1915:
594:
1864:
1850:
4282:
6397:
2003:
1964:, were also found with gastroliths, though in those cases the stones were much more numerous and smaller in size. Ji and colleagues speculated that, since the later two dinosaurs were probably primarily herbivores, the number and size of gastroliths may correspond with diet; that is, herbivores ingested many small stones, while carnivores ingested only a few larger stones to aid in digestion. However, the 2012 study could find no gastroliths with the second specimen and therefore concluded that the stones with the holotype were swallowed by accident. In any case no special
4275:
2779:
3604:
3590:
118:
5849:
5070:
5835:
3583:
5842:
2785:
4289:
139:
3597:
6410:
5088:
5076:
477:
5095:
6404:
5829:
353:
446:, about 125 million years old. It consists of a nearly complete skeleton with skull, compressed on a single plate, of an adult individual. Extensive remains of protofeathers have been preserved. In 2012 a second specimen was described, CAGS-IG-T1, of an individual larger than the holotype. The skull was about 10% longer, the feet about a third, a difference explained by positive
5083:
2208:
1884:
within the abdominal cavity, comprising a complete lower leg and foot with toes and claws in their natural, articulated position. While the leg part, about one foot long, is very large in relation to the abdominal cavity, it is clearly situated within it, lying between the ribs. Ji and colleagues in
572:
differ in length across the body, with the longest protofeathers covering the hips, base of the tail, and back of the thighs. These longest protofeathers measured up to ten centimeters (4 in) in length. Protofeathers were also found on the metatarsus (upper part of the foot). While these were
1896:
with a length of 1.2 metres (3.9 ft). The 2012 study also reported newly discovered remains of prey animals. Above the dromaeosaurid leg, feathers are visible. Below it two clusters of digested food can be seen. It was suggested that the feathers had belonged to a bird and were with the leg
1930:
In addition to the dromaeosaurid leg, four irregularly-shaped stones with a diameter between fifteen and twenty millimetres were found in the abdomen, with no similar stones present in any other portions of the skeleton or embedded in the surrounding rock. The authors interpreted these as
2097:
Lida Xing, Phil R. Bell, W. Scott
Persons IV, Shuan Ji, Tetsuto Miyashita, Michael E. Burns, Qiang Ji, Philip J. Currie, 2012, "Abdominal Contents from Two Large Early Cretaceous Compsognathids (Dinosauria: Theropoda) Demonstrate Feeding on Confuciusornithids and Dromaeosaurids",
1805:
compared to its relatives is also notable, and may indicate a trend towards large size among compsognathids (a group well known for their small size compared to other, giant theropod dinosaurs), similar to the trends towards larger sizes in other dinosaurian lineages.
1889:
may have been an agile, active, "fierce" predator, especially since other compsognathids have been found with (presumably fast-moving) lizards and small mammals in their abdominal cavities. In 2012 the dromaeosaurid was tentatively identified as an individual of
1994:
species. The bone surface of the scapula looked as if it had been etched by stomach acid for about thirteen days, leading to the conclusion the birds were swallowed later and in quick succession. This again would indicate a high metabolism for
2188:
Dal Sasso C, Maganuco S (2011) Scipionyx samniticus (Theropoda: Compsognathidae) from the Lower
Cretaceous of Italy; osteology, ontogenetic assessment, phylogeny, soft tissue anatomy taphonomy and palaeobiology. Memorie 37: 1–281.
508:
The vertebral column consisted of eleven cervical, twelve dorsal, five sacral and at least forty-nine caudal vertebrae. The point of the tail is missing. In the tail, the spines and chevrons strongly inclined to the back. The
473:, as well as other compsognathids, by its relatively long hands in relation to its arms. The arms and hindlimbs were also longer overall than in other compsognathids, a feature possibly related to its size.
2028:
was in all likelihood cursorial, or ground-dwelling, and yet apparently quite capable of catching birds, e.g. by using stealth attack, a method employed by many modern cursorial predators of fowl.
1971:
The second specimen, CAGS-IG-T1, also preserves the remains of several meals. Disarticulated bones were found, in front of the lower ischia, identified as belonging to at least two individuals of
501:
specimen but the number of its tooth positions could not be adequately determined. Specimen CAGS-IG-T1 preserves ten maxillary tooth sockets with room for one or two more in damaged areas. The
1914:
593:
505:
was a strongly built element with a high-rising front branch that formed part of the lower front edge of the eye socket. The lower jaw lacked an opening in its outer side.
1863:
543:
curved downwards. The hindlimb was elongated, largely because of a long lower leg, having 90% of the length of the thighbone. The foot was also long, especially in the
1978:, a basal bird that is very common in the formation. Also a 13.5 centimetres long scapula was discovered, belonging to some 1.5 metres (4.9 ft) long herbivorous
524:
showing only a weakly developed olecranon process on its back upper end. The hand was very elongated though, as long as the ulna and upper arm combined. The second
2138:(Theropoda: Compsognathidae) from the Lower Cretaceous of Italy — Osteology, ontogenetic assessment, phylogeny, soft tissue anatomy, taphonomy and palaeobiology.
2010:
That the second specimen in a short period of time caught two birds, together with the bird feather remains in the holotype stomach, might be an indication that
2067:
Ji, S., Ji, Q., Lu J., and Yuan, C. (2007). "A new giant compsognathid dinosaur with long filamentous integuments from Lower
Cretaceous of Northeastern China."
2024:
was reported with a bird in its stomach and seen as proof that the former were arboreal. The 2012 study denied the relevance of flying prey for this issue as
6533:
6474:
344:, at 2.37 meters (7.78 ft) in length, in 2007 was the largest known compsognathid exemplar. In 2012 an even larger specimen was reported.
532:
more compact. The second claw was elongated, as long as the thumb claw. The third metacarpal was rather short and carried a thin, third, finger.
4281:
6548:
4274:
6396:
6553:
585:, they do indicate that foot-feathers or similar structures first arose in dinosaurs much more basal or "primitive" than previously known.
2241:
3603:
5069:
3589:
5834:
1849:
2778:
3582:
138:
5848:
2784:
2002:
6275:
5087:
6543:
4288:
5841:
3596:
1885:
2007 suggested that this could indicate it preyed upon the smaller, bird-like dinosaur. This discovery indicated that
461:
was a bipedal predator. The preserved length of the holotype specimen is 237 centimetres. Its weight was estimated by
1911:. The C-shaped abdominal contents of this specimen appear to reflect of the original contour of the digestive tract.
6409:
493:
which were small but exceptionally had denticles on their front edges. Only six, larger, teeth were present in the
5094:
2156:
2329:
5075:
6403:
6528:
2234:
404:
1809:
In 2024, Andrea Cau published a study on the phylogenetics of
Compsognathids that called the assessment of
450:. The specimen consists of a partial skull, tail, hands, feet and ribcage. It shows filaments on the tail.
6429:
2018:
might conform to this pattern as the authors considered it a possibly volant form. In 2011 a specimen of
628:
489:
had an elongated head with a pointed snout, showing a convex upper profile. There were four teeth in the
6538:
6500:
4800:
4712:
267:
117:
5828:
431:
6461:
6361:
4489:
4457:
4813:
4600:
2140:
Memorie della Società Italiana de
Scienze Naturali e del Museo Civico di Storia Naturale di Milano
368:
5082:
380:
6268:
4789:
4649:
4505:
4445:
3880:
3790:
3673:
2537:
2227:
1138:
133:
6505:
419:
6487:
6150:
6067:
6041:
5880:
5867:
5745:
5303:
5255:
4087:
3437:
2516:
1647:
891:
573:
not nearly as long or modern as the corresponding feathers of "four-winged" dinosaurs such as
427:
372:
125:
6492:
423:
6254:
5921:
5710:
5642:
5505:
5424:
5396:
5356:
5237:
5207:
4725:
4478:
4342:
4199:
4099:
3948:
3855:
3813:
3380:
2690:
2213:
2190:
2171:
2106:
1892:
1087:
1019:
651:
602:
480:
435:
320:
317:
30:
6317:
6055:
6018:
5950:
5805:
5798:
5626:
5568:
5541:
5531:
5491:
5326:
5185:
5169:
4987:
4959:
4937:
4824:
4669:
4376:
4150:
4033:
4011:
3932:
3761:
3738:
3731:
3709:
3695:
3688:
3649:
3500:
3347:
3274:
3263:
3199:
3087:
2974:
2551:
2460:
2314:
2289:
1828:
1814:
1733:
1555:
813:
670:
613:
564:
462:
222:
6282:
6245:
6226:
6188:
6177:
5974:
5687:
5653:
5635:
5615:
5512:
5226:
4971:
4833:
4660:
4642:
4470:
4437:
4423:
4384:
4158:
4019:
3976:
3969:
3891:
3867:
3820:
3779:
3716:
3702:
3430:
3415:
3242:
3076:
2940:
2908:
2897:
2874:
2836:
2697:
2667:
2591:
2530:
2429:
1990:
1973:
1923:
1823:
1634:
1172:
1070:
1053:
1036:
638:
476:
376:
6522:
6324:
6170:
6131:
6110:
5989:
5982:
5909:
5703:
5676:
5550:
5467:
5445:
5438:
5431:
5374:
5291:
5154:
5126:
5113:
5037:
4914:
4754:
4588:
4519:
4405:
4392:
4137:
4115:
4108:
4072:
4045:
3997:
3921:
3657:
3448:
3422:
3338:
3319:
3312:
3249:
3228:
3207:
3192:
3178:
3137:
3103:
3096:
3015:
2867:
2754:
2747:
2674:
2644:
2633:
2523:
2494:
2487:
2413:
1881:
1818:
1700:
1683:
1604:
1541:
1522:
1452:
1437:
1397:
1305:
1288:
976:
789:
765:
341:
332:
307:
68:
6296:
6234:
6204:
6124:
6117:
6095:
6087:
5967:
5959:
5939:
5662:
5581:
5519:
5410:
5381:
5333:
5162:
5147:
4922:
4854:
4782:
4761:
4740:
4556:
4512:
4320:
4307:
4237:
4190:
4065:
3955:
3907:
3900:
3837:
3806:
3799:
3680:
3507:
3359:
3256:
3158:
3151:
3046:
3030:
2966:
2917:
2849:
2626:
2509:
2502:
2473:
2436:
2405:
1979:
1961:
1941:
1509:
1356:
1247:
1213:
1155:
1121:
1104:
536:
361:
2194:
2110:
6350:
6331:
6261:
6211:
6197:
6139:
6080:
6025:
5902:
5696:
5588:
5557:
5474:
5460:
5192:
5140:
5008:
4930:
4893:
4775:
4747:
4700:
4620:
4498:
4355:
4206:
4179:
4079:
4026:
3962:
3941:
3914:
3665:
3478:
3471:
3457:
3283:
3130:
3121:
3023:
2994:
2924:
2889:
2882:
2682:
2660:
2653:
2612:
2604:
2580:
2565:
2544:
2452:
2444:
2020:
1953:
1932:
1527:
1414:
1339:
1322:
1264:
1230:
1196:
830:
575:
43:
528:
was expanded at the top at the side of the first metacarpal, making the entire
352:
6289:
6218:
6163:
5784:
5599:
5417:
5403:
5388:
5340:
5274:
5200:
5022:
5015:
5001:
4907:
4840:
4768:
4732:
4685:
4627:
4613:
4580:
4572:
4565:
4546:
4369:
4334:
4122:
3723:
3514:
3464:
3401:
3372:
3327:
3305:
3298:
3290:
3214:
3110:
3069:
3062:
3054:
3038:
2958:
2829:
2705:
2558:
2421:
2280:
2203:
1984:
1717:
959:
708:
559:
544:
529:
525:
490:
196:
88:
53:
6452:
1813:
into question. The paper recovered the taxon, along with four other proposed
6310:
6303:
6003:
5996:
5669:
5608:
5262:
4980:
4526:
4430:
4165:
3750:
3635:
3622:
3549:
3485:
3408:
3387:
3144:
3007:
2815:
2802:
2720:
2572:
2391:
2378:
2341:
2250:
1903:
1587:
1380:
998:
938:
691:
620:
581:
510:
447:
439:
209:
150:
93:
37:
6446:
5894:
5319:
5312:
5284:
5177:
4677:
4058:
3394:
3221:
2982:
2619:
2298:
2274:
2175:
1958:
1947:
1908:
551:
498:
415:
313:
310:
183:
170:
83:
78:
63:
58:
48:
6479:
5452:
5214:
4944:
4847:
4172:
3493:
1965:
1936:
1898:
1621:
562:(hairlike structures covering the skin), very similar to that found in
540:
517:
494:
98:
73:
6103:
5791:
3235:
2268:
443:
160:
6423:
4951:
4692:
4634:
2001:
1913:
1901:. The digested food would, using the preserved digestive tract of
592:
502:
475:
388:
351:
324:
303:
1869:
Undigested integument within the abdominal cavity of the holotype
6466:
6367:
4994:
535:
In the pelvis there was a little notch in the front edge of the
521:
6427:
5865:
5111:
4305:
3620:
2800:
2376:
2261:
2223:
399:, "feather". The large size of this "giant compsognathid" lent
2219:
2157:"A Unified Framework for Predatory Dinosaur Macroevolution"
558:
was preserved with "protofeathers," simple filamentous
520:
and also the lower arm was short and elegant with an
6436:
6349:
6244:
6187:
6149:
6066:
6039:
5949:
5919:
5878:
5776:
5744:
5686:
5652:
5625:
5598:
5567:
5540:
5529:
5489:
5354:
5302:
5273:
5235:
5224:
5124:
5036:
4970:
4892:
4823:
4799:
4711:
4659:
4599:
4555:
4544:
4488:
4456:
4403:
4353:
4318:
4236:
4189:
4136:
4098:
4044:
3995:
3931:
3890:
3866:
3835:
3789:
3759:
3748:
3633:
3548:
3447:
3358:
3337:
3273:
3176:
3120:
3086:
2993:
2938:
2907:
2847:
2813:
2739:
2719:
2643:
2590:
2471:
2389:
2340:
2313:
2134:Cristiano dal Sasso & Simone Maganuco, 2011,
2164:Bollettino della Società Paleontologica Italiana
1827:proper, which would make none of these species
627:in the Compsognathidae according to a study by
2063:
2061:
2235:
2059:
2057:
2055:
2053:
2051:
2049:
2047:
2045:
2043:
2041:
8:
1907:as a reference, have been positioned in the
302:(meaning 'Chinese beautiful feather') is a
6424:
6155:
6072:
6047:
5931:
5927:
5886:
5875:
5862:
5773:
5741:
5573:
5537:
5497:
5366:
5362:
5247:
5243:
5232:
5132:
5121:
5108:
4899:
4889:
4805:
4717:
4605:
4552:
4462:
4415:
4411:
4361:
4326:
4315:
4302:
4233:
4142:
4050:
4003:
3872:
3847:
3843:
3771:
3767:
3756:
3641:
3630:
3617:
3545:
3364:
3355:
3184:
2999:
2950:
2946:
2859:
2855:
2821:
2810:
2797:
2596:
2479:
2397:
2386:
2373:
2319:
2310:
2258:
2242:
2228:
2220:
2093:
2091:
2089:
2087:
2085:
2083:
2081:
116:
20:
1880:holotype contained the partial leg of a
1821:. This polytomy notably did not include
327:(Jianshangou Beds, dating to 124.6 Ma).
2037:
1999:, necessitating a regular food intake.
2123:The Princeton Field Guide to Dinosaurs
612:was by its describers assigned to the
7:
1939:stones) similar to those found with
2125:, Princeton University Press p. 121
2014:specialised in such prey. Even the
1876:The well-preserved skeleton of the
1662:
1640:
1547:
1515:
1430:
1372:
1280:
1188:
1011:
1004:
951:
944:
934:
805:
781:
757:
733:
683:
676:
666:
644:
634:
539:. The relatively long shaft of the
418:, JMP-V-05-8-01, was discovered at
383:. The generic name is derived from
367:was named and described in 2007 by
6534:Early Cretaceous dinosaurs of Asia
1855:Abdominal contents of the holotype
14:
513:had very short lateral segments.
391:for the Chinese and Greek καλός,
6408:
6402:
6395:
5847:
5840:
5833:
5827:
5093:
5086:
5081:
5074:
5068:
4287:
4280:
4273:
3602:
3595:
3588:
3581:
2783:
2777:
2206:
1862:
1848:
923:Here is a simplified version of
137:
465:in 2010 at twenty kilogrammes.
2006:Stomach contents of CAGS-IG-T1
1531:at an indeterminate position)
1:
6549:Fossil taxa described in 2007
330:While similar to the related
2195:10.1080/08912963.2012.654705
2111:10.1371/journal.pone.0044012
356:Assigned specimen CAGS-IG-T1
1951:. Other theropods, such as
1817:in a polytomy within basal
395:, "beautiful", and πτέρυξ,
6570:
1306:Compsognathus corallestris
16:Extinct genus of dinosaurs
6554:Taxa named by Lü Junchang
6392:
6158:
6075:
6050:
5934:
5930:
5889:
5874:
5861:
5825:
5772:
5740:
5576:
5500:
5369:
5365:
5250:
5246:
5135:
5120:
5107:
5066:
4902:
4888:
4808:
4720:
4608:
4465:
4418:
4414:
4364:
4329:
4314:
4301:
4270:
4232:
4145:
4053:
4006:
3875:
3850:
3846:
3774:
3770:
3644:
3629:
3616:
3578:
3544:
3367:
3187:
3002:
2953:
2949:
2862:
2858:
2824:
2809:
2796:
2774:
2599:
2482:
2400:
2385:
2372:
2322:
2309:
2257:
1968:would have been present.
1731:
1714:
1697:
1680:
1667:
1660:
1645:
1638:
1618:
1601:
1584:
1569:
1552:
1545:
1520:
1513:
1450:
1435:
1428:
1411:
1394:
1377:
1370:
1353:
1336:
1319:
1302:
1285:
1278:
1261:
1244:
1227:
1210:
1193:
1186:
1169:
1152:
1135:
1118:
1101:
1084:
1067:
1050:
1033:
1016:
1009:
1002:
973:
956:
949:
942:
889:
827:
810:
803:
786:
779:
762:
755:
738:
731:
705:
688:
681:
674:
664:
649:
642:
554:of the Yixian Formation,
273:
266:
134:Scientific classification
132:
124:
115:
23:
631:& Maganuco in 2011:
2007:
1927:
1289:Compsognathus longipes
623:shows the position of
606:
483:
469:is distinguished from
365:Sinocalliopteryx gigas
357:
280:Sinocalliopteryx gigas
6501:Paleobiology Database
4801:Carcharodontosaurinae
4713:Carcharodontosauridae
2069:Acta Geologica Sinica
2005:
1917:
1897:still present in the
596:
479:
355:
5777:Dubious coelurosaurs
4490:Metriacanthosaurinae
4458:Metriacanthosauridae
2740:Dubious neotheropods
2176:10.4435/BSPI.2024.08
2155:Cau, Andrea (2024).
2136:Scipionyx samniticus
568:. The integument of
516:The arm had a short
6544:Feathered dinosaurs
4814:Carcharodontosaurus
4601:Carcharodontosauria
1340:Torvosaurus gurneyi
1323:Torvosaurus tanneri
411:, meaning 'giant'.
6269:Archaeornithomimus
4790:Veterupristisaurus
4650:Veterupristisaurus
4506:Metriacanthosaurus
4446:Piatnitzkysauridae
3881:Eustreptospondylus
3791:Piatnitzkysauridae
3674:Chuandongocoelurus
2538:Notatesseraeraptor
2121:Paul, G.S., 2010,
2008:
1928:
1801:The large size of
1139:Eustreptospondylus
607:
484:
358:
6516:
6515:
6488:Open Tree of Life
6430:Taxon identifiers
6421:
6420:
6417:
6416:
6390:
6389:
6386:
6385:
6382:
6381:
6378:
6377:
6345:
6344:
6341:
6340:
6151:Macrocheiriformes
6068:Ornithomimosauria
6042:Maniraptoriformes
6035:
6034:
5881:Maniraptoromorpha
5868:Maniraptoromorpha
5857:
5856:
5823:
5822:
5819:
5818:
5815:
5814:
5768:
5767:
5764:
5763:
5746:Maniraptoromorpha
5736:
5735:
5732:
5731:
5728:
5727:
5724:
5723:
5720:
5719:
5485:
5484:
5350:
5349:
5304:Proceratosauridae
5256:Chingkankousaurus
5103:
5102:
5064:
5063:
5060:
5059:
5056:
5055:
5032:
5031:
4884:
4883:
4880:
4879:
4876:
4875:
4872:
4871:
4868:
4867:
4864:
4863:
4540:
4539:
4536:
4535:
4297:
4296:
4268:
4267:
4264:
4263:
4260:
4259:
4256:
4255:
4228:
4227:
4224:
4223:
4220:
4219:
4216:
4215:
4132:
4131:
4088:Vallibonavenatrix
3991:
3990:
3987:
3986:
3831:
3830:
3612:
3611:
3576:
3575:
3572:
3571:
3568:
3567:
3540:
3539:
3536:
3535:
3532:
3531:
3528:
3527:
3524:
3523:
3172:
3171:
3168:
3167:
2934:
2933:
2792:
2791:
2772:
2771:
2768:
2767:
2764:
2763:
2715:
2714:
2368:
2367:
2364:
2363:
2360:
2359:
1798:
1797:
1789:
1788:
1780:
1779:
1771:
1770:
1762:
1761:
1753:
1752:
1744:
1743:
1648:Maniraptoromorpha
1499:
1498:
1490:
1489:
1481:
1480:
1472:
1471:
1463:
1462:
988:
987:
920:
919:
911:
910:
902:
901:
892:Maniraptoriformes
878:
877:
869:
868:
860:
859:
851:
850:
842:
841:
720:
719:
428:Liaoning Province
340:were larger. The
295:
294:
247:
126:Holotype specimen
6561:
6509:
6508:
6496:
6495:
6483:
6482:
6470:
6469:
6457:
6456:
6455:
6438:Sinocalliopteryx
6425:
6412:
6406:
6399:
6255:Aepyornithomimus
6156:
6073:
6048:
6012:Sinocalliopteryx
5932:
5928:
5922:Neocoelurosauria
5887:
5876:
5863:
5851:
5844:
5837:
5831:
5774:
5742:
5711:Zhuchengtyrannus
5643:Thanatotheristes
5574:
5538:
5506:Appalachiosaurus
5498:
5367:
5363:
5357:Pantyrannosauria
5248:
5244:
5238:Tyrannosauroidea
5233:
5208:Xinjiangovenator
5133:
5122:
5109:
5097:
5090:
5085:
5078:
5072:
4900:
4890:
4806:
4726:Acrocanthosaurus
4718:
4606:
4553:
4479:Yangchuanosaurus
4463:
4416:
4412:
4362:
4343:Lourinhanosaurus
4327:
4316:
4303:
4291:
4284:
4277:
4234:
4200:Sigilmassasaurus
4143:
4100:Ceratosuchopsini
4051:
4004:
3949:Dubreuillosaurus
3873:
3856:Streptospondylus
3848:
3844:
3814:Piatnitzkysaurus
3772:
3768:
3757:
3642:
3631:
3618:
3606:
3599:
3592:
3585:
3546:
3381:Ekrixinatosaurus
3365:
3356:
3185:
3000:
2951:
2947:
2860:
2856:
2822:
2811:
2798:
2787:
2781:
2691:Procompsognathus
2597:
2480:
2398:
2387:
2374:
2320:
2311:
2304:
2303:
2259:
2244:
2237:
2230:
2221:
2216:
2214:Dinosaurs portal
2211:
2210:
2209:
2198:
2186:
2180:
2179:
2161:
2152:
2146:
2132:
2126:
2119:
2113:
2095:
2076:
2065:
2026:Sinocalliopteryx
2016:Sinornithosaurus
2012:Sinocalliopteryx
1997:Sinocalliopteryx
1919:Sinocalliopteryx
1893:Sinornithosaurus
1887:Sinocalliopteryx
1878:Sinocalliopteryx
1866:
1852:
1840:Diet and feeding
1811:Sinocalliopteryx
1803:Sinocalliopteryx
1663:
1641:
1572:Sinocalliopteryx
1548:
1516:
1431:
1373:
1281:
1189:
1088:Piatnitzkysaurus
1020:Streptospondylus
1012:
1005:
952:
945:
935:
929:Sinocalliopteryx
806:
782:
758:
742:Sinocalliopteryx
734:
684:
677:
667:
652:Tyrannosauroidea
645:
635:
625:Sinocalliopteryx
610:Sinocalliopteryx
603:Sinornithosaurus
598:Sinocalliopteryx
570:Sinocalliopteryx
556:Sinocalliopteryx
550:Like many other
487:Sinocalliopteryx
481:Life restoration
467:Sinocalliopteryx
459:Sinocalliopteryx
438:dating from the
436:Yixian Formation
432:Jianshangou Beds
401:Sinocalliopteryx
338:Sinocalliopteryx
321:Yixian Formation
318:Lower Cretaceous
299:Sinocalliopteryx
282:
278:
241:
237:Sinocalliopteryx
234:
221:
142:
141:
120:
110:
40:
31:Early Cretaceous
29:Temporal range:
25:Sinocalliopteryx
21:
6569:
6568:
6564:
6563:
6562:
6560:
6559:
6558:
6519:
6518:
6517:
6512:
6504:
6499:
6491:
6486:
6478:
6473:
6465:
6460:
6451:
6450:
6445:
6432:
6422:
6413:
6400:
6374:
6337:
6318:Sinornithomimus
6240:
6183:
6145:
6062:
6056:Compsognathidae
6044:
6031:
6019:Sinosauropteryx
5951:Compsognathidae
5945:
5924:
5915:
5883:
5870:
5853:
5852:
5838:
5811:
5806:Shanyangosaurus
5799:Phaedrolosaurus
5760:
5716:
5682:
5648:
5627:Daspletosaurini
5621:
5594:
5569:Tyrannosaurinae
5563:
5542:Albertosaurinae
5534:
5532:Tyrannosauridae
5525:
5494:
5492:Eutyrannosauria
5481:
5359:
5346:
5327:Proceratosaurus
5298:
5269:
5240:
5229:
5220:
5186:Richardoestesia
5170:Chilantaisaurus
5129:
5116:
5099:
5098:
5079:
5052:
5028:
4988:Australovenator
4966:
4960:Chilantaisaurus
4938:Phuwiangvenator
4860:
4825:Giganotosaurini
4819:
4795:
4707:
4670:Chilantaisaurus
4655:
4595:
4549:
4532:
4484:
4452:
4408:
4399:
4377:Monolophosaurus
4358:
4349:
4323:
4310:
4293:
4292:
4278:
4252:
4212:
4185:
4151:Camarillasaurus
4128:
4094:
4040:
4034:Ostafrikasaurus
4012:Camarillasaurus
4000:
3983:
3933:Afrovenatorinae
3927:
3886:
3862:
3840:
3827:
3785:
3764:
3762:Megalosauroidea
3753:
3744:
3739:Vectaerovenator
3732:Szechuanosaurus
3710:Monolophosaurus
3696:Kaijiangosaurus
3689:Cryolophosaurus
3650:Calamospondylus
3638:
3625:
3608:
3607:
3600:
3593:
3586:
3564:
3520:
3501:Pycnonemosaurus
3443:
3354:
3348:Majungasaurinae
3333:
3275:Majungasaurinae
3269:
3264:Xenotarsosaurus
3200:Dryptosauroides
3181:
3164:
3116:
3088:Elaphrosaurinae
3082:
2989:
2975:Ornithomimoides
2943:
2930:
2903:
2852:
2843:
2818:
2805:
2788:
2782:
2760:
2735:
2711:
2639:
2586:
2552:Shuangbaisaurus
2476:
2467:
2394:
2381:
2356:
2336:
2330:Avemetatarsalia
2315:Avemetatarsalia
2305:
2290:Avemetatarsalia
2264:
2263:
2253:
2248:
2212:
2207:
2205:
2202:
2201:
2187:
2183:
2159:
2154:
2153:
2149:
2133:
2129:
2120:
2116:
2096:
2079:
2066:
2039:
2034:
1874:
1873:
1872:
1871:
1870:
1867:
1858:
1857:
1856:
1853:
1842:
1837:
1799:
1790:
1781:
1772:
1763:
1754:
1745:
1734:Eutyrannosauria
1556:Sinosauropteryx
1500:
1491:
1482:
1473:
1464:
989:
921:
912:
903:
879:
870:
861:
852:
843:
814:Sinosauropteryx
721:
671:Compsognathidae
614:Compsognathidae
591:
565:Sinosauropteryx
463:Gregory S. Paul
456:
350:
306:of carnivorous
291:
284:
276:
275:
262:
240:
232:
223:Compsognathidae
219:
136:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
27:
17:
12:
11:
5:
6567:
6565:
6557:
6556:
6551:
6546:
6541:
6536:
6531:
6529:Compsognathids
6521:
6520:
6514:
6513:
6511:
6510:
6497:
6484:
6471:
6458:
6442:
6440:
6434:
6433:
6428:
6419:
6418:
6415:
6414:
6394:
6393:
6391:
6388:
6387:
6384:
6383:
6380:
6379:
6376:
6375:
6373:
6372:
6371:
6370:
6364:
6355:
6353:
6347:
6346:
6343:
6342:
6339:
6338:
6336:
6335:
6328:
6321:
6314:
6307:
6300:
6293:
6286:
6283:Dromiceiomimus
6279:
6272:
6265:
6258:
6250:
6248:
6246:Ornithomimidae
6242:
6241:
6239:
6238:
6231:
6227:Paraxenisaurus
6223:
6215:
6208:
6201:
6193:
6191:
6189:Deinocheiridae
6185:
6184:
6182:
6181:
6178:Shenzhousaurus
6174:
6167:
6159:
6153:
6147:
6146:
6144:
6143:
6136:
6128:
6121:
6114:
6107:
6100:
6092:
6084:
6076:
6070:
6064:
6063:
6061:
6060:
6051:
6045:
6040:
6037:
6036:
6033:
6032:
6030:
6029:
6022:
6015:
6008:
6000:
5993:
5986:
5979:
5975:Beipiaognathus
5971:
5964:
5955:
5953:
5947:
5946:
5944:
5943:
5935:
5925:
5920:
5917:
5916:
5914:
5913:
5906:
5899:
5890:
5884:
5879:
5872:
5871:
5866:
5859:
5858:
5855:
5854:
5846:
5832:
5826:
5824:
5821:
5820:
5817:
5816:
5813:
5812:
5810:
5809:
5802:
5795:
5788:
5780:
5778:
5770:
5769:
5766:
5765:
5762:
5761:
5759:
5758:
5757:
5756:
5750:
5748:
5738:
5737:
5734:
5733:
5730:
5729:
5726:
5725:
5722:
5721:
5718:
5717:
5715:
5714:
5707:
5700:
5692:
5690:
5688:Tyrannosaurini
5684:
5683:
5681:
5680:
5673:
5666:
5658:
5656:
5654:Teratophoneini
5650:
5649:
5647:
5646:
5639:
5636:Daspletosaurus
5631:
5629:
5623:
5622:
5620:
5619:
5616:Qianzhousaurus
5612:
5604:
5602:
5596:
5595:
5593:
5592:
5585:
5577:
5571:
5565:
5564:
5562:
5561:
5554:
5546:
5544:
5535:
5530:
5527:
5526:
5524:
5523:
5516:
5513:Bistahieversor
5509:
5501:
5495:
5490:
5487:
5486:
5483:
5482:
5480:
5479:
5471:
5464:
5457:
5449:
5442:
5435:
5428:
5421:
5414:
5407:
5400:
5393:
5385:
5378:
5370:
5360:
5355:
5352:
5351:
5348:
5347:
5345:
5344:
5337:
5330:
5323:
5316:
5308:
5306:
5300:
5299:
5297:
5296:
5288:
5280:
5278:
5271:
5270:
5268:
5267:
5259:
5251:
5241:
5236:
5230:
5227:Tyrannoraptora
5225:
5222:
5221:
5219:
5218:
5211:
5204:
5197:
5189:
5182:
5174:
5166:
5159:
5151:
5144:
5136:
5130:
5125:
5118:
5117:
5112:
5105:
5104:
5101:
5100:
5092:
5073:
5067:
5065:
5062:
5061:
5058:
5057:
5054:
5053:
5051:
5050:
5049:
5048:
5042:
5040:
5034:
5033:
5030:
5029:
5027:
5026:
5019:
5012:
5005:
4998:
4991:
4984:
4976:
4974:
4972:Megaraptoridae
4968:
4967:
4965:
4964:
4956:
4948:
4941:
4934:
4927:
4919:
4911:
4903:
4897:
4886:
4885:
4882:
4881:
4878:
4877:
4874:
4873:
4870:
4869:
4866:
4865:
4862:
4861:
4859:
4858:
4851:
4844:
4837:
4834:Giganotosaurus
4829:
4827:
4821:
4820:
4818:
4817:
4809:
4803:
4797:
4796:
4794:
4793:
4786:
4779:
4772:
4765:
4758:
4751:
4744:
4737:
4729:
4721:
4715:
4709:
4708:
4706:
4705:
4697:
4689:
4682:
4674:
4665:
4663:
4661:Neovenatoridae
4657:
4656:
4654:
4653:
4646:
4643:Ulughbegsaurus
4639:
4631:
4624:
4617:
4609:
4603:
4597:
4596:
4594:
4593:
4585:
4577:
4569:
4561:
4559:
4550:
4545:
4542:
4541:
4538:
4537:
4534:
4533:
4531:
4530:
4523:
4516:
4509:
4502:
4494:
4492:
4486:
4485:
4483:
4482:
4475:
4471:Xuanhanosaurus
4466:
4460:
4454:
4453:
4451:
4450:
4442:
4438:Xuanhanosaurus
4434:
4427:
4424:Asfaltovenator
4419:
4409:
4404:
4401:
4400:
4398:
4397:
4389:
4385:Megalosauridae
4381:
4373:
4365:
4359:
4354:
4351:
4350:
4348:
4347:
4339:
4330:
4324:
4319:
4312:
4311:
4306:
4299:
4298:
4295:
4294:
4286:
4272:
4271:
4269:
4266:
4265:
4262:
4261:
4258:
4257:
4254:
4253:
4251:
4250:
4249:
4248:
4242:
4240:
4230:
4229:
4226:
4225:
4222:
4221:
4218:
4217:
4214:
4213:
4211:
4210:
4203:
4195:
4193:
4187:
4186:
4184:
4183:
4176:
4169:
4162:
4159:Ichthyovenator
4155:
4146:
4140:
4134:
4133:
4130:
4129:
4127:
4126:
4119:
4112:
4104:
4102:
4096:
4095:
4093:
4092:
4084:
4076:
4069:
4062:
4054:
4048:
4042:
4041:
4039:
4038:
4030:
4023:
4020:Cristatusaurus
4016:
4007:
4001:
3996:
3993:
3992:
3989:
3988:
3985:
3984:
3982:
3981:
3977:Poekilopleuron
3973:
3970:Piveteausaurus
3966:
3959:
3952:
3945:
3937:
3935:
3929:
3928:
3926:
3925:
3918:
3911:
3904:
3896:
3894:
3892:Megalosaurinae
3888:
3887:
3885:
3884:
3876:
3870:
3868:Megalosauridae
3864:
3863:
3861:
3860:
3851:
3841:
3836:
3833:
3832:
3829:
3828:
3826:
3825:
3821:Xuanhanosaurus
3817:
3810:
3803:
3795:
3793:
3787:
3786:
3784:
3783:
3780:Yunyangosaurus
3775:
3765:
3760:
3754:
3749:
3746:
3745:
3743:
3742:
3735:
3728:
3720:
3717:Pandoravenator
3713:
3706:
3703:Kayentavenator
3699:
3692:
3685:
3677:
3670:
3662:
3654:
3645:
3639:
3634:
3627:
3626:
3621:
3614:
3613:
3610:
3609:
3601:
3594:
3587:
3580:
3579:
3577:
3574:
3573:
3570:
3569:
3566:
3565:
3563:
3562:
3561:
3560:
3554:
3552:
3542:
3541:
3538:
3537:
3534:
3533:
3530:
3529:
3526:
3525:
3522:
3521:
3519:
3518:
3511:
3504:
3497:
3490:
3482:
3475:
3468:
3461:
3453:
3451:
3445:
3444:
3442:
3441:
3434:
3431:Skorpiovenator
3427:
3419:
3412:
3405:
3398:
3391:
3384:
3377:
3368:
3362:
3353:
3352:
3343:
3341:
3335:
3334:
3332:
3331:
3324:
3316:
3309:
3302:
3295:
3287:
3279:
3277:
3271:
3270:
3268:
3267:
3260:
3253:
3246:
3243:Spectrovenator
3239:
3232:
3225:
3218:
3211:
3204:
3196:
3188:
3182:
3177:
3174:
3173:
3170:
3169:
3166:
3165:
3163:
3162:
3155:
3148:
3141:
3134:
3126:
3124:
3118:
3117:
3115:
3114:
3107:
3100:
3092:
3090:
3084:
3083:
3081:
3080:
3077:Spinostropheus
3073:
3066:
3059:
3051:
3043:
3035:
3027:
3020:
3012:
3003:
2997:
2991:
2990:
2988:
2987:
2979:
2971:
2963:
2954:
2944:
2941:Abelisauroidea
2939:
2936:
2935:
2932:
2931:
2929:
2928:
2921:
2913:
2911:
2909:Ceratosauridae
2905:
2904:
2902:
2901:
2898:Saltriovenator
2894:
2886:
2879:
2875:Fosterovenator
2871:
2863:
2853:
2848:
2845:
2844:
2842:
2841:
2837:Lophostropheus
2833:
2825:
2819:
2814:
2807:
2806:
2801:
2794:
2793:
2790:
2789:
2776:
2775:
2773:
2770:
2769:
2766:
2765:
2762:
2761:
2759:
2758:
2751:
2743:
2741:
2737:
2736:
2734:
2733:
2732:
2731:
2725:
2723:
2717:
2716:
2713:
2712:
2710:
2709:
2702:
2698:Pterospondylus
2694:
2687:
2679:
2671:
2668:Lucianovenator
2664:
2657:
2649:
2647:
2641:
2640:
2638:
2637:
2630:
2623:
2616:
2609:
2600:
2594:
2592:Coelophysoidea
2588:
2587:
2585:
2584:
2577:
2569:
2562:
2555:
2548:
2541:
2534:
2531:Lophostropheus
2527:
2520:
2513:
2506:
2499:
2491:
2483:
2477:
2472:
2469:
2468:
2466:
2465:
2457:
2449:
2441:
2433:
2430:Erythrovenator
2426:
2418:
2410:
2401:
2395:
2390:
2383:
2382:
2377:
2370:
2369:
2366:
2365:
2362:
2361:
2358:
2357:
2355:
2354:
2353:
2352:
2346:
2344:
2338:
2337:
2335:
2334:
2333:
2332:
2323:
2317:
2307:
2306:
2302:
2301:
2292:
2283:
2277:
2271:
2262:
2255:
2254:
2249:
2247:
2246:
2239:
2232:
2224:
2218:
2217:
2200:
2199:
2181:
2147:
2127:
2114:
2077:
2036:
2035:
2033:
2030:
1991:Psittacosaurus
1974:Confuciusornis
1924:Confuciusornis
1868:
1861:
1860:
1859:
1854:
1847:
1846:
1845:
1844:
1843:
1841:
1838:
1836:
1833:
1829:Compsognathids
1824:Composognathus
1815:Compsognathids
1796:
1795:
1792:
1791:
1787:
1786:
1783:
1782:
1778:
1777:
1774:
1773:
1769:
1768:
1765:
1764:
1760:
1759:
1756:
1755:
1751:
1750:
1747:
1746:
1742:
1741:
1738:
1737:
1730:
1727:
1726:
1723:
1722:
1713:
1710:
1709:
1706:
1705:
1696:
1693:
1692:
1689:
1688:
1679:
1676:
1675:
1672:
1671:
1666:
1661:
1659:
1656:
1655:
1652:
1651:
1644:
1639:
1637:
1635:Tyrannoraptora
1631:
1630:
1627:
1626:
1617:
1614:
1613:
1610:
1609:
1600:
1597:
1596:
1593:
1592:
1583:
1580:
1579:
1576:
1575:
1568:
1565:
1564:
1561:
1560:
1551:
1546:
1544:
1538:
1537:
1534:
1533:
1519:
1514:
1512:
1506:
1505:
1502:
1501:
1497:
1496:
1493:
1492:
1488:
1487:
1484:
1483:
1479:
1478:
1475:
1474:
1470:
1469:
1466:
1465:
1461:
1460:
1457:
1456:
1449:
1446:
1445:
1442:
1441:
1434:
1429:
1427:
1424:
1423:
1420:
1419:
1410:
1407:
1406:
1403:
1402:
1393:
1390:
1389:
1386:
1385:
1376:
1371:
1369:
1366:
1365:
1362:
1361:
1352:
1349:
1348:
1345:
1344:
1335:
1332:
1331:
1328:
1327:
1318:
1315:
1314:
1311:
1310:
1301:
1298:
1297:
1294:
1293:
1284:
1279:
1277:
1274:
1273:
1270:
1269:
1260:
1257:
1256:
1253:
1252:
1243:
1240:
1239:
1236:
1235:
1226:
1223:
1222:
1219:
1218:
1209:
1206:
1205:
1202:
1201:
1192:
1187:
1185:
1182:
1181:
1178:
1177:
1173:Asfaltovenator
1168:
1165:
1164:
1161:
1160:
1151:
1148:
1147:
1144:
1143:
1134:
1131:
1130:
1127:
1126:
1117:
1114:
1113:
1110:
1109:
1100:
1097:
1096:
1093:
1092:
1083:
1080:
1079:
1076:
1075:
1071:Piveteausaurus
1066:
1063:
1062:
1059:
1058:
1054:Poekilopleuron
1049:
1046:
1045:
1042:
1041:
1037:Xuanhanosaurus
1032:
1029:
1028:
1025:
1024:
1015:
1010:
1008:
1003:
1001:
995:
994:
991:
990:
986:
985:
982:
981:
972:
969:
968:
965:
964:
955:
950:
948:
943:
941:
933:
918:
917:
914:
913:
909:
908:
905:
904:
900:
899:
896:
895:
888:
885:
884:
881:
880:
876:
875:
872:
871:
867:
866:
863:
862:
858:
857:
854:
853:
849:
848:
845:
844:
840:
839:
836:
835:
826:
823:
822:
819:
818:
809:
804:
802:
799:
798:
795:
794:
785:
780:
778:
775:
774:
771:
770:
761:
756:
754:
751:
750:
747:
746:
737:
732:
730:
727:
726:
723:
722:
718:
717:
714:
713:
704:
701:
700:
697:
696:
687:
682:
680:
675:
673:
665:
663:
660:
659:
656:
655:
648:
643:
641:
639:Tyrannoraptora
633:
590:
589:Classification
587:
455:
452:
349:
346:
293:
292:
285:
271:
270:
264:
263:
255:
253:
249:
248:
230:
226:
225:
217:
213:
212:
207:
200:
199:
194:
187:
186:
181:
174:
173:
168:
164:
163:
158:
154:
153:
148:
144:
143:
130:
129:
128:JMP-V-05-8-01
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
6566:
6555:
6552:
6550:
6547:
6545:
6542:
6540:
6537:
6535:
6532:
6530:
6527:
6526:
6524:
6507:
6502:
6498:
6494:
6489:
6485:
6481:
6476:
6472:
6468:
6463:
6459:
6454:
6448:
6444:
6443:
6441:
6439:
6435:
6431:
6426:
6411:
6407:
6405:
6398:
6369:
6365:
6363:
6359:
6358:
6357:
6356:
6354:
6352:
6348:
6334:
6333:
6329:
6327:
6326:
6325:Struthiomimus
6322:
6320:
6319:
6315:
6313:
6312:
6308:
6306:
6305:
6301:
6299:
6298:
6294:
6292:
6291:
6287:
6285:
6284:
6280:
6278:
6277:
6276:"Coelosaurus"
6273:
6271:
6270:
6266:
6264:
6263:
6259:
6257:
6256:
6252:
6251:
6249:
6247:
6243:
6237:
6236:
6232:
6229:
6228:
6224:
6221:
6220:
6216:
6214:
6213:
6209:
6207:
6206:
6202:
6200:
6199:
6195:
6194:
6192:
6190:
6186:
6180:
6179:
6175:
6173:
6172:
6171:Pelecanimimus
6168:
6166:
6165:
6161:
6160:
6157:
6154:
6152:
6148:
6142:
6141:
6137:
6134:
6133:
6132:Thecocoelurus
6129:
6127:
6126:
6122:
6120:
6119:
6115:
6113:
6112:
6111:Kinnareemimus
6108:
6106:
6105:
6101:
6098:
6097:
6093:
6090:
6089:
6085:
6083:
6082:
6078:
6077:
6074:
6071:
6069:
6065:
6058:
6057:
6053:
6052:
6049:
6046:
6043:
6038:
6028:
6027:
6023:
6021:
6020:
6016:
6014:
6013:
6009:
6006:
6005:
6001:
5999:
5998:
5994:
5992:
5991:
5990:Huaxiagnathus
5987:
5985:
5984:
5983:Compsognathus
5980:
5977:
5976:
5972:
5970:
5969:
5965:
5962:
5961:
5957:
5956:
5954:
5952:
5948:
5942:
5941:
5937:
5936:
5933:
5929:
5926:
5923:
5918:
5912:
5911:
5910:Ornitholestes
5907:
5905:
5904:
5900:
5897:
5896:
5892:
5891:
5888:
5885:
5882:
5877:
5873:
5869:
5864:
5860:
5850:
5845:
5843:
5836:
5830:
5808:
5807:
5803:
5801:
5800:
5796:
5794:
5793:
5789:
5787:
5786:
5782:
5781:
5779:
5775:
5771:
5754:
5753:
5752:
5751:
5749:
5747:
5743:
5739:
5713:
5712:
5708:
5706:
5705:
5704:Tyrannosaurus
5701:
5699:
5698:
5694:
5693:
5691:
5689:
5685:
5679:
5678:
5677:Teratophoneus
5674:
5672:
5671:
5667:
5665:
5664:
5660:
5659:
5657:
5655:
5651:
5645:
5644:
5640:
5638:
5637:
5633:
5632:
5630:
5628:
5624:
5618:
5617:
5613:
5611:
5610:
5606:
5605:
5603:
5601:
5597:
5591:
5590:
5586:
5584:
5583:
5579:
5578:
5575:
5572:
5570:
5566:
5560:
5559:
5555:
5553:
5552:
5551:Albertosaurus
5548:
5547:
5545:
5543:
5539:
5536:
5533:
5528:
5522:
5521:
5517:
5515:
5514:
5510:
5508:
5507:
5503:
5502:
5499:
5496:
5493:
5488:
5477:
5476:
5472:
5470:
5469:
5468:Xiongguanlong
5465:
5463:
5462:
5458:
5455:
5454:
5450:
5448:
5447:
5446:Suskityrannus
5443:
5441:
5440:
5439:Stokesosaurus
5436:
5434:
5433:
5432:Santanaraptor
5429:
5427:
5426:
5422:
5420:
5419:
5415:
5413:
5412:
5408:
5406:
5405:
5401:
5399:
5398:
5394:
5391:
5390:
5386:
5384:
5383:
5379:
5377:
5376:
5375:Alectrosaurus
5372:
5371:
5368:
5364:
5361:
5358:
5353:
5343:
5342:
5338:
5336:
5335:
5331:
5329:
5328:
5324:
5322:
5321:
5317:
5315:
5314:
5310:
5309:
5307:
5305:
5301:
5294:
5293:
5292:Tanycolagreus
5289:
5287:
5286:
5282:
5281:
5279:
5276:
5272:
5265:
5264:
5260:
5258:
5257:
5253:
5252:
5249:
5245:
5242:
5239:
5234:
5231:
5228:
5223:
5217:
5216:
5212:
5210:
5209:
5205:
5203:
5202:
5198:
5195:
5194:
5190:
5188:
5187:
5183:
5180:
5179:
5175:
5172:
5171:
5167:
5165:
5164:
5160:
5157:
5156:
5155:Bahariasaurus
5152:
5150:
5149:
5145:
5143:
5142:
5138:
5137:
5134:
5131:
5128:
5127:Coelurosauria
5123:
5119:
5115:
5114:Coelurosauria
5110:
5106:
5096:
5091:
5089:
5084:
5077:
5071:
5046:
5045:
5044:
5043:
5041:
5039:
5038:Coelurosauria
5035:
5025:
5024:
5020:
5018:
5017:
5013:
5011:
5010:
5006:
5004:
5003:
4999:
4997:
4996:
4992:
4990:
4989:
4985:
4983:
4982:
4978:
4977:
4975:
4973:
4969:
4963:
4961:
4957:
4955:
4953:
4949:
4947:
4946:
4942:
4940:
4939:
4935:
4933:
4932:
4928:
4925:
4924:
4920:
4917:
4916:
4915:Bahariasaurus
4912:
4910:
4909:
4905:
4904:
4901:
4898:
4895:
4891:
4887:
4857:
4856:
4852:
4850:
4849:
4845:
4843:
4842:
4838:
4836:
4835:
4831:
4830:
4828:
4826:
4822:
4816:
4815:
4811:
4810:
4807:
4804:
4802:
4798:
4792:
4791:
4787:
4785:
4784:
4780:
4778:
4777:
4773:
4771:
4770:
4766:
4764:
4763:
4759:
4757:
4756:
4755:Kelmayisaurus
4752:
4750:
4749:
4745:
4743:
4742:
4738:
4735:
4734:
4730:
4728:
4727:
4723:
4722:
4719:
4716:
4714:
4710:
4703:
4702:
4698:
4695:
4694:
4690:
4688:
4687:
4683:
4680:
4679:
4675:
4672:
4671:
4667:
4666:
4664:
4662:
4658:
4652:
4651:
4647:
4645:
4644:
4640:
4637:
4636:
4632:
4630:
4629:
4625:
4623:
4622:
4618:
4616:
4615:
4611:
4610:
4607:
4604:
4602:
4598:
4591:
4590:
4589:Saurophaganax
4586:
4583:
4582:
4578:
4575:
4574:
4570:
4568:
4567:
4563:
4562:
4560:
4558:
4554:
4551:
4548:
4543:
4529:
4528:
4524:
4522:
4521:
4520:Siamotyrannus
4517:
4515:
4514:
4510:
4508:
4507:
4503:
4501:
4500:
4496:
4495:
4493:
4491:
4487:
4481:
4480:
4476:
4473:
4472:
4468:
4467:
4464:
4461:
4459:
4455:
4448:
4447:
4443:
4440:
4439:
4435:
4433:
4432:
4428:
4426:
4425:
4421:
4420:
4417:
4413:
4410:
4407:
4406:Allosauroidea
4402:
4395:
4394:
4393:Spinosauridae
4390:
4387:
4386:
4382:
4379:
4378:
4374:
4372:
4371:
4367:
4366:
4363:
4360:
4357:
4352:
4345:
4344:
4340:
4337:
4336:
4332:
4331:
4328:
4325:
4322:
4317:
4313:
4309:
4304:
4300:
4290:
4285:
4283:
4276:
4246:
4245:
4244:
4243:
4241:
4239:
4235:
4231:
4209:
4208:
4204:
4202:
4201:
4197:
4196:
4194:
4192:
4188:
4182:
4181:
4177:
4175:
4174:
4170:
4168:
4167:
4163:
4161:
4160:
4156:
4153:
4152:
4148:
4147:
4144:
4141:
4139:
4138:Spinosaurinae
4135:
4125:
4124:
4120:
4118:
4117:
4116:Riparovenator
4113:
4111:
4110:
4109:Ceratosuchops
4106:
4105:
4103:
4101:
4097:
4090:
4089:
4085:
4082:
4081:
4077:
4075:
4074:
4073:Riojavenatrix
4070:
4068:
4067:
4063:
4061:
4060:
4056:
4055:
4052:
4049:
4047:
4046:Baryonychinae
4043:
4036:
4035:
4031:
4029:
4028:
4024:
4022:
4021:
4017:
4014:
4013:
4009:
4008:
4005:
4002:
3999:
3998:Spinosauridae
3994:
3979:
3978:
3974:
3972:
3971:
3967:
3965:
3964:
3960:
3958:
3957:
3953:
3951:
3950:
3946:
3944:
3943:
3939:
3938:
3936:
3934:
3930:
3924:
3923:
3922:Wiehenvenator
3919:
3917:
3916:
3912:
3910:
3909:
3905:
3903:
3902:
3898:
3897:
3895:
3893:
3889:
3883:
3882:
3878:
3877:
3874:
3871:
3869:
3865:
3858:
3857:
3853:
3852:
3849:
3845:
3842:
3839:
3834:
3823:
3822:
3818:
3816:
3815:
3811:
3809:
3808:
3804:
3802:
3801:
3797:
3796:
3794:
3792:
3788:
3782:
3781:
3777:
3776:
3773:
3769:
3766:
3763:
3758:
3755:
3752:
3747:
3741:
3740:
3736:
3734:
3733:
3729:
3726:
3725:
3721:
3719:
3718:
3714:
3712:
3711:
3707:
3705:
3704:
3700:
3698:
3697:
3693:
3691:
3690:
3686:
3683:
3682:
3678:
3676:
3675:
3671:
3668:
3667:
3663:
3660:
3659:
3658:Chienkosaurus
3655:
3652:
3651:
3647:
3646:
3643:
3640:
3637:
3632:
3628:
3624:
3619:
3615:
3605:
3598:
3591:
3584:
3558:
3557:
3556:
3555:
3553:
3551:
3547:
3543:
3517:
3516:
3512:
3510:
3509:
3505:
3503:
3502:
3498:
3496:
3495:
3491:
3488:
3487:
3483:
3481:
3480:
3476:
3474:
3473:
3469:
3467:
3466:
3462:
3460:
3459:
3455:
3454:
3452:
3450:
3449:Furileusauria
3446:
3440:
3439:
3435:
3433:
3432:
3428:
3425:
3424:
3423:Rahiolisaurus
3420:
3418:
3417:
3413:
3411:
3410:
3406:
3404:
3403:
3399:
3397:
3396:
3392:
3390:
3389:
3385:
3383:
3382:
3378:
3375:
3374:
3370:
3369:
3366:
3363:
3361:
3357:
3350:
3349:
3345:
3344:
3342:
3340:
3339:Carnotaurinae
3336:
3330:
3329:
3325:
3322:
3321:
3320:Rahiolisaurus
3317:
3315:
3314:
3313:Majungasaurus
3310:
3308:
3307:
3303:
3301:
3300:
3296:
3293:
3292:
3288:
3286:
3285:
3281:
3280:
3278:
3276:
3272:
3266:
3265:
3261:
3259:
3258:
3254:
3252:
3251:
3250:Tarascosaurus
3247:
3245:
3244:
3240:
3238:
3237:
3233:
3231:
3230:
3226:
3224:
3223:
3219:
3217:
3216:
3212:
3210:
3209:
3208:Eoabelisaurus
3205:
3202:
3201:
3197:
3195:
3194:
3193:Chenanisaurus
3190:
3189:
3186:
3183:
3180:
3179:Abelisauridae
3175:
3161:
3160:
3156:
3154:
3153:
3149:
3147:
3146:
3142:
3140:
3139:
3138:Masiakasaurus
3135:
3133:
3132:
3128:
3127:
3125:
3123:
3119:
3113:
3112:
3108:
3106:
3105:
3104:Huinculsaurus
3101:
3099:
3098:
3097:Elaphrosaurus
3094:
3093:
3091:
3089:
3085:
3079:
3078:
3074:
3072:
3071:
3067:
3065:
3064:
3060:
3057:
3056:
3052:
3049:
3048:
3044:
3041:
3040:
3036:
3033:
3032:
3028:
3026:
3025:
3021:
3018:
3017:
3016:Austrocheirus
3013:
3010:
3009:
3005:
3004:
3001:
2998:
2996:
2992:
2985:
2984:
2980:
2977:
2976:
2972:
2969:
2968:
2964:
2961:
2960:
2956:
2955:
2952:
2948:
2945:
2942:
2937:
2927:
2926:
2922:
2920:
2919:
2915:
2914:
2912:
2910:
2906:
2900:
2899:
2895:
2892:
2891:
2887:
2885:
2884:
2880:
2877:
2876:
2872:
2870:
2869:
2868:Berberosaurus
2865:
2864:
2861:
2857:
2854:
2851:
2846:
2839:
2838:
2834:
2832:
2831:
2827:
2826:
2823:
2820:
2817:
2812:
2808:
2804:
2799:
2795:
2786:
2780:
2757:
2756:
2755:Halticosaurus
2752:
2750:
2749:
2748:Teinurosaurus
2745:
2744:
2742:
2738:
2729:
2728:
2727:
2726:
2724:
2722:
2718:
2708:
2707:
2703:
2700:
2699:
2695:
2693:
2692:
2688:
2685:
2684:
2680:
2677:
2676:
2675:Megapnosaurus
2672:
2670:
2669:
2665:
2663:
2662:
2658:
2656:
2655:
2651:
2650:
2648:
2646:
2645:Coelophysidae
2642:
2636:
2635:
2634:Powellvenator
2631:
2629:
2628:
2624:
2622:
2621:
2617:
2615:
2614:
2610:
2607:
2606:
2602:
2601:
2598:
2595:
2593:
2589:
2583:
2582:
2578:
2575:
2574:
2570:
2568:
2567:
2563:
2561:
2560:
2556:
2554:
2553:
2549:
2547:
2546:
2542:
2540:
2539:
2535:
2533:
2532:
2528:
2526:
2525:
2524:Liliensternus
2521:
2519:
2518:
2514:
2512:
2511:
2507:
2505:
2504:
2500:
2497:
2496:
2495:Dolichosuchus
2492:
2490:
2489:
2488:Dilophosaurus
2485:
2484:
2481:
2478:
2475:
2470:
2463:
2462:
2461:Herrerasauria
2458:
2455:
2454:
2450:
2447:
2446:
2442:
2439:
2438:
2434:
2432:
2431:
2427:
2424:
2423:
2419:
2416:
2415:
2414:Daemonosaurus
2411:
2408:
2407:
2403:
2402:
2399:
2396:
2393:
2388:
2384:
2380:
2375:
2371:
2350:
2349:
2348:
2347:
2345:
2343:
2339:
2331:
2327:
2326:
2325:
2324:
2321:
2318:
2316:
2312:
2308:
2300:
2296:
2293:
2291:
2287:
2284:
2282:
2278:
2276:
2272:
2270:
2266:
2265:
2260:
2256:
2252:
2245:
2240:
2238:
2233:
2231:
2226:
2225:
2222:
2215:
2204:
2196:
2192:
2185:
2182:
2177:
2173:
2169:
2165:
2158:
2151:
2148:
2144:
2141:
2137:
2131:
2128:
2124:
2118:
2115:
2112:
2108:
2105:(8): e44012.
2104:
2101:
2094:
2092:
2090:
2088:
2086:
2084:
2082:
2078:
2074:
2070:
2064:
2062:
2060:
2058:
2056:
2054:
2052:
2050:
2048:
2046:
2044:
2042:
2038:
2031:
2029:
2027:
2023:
2022:
2017:
2013:
2004:
2000:
1998:
1993:
1992:
1987:
1986:
1981:
1980:ornithischian
1977:
1975:
1969:
1967:
1963:
1960:
1956:
1955:
1950:
1949:
1944:
1943:
1938:
1934:
1926:
1925:
1920:
1916:
1912:
1910:
1906:
1905:
1900:
1895:
1894:
1888:
1883:
1882:dromaeosaurid
1879:
1865:
1851:
1839:
1834:
1832:
1830:
1826:
1825:
1820:
1819:Coelurosauria
1816:
1812:
1807:
1804:
1794:
1793:
1785:
1784:
1776:
1775:
1767:
1766:
1758:
1757:
1749:
1748:
1740:
1739:
1736:
1735:
1729:
1728:
1725:
1724:
1721:
1720:
1719:
1712:
1711:
1708:
1707:
1704:
1703:
1702:
1701:Stokesosaurus
1695:
1694:
1691:
1690:
1687:
1686:
1685:
1684:Tanycolagreus
1678:
1677:
1674:
1673:
1670:
1665:
1664:
1658:
1657:
1654:
1653:
1650:
1649:
1643:
1642:
1636:
1633:
1632:
1629:
1628:
1625:
1624:
1623:
1616:
1615:
1612:
1611:
1608:
1607:
1606:
1605:Huaxiagnathus
1599:
1598:
1595:
1594:
1591:
1590:
1589:
1582:
1581:
1578:
1577:
1574:
1573:
1567:
1566:
1563:
1562:
1559:
1558:
1557:
1550:
1549:
1543:
1542:Coelurosauria
1540:
1539:
1536:
1535:
1532:
1530:
1529:
1524:
1523:Allosauroidea
1518:
1517:
1511:
1508:
1507:
1504:
1503:
1495:
1494:
1486:
1485:
1477:
1476:
1468:
1467:
1459:
1458:
1455:
1454:
1453:Spinosaurinae
1448:
1447:
1444:
1443:
1440:
1439:
1438:Baryonychinae
1433:
1432:
1426:
1425:
1422:
1421:
1418:
1417:
1416:
1409:
1408:
1405:
1404:
1401:
1400:
1399:
1398:Wiehenvenator
1392:
1391:
1388:
1387:
1384:
1383:
1382:
1375:
1374:
1368:
1367:
1364:
1363:
1360:
1359:
1358:
1351:
1350:
1347:
1346:
1343:
1342:
1341:
1334:
1333:
1330:
1329:
1326:
1325:
1324:
1317:
1316:
1313:
1312:
1309:
1308:
1307:
1300:
1299:
1296:
1295:
1292:
1291:
1290:
1283:
1282:
1276:
1275:
1272:
1271:
1268:
1267:
1266:
1259:
1258:
1255:
1254:
1251:
1250:
1249:
1242:
1241:
1238:
1237:
1234:
1233:
1232:
1225:
1224:
1221:
1220:
1217:
1216:
1215:
1208:
1207:
1204:
1203:
1200:
1199:
1198:
1191:
1190:
1184:
1183:
1180:
1179:
1176:
1175:
1174:
1167:
1166:
1163:
1162:
1159:
1158:
1157:
1150:
1149:
1146:
1145:
1142:
1141:
1140:
1133:
1132:
1129:
1128:
1125:
1124:
1123:
1116:
1115:
1112:
1111:
1108:
1107:
1106:
1099:
1098:
1095:
1094:
1091:
1090:
1089:
1082:
1081:
1078:
1077:
1074:
1073:
1072:
1065:
1064:
1061:
1060:
1057:
1056:
1055:
1048:
1047:
1044:
1043:
1040:
1039:
1038:
1031:
1030:
1027:
1026:
1023:
1022:
1021:
1014:
1013:
1007:
1006:
1000:
997:
996:
993:
992:
984:
983:
980:
979:
978:
977:Siamotyrannus
971:
970:
967:
966:
963:
962:
961:
954:
953:
947:
946:
940:
937:
936:
932:
930:
926:
916:
915:
907:
906:
898:
897:
894:
893:
887:
886:
883:
882:
874:
873:
865:
864:
856:
855:
847:
846:
838:
837:
834:
833:
832:
825:
824:
821:
820:
817:
816:
815:
808:
807:
801:
800:
797:
796:
793:
792:
791:
790:Huaxiagnathus
784:
783:
777:
776:
773:
772:
769:
768:
767:
766:Compsognathus
760:
759:
753:
752:
749:
748:
745:
744:
743:
736:
735:
729:
728:
725:
724:
716:
715:
712:
711:
710:
703:
702:
699:
698:
695:
694:
693:
686:
685:
679:
678:
672:
669:
668:
662:
661:
658:
657:
654:
653:
647:
646:
640:
637:
636:
632:
630:
626:
622:
617:
615:
611:
605:
604:
599:
595:
588:
586:
584:
583:
578:
577:
571:
567:
566:
561:
557:
553:
548:
546:
542:
538:
533:
531:
527:
523:
519:
514:
512:
506:
504:
500:
496:
492:
488:
482:
478:
474:
472:
471:Huaxiagnathus
468:
464:
460:
453:
451:
449:
445:
441:
437:
433:
429:
425:
421:
417:
412:
410:
406:
405:specific name
402:
398:
394:
390:
386:
382:
378:
374:
370:
366:
363:
354:
347:
345:
343:
342:type specimen
339:
335:
334:
333:Huaxiagnathus
328:
326:
322:
319:
315:
312:
309:
308:compsognathid
305:
301:
300:
289:
283:
281:
272:
269:
268:Binomial name
265:
261:
260:
259:S. gigas
254:
251:
250:
245:
239:
238:
231:
228:
227:
224:
218:
215:
214:
211:
208:
205:
202:
201:
198:
195:
192:
189:
188:
185:
182:
179:
176:
175:
172:
169:
166:
165:
162:
159:
156:
155:
152:
149:
146:
145:
140:
135:
131:
127:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
6539:Yixian fauna
6437:
6401:
6330:
6323:
6316:
6309:
6302:
6297:Ornithomimus
6295:
6288:
6281:
6274:
6267:
6260:
6253:
6235:Tyrannomimus
6233:
6225:
6217:
6210:
6205:Deinocheirus
6203:
6196:
6176:
6169:
6162:
6138:
6130:
6125:Nqwebasaurus
6123:
6118:Nedcolbertia
6116:
6109:
6102:
6096:Calamosaurus
6094:
6088:Aviatyrannis
6086:
6079:
6054:
6024:
6017:
6011:
6010:
6002:
5995:
5988:
5981:
5973:
5968:Aristosuchus
5966:
5960:Aniksosaurus
5958:
5940:Aniksosaurus
5938:
5908:
5901:
5893:
5839:
5804:
5797:
5790:
5783:
5709:
5702:
5695:
5675:
5668:
5663:Dynamoterror
5661:
5641:
5634:
5614:
5607:
5587:
5582:Asiatyrannus
5580:
5556:
5549:
5520:Dryptosaurus
5518:
5511:
5504:
5473:
5466:
5459:
5451:
5444:
5437:
5430:
5423:
5416:
5411:Jinbeisaurus
5409:
5402:
5395:
5387:
5382:Aviatyrannis
5380:
5373:
5339:
5334:Sinotyrannus
5332:
5325:
5318:
5311:
5290:
5283:
5261:
5254:
5213:
5206:
5199:
5191:
5184:
5176:
5168:
5163:Bicentenaria
5161:
5153:
5148:Asiamericana
5146:
5139:
5080:
5021:
5014:
5007:
5000:
4993:
4986:
4979:
4958:
4950:
4943:
4936:
4929:
4923:Deltadromeus
4921:
4913:
4906:
4855:Tyrannotitan
4853:
4846:
4839:
4832:
4812:
4788:
4783:Taurovenator
4781:
4774:
4767:
4762:Lajasvenator
4760:
4753:
4746:
4741:Concavenator
4739:
4731:
4724:
4699:
4691:
4684:
4676:
4668:
4648:
4641:
4633:
4626:
4619:
4612:
4587:
4579:
4571:
4564:
4557:Allosauridae
4525:
4518:
4513:Shidaisaurus
4511:
4504:
4497:
4477:
4469:
4444:
4436:
4429:
4422:
4391:
4383:
4375:
4368:
4341:
4333:
4321:Avetheropoda
4308:Avetheropoda
4279:
4238:Avetheropoda
4205:
4198:
4191:Spinosaurini
4178:
4171:
4164:
4157:
4149:
4121:
4114:
4107:
4086:
4078:
4071:
4066:Protathlitis
4064:
4057:
4032:
4025:
4018:
4010:
3975:
3968:
3961:
3956:Leshansaurus
3954:
3947:
3940:
3920:
3913:
3908:Megalosaurus
3906:
3901:Duriavenator
3899:
3879:
3854:
3838:Megalosauria
3819:
3812:
3807:Marshosaurus
3805:
3800:Condorraptor
3798:
3778:
3737:
3730:
3722:
3715:
3708:
3701:
3694:
3687:
3681:Cruxicheiros
3679:
3672:
3664:
3656:
3648:
3513:
3508:Quilmesaurus
3506:
3499:
3492:
3484:
3477:
3470:
3463:
3456:
3436:
3429:
3421:
3414:
3407:
3400:
3393:
3386:
3379:
3371:
3360:Brachyrostra
3346:
3326:
3318:
3311:
3304:
3297:
3289:
3282:
3262:
3257:Tralkasaurus
3255:
3248:
3241:
3234:
3227:
3220:
3213:
3206:
3198:
3191:
3159:Vespersaurus
3157:
3152:Velocisaurus
3150:
3143:
3136:
3129:
3109:
3102:
3095:
3075:
3068:
3061:
3053:
3047:Deltadromeus
3045:
3037:
3031:Compsosuchus
3029:
3022:
3014:
3006:
2981:
2973:
2967:Lametasaurus
2965:
2957:
2923:
2918:Ceratosaurus
2916:
2896:
2888:
2881:
2873:
2866:
2850:Ceratosauria
2835:
2828:
2753:
2746:
2704:
2696:
2689:
2681:
2673:
2666:
2659:
2652:
2632:
2627:Podokesaurus
2625:
2618:
2611:
2603:
2579:
2571:
2564:
2557:
2550:
2543:
2536:
2529:
2522:
2515:
2510:Gojirasaurus
2508:
2503:Dracovenator
2501:
2493:
2486:
2474:Neotheropoda
2459:
2451:
2443:
2437:Guaibasaurus
2435:
2428:
2420:
2412:
2406:Chindesaurus
2404:
2294:
2285:
2184:
2167:
2163:
2150:
2142:
2139:
2135:
2130:
2122:
2117:
2102:
2099:
2072:
2068:
2025:
2019:
2015:
2011:
2009:
1996:
1989:
1983:
1972:
1970:
1962:ornithomimid
1952:
1946:
1942:Nqwebasaurus
1940:
1929:
1922:
1918:
1902:
1891:
1886:
1877:
1875:
1835:Paleobiology
1822:
1810:
1808:
1802:
1800:
1732:
1716:
1715:
1699:
1698:
1682:
1681:
1668:
1646:
1620:
1619:
1603:
1602:
1586:
1585:
1571:
1570:
1554:
1553:
1526:
1521:
1510:Avetheropoda
1451:
1436:
1413:
1412:
1396:
1395:
1379:
1378:
1357:Megalosaurus
1355:
1354:
1338:
1337:
1321:
1320:
1304:
1303:
1287:
1286:
1263:
1262:
1248:Duriavenator
1246:
1245:
1229:
1228:
1214:Nedcolbertia
1212:
1211:
1195:
1194:
1171:
1170:
1156:Condorraptor
1154:
1153:
1137:
1136:
1122:Leshansaurus
1120:
1119:
1105:Marshosaurus
1103:
1102:
1086:
1085:
1069:
1068:
1052:
1051:
1035:
1034:
1018:
1017:
975:
974:
958:
957:
928:
927:(2024) with
924:
922:
890:
829:
828:
812:
811:
788:
787:
764:
763:
741:
740:
739:
707:
706:
690:
689:
650:
624:
618:
609:
608:
601:
597:
580:
574:
569:
563:
555:
549:
534:
515:
507:
486:
485:
470:
466:
458:
457:
413:
408:
400:
396:
392:
384:
381:Yuan Chongxi
364:
362:type species
359:
337:
331:
329:
298:
297:
296:
287:
279:
274:
258:
257:
243:
236:
235:
203:
190:
177:
24:
18:
6362:Maniraptora
6351:Maniraptora
6332:Tototlmimus
6262:Anserimimus
6212:Garudimimus
6198:Beishanlong
6140:Valdoraptor
6081:Arkansaurus
6026:Xunmenglong
5903:Juravenator
5697:Tarbosaurus
5589:Nanuqsaurus
5558:Gorgosaurus
5475:Megaraptora
5461:Timurlengia
5193:Sciurumimus
5141:Aratasaurus
5009:Murusraptor
4931:Fukuiraptor
4894:Megaraptora
4776:Shaochilong
4748:Eocarcharia
4701:Megaraptora
4621:Lusovenator
4499:Alpkarakush
4356:Carnosauria
4207:Spinosaurus
4180:Siamosaurus
4080:Suchosaurus
4027:Iberospinus
3963:Magnosaurus
3942:Afrovenator
3915:Torvosaurus
3666:Chilesaurus
3479:Carnotaurus
3472:Caletodraco
3458:Abelisaurus
3284:Arcovenator
3131:Laevisuchus
3122:Noasaurinae
3024:Berthasaura
2995:Noasauridae
2925:Genyodectes
2890:Lukousaurus
2883:Jubbulpuria
2683:Panguraptor
2661:Coelophysis
2654:Camposaurus
2613:Panguraptor
2605:Dracoraptor
2581:Zupaysaurus
2566:Tachiraptor
2545:Sarcosaurus
2445:Nhandumirim
2170:(1): 1-19.
2021:Microraptor
1954:Caudipteryx
1933:gastroliths
1921:feeding on
1528:Juravenator
1415:Iberospinus
1265:Afrovenator
1231:Magnosaurus
1197:Sciurumimus
831:Juravenator
600:feeding on
576:Microraptor
454:Description
430:, from the
377:Lü Junchang
36:124.6
6523:Categories
6290:Gallimimus
6219:Harpymimus
6164:Harpymimus
5785:Iliosuchus
5755:see below↓
5600:Alioramini
5418:Juratyrant
5404:Eotyrannus
5389:Bagaraatan
5341:Yutyrannus
5275:Coeluridae
5201:Vayuraptor
5047:see below↓
5023:Tratayenia
5016:Orkoraptor
5002:Megaraptor
4908:Aoniraptor
4841:Mapusaurus
4769:Sauroniops
4733:Altispinax
4686:Neovenator
4628:Siamraptor
4614:Datanglong
4581:Epanterias
4573:Antrodemus
4566:Allosaurus
4547:Allosauria
4370:Altispinax
4335:Gasosaurus
4247:see below↓
4123:Suchomimus
3724:Sinosaurus
3559:see below↓
3515:Viavenator
3465:Aucasaurus
3402:Ilokelesia
3373:Dahalokely
3328:Rajasaurus
3306:Indosaurus
3299:Genusaurus
3291:Dahalokely
3215:Indosuchus
3111:Limusaurus
3070:Ligabueino
3063:Kiyacursor
3055:Genusaurus
3039:Dahalokely
2959:Betasuchus
2830:Dornraptor
2730:see below↓
2706:Segisaurus
2559:Sinosaurus
2422:Eodromaeus
2351:see below↓
2299:Dinosauria
2281:Sauropsida
2145:(I): 1-281
2032:References
1985:Yueosaurus
1982:, perhaps
1718:Juratyrant
1669:NGMC 2124
960:Siamraptor
709:Orkoraptor
560:integument
545:metatarsus
530:metacarpus
526:metacarpal
491:premaxilla
197:Saurischia
184:Dinosauria
6366:includes
6311:Rativates
6304:Qiupalong
6004:Scipionyx
5997:Mirischia
5670:Lythronax
5609:Alioramus
5263:Labocania
4981:Aerosteon
4527:Sinraptor
4431:Erectopus
4166:Irritator
3751:Orionides
3636:Tetanurae
3623:Tetanurae
3550:Tetanurae
3486:Elemgasem
3409:Llukalkan
3388:Elemgasem
3145:Noasaurus
3008:Afromimus
2816:Averostra
2803:Averostra
2721:Averostra
2573:Velocipes
2392:Theropoda
2379:Theropoda
2342:Theropoda
2267:Kingdom:
2251:Theropoda
2075:(1): 8-15
1959:Mongolian
1904:Scipionyx
1588:Mirischia
1381:Scipionyx
999:Orionides
939:Tetanurae
931:in bold.
692:Scipionyx
629:dal Sasso
621:cladogram
582:Pedopenna
552:theropods
511:gastralia
448:allometry
440:Barremian
420:Hengdaozi
369:Ji Shu'an
348:Discovery
316:from the
314:dinosaurs
252:Species:
210:Theropoda
157:Kingdom:
151:Eukaryota
6447:Wikidata
5895:Coelurus
5320:Kileskus
5313:Guanlong
5285:Coelurus
5178:Gualicho
4678:Gualicho
4059:Baryonyx
3395:Guemesia
3222:Kryptops
2983:Ozraptor
2620:Pendraig
2275:Chordata
2273:Phylum:
2269:Animalia
2100:PLoS ONE
1948:Baryonyx
1909:duodenum
499:holotype
416:holotype
373:Ji Qiang
311:theropod
216:Family:
171:Chordata
167:Phylum:
161:Animalia
147:Domain:
6493:4126514
6480:4966955
6467:4433891
6453:Q133117
5453:Timimus
5215:Zuolong
4945:Rapator
4848:Meraxes
4173:Oxalaia
3494:Koleken
2517:Lepidus
2279:Class:
1976:sanctus
1966:gizzard
1937:gizzard
1899:stomach
1622:Zuolong
1525:(Incl.
541:ischium
518:humerus
497:of the
495:maxilla
434:of the
424:Sihetun
277:†
256:†
229:Genus:
105:↓
6104:Hexing
5792:Kakuru
5397:Dilong
3438:Thanos
3416:Niebla
3236:Rugops
3229:Kurupi
2143:XXXVII
1957:and a
444:Aptian
397:pteryx
290:, 2007
288:et al.
246:, 2007
244:et al.
6506:98810
6368:birds
5425:Moros
4952:Siats
4693:Siats
4635:Siats
2295:Clade
2286:Clade
2160:(PDF)
1988:or a
619:This
537:ilium
503:jugal
426:, in
422:, in
409:gigas
393:kalos
389:Latin
385:Sinae
325:China
304:genus
204:Clade
191:Clade
178:Clade
6475:GBIF
6360:see
4995:Maip
2453:Tawa
2328:see
1945:and
579:and
522:ulna
414:The
403:its
379:and
360:The
44:PreꞒ
6462:EoL
2191:doi
2172:doi
2107:doi
925:Cau
323:of
286:Ji
242:Ji
6525::
6503::
6490::
6477::
6464::
6449::
2297::
2288::
2168:63
2166:.
2162:.
2080:^
2073:81
2071:,
2040:^
1831:.
616:.
547:.
407:,
387:,
375:,
371:,
336:,
206::
193::
180::
94:Pg
38:Ma
33:,
6230:?
6222:?
6135:?
6099:?
6091:?
6059:?
6007:?
5978:?
5963:?
5898:?
5478:?
5456:?
5392:?
5295:?
5277:?
5266:?
5196:?
5181:?
5173:?
5158:?
4962:?
4954:?
4926:?
4918:?
4896:?
4736:?
4704:?
4696:?
4681:?
4673:?
4638:?
4592:?
4584:?
4576:?
4474:?
4449:?
4441:?
4396:?
4388:?
4380:?
4346:?
4338:?
4154:?
4091:?
4083:?
4037:?
4015:?
3980:?
3859:?
3824:?
3727:?
3684:?
3669:?
3661:?
3653:?
3489:?
3426:?
3376:?
3351:?
3323:?
3294:?
3203:?
3058:?
3050:?
3042:?
3034:?
3019:?
3011:?
2986:?
2978:?
2970:?
2962:?
2893:?
2878:?
2840:?
2701:?
2686:?
2678:?
2608:?
2576:?
2498:?
2464:?
2456:?
2448:?
2440:?
2425:?
2417:?
2409:?
2243:e
2236:t
2229:v
2197:.
2193::
2178:.
2174::
2109::
2103:7
1935:(
442:-
233:†
220:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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