Knowledge (XXG)

Sapeornis

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in the proportions of the foot and number of wing and foot bones. However, the relevant portions of the specimen were badly crushed, and later authors concluded that these differences were based on misinterpretation of the poorly preserved specimen. In a 2010 survey of Chinese avialan fossils, Li and
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for "from Chaoyang". Soon after this, two more, nearly complete specimens were discovered in the Dapingfang area, about 40 kilometres (25 mi) from the original fossil site. None of these first three specimens preserved traces of feathers, but based on the size of their skeletons alone, they were
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probably was a glider and/or soarer that preferred more open country compared to the Enantiornithes and predominantly woodland birds, although it was able to perch on branches. The small gastroliths, overall large size, and the inferred habitat indicate that
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Gao, C.; Chiappe, L.M.; Zhang, F.; Pomeroy, D.L.; Shen, C.; Chinsamy, A.; Walsh, M.O. (2012). "A subadult specimen of the Early Cretaceous bird
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Zhou, Z.; Zhang, F. (2002). "Largest Bird from the Early Cretaceous and Its Implications for the Earliest Avian Ecological Diversification".
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Erickson, Gregory M.; Rauhut, Oliver W. M.; Zhou, Zhonghe; Turner, Alan H.; Inouye, Brian D.; Hu, Dongyu; Norell, Mark A. (2009).
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was long and reached the distal point of the tarsal joint, not reduced as in more modern birds (and some non-avian theropods like
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Zhou, Zhonghe; Zhang, Fucheng (2006). "A beaked basal ornithurine bird (Aves, Ornithurae) from the Lower Cretaceous of China".
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epochs of the Cretaceous period, and are about 125-120 million years old. Several nearly complete skeletons have been found.
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consisted of rectrices that formed a graded, fan-like structure. The reduced fingers suggest that it might have had an
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was large for an early avialan, about 30–33 centimetres (0.98–1.08 ft) long in life, excluding the tail feathers.
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Hu, D.; et al. (2010). "A new sapeornithid bird from China and its implication for early avian evolution".
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Yuan, C. (2008). "A new genus and species of Sapeornithidae from Lower Cretaceous in western Liaoning, China".
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Schmitz, L.; Motani, R. (2011). "Nocturnality in Dinosaurs Inferred from Scleral Ring and Orbit Morphology".
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Foth, C. (2014). Comment on the absence of ossified sternal elements in basal paravian dinosaurs.
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was an incomplete skeleton dug up from Jiufotang Formation rocks in the area of Shangheshou, near
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and small carnivorous dinosaurs, rather than the explosively fast growth seen in modern birds.
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Li, D.; Sullivan, C.; Zhou, Z.; Zhang, Z. (2010). "Basal birds from China: a brief review".
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in Liaoning Province, China in the summer of 2000. It was discovered by a team from the
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Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,
847:"Scapular orientation in theropods and basal birds, and the origin of flapping flight" 4533: 4388: 4383: 4252: 4112: 3996: 3940: 3619: 3538: 3482: 3452: 3399: 3392: 3278: 3269: 3053: 3033: 2576: 2565: 2451: 2420: 2377: 2326: 2312: 2006: 1738: 1716: 1701: 1619: 973: 823: 707: 695: 687: 586: 487: 283: 79: 1270: 1223: 1188: 1063: 981: 831: 4398: 4260: 3987: 3861: 3854: 3793: 3711: 3444: 3323: 3235: 3184: 3154: 3103: 3043: 2861: 2714: 2699: 2684: 2618: 2506: 2458: 2370: 2260: 2115: 2075: 2054: 2047: 2027: 2020: 2013: 1960: 1945: 1881: 1838: 1799: 1669: 1648: 1579: 1557: 1114: 279: 1262: 925: 4457: 4223: 4133: 4068: 3809: 3759: 3690: 3642: 3467: 3413: 3361: 3308: 3194: 3164: 3083: 2847: 2597: 2587: 2250: 2089: 1854: 1813: 1784: 1777: 1745: 1655: 1589: 1529: 1516: 1475: 1354: 507:
similar to more advanced avialans but a general anatomy even more basal than in
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Zhou, Zhonghe & Zhang, Fucheng (2003): Anatomy of the primitive bird
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recognized as the largest early Cretaceous avialans known at the time.
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Zhou, Z., & Zhang, F. (2003). Anatomy of the primitive bird
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rocks in western Liaoning, China. These rocks formed during the
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was large and bore holes, apparently to save weight, as in the
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and modern birds and reptiles indicate that it may have been
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The skull has a handful of teeth in the upper jawtip only.
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Institute of Vertebrate Paleontology and Paleoanthropology
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In absolute number of features shared with modern birds,
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Analysis of its skeletal bones suggest that it had an 4183: 4144: 4100: 4084: 4074: 4064: 3835: 3781: 3660: 3630: 3615: 3605: 3595: 3585: 3575: 3565: 3555: 3545: 3534: 3479: 3424: 3373: 3334: 3296: 3266: 3251: 3241: 3231: 3220: 3210: 3200: 3190: 3180: 3170: 3160: 3150: 3140: 3130: 3120: 3110: 3099: 3089: 3079: 3069: 3059: 3049: 3039: 3029: 3019: 3009: 2881: 2835: 2791: 2760: 2743: 2672: 2656: 2645: 2634: 2624: 2614: 2603: 2593: 2583: 2572: 2561: 2551: 2503: 2477: 2431: 2337: 2292: 2277: 2267: 2256: 2246: 2124: 1929: 1602: 1585: 1575: 1564: 1553: 1542: 993: 991: 4432: 4182: 4143: 4099: 4049: 4018: 3995: 3986: 3962: 3939: 3872: 3834: 3780: 3728: 3659: 3629: 3519: 3478: 3423: 3372: 3333: 3295: 3265: 2990: 2906: 2880: 2871: 2834: 2790: 2759: 2728: 2671: 2532: 2502: 2476: 2430: 2336: 2291: 2227: 2149: 2123: 2114: 2037: 1985: 1928: 1871: 1837: 1756: 1687: 1600: 1527: 1474: 1447: 1420: 1303:Early Cretaceous birds with crops found in China 1241:and a taxonomic reassessment of sapeornithids". 572:). The first toe pointed backwards. In specimen 872:Proceedings of the National Academy of Sciences 1134:from the Early Cretaceous of Liaoning, China. 1001:from the Early Cretaceous of Liaoning, China. 726:In 2008, Yuan named a new specimen related to 1339: 617:. Not being well-adapted to flapping flight, 576:V12375, the stomach contained numerous small 8: 1126: 1124: 704:Society of Avian Paleontology and Evolution 4420: 4057: 3992: 3968: 3936: 3923: 3668: 3527: 3002: 2998: 2987: 2974: 2877: 2736: 2544: 2540: 2239: 2235: 2224: 2211: 2120: 1991: 1937: 1925: 1762: 1693: 1611: 1535: 1524: 1511: 1453: 1426: 1417: 1362: 1346: 1332: 1324: 714:(ornis), meaning "bird". The species name 630:, possibly eating plant seeds and fruits. 127: 31: 934: 924: 794: 374: 347: 7: 767:and the supposed second species of 4545:Early Cretaceous dinosaurs of Asia 1243:Journal of Vertebrate Paleontology 1181:10.1111/j.1755-6724.2008.tb00323.x 1136:Canadian Journal of Earth Sciences 1003:Canadian Journal of Earth Sciences 485:was far more derived than that of 138:National Museum of Natural Science 25: 4404: 4403: 4349: 4343: 3909: 3904: 2960: 2953: 2946: 2197: 2191: 2185: 1310: 974:10.1111/j.1463-6409.2006.00234.x 824:10.1111/j.1755-6724.2010.00188.x 645:, similar to most modern birds. 148: 781:is also likely synonymous with 759:was indeed a junior synonym of 706:, which they combined with the 667:is known from fossils found in 659:Paleozoological Museum of China 686:The first known specimen (the 1: 4550:Fossil taxa described in 2002 854:Acta Palaeontologica Polonica 429:which lived during the early 1283:Holtz, Thomas R. Jr. (2011) 1263:10.1080/02724634.2012.693865 926:10.1371/journal.pone.0007390 605:were largely different from 449:contains only one species, 4566: 379:Omnivoropteryx sinousaorum 27:Extinct genus of dinosaurs 4363: 4341: 4060: 3971: 3935: 3922: 3902: 3671: 3530: 3005: 3001: 2986: 2973: 2944: 2739: 2547: 2543: 2242: 2238: 2223: 2210: 2183: 1994: 1940: 1924: 1765: 1696: 1614: 1538: 1523: 1510: 1456: 1429: 1416: 1361: 1099:10.1007/s00114-001-0276-9 338: 331: 313: 306: 145:Scientific classification 143: 135: 126: 34: 1216:10.5122/cbirds.2010.0002 657:Holotype (IVPP V12698), 633:Comparisons between the 558:were more fused than in 422:is a monotypic genus of 4464:Sapeornis chaoyangensis 4434:Sapeornis chaoyangensis 1239:Sapeornis chaoyangensis 1132:Sapeornis chaoyangensis 1048:10.1126/science.1200043 999:Sapeornis chaoyangensis 884:10.1073/pnas.1419023111 761:Sapeornis chaoyangensis 751:. In a 2012 study, Gao 749:Sapeornis chaoyangensis 597:is about as derived as 530:but no (or unossified) 452:Sapeornis chaoyangensis 320:Sapeornis chaoyangensis 743:colleagues considered 734:. Yuan concluded that 661: 472: 391:Zhou & Zhang, 2009 383:Czerkas & Ji, 2002 326:Zhou & Zhang, 2002 4512:Paleobiology Database 2083:Paralitherizinosaurus 1288:Winter 2010 Appendix. 1161:Acta Geologica Sinica 845:Senter, Phil (2006). 804:Acta Geologica Sinica 656: 649:Discovery and history 584:and slow growth like 471:compared with a human 466: 299:S. chaoyangensis 247:Omnivoropterygiformes 503:was unusual, with a 404:Shenshiornis primita 3300:Patagopterygiformes 2795:Confuciusornithidae 2661:Scansoriopterygidae 2296:Scansoriopterygidae 1318:Paleontology portal 1255:2012JVPal..32.1103G 1173:2008AcGlS..82...48Y 1091:2002NW.....89...34Z 1079:Naturwissenschaften 1040:2011Sci...332..705S 917:2009PLoSO...4.7390E 878:(50): E5334-E5334. 816:2010AcGlS..84..472H 669:Jiufotang Formation 542:was rod-like as in 517:Confuciusornithidae 495:, and a well-fused 3428:Songlingornithidae 3377:Hongshanornithidae 2764:Omnivoropterygidae 2676:Jeholornithiformes 1987:Therizinosauroidea 662: 626:was most likely a 534:. The breastbone ( 532:uncinate processes 473: 441:, roughly 125-120 388:Sapeornis angustis 260:Omnivoropterygidae 4527: 4526: 4499:Open Tree of Life 4426:Taxon identifiers 4417: 4416: 4359: 4358: 4339: 4338: 4335: 4334: 4239:Lutetodontopteris 4148:Gastornithiformes 4045: 4044: 4041: 4040: 3929:Aves / Neornithes 3918: 3917: 3900: 3899: 3896: 3895: 3892: 3891: 3874:Aves / Neornithes 3776: 3775: 3515: 3514: 3407:Parahongshanornis 2969: 2968: 2942: 2941: 2938: 2937: 2934: 2933: 2930: 2929: 2926: 2925: 2724: 2723: 2650:Archaeopterygidae 2528: 2527: 2435:Archaeopterygidae 2206: 2205: 2181: 2180: 2177: 2176: 2173: 2172: 2169: 2168: 2110: 2109: 2106: 2105: 2039:Therizinosauridae 1920: 1919: 1916: 1915: 1912: 1911: 1506: 1505: 1502: 1501: 1498: 1497: 1494: 1493: 1406:Maniraptoriformes 732:Didactylornis jii 431:Cretaceous period 415: 414: 408: 400: 396:Didactylornis jii 392: 384: 371: 362: 354: 344: 287: 136:Fossil specimen, 16:(Redirected from 4557: 4520: 4519: 4507: 4506: 4494: 4493: 4481: 4480: 4468: 4467: 4466: 4453: 4452: 4451: 4421: 4407: 4406: 4379:Deinonychosauria 4354: 4353: 4347: 4304:Protodontopteryx 4246:Macrodontopteryx 4185: 4146: 4102: 4086: 4076: 4066: 4058: 3993: 3969: 3937: 3924: 3913: 3908: 3839:Cimolopterygidae 3837: 3783: 3730:Hesperornithidae 3669: 3662: 3632: 3617: 3607: 3597: 3587: 3577: 3567: 3557: 3547: 3536: 3528: 3481: 3426: 3375: 3336: 3298: 3268: 3253: 3243: 3233: 3222: 3212: 3202: 3192: 3182: 3172: 3162: 3152: 3142: 3132: 3122: 3112: 3101: 3091: 3081: 3071: 3061: 3051: 3041: 3031: 3021: 3011: 3003: 2999: 2988: 2975: 2964: 2958: 2957: 2950: 2883: 2878: 2839:Jinguofortisidae 2837: 2818:Eoconfuciusornis 2793: 2762: 2745: 2737: 2674: 2658: 2647: 2636: 2626: 2616: 2605: 2595: 2585: 2574: 2563: 2553: 2545: 2541: 2505: 2479: 2433: 2339: 2294: 2279: 2269: 2258: 2248: 2240: 2236: 2225: 2212: 2201: 2195: 2189: 2139:Oviraptorosauria 2128:Oviraptorosauria 2126: 2121: 2062:Nanshiungosaurus 1992: 1969:Jianchangosaurus 1938: 1931: 1926: 1763: 1694: 1612: 1606:Alvarezsauroidea 1604: 1587: 1577: 1566: 1555: 1544: 1536: 1525: 1512: 1454: 1427: 1418: 1411: 1410: 1363: 1348: 1341: 1334: 1325: 1320: 1315: 1314: 1313: 1290: 1281: 1275: 1274: 1249:(5): 1103–1112. 1234: 1228: 1227: 1199: 1193: 1192: 1156: 1150: 1128: 1119: 1118: 1074: 1068: 1067: 1023: 1017: 995: 986: 985: 955: 949: 948: 938: 928: 892: 886: 868: 862: 861: 851: 842: 836: 835: 799: 673:Yixian Formation 595:S. chaoyangensis 469:S. chaoyangensis 406: 398: 390: 382: 370:Species synonymy 369: 360: 352: 342: 322: 318: 278: 271: 258: 245: 153: 152: 131: 121: 51: 42:Early Cretaceous 40:Temporal range: 32: 21: 4565: 4564: 4560: 4559: 4558: 4556: 4555: 4554: 4530: 4529: 4528: 4523: 4515: 4510: 4502: 4497: 4489: 4484: 4476: 4471: 4462: 4461: 4456: 4447: 4446: 4441: 4428: 4418: 4413: 4355: 4348: 4331: 4326:Tympanonesiotes 4319:Pseudodontornis 4282:Palaeochenoides 4203:Caspiodontornis 4187:Pelagornithidae 4178: 4139: 4095: 4037: 4014: 3982: 3958: 3931: 3914: 3888: 3868: 3830: 3772: 3767:Parahesperornis 3739:Asiahesperornis 3724: 3655: 3625: 3524: 3511: 3474: 3419: 3386:Archaeornithura 3368: 3329: 3291: 3261: 3014:Archaeorhynchus 2995: 2982: 2965: 2952: 2922: 2902: 2873:Ornithothoraces 2867: 2830: 2804:Changchengornis 2786: 2755: 2733: 2720: 2667: 2639:Anchiornithidae 2537: 2524: 2498: 2492:Dromaeosauridae 2481:Dromaeosauridae 2472: 2466:Anchiornithidae 2426: 2385:Liaoningvenator 2341:Anchiornithidae 2332: 2320:Scansoriopteryx 2287: 2232: 2219: 2202: 2165: 2145: 2102: 2097:Therizinosaurus 2033: 1981: 1976:Lingyuanosaurus 1933:Therizinosauria 1908: 1867: 1833: 1758:Parvicursorinae 1752: 1689:Alvarezsauridae 1683: 1608: 1596: 1532: 1519: 1490: 1470: 1443: 1437:Avemetatarsalia 1422:Avemetatarsalia 1412: 1368: 1367: 1357: 1352: 1316: 1311: 1309: 1299: 1294: 1293: 1282: 1278: 1236: 1235: 1231: 1201: 1200: 1196: 1158: 1157: 1153: 1148:10.1139/E03-011 1129: 1122: 1076: 1075: 1071: 1034:(6030): 705–8. 1025: 1024: 1020: 1016:(HTML abstract) 1014:10.1139/E03-011 996: 989: 957: 956: 952: 894: 893: 889: 869: 865: 849: 844: 843: 839: 801: 800: 796: 791: 755:concluded that 651: 601:. However, the 497:carpometacarpus 461: 411: 373: 372: 365: 346: 345: 327: 324: 316: 315: 302: 277: 269: 256: 243: 147: 122: 120: 119: 118: 117: 112: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 47:124.5–120  46: 45: 38: 28: 23: 22: 15: 12: 11: 5: 4563: 4561: 4553: 4552: 4547: 4542: 4532: 4531: 4525: 4524: 4522: 4521: 4508: 4495: 4482: 4469: 4454: 4438: 4436: 4430: 4429: 4424: 4415: 4414: 4412: 4411: 4401: 4396: 4391: 4386: 4381: 4376: 4371: 4369:Archaeornithes 4364: 4361: 4360: 4357: 4356: 4342: 4340: 4337: 4336: 4333: 4332: 4330: 4329: 4322: 4315: 4311:Protopelicanus 4307: 4300: 4292: 4285: 4278: 4275:Osteodontornis 4271: 4264: 4257: 4249: 4242: 4235: 4227: 4220: 4213: 4206: 4199: 4191: 4189: 4180: 4179: 4177: 4176: 4172:Dromornithidae 4168: 4161: 4152: 4150: 4141: 4140: 4138: 4137: 4130: 4123: 4116: 4108: 4106: 4104:Dromornithidae 4097: 4096: 4094: 4093: 4082: 4079:Perplexicervix 4072: 4061: 4055: 4052:Incertae sedis 4047: 4046: 4043: 4042: 4039: 4038: 4036: 4035: 4034: 4033: 4031:Pangalliformes 4024: 4022: 4020:Pangalliformes 4016: 4015: 4013: 4012: 4011: 4010: 4001: 3999: 3990: 3984: 3983: 3981: 3980: 3972: 3966: 3960: 3959: 3957: 3956: 3955: 3954: 3945: 3943: 3933: 3932: 3927: 3920: 3919: 3916: 3915: 3903: 3901: 3898: 3897: 3894: 3893: 3890: 3889: 3887: 3886: 3885: 3884: 3878: 3876: 3870: 3869: 3867: 3866: 3858: 3851: 3843: 3841: 3832: 3831: 3829: 3828: 3821: 3814: 3806: 3798: 3789: 3787: 3778: 3777: 3774: 3773: 3771: 3770: 3763: 3756: 3749: 3742: 3734: 3732: 3726: 3725: 3723: 3722: 3715: 3708: 3701: 3694: 3687: 3680: 3672: 3666: 3664:Hesperornithes 3657: 3656: 3654: 3653: 3646: 3638: 3636: 3634:Ichthyornithes 3627: 3626: 3624: 3623: 3613: 3603: 3593: 3583: 3573: 3563: 3553: 3543: 3531: 3525: 3520: 3517: 3516: 3513: 3512: 3510: 3509: 3502: 3499:Similiyanornis 3495: 3487: 3485: 3476: 3475: 3473: 3472: 3464: 3456: 3449: 3441: 3432: 3430: 3421: 3420: 3418: 3417: 3410: 3403: 3396: 3389: 3381: 3379: 3370: 3369: 3367: 3366: 3358: 3350: 3342: 3340: 3338:Ambiortiformes 3331: 3330: 3328: 3327: 3320: 3312: 3304: 3302: 3293: 3292: 3290: 3289: 3282: 3274: 3272: 3263: 3262: 3260: 3259: 3256:Zhongjianornis 3249: 3239: 3229: 3218: 3208: 3198: 3188: 3178: 3168: 3158: 3148: 3145:Jianchangornis 3138: 3128: 3118: 3108: 3097: 3087: 3077: 3067: 3057: 3047: 3037: 3027: 3017: 3006: 2996: 2991: 2984: 2983: 2978: 2971: 2970: 2967: 2966: 2945: 2943: 2940: 2939: 2936: 2935: 2932: 2931: 2928: 2927: 2924: 2923: 2921: 2920: 2919: 2918: 2912: 2910: 2904: 2903: 2901: 2900: 2899: 2898: 2896:Enantiornithes 2889: 2887: 2885:Enantiornithes 2875: 2869: 2868: 2866: 2865: 2858: 2851: 2843: 2841: 2832: 2831: 2829: 2828: 2821: 2814: 2811:Confuciusornis 2807: 2799: 2797: 2788: 2787: 2785: 2784: 2777: 2774:Omnivoropteryx 2769: 2767: 2757: 2756: 2754: 2753: 2740: 2734: 2729: 2726: 2725: 2722: 2721: 2719: 2718: 2711: 2704: 2696: 2689: 2680: 2678: 2669: 2668: 2666: 2665: 2654: 2643: 2632: 2622: 2612: 2601: 2591: 2581: 2570: 2559: 2548: 2538: 2533: 2530: 2529: 2526: 2525: 2523: 2522: 2521: 2520: 2511: 2509: 2500: 2499: 2497: 2496: 2495: 2494: 2485: 2483: 2474: 2473: 2471: 2470: 2462: 2455: 2448: 2439: 2437: 2428: 2427: 2425: 2424: 2417: 2410: 2403: 2396: 2389: 2381: 2374: 2367: 2360: 2353: 2345: 2343: 2334: 2333: 2331: 2330: 2323: 2316: 2309: 2301: 2299: 2289: 2288: 2286: 2285: 2275: 2272:Pneumatoraptor 2265: 2254: 2243: 2233: 2228: 2221: 2220: 2215: 2208: 2207: 2204: 2203: 2184: 2182: 2179: 2178: 2175: 2174: 2171: 2170: 2167: 2166: 2164: 2163: 2162: 2161: 2155: 2153: 2147: 2146: 2144: 2143: 2142: 2141: 2132: 2130: 2118: 2112: 2111: 2108: 2107: 2104: 2103: 2101: 2100: 2093: 2086: 2079: 2072: 2069:Neimongosaurus 2065: 2058: 2051: 2043: 2041: 2035: 2034: 2032: 2031: 2024: 2017: 2010: 2003: 1995: 1989: 1983: 1982: 1980: 1979: 1972: 1965: 1957: 1950: 1941: 1935: 1922: 1921: 1918: 1917: 1914: 1913: 1910: 1909: 1907: 1906: 1899: 1892: 1885: 1877: 1875: 1869: 1868: 1866: 1865: 1858: 1851: 1843: 1841: 1835: 1834: 1832: 1831: 1828:Trierarchuncus 1824: 1817: 1810: 1803: 1796: 1788: 1781: 1774: 1766: 1760: 1754: 1753: 1751: 1750: 1742: 1735: 1728: 1724:Bonapartenykus 1720: 1713: 1706: 1697: 1691: 1685: 1684: 1682: 1681: 1674: 1666: 1663:Shishugounykus 1659: 1652: 1645: 1642:Bonapartenykus 1638: 1631: 1623: 1615: 1609: 1601: 1598: 1597: 1595: 1594: 1583: 1573: 1562: 1551: 1539: 1533: 1528: 1521: 1520: 1515: 1508: 1507: 1504: 1503: 1500: 1499: 1496: 1495: 1492: 1491: 1489: 1488: 1487: 1486: 1480: 1478: 1472: 1471: 1469: 1468: 1467: 1466: 1457: 1451: 1445: 1444: 1442: 1441: 1440: 1439: 1430: 1424: 1414: 1413: 1409: 1408: 1399: 1390: 1381: 1375: 1366: 1359: 1358: 1353: 1351: 1350: 1343: 1336: 1328: 1322: 1321: 1306: 1305: 1298: 1297:External links 1295: 1292: 1291: 1276: 1229: 1194: 1151: 1142:(5): 731-747. 1120: 1069: 1018: 1008:(5): 731–747. 987: 968:(4): 363–373. 950: 887: 863: 837: 810:(3): 472–482. 793: 792: 790: 787: 778:Omnivoropteryx 738:differed from 679:through early 650: 647: 607:Confuciusornis 599:Confuciusornis 556:tarsometatarsi 545:Confuciusornis 460: 457: 435:late Barremian 413: 412: 410: 409: 401: 393: 385: 368: 367: 366: 364: 363: 355: 343:Genus synonymy 341: 340: 339: 336: 335: 329: 328: 325: 311: 310: 304: 303: 295: 293: 289: 288: 267: 263: 262: 254: 250: 249: 241: 237: 236: 231: 224: 223: 218: 211: 210: 205: 198: 197: 192: 185: 184: 179: 175: 174: 169: 165: 164: 159: 155: 154: 141: 140: 133: 132: 124: 123: 115: 114: 113: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 52: 39: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 4562: 4551: 4548: 4546: 4543: 4541: 4538: 4537: 4535: 4518: 4513: 4509: 4505: 4500: 4496: 4492: 4487: 4483: 4479: 4474: 4470: 4465: 4459: 4455: 4450: 4444: 4440: 4439: 4437: 4435: 4431: 4427: 4422: 4410: 4402: 4400: 4397: 4395: 4392: 4390: 4389:Odontornithes 4387: 4385: 4384:Odontognathae 4382: 4380: 4377: 4375: 4372: 4370: 4366: 4365: 4362: 4352: 4346: 4328: 4327: 4323: 4321: 4320: 4316: 4313: 4312: 4308: 4306: 4305: 4301: 4298: 4297: 4293: 4291: 4290: 4286: 4284: 4283: 4279: 4277: 4276: 4272: 4270: 4269: 4265: 4263: 4262: 4258: 4255: 4254: 4250: 4248: 4247: 4243: 4241: 4240: 4236: 4233: 4232: 4228: 4226: 4225: 4221: 4219: 4218: 4214: 4212: 4211: 4207: 4205: 4204: 4200: 4198: 4197: 4193: 4192: 4190: 4188: 4181: 4174: 4173: 4169: 4167: 4166: 4162: 4159: 4158: 4154: 4153: 4151: 4149: 4142: 4136: 4135: 4131: 4129: 4128: 4124: 4122: 4121: 4117: 4115: 4114: 4113:Barawertornis 4110: 4109: 4107: 4105: 4098: 4091: 4090: 4083: 4081: 4080: 4073: 4071: 4070: 4063: 4062: 4059: 4056: 4054: 4053: 4048: 4032: 4028: 4027: 4026: 4025: 4023: 4021: 4017: 4009: 4008:Anserimorphae 4005: 4004: 4003: 4002: 4000: 3998: 3997:Anserimorphae 3994: 3991: 3989: 3985: 3979: 3978: 3974: 3973: 3970: 3967: 3965: 3961: 3953: 3952:Palaeognathae 3949: 3948: 3947: 3946: 3944: 3942: 3941:Palaeognathae 3938: 3934: 3930: 3925: 3921: 3912: 3907: 3882: 3881: 3880: 3879: 3877: 3875: 3871: 3864: 3863: 3859: 3857: 3856: 3852: 3850: 3849: 3845: 3844: 3842: 3840: 3833: 3827: 3826: 3822: 3820: 3819: 3815: 3812: 3811: 3807: 3804: 3803: 3799: 3796: 3795: 3791: 3790: 3788: 3786: 3779: 3769: 3768: 3764: 3762: 3761: 3757: 3755: 3754: 3750: 3748: 3747: 3743: 3741: 3740: 3736: 3735: 3733: 3731: 3727: 3721: 3720: 3716: 3714: 3713: 3709: 3707: 3706: 3702: 3700: 3699: 3695: 3693: 3692: 3688: 3686: 3685: 3681: 3679: 3678: 3674: 3673: 3670: 3667: 3665: 3658: 3652: 3651: 3647: 3645: 3644: 3640: 3639: 3637: 3635: 3628: 3622: 3621: 3620:Tingmiatornis 3614: 3612: 3611: 3604: 3602: 3601: 3594: 3592: 3591: 3584: 3582: 3581: 3574: 3572: 3571: 3564: 3562: 3561: 3554: 3552: 3551: 3544: 3541: 3540: 3539:Antarcticavis 3533: 3532: 3529: 3526: 3523: 3518: 3508: 3507: 3503: 3501: 3500: 3496: 3494: 3493: 3489: 3488: 3486: 3484: 3483:Yanornithidae 3477: 3470: 3469: 3465: 3462: 3461: 3457: 3455: 3454: 3453:Songlingornis 3450: 3447: 3446: 3442: 3439: 3438: 3434: 3433: 3431: 3429: 3422: 3416: 3415: 3411: 3409: 3408: 3404: 3402: 3401: 3400:Longicrusavis 3397: 3395: 3394: 3393:Hongshanornis 3390: 3388: 3387: 3383: 3382: 3380: 3378: 3371: 3364: 3363: 3359: 3356: 3355: 3351: 3349: 3348: 3344: 3343: 3341: 3339: 3332: 3326: 3325: 3321: 3318: 3317: 3313: 3311: 3310: 3306: 3305: 3303: 3301: 3294: 3288: 3287: 3283: 3281: 3280: 3279:Mengciusornis 3276: 3275: 3273: 3271: 3270:Schizoouridae 3264: 3258: 3257: 3250: 3248: 3247: 3240: 3238: 3237: 3230: 3227: 3226: 3219: 3217: 3216: 3209: 3207: 3206: 3199: 3197: 3196: 3189: 3187: 3186: 3179: 3177: 3176: 3169: 3167: 3166: 3159: 3157: 3156: 3149: 3147: 3146: 3139: 3137: 3136: 3129: 3127: 3126: 3119: 3117: 3116: 3109: 3106: 3105: 3098: 3096: 3095: 3088: 3086: 3085: 3078: 3076: 3075: 3068: 3066: 3065: 3058: 3056: 3055: 3054:Changzuiornis 3048: 3046: 3045: 3038: 3036: 3035: 3034:Brevidentavis 3028: 3026: 3025: 3018: 3016: 3015: 3008: 3007: 3004: 3000: 2997: 2994: 2989: 2985: 2981: 2976: 2972: 2963: 2959: 2956: 2949: 2916: 2915: 2914: 2913: 2911: 2909: 2905: 2897: 2893: 2892: 2891: 2890: 2888: 2886: 2879: 2876: 2874: 2870: 2864: 2863: 2859: 2857: 2856: 2852: 2850: 2849: 2845: 2844: 2842: 2840: 2833: 2827: 2826: 2822: 2820: 2819: 2815: 2813: 2812: 2808: 2806: 2805: 2801: 2800: 2798: 2796: 2789: 2783: 2782: 2778: 2776: 2775: 2771: 2770: 2768: 2765: 2758: 2752: 2750: 2742: 2741: 2738: 2735: 2732: 2727: 2717: 2716: 2712: 2710: 2709: 2705: 2702: 2701: 2697: 2695: 2694: 2690: 2687: 2686: 2682: 2681: 2679: 2677: 2670: 2663: 2662: 2655: 2652: 2651: 2644: 2641: 2640: 2633: 2631: 2630: 2623: 2621: 2620: 2613: 2610: 2609: 2602: 2600: 2599: 2592: 2590: 2589: 2582: 2579: 2578: 2577:Cretaaviculus 2571: 2568: 2567: 2560: 2558: 2557: 2550: 2549: 2546: 2542: 2539: 2536: 2531: 2519: 2515: 2514: 2513: 2512: 2510: 2508: 2501: 2493: 2489: 2488: 2487: 2486: 2484: 2482: 2475: 2468: 2467: 2463: 2461: 2460: 2456: 2454: 2453: 2452:Archaeopteryx 2449: 2446: 2445: 2441: 2440: 2438: 2436: 2429: 2423: 2422: 2421:Yixianosaurus 2418: 2416: 2415: 2411: 2409: 2408: 2404: 2402: 2401: 2397: 2395: 2394: 2390: 2387: 2386: 2382: 2380: 2379: 2378:Fujianvenator 2375: 2373: 2372: 2368: 2366: 2365: 2361: 2359: 2358: 2354: 2352: 2351: 2347: 2346: 2344: 2342: 2335: 2329: 2328: 2324: 2322: 2321: 2317: 2315: 2314: 2313:Epidexipteryx 2310: 2308: 2307: 2303: 2302: 2300: 2297: 2290: 2284: 2283: 2276: 2274: 2273: 2266: 2263: 2262: 2255: 2253: 2252: 2245: 2244: 2241: 2237: 2234: 2231: 2226: 2222: 2218: 2213: 2209: 2200: 2196: 2194: 2188: 2159: 2158: 2157: 2156: 2154: 2152: 2148: 2140: 2136: 2135: 2134: 2133: 2131: 2129: 2122: 2119: 2117: 2113: 2099: 2098: 2094: 2092: 2091: 2087: 2085: 2084: 2080: 2078: 2077: 2073: 2071: 2070: 2066: 2064: 2063: 2059: 2057: 2056: 2052: 2050: 2049: 2045: 2044: 2042: 2040: 2036: 2030: 2029: 2025: 2023: 2022: 2018: 2016: 2015: 2011: 2009: 2008: 2007:Beipiaosaurus 2004: 2002: 2001: 1997: 1996: 1993: 1990: 1988: 1984: 1978: 1977: 1973: 1971: 1970: 1966: 1963: 1962: 1958: 1956: 1955: 1951: 1948: 1947: 1943: 1942: 1939: 1936: 1934: 1927: 1923: 1905: 1904: 1900: 1898: 1897: 1893: 1891: 1890: 1886: 1884: 1883: 1879: 1878: 1876: 1874: 1870: 1864: 1863: 1859: 1857: 1856: 1852: 1850: 1849: 1845: 1844: 1842: 1840: 1836: 1830: 1829: 1825: 1823: 1822: 1818: 1816: 1815: 1811: 1809: 1808: 1804: 1802: 1801: 1797: 1794: 1793: 1789: 1787: 1786: 1782: 1780: 1779: 1775: 1773: 1772: 1768: 1767: 1764: 1761: 1759: 1755: 1748: 1747: 1743: 1741: 1740: 1739:Heptasteornis 1736: 1734: 1733: 1729: 1726: 1725: 1721: 1719: 1718: 1717:Alvarezsaurus 1714: 1712: 1711: 1707: 1704: 1703: 1702:Achillesaurus 1699: 1698: 1695: 1692: 1690: 1686: 1680: 1679: 1675: 1672: 1671: 1667: 1665: 1664: 1660: 1658: 1657: 1653: 1651: 1650: 1646: 1644: 1643: 1639: 1637: 1636: 1632: 1629: 1628: 1624: 1622: 1621: 1620:Achillesaurus 1617: 1616: 1613: 1610: 1607: 1599: 1592: 1591: 1584: 1582: 1581: 1574: 1571: 1570: 1563: 1560: 1559: 1552: 1549: 1548: 1541: 1540: 1537: 1534: 1531: 1526: 1522: 1518: 1513: 1509: 1484: 1483: 1482: 1481: 1479: 1477: 1473: 1465: 1461: 1460: 1459: 1458: 1455: 1452: 1450: 1446: 1438: 1434: 1433: 1432: 1431: 1428: 1425: 1423: 1419: 1415: 1407: 1403: 1400: 1398: 1394: 1391: 1389: 1385: 1382: 1380: 1376: 1374: 1370: 1369: 1364: 1360: 1356: 1349: 1344: 1342: 1337: 1335: 1330: 1329: 1326: 1319: 1308: 1304: 1301: 1300: 1296: 1289: 1286: 1280: 1277: 1272: 1268: 1264: 1260: 1256: 1252: 1248: 1244: 1240: 1233: 1230: 1225: 1221: 1217: 1213: 1209: 1205: 1204:Chinese Birds 1198: 1195: 1190: 1186: 1182: 1178: 1174: 1170: 1166: 1162: 1155: 1152: 1149: 1145: 1141: 1137: 1133: 1127: 1125: 1121: 1116: 1112: 1108: 1104: 1100: 1096: 1092: 1088: 1084: 1080: 1073: 1070: 1065: 1061: 1057: 1053: 1049: 1045: 1041: 1037: 1033: 1029: 1022: 1019: 1015: 1011: 1007: 1004: 1000: 994: 992: 988: 983: 979: 975: 971: 967: 963: 962: 961:Zool. Scripta 954: 951: 946: 942: 937: 932: 927: 922: 918: 914: 911:(10): e7390. 910: 906: 902: 900: 899:Archaeopteryx 891: 888: 885: 881: 877: 873: 867: 864: 860:(2): 305–313. 859: 855: 848: 841: 838: 833: 829: 825: 821: 817: 813: 809: 805: 798: 795: 788: 786: 784: 780: 779: 774: 770: 766: 762: 758: 757:Didactylornis 754: 750: 747:a synonym of 746: 745:Didactylornis 741: 737: 736:Didactylornis 733: 729: 724: 721: 717: 716:chaoyangensis 713: 709: 708:Ancient Greek 705: 701: 697: 696:Chaoyang City 693: 689: 688:type specimen 684: 682: 678: 674: 670: 666: 660: 655: 648: 646: 644: 640: 636: 635:scleral rings 631: 629: 625: 620: 616: 612: 608: 604: 600: 596: 591: 589: 588: 587:Archaeopteryx 583: 579: 575: 571: 570: 565: 561: 560:Archaeopteryx 557: 553: 552: 547: 546: 541: 537: 533: 529: 525: 520: 518: 514: 510: 509:Archaeopteryx 506: 502: 498: 494: 490: 489: 488:Archaeopteryx 484: 479: 477: 470: 465: 458: 456: 454: 453: 448: 444: 440: 436: 432: 428: 425: 421: 420: 405: 402: 397: 394: 389: 386: 381: 380: 375: 359: 356: 351: 350:Didactylornis 348: 337: 334: 330: 323: 321: 312: 309: 308:Binomial name 305: 301: 300: 294: 291: 290: 285: 281: 276: 275: 268: 265: 264: 261: 255: 252: 251: 248: 242: 239: 238: 235: 232: 229: 226: 225: 222: 219: 216: 213: 212: 209: 206: 203: 200: 199: 196: 193: 190: 187: 186: 183: 180: 177: 176: 173: 170: 167: 166: 163: 160: 157: 156: 151: 146: 142: 139: 134: 130: 125: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 50: 43: 37: 33: 30: 19: 4540:Pygostylians 4433: 4399:Unenlagiidae 4324: 4317: 4309: 4302: 4294: 4287: 4280: 4273: 4266: 4261:Odontopteryx 4259: 4251: 4244: 4237: 4229: 4222: 4215: 4208: 4201: 4194: 4170: 4163: 4155: 4132: 4125: 4118: 4111: 4087: 4077: 4067: 4050: 3988:Galloanserae 3975: 3862:Lamarqueavis 3860: 3855:Cimolopteryx 3853: 3846: 3823: 3816: 3808: 3800: 3794:Australornis 3792: 3765: 3758: 3751: 3744: 3737: 3717: 3712:Pasquiaornis 3710: 3703: 3696: 3689: 3682: 3675: 3648: 3641: 3618: 3608: 3598: 3588: 3578: 3568: 3558: 3548: 3537: 3504: 3497: 3490: 3466: 3458: 3451: 3445:Piscivoravis 3443: 3435: 3412: 3405: 3398: 3391: 3384: 3360: 3352: 3345: 3324:Patagopteryx 3322: 3314: 3307: 3284: 3277: 3254: 3244: 3236:Xinghaiornis 3234: 3223: 3213: 3203: 3193: 3185:Khinganornis 3183: 3173: 3163: 3155:Jiuquanornis 3153: 3143: 3133: 3123: 3113: 3104:Gargantuavis 3102: 3092: 3082: 3072: 3062: 3052: 3044:Changmaornis 3042: 3032: 3022: 3012: 2951: 2862:Jinguofortis 2860: 2853: 2846: 2823: 2816: 2809: 2802: 2780: 2779: 2772: 2746: 2715:Neimengornis 2713: 2706: 2700:Jixiangornis 2698: 2691: 2685:Dalianraptor 2683: 2659: 2648: 2637: 2627: 2619:Yandangornis 2617: 2606: 2596: 2586: 2575: 2564: 2554: 2518:Troodontidae 2507:Troodontidae 2464: 2459:Wellnhoferia 2457: 2450: 2442: 2419: 2412: 2405: 2398: 2391: 2383: 2376: 2371:Eosinopteryx 2369: 2362: 2355: 2348: 2325: 2318: 2311: 2304: 2280: 2270: 2261:Palaeopteryx 2259: 2249: 2190: 2116:Pennaraptora 2095: 2088: 2081: 2076:Nothronychus 2074: 2067: 2060: 2055:Erlikosaurus 2053: 2048:Erliansaurus 2046: 2028:Suzhousaurus 2026: 2021:Martharaptor 2019: 2014:Enigmosaurus 2012: 2005: 1998: 1974: 1967: 1961:Fukuivenator 1959: 1952: 1946:Eshanosaurus 1944: 1901: 1894: 1887: 1882:Albertonykus 1880: 1860: 1853: 1846: 1839:Ceratonykini 1826: 1819: 1812: 1805: 1800:Nemegtonykus 1798: 1790: 1783: 1776: 1769: 1744: 1737: 1730: 1722: 1715: 1708: 1700: 1676: 1670:Tugulusaurus 1668: 1661: 1654: 1649:Haplocheirus 1647: 1640: 1633: 1625: 1618: 1588: 1580:Migmanychion 1578: 1567: 1558:Fukuivenator 1556: 1545: 1401: 1392: 1383: 1284: 1279: 1246: 1242: 1238: 1232: 1210:(2): 83–96. 1207: 1203: 1197: 1167:(1): 48–55. 1164: 1160: 1154: 1139: 1135: 1131: 1085:(1): 34–38. 1082: 1078: 1072: 1031: 1027: 1021: 1005: 1002: 998: 965: 959: 953: 908: 904: 898: 890: 875: 871: 866: 857: 853: 840: 807: 803: 797: 782: 776: 772: 768: 765:Shenshiornis 764: 760: 756: 752: 748: 744: 739: 735: 731: 727: 725: 715: 711: 691: 685: 664: 663: 638: 632: 623: 618: 610: 606: 598: 594: 592: 585: 567: 559: 549: 543: 523: 521: 508: 486: 482: 481:The hand of 480: 475: 474: 468: 451: 450: 446: 418: 417: 416: 403: 395: 387: 377: 358:Shenshiornis 357: 349: 319: 314: 298: 297: 273: 272: 227: 214: 201: 188: 35: 29: 18:Shenshiornis 4458:Wikispecies 4296:Proceriavis 4268:Odontoptila 4224:Gigantornis 4134:Ilbandornis 4069:Asteriornis 3810:Neogaeornis 3760:Hesperornis 3691:Chupkaornis 3643:Ichthyornis 3468:Yixianornis 3414:Tianyuornis 3362:Palintropus 3309:Alamitornis 3195:Meemannavis 3165:Juehuaornis 3084:Eogranivora 3024:Bellulornis 2848:Chongmingia 2598:Overoraptor 2588:Fukuipteryx 2251:Imperobator 2090:Segnosaurus 1855:Ceratonykus 1814:Parvicursor 1785:Khulsanurus 1778:Jaculinykus 1746:Patagonykus 1656:Patagonykus 1590:Yaverlandia 1530:Maniraptora 1517:Maniraptora 1476:Maniraptora 1355:Maniraptora 773:S. angustis 677:late Aptian 603:apomorphies 578:gastroliths 505:hypocleidum 459:Description 4534:Categories 4367:See also: 4289:Pelagornis 4157:Brontornis 3964:Neognathae 3883:see below↓ 3848:Ceramornis 3818:Polarornis 3785:Vegaviidae 3753:Fumicollis 3719:Potamornis 3705:Judinornis 3698:Enaliornis 3522:Ornithurae 3492:Abitusavis 3286:Schizooura 3246:Yumenornis 3205:Platanavis 3175:Kaririavis 3125:Horezmavis 3064:Chaoyangia 2993:Euornithes 2980:Euornithes 2917:see below↓ 2908:Euornithes 2855:Cratonavis 2731:Pygostylia 2708:Kompsornis 2693:Jeholornis 2629:Zhongornis 2556:Alcmonavis 2444:Alcmonavis 2414:Xiaotingia 2407:Serikornis 2350:Anchiornis 2306:Ambopteryx 2160:see below↓ 2000:Alxasaurus 1889:Linhenykus 1873:Mononykini 1862:Xixianykus 1821:Qiupanykus 1807:Ondogurvel 1771:Dzharaonyx 1732:Bradycneme 1710:Alnashetri 1485:see below↓ 1388:Dinosauria 789:References 763:, as were 399:Yuan, 2008 353:Yuan, 2008 208:Saurischia 195:Dinosauria 4394:Sauriurae 4374:Carinatae 4210:Cyphornis 4196:Aequornis 4165:Gastornis 4127:Genyornis 4120:Dromornis 3677:Baptornis 3610:Limenavis 3590:Iaceornis 3580:Guildavis 3570:Gallornis 3560:Cerebavis 3550:Apatornis 3354:Apsaravis 3347:Ambiortus 3316:Kuszholia 2781:Sapeornis 2608:Rahonavis 2400:Pedopenna 2282:Rahonavis 1954:Falcarius 1896:Mononykus 1848:Albinykus 1678:Xiyunykus 1547:Elopteryx 1464:Theropoda 1449:Theropoda 1397:Theropoda 1371:Kingdom: 783:Sapeornis 769:Sapeornis 740:Sapeornis 728:Sapeornis 692:Sapeornis 665:Sapeornis 639:Sapeornis 628:herbivore 624:Sapeornis 619:Sapeornis 611:Sapeornis 540:pygostyle 528:gastralia 524:Sapeornis 493:phalanges 483:Sapeornis 476:Sapeornis 447:Sapeornis 437:to early 427:dinosaurs 419:Sapeornis 292:Species: 274:Sapeornis 221:Theropoda 168:Kingdom: 162:Eukaryota 36:Sapeornis 4478:52571602 4443:Wikidata 4409:Category 4217:Dasornis 4089:Qinornis 3746:Canadaga 3684:Brodavis 3506:Yanornis 3460:Yanornis 3437:Hollanda 3135:Iteravis 3115:Hollanda 3074:Dingavis 2825:Yangavis 2749:Proornis 2393:Ostromia 2357:Aurornis 1903:Shuvuuia 1635:Bannykus 1379:Chordata 1377:Phylum: 1373:Animalia 1271:86195304 1224:84976296 1189:86691693 1107:12008971 1064:33253407 1056:21493820 982:85222311 945:19816582 905:PLOS ONE 832:86441777 582:ontogeny 569:Avimimus 551:Nomingia 467:Size of 407:Hu, 2010 361:Hu, 2010 333:Synonyms 253:Family: 182:Chordata 178:Phylum: 172:Animalia 158:Domain: 4504:3604529 4491:4966829 4449:Q141637 4231:Laornis 3977:Neoaves 3825:Vegavis 3650:Janavis 3225:Wyleyia 2535:Avialae 2364:Caihong 2230:Paraves 2217:Paraves 2151:Paraves 1251:Bibcode 1169:Bibcode 1115:1116829 1087:Bibcode 1036:Bibcode 1028:Science 936:2756958 913:Bibcode 812:Bibcode 643:diurnal 536:sternum 513:humerus 501:furcula 424:avialan 376: ? 317:† 296:† 266:Genus: 240:Order: 234:Avialae 116:↓ 4517:119249 3802:Maaqwi 3600:Kookne 3215:Vorona 3094:Gansus 2566:Balaur 1569:Kakuru 1269:  1222:  1187:  1113:  1105:  1062:  1054:  980:  943:  933:  830:  753:et al. 681:Albian 564:fibula 562:, the 511:. The 439:Aptian 286:, 2002 282:& 1627:Aorun 1402:Clade 1393:Clade 1384:Clade 1267:S2CID 1220:S2CID 1185:S2CID 1111:S2CID 1060:S2CID 978:S2CID 850:(PDF) 828:S2CID 720:Latin 712:όρνις 710:word 690:) of 615:alula 284:Zhang 228:Clade 215:Clade 202:Clade 189:Clade 4486:GBIF 4253:Manu 4029:see 4006:see 3950:see 2894:see 2516:see 2490:see 2137:see 1462:see 1435:see 1103:PMID 1052:PMID 941:PMID 671:and 574:IVPP 548:and 526:had 280:Zhou 55:PreꞒ 4473:EoL 1792:Kol 1259:doi 1212:doi 1177:doi 1144:doi 1095:doi 1044:doi 1032:332 1010:doi 970:doi 931:PMC 921:doi 880:doi 876:111 820:doi 730:as 718:is 637:of 445:). 443:mya 4536:: 4514:: 4501:: 4488:: 4475:: 4460:: 4445:: 2327:Yi 1404:: 1395:: 1386:: 1265:. 1257:. 1247:32 1245:. 1218:. 1206:. 1183:. 1175:. 1165:82 1163:. 1140:40 1138:, 1123:^ 1109:. 1101:. 1093:. 1083:89 1081:. 1058:. 1050:. 1042:. 1030:. 1006:40 990:^ 976:. 966:35 964:. 939:. 929:. 919:. 907:. 903:. 874:, 858:51 856:. 852:. 826:. 818:. 808:84 806:. 785:. 775:. 771:, 519:. 455:. 230:: 217:: 204:: 191:: 105:Pg 49:Ma 44:, 4314:? 4299:? 4256:? 4234:? 4184:† 4175:? 4160:? 4145:† 4101:† 4092:? 4085:† 4075:† 4065:† 3865:? 3836:† 3813:? 3805:? 3797:? 3782:† 3661:† 3631:† 3616:† 3606:† 3596:† 3586:† 3576:† 3566:† 3556:† 3546:† 3542:? 3535:† 3480:† 3471:? 3463:? 3448:? 3440:? 3425:† 3374:† 3365:? 3357:? 3335:† 3319:? 3297:† 3267:† 3252:† 3242:† 3232:† 3228:? 3221:† 3211:† 3201:† 3191:† 3181:† 3171:† 3161:† 3151:† 3141:† 3131:† 3121:† 3111:† 3107:? 3100:† 3090:† 3080:† 3070:† 3060:† 3050:† 3040:† 3030:† 3020:† 3010:† 2882:† 2836:† 2792:† 2766:? 2761:† 2751:" 2747:" 2744:† 2703:? 2688:? 2673:† 2664:? 2657:† 2653:? 2646:† 2642:? 2635:† 2625:† 2615:† 2611:? 2604:† 2594:† 2584:† 2580:? 2573:† 2569:? 2562:† 2552:† 2504:† 2478:† 2469:? 2447:? 2432:† 2388:? 2338:† 2298:? 2293:† 2278:† 2268:† 2264:? 2257:† 2247:† 2125:† 1964:? 1949:? 1930:† 1795:? 1749:? 1727:? 1705:? 1673:? 1630:? 1603:† 1593:? 1586:† 1576:† 1572:? 1565:† 1561:? 1554:† 1550:? 1543:† 1347:e 1340:t 1333:v 1273:. 1261:: 1253:: 1226:. 1214:: 1208:1 1191:. 1179:: 1171:: 1146:: 1117:. 1097:: 1089:: 1066:. 1046:: 1038:: 1012:: 984:. 972:: 947:. 923:: 915:: 909:4 901:" 882:: 834:. 822:: 814:: 433:( 270:† 257:† 244:† 110:N 100:K 95:J 90:T 85:P 80:C 75:D 70:S 65:O 60:Ꞓ 20:)

Index

Shenshiornis
Early Cretaceous
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

National Museum of Natural Science
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Saurischia
Theropoda
Avialae
Omnivoropterygiformes
Omnivoropterygidae
Sapeornis
Zhou

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