2301:
2293:
2285:
2278:
2270:
2262:
2255:
2247:
2239:
2231:
2224:
2216:
2209:
2206:Carolinochelys
2201:
2194:
2185:
2183:
2174:
2163:
2157:
2156:
2153:
2152:
2149:
2148:
2145:
2144:
2137:
2131:
2122:
2113:
2107:
2106:
2104:
2103:
2097:
2088:
2082:
2076:
2069:
2066:
2065:
2060:
2058:
2057:
2050:
2043:
2035:
2026:
2025:
2018:
2015:
2014:
2012:
2011:
2005:
2000:
1995:
1990:
1984:
1982:
1976:
1975:
1973:
1972:
1966:
1960:
1954:
1948:
1942:
1936:
1930:
1924:
1918:
1916:Stylemys copei
1912:
1906:
1904:Stylemys capax
1900:
1894:
1888:
1886:Stylemys botti
1881:
1879:
1872:
1871:
1869:
1868:
1862:
1860:
1854:
1853:
1851:
1850:
1845:
1839:
1837:
1831:
1830:
1828:
1827:
1822:
1817:
1811:
1809:
1803:
1802:
1800:
1799:
1793:
1787:
1781:
1774:
1772:
1765:
1764:
1762:
1761:
1756:
1750:
1748:
1742:
1741:
1739:
1738:
1732:
1730:
1724:
1723:
1721:
1720:
1715:
1710:
1705:
1700:
1695:
1689:
1687:
1681:
1680:
1678:
1677:
1672:
1667:
1661:
1659:
1653:
1652:
1650:
1649:
1644:
1638:
1636:
1630:
1629:
1627:
1626:
1619:
1617:
1615:Hesperotestudo
1610:
1609:
1607:
1606:
1600:
1594:
1588:
1582:
1576:
1570:
1563:
1561:
1554:
1553:
1551:
1550:
1548:Texas tortoise
1545:
1540:
1535:
1530:
1525:
1519:
1517:
1511:
1510:
1508:
1507:
1502:
1496:
1494:
1488:
1487:
1485:
1484:
1478:
1472:
1466:
1460:
1453:
1451:
1444:
1443:
1441:
1440:
1435:
1433:Karoo padloper
1430:
1424:
1422:
1416:
1415:
1413:
1412:
1406:
1404:
1398:
1397:
1395:
1394:
1388:
1382:
1375:
1373:
1366:
1365:
1363:
1362:
1357:
1352:
1347:
1342:
1337:
1332:
1327:
1316:
1311:
1301:
1295:
1290:
1285:
1280:
1274:
1272:
1271:
1270:
1259:
1258:
1256:
1255:
1249:
1243:
1237:
1231:
1225:
1219:
1214:
1209:
1207:Chaco tortoise
1203:
1201:
1195:
1194:
1192:
1191:
1185:
1179:
1173:
1167:
1165:
1159:
1158:
1156:
1155:
1147:
1142:
1136:
1134:
1128:
1127:
1125:
1124:
1116:
1108:
1100:
1097:A. g. daudinii
1092:
1085:
1078:
1076:
1070:
1069:
1067:
1066:
1060:
1058:
1052:
1051:
1044:
1042:
1039:
1038:
1029:
1023:
1019:
1018:
1016:
1015:
1009:
1003:
997:
991:
985:
979:
973:
966:
963:
962:
956:
954:
953:
946:
939:
931:
923:
922:
901:
880:
859:
835:
811:
787:
763:
738:
707:
658:
637:
609:
590:
589:
587:
584:
549:
546:
530:giant tortoise
525:
522:
505:
502:
460:
457:
449:Frégate Island
443:
440:
436:A. g. gigantea
400:
397:
345:
344:
343:
342:
334:
326:
312:
304:
296:
288:
274:
260:
246:
232:
218:
207:
206:
200:
199:
192:
181:
180:
178:Trinomial name
174:
173:
166:
164:
160:
159:
152:
150:
146:
145:
138:
134:
133:
128:
124:
123:
118:
114:
113:
108:
104:
103:
98:
94:
93:
88:
84:
83:
78:
74:
73:
68:
64:
63:
58:
54:
53:
40:
39:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
4380:
4369:
4366:
4364:
4361:
4359:
4356:
4354:
4353:Aldabrachelys
4351:
4350:
4348:
4331:
4326:
4322:
4318:
4313:
4309:
4305:
4300:
4296:
4292:
4287:
4283:
4278:
4272:
4268:
4263:
4257:
4253:
4252:
4250:
4248:
4244:
4240:
4235:
4223:
4219:
4218:
4214:
4213:
4208:
4207:
4204:
4190:
4189:
4188:Tacuarembemys
4184:
4182:
4181:
4176:
4174:
4173:
4168:
4167:
4165:
4161:
4158:
4154:
4151:
4147:
4133:
4132:
4127:
4125:
4124:
4123:Sahonachelys
4119:
4118:
4116:
4114:
4109:
4103:
4102:
4097:
4095:
4094:
4090:
4088:
4087:
4086:Peltocephalus
4083:
4081:
4080:
4075:
4073:
4072:
4071:Lapparentemys
4067:
4065:
4064:
4060:
4058:
4057:
4052:
4050:
4049:
4044:
4042:
4041:
4036:
4034:
4033:
4028:
4026:
4025:
4020:
4018:
4017:
4012:
4010:
4009:
4004:
4003:
4001:
3999:
3995:
3989:
3988:
3984:
3982:
3981:
3977:
3976:
3974:
3972:
3971:Pelomedusidae
3968:
3962:
3961:
3960:Yaminuechelys
3956:
3954:
3953:
3949:
3947:
3946:
3942:
3940:
3939:
3935:
3933:
3932:
3931:Prochelidella
3927:
3925:
3924:
3920:
3918:
3917:
3913:
3911:
3910:
3906:
3904:
3903:
3899:
3897:
3896:
3891:
3889:
3888:
3884:
3882:
3881:
3877:
3875:
3874:
3870:
3868:
3867:
3863:
3861:
3860:
3856:
3854:
3853:
3849:
3847:
3846:
3845:Acanthochelys
3842:
3841:
3839:
3837:
3833:
3827:
3826:
3821:
3819:
3818:
3813:
3811:
3810:
3805:
3803:
3802:
3797:
3795:
3794:
3789:
3787:
3786:
3781:
3779:
3778:
3773:
3771:
3770:
3765:
3763:
3762:
3757:
3755:
3754:
3749:
3747:
3746:
3741:
3739:
3738:
3733:
3731:
3730:
3729:Kinkonychelys
3725:
3723:
3722:
3717:
3715:
3714:
3713:Itapecuruemys
3709:
3707:
3706:
3701:
3699:
3698:
3693:
3691:
3690:
3685:
3683:
3682:
3677:
3675:
3674:
3673:Eotaphrosphys
3669:
3667:
3666:
3661:
3659:
3658:
3653:
3651:
3650:
3645:
3643:
3642:
3637:
3635:
3634:
3629:
3627:
3626:
3621:
3619:
3618:
3613:
3612:
3610:
3608:
3607:Bothremydidae
3603:
3597:
3596:
3591:
3590:
3588:
3586:
3581:
3578:
3574:
3571:
3569:
3565:
3551:
3550:
3545:
3543:
3542:
3537:
3535:
3534:
3529:
3527:
3526:
3521:
3519:
3518:
3517:Argillochelys
3513:
3511:
3510:
3505:
3504:
3502:
3498:
3492:
3491:
3486:
3484:
3483:
3482:Santanachelys
3478:
3476:
3475:
3470:
3468:
3467:
3462:
3460:
3459:
3454:
3452:
3451:
3446:
3444:
3443:
3438:
3436:
3435:
3430:
3428:
3427:
3426:Desmatochelys
3422:
3420:
3419:
3414:
3412:
3411:
3410:Calcarichelys
3406:
3404:
3403:
3398:
3396:
3395:
3394:Atlantochelys
3390:
3388:
3387:
3382:
3380:
3379:
3374:
3373:
3371:
3369:
3368:Protostegidae
3364:
3358:
3357:
3356:Jiangxichelys
3352:
3350:
3349:
3344:
3342:
3341:
3340:Anomalochelys
3336:
3335:
3333:
3331:
3326:
3320:
3319:
3318:Protochelydra
3314:
3312:
3311:
3310:Planiplastron
3306:
3304:
3303:
3299:
3297:
3296:
3291:
3289:
3288:
3283:
3281:
3280:
3275:
3273:
3272:
3267:
3265:
3264:
3260:
3259:
3257:
3255:
3251:
3248:
3244:
3234:
3233:
3228:
3226:
3225:
3220:
3219:
3217:
3213:
3207:
3206:
3202:
3200:
3199:
3195:
3193:
3192:
3188:
3186:
3185:
3181:
3179:
3178:
3173:
3171:
3170:
3166:
3164:
3163:
3159:
3157:
3156:
3152:
3150:
3149:
3144:
3142:
3141:
3136:
3134:
3133:
3128:
3126:
3125:
3121:
3119:
3118:
3117:Drazinderetes
3113:
3111:
3110:
3106:
3104:
3103:
3099:
3097:
3096:
3092:
3090:
3089:
3084:
3082:
3081:
3077:
3075:
3074:
3070:
3069:
3067:
3065:
3061:
3055:
3054:
3053:Carettochelys
3050:
3048:
3047:
3042:
3040:
3039:
3034:
3033:
3031:
3029:
3025:
3022:
3020:
3016:
3006:
3005:
3001:
2999:
2998:
2993:
2991:
2990:
2986:
2984:
2983:
2978:
2976:
2975:
2971:
2969:
2968:
2964:
2962:
2961:
2956:
2954:
2953:
2948:
2946:
2945:
2941:
2939:
2938:
2937:Malacochersus
2934:
2932:
2931:
2927:
2925:
2924:
2920:
2918:
2917:
2913:
2911:
2910:
2905:
2903:
2902:
2897:
2895:
2894:
2890:
2888:
2887:
2883:
2881:
2880:
2875:
2873:
2872:
2867:
2865:
2864:
2860:
2858:
2857:
2853:
2851:
2850:
2846:
2844:
2843:
2839:
2837:
2836:
2832:
2830:
2829:
2828:Aldabrachelys
2825:
2824:
2822:
2820:
2816:
2810:
2809:
2805:
2804:
2802:
2800:
2795:
2789:
2788:
2784:
2782:
2781:
2777:
2775:
2774:
2770:
2768:
2767:
2763:
2761:
2760:
2756:
2754:
2753:
2749:
2747:
2746:
2742:
2740:
2739:
2735:
2733:
2732:
2728:
2726:
2725:
2721:
2719:
2718:
2714:
2712:
2711:
2710:Leucocephalon
2707:
2705:
2704:
2700:
2698:
2697:
2693:
2691:
2690:
2686:
2684:
2683:
2679:
2677:
2676:
2672:
2670:
2669:
2665:
2663:
2662:
2661:Banhxeochelys
2657:
2655:
2654:
2650:
2649:
2647:
2645:
2641:
2635:
2634:
2629:
2627:
2626:
2622:
2620:
2619:
2615:
2613:
2612:
2608:
2606:
2605:
2601:
2599:
2598:
2594:
2592:
2591:
2587:
2585:
2584:
2580:
2578:
2577:
2573:
2571:
2570:
2566:
2564:
2563:
2559:
2557:
2556:
2552:
2550:
2549:
2545:
2543:
2542:
2537:
2536:
2534:
2532:
2528:
2525:
2523:
2522:Testudinoidea
2519:
2509:
2508:
2504:
2502:
2501:
2497:
2495:
2494:
2490:
2488:
2487:
2482:
2480:
2479:
2475:
2474:
2472:
2470:
2469:Kinosternidae
2466:
2460:
2459:
2455:
2454:
2452:
2450:
2446:
2443:
2441:
2437:
2427:
2426:
2421:
2419:
2418:
2417:Protosphargis
2413:
2411:
2410:
2405:
2403:
2402:
2397:
2395:
2394:
2389:
2388:
2386:
2382:
2376:
2375:
2370:
2368:
2367:
2362:
2360:
2359:
2354:
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1428:Nama padloper
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1090:
1089:A. g. arnoldi
1086:
1083:
1080:
1079:
1077:
1075:
1074:Aldabrachelys
1071:
1065:
1062:
1061:
1059:
1057:
1053:
1035:
1034:
1027:
1024:
1020:
1014:
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1007:Testudinoidea
1005:Superfamily:
1004:
1002:
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986:
984:
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974:
972:
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498:Australia Zoo
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373:Aldabrachelys
370:in the genus
369:
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216:GÜNTHER, 1877
214:
211:
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196:
190:
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182:
179:
175:
171:
170:
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157:
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148:
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144:
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142:Aldabrachelys
139:
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121:Testudinoidea
119:
117:Superfamily:
116:
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109:
106:
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89:
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59:
56:
55:
50:
45:
41:
36:
32:
27:
22:
19:
4246:
4211:
4186:
4178:
4170:
4129:
4121:
4099:
4091:
4084:
4077:
4069:
4063:Erymnochelys
4061:
4054:
4048:Cerrejonemys
4046:
4038:
4030:
4024:Brontochelys
4022:
4014:
4006:
3985:
3978:
3958:
3950:
3943:
3938:Pseudemydura
3936:
3929:
3921:
3914:
3907:
3900:
3893:
3885:
3878:
3871:
3864:
3857:
3850:
3843:
3823:
3815:
3807:
3799:
3791:
3783:
3775:
3767:
3759:
3751:
3743:
3735:
3727:
3719:
3711:
3703:
3695:
3687:
3679:
3671:
3663:
3655:
3647:
3639:
3631:
3623:
3615:
3593:
3547:
3541:Prionochelys
3539:
3531:
3523:
3515:
3507:
3488:
3480:
3472:
3464:
3456:
3448:
3440:
3432:
3424:
3418:Cratochelone
3416:
3408:
3402:Bouliachelys
3400:
3392:
3384:
3378:Alienochelys
3376:
3354:
3346:
3338:
3316:
3308:
3300:
3293:
3285:
3279:Chelydropsis
3277:
3269:
3261:
3230:
3224:Basilochelys
3222:
3203:
3196:
3189:
3182:
3175:
3167:
3160:
3153:
3148:Khunnuchelys
3146:
3138:
3130:
3122:
3115:
3107:
3100:
3093:
3086:
3078:
3071:
3064:Trionychidae
3051:
3044:
3038:Allaeochelys
3036:
3002:
2995:
2989:Stigmochelys
2987:
2980:
2972:
2965:
2958:
2952:Megalochelys
2950:
2942:
2935:
2928:
2921:
2914:
2907:
2899:
2891:
2884:
2877:
2869:
2863:Cylindraspis
2861:
2854:
2847:
2842:Centrochelys
2840:
2833:
2826:
2819:Testudinidae
2808:Platysternon
2806:
2787:Vijayachelys
2785:
2778:
2771:
2766:Rhinoclemmys
2764:
2757:
2750:
2743:
2736:
2731:Melanochelys
2729:
2722:
2715:
2708:
2701:
2694:
2687:
2680:
2673:
2666:
2659:
2651:
2631:
2623:
2616:
2609:
2602:
2595:
2588:
2581:
2574:
2567:
2560:
2553:
2546:
2541:Acherontemys
2539:
2507:Sternotherus
2505:
2498:
2491:
2484:
2476:
2456:
2423:
2415:
2407:
2399:
2391:
2374:Psephophorus
2372:
2364:
2356:
2348:
2341:
2333:
2325:
2304:
2296:
2290:Pacifichelys
2288:
2280:
2273:
2265:
2259:Lepidochelys
2257:
2250:
2242:
2236:Gigantatypus
2234:
2228:Eretmochelys
2226:
2219:
2211:
2204:
2196:
2189:
2170:Chelonioidea
2138:
2090:
1858:Stigmochelys
1770:Megalochelys
1449:Cylindraspis
1371:Cheirogaster
1163:Centrochelys
1150:
1119:
1111:
1102:
1095:
1087:
1084:(subspecies:
1031:
1013:Testudinidae
913:. Retrieved
904:
892:. Retrieved
883:
871:. Retrieved
862:
850:. Retrieved
826:. Retrieved
802:. Retrieved
778:. Retrieved
766:
751:
741:
730:. Retrieved
720:
710:
675:
671:
661:
649:. Retrieved
640:
628:. Retrieved
599:
594:
571:
567:
559:
548:Conservation
534:
527:
518:
514:
507:
490:Saint Helena
478:Calcutta Zoo
470:Robert Clive
462:
445:
435:
423:
421:
410:
393:
378:
371:
360:
355:
354:
350:
348:
336:
328:
319:
314:
306:
302:LÜCKER, 2000
298:
290:
281:
276:
267:
262:
253:
248:
239:
234:
225:
220:
212:
186:
184:
168:
167:
163:Subspecies:
153:
141:
131:Testudinidae
18:
4312:iNaturalist
4271:Wikispecies
4101:Stupendemys
4056:Cordichelys
3902:Mesoclemmys
3887:Hydromedusa
3809:Taphrosphys
3801:Sankuchemys
3793:Rhothonemys
3769:Polysternon
3649:Cearachelys
3633:Azabbaremys
3617:Araiochelys
3533:Ctenochelys
3525:Bashuchelys
3474:Rhinochelys
3450:Ocepechelon
3442:Notochelone
3434:Iserosaurus
3302:Macrochelys
3254:Chelydridae
3177:Palaeoamyda
3102:Cyclanorbis
2967:Psammobates
2960:Oligopherus
2923:Indotestudo
2871:Cymatholcus
2849:Chelonoidis
2835:Astrochelys
2644:Geoemydidae
2562:Deirochelys
2500:Staurotypus
2493:Kinosternon
2486:Hoplochelys
2401:Peritresius
2350:Dermochelys
2343:Cosmochelys
2335:Corsochelys
2252:Itilochelys
2244:Glarichelys
2191:Allopleuron
2181:Cheloniidae
2120:Superfamily
1807:Psammobates
1657:Indotestudo
1199:Chelonoidis
1132:Astrochelys
889:"Hatchings"
799:"Tortoises"
577:already on
399:Description
227:nomen nudum
4347:Categories
4180:Caririemys
4093:Podocnemis
4079:Latentemys
4040:Carbonemys
3980:Pelomedusa
3817:Ummulisani
3737:Kurmademys
3705:Inaechelys
3689:Galianemys
3657:Chedighaii
3595:Araripemys
3568:Pleurodira
3549:Toxochelys
3466:Protostega
3271:Chelydrops
3191:Pelodiscus
3184:Pelochelys
3132:Gilmoremys
3109:Cycloderma
3019:Trionychia
2886:Geochelone
2879:Floridemys
2745:Notochelys
2633:Wilburemys
2604:Malaclemys
2458:Dermatemys
2425:Puppigerus
2358:Eosphargis
2275:Miocaretta
2267:Mexichelys
2161:Cryptodira
2101:Testudines
2062:Testudines
1492:Geochelone
1420:Chersobius
1056:Agrionemys
1001:Cryptodira
999:Suborder:
995:Testudines
987:Subclass:
732:2014-01-23
727:BBC Online
586:References
510:morphotype
504:Morphotype
453:noddy tern
381:Seychelles
368:subspecies
340:TTWG, 2012
332:TTWG, 2010
111:Cryptodira
107:Suborder:
101:Testudines
4262:Q20721641
4220:See also
4172:Caribemys
4032:Caninemys
4016:Bauruemys
3945:Rheodytes
3909:Myuchelys
3852:Chelodina
3777:Puentemys
3753:Nigeremys
3697:Ilatardia
3641:Bothremys
3348:Basilemys
3162:Nilssonia
3140:Hutchemys
3088:Axestemys
3046:Anosteira
2901:Hadrianus
2759:Pangshura
2717:Malayemys
2682:Geoclemys
2625:Trachemys
2618:Terrapene
2611:Pseudemys
2597:Graptemys
2590:Glyptemys
2583:Emydoidea
2576:Actinemys
2548:Chrysemys
2393:Euclastes
2221:Eochelone
2072:Kingdom:
1559:Hadrianus
969:Kingdom:
482:life span
283:ex errore
269:ex errore
241:ex errore
149:Species:
67:Kingdom:
61:Eukaryota
4256:Wikidata
3987:Pelusios
3952:Rhinemys
3923:Platemys
3916:Phrynops
3836:Chelidae
3825:Zolhafah
3721:Jainemys
3386:Archelon
3263:Chelydra
3155:Lissemys
2997:Stylemys
2982:Solitudo
2944:Manouria
2893:Gopherus
2856:Chersina
2724:Mauremys
2703:Heosemys
2696:Hardella
2689:Geoemyda
2675:Cyclemys
2531:Emydidae
2478:Claudius
2327:Arabemys
2306:Tasbacka
2298:Syllomus
2213:Chelonia
2111:Suborder
2095:Diapsida
2086:Reptilia
2080:Chordata
2078:Phylum:
2074:Animalia
1877:Stylemys
1746:Manouria
1515:Gopherus
1402:Chersina
1033:tortoise
1011:Family:
989:Anapsida
983:Reptilia
977:Chordata
975:Phylum:
971:Animalia
958:Tortoise
758:Archived
753:BBC News
722:BBC News
702:34428417
486:Jonathan
459:Lifespan
413:carapace
365:tortoise
318:BONIN, (
204:Synonyms
127:Family:
91:Reptilia
81:Chordata
77:Phylum:
71:Animalia
57:Domain:
4304:9240911
4131:Sokatra
3880:Emydura
3785:Rosasia
3681:Foxemys
3625:Arenila
3205:Trionyx
3198:Rafetus
3124:Dogania
3080:Apalone
3004:Testudo
2930:Kinixys
2916:Homopus
2773:Sacalia
2752:Orlitia
2738:Morenia
2653:Batagur
2555:Clemmys
2282:Natator
2198:Caretta
2099:Order:
2084:Class:
1980:Testudo
1685:Kinixys
1634:Homopus
993:Order:
981:Class:
680:Bibcode
603:at the
579:Cousine
574:Cousine
494:Harriet
472:of the
466:Adwaita
455:chick.
417:captive
385:extinct
363:, is a
197:, 1877)
195:Günther
137:Genus:
97:Order:
87:Class:
4330:936915
4317:540339
4163:
4156:
4149:
3873:Elusor
3866:Elseya
3859:Chelus
3576:
3509:Adocus
3500:
3246:
3215:
3095:Chitra
2797:
2384:
2129:Family
1036:family
1022:Genera
960:family
915:7 June
894:7 June
873:5 June
852:7 June
828:5 June
804:6 June
780:7 June
700:
651:5 June
630:5 June
428:scutes
324:) 2006
4291:5DFY3
3169:Palea
3073:Amyda
2974:Pyxis
2668:Cuora
2140:Genus
2091:Clade
1835:Pyxis
1308:Diego
775:(PDF)
560:With
321:et al
255:part.
4325:ITIS
4299:GBIF
2569:Emys
917:2015
896:2015
875:2015
854:2015
830:2015
806:2015
782:2015
698:PMID
653:2015
632:2015
442:Diet
349:The
4286:CoL
688:doi
562:DNA
4349::
4327::
4314::
4301::
4288::
4273::
4258::
2093::
1322:†(
838:^
814:^
790:^
756:.
750:.
725:.
719:.
696:.
686:.
676:31
674:.
670:.
612:^
376:.
4185:†
4177:†
4169:†
4128:†
4120:†
4111:†
4098:†
4076:†
4068:†
4053:†
4045:†
4037:†
4029:†
4021:†
4013:†
4005:†
3957:†
3928:†
3892:†
3822:†
3814:†
3806:†
3798:†
3790:†
3782:†
3774:†
3766:†
3758:†
3750:†
3742:†
3734:†
3726:†
3718:†
3710:†
3702:†
3694:†
3686:†
3678:†
3670:†
3662:†
3654:†
3646:†
3638:†
3630:†
3622:†
3614:†
3605:†
3592:†
3583:†
3546:†
3538:†
3530:†
3522:†
3514:†
3506:†
3487:†
3479:†
3471:†
3463:†
3455:†
3447:†
3439:†
3431:†
3423:†
3415:†
3407:†
3399:†
3391:†
3383:†
3375:†
3366:†
3353:†
3345:†
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