283:
82:
31:
350:
enlarged vocal sacs and large tracheal cartilages which act as resonators to amplify the sounds. The diet of these monkeys consists mostly of leaves, supplemented by flowers and fruit, and these foods being plentiful in the rainforest, the territories defended by the troops are small. Rival troops howl at each other but territorial disputes are mostly settled vocally, without the monkeys needing to meet face to face.
57:
409:
in Brazil. The animal has a wide range and is present in a number of protected areas, but the rainforest in which it lives is being cleared for cattle ranching and agriculture, resulting in fragmentation of the forest. The subpopulations in these fragments are likely to suffer from inbreeding. They
349:
Howlers move around in small troops, consisting of a dominant male, several females and a number of juveniles. The monkeys are so called because of the loud calls made by males, more roar than howl, which can be heard a mile (1.5 km) or more away, mostly at dawn and dusk. The throats of males have
848:
394:
767:
853:
843:
838:
808:
405:; during the period 2007 to 2009, an outbreak of yellow fever killed large numbers of howlers in the Misiones Province in Argentina, and the State of
702:
728:
863:
828:
583:
833:
868:
522:
290:
341:. Habitats include lowland, sub-montane and montane forests, and it is tolerant of disturbed habitats and human encroachment.
759:
733:
680:
818:
226:
823:
754:
547:
858:
772:
81:
297:
The southern brown howler is found in the
Atlantic forests of eastern Brazil where it occurs in the states of
813:
642:
318:
302:
210:
282:
310:
338:
46:
504:
398:
76:
61:
689:
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Miranda, João M. D.; Bernardi, Itiberê P.; Abreu, Kauê C. & Passos, Fernando C. (2005).
494:
326:
746:
780:
30:
483:
in the South of Porto Alegre and its
Influence on Land use Policy between 1997 and 2007"
802:
741:
450:
437:
173:
66:
625:
600:
508:
694:
436:
Mendes, S. L.; Rylands, A. B.; Kierulff, M. C. M. & de
Oliveira, M. M. (2020).
402:
382:
306:
270:). Gregorin, 2006, considered the southern brown howler to be a separate species,
255:
247:
187:
715:
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665:
334:
143:
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93:
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333:
in
Argentina. Its northern extent is unclear, but may be the
353:
In the
Araucaria pine forests of southern Brazil, the chief
410:
are also impacted by urbanisation and are hunted for food.
401:. This is partly because of the monkey's susceptibility to
397:
has assessed this monkey's conservation status as being
649:
477:Printes, Rodrigo C.; et al. (December 2010).
578:. Princeton University Press. pp. 326–327.
293:, which provides a habitat for this subspecies.
395:International Union for Conservation of Nature
274:, but this has not been universally accepted.
325:. Its range extends southwards as far as the
8:
637:
55:
29:
20:
624:
498:
479:"The Urban Monkeys Program: A Survey of
419:
431:
429:
427:
425:
423:
849:Environment of Santa Catarina (state)
7:
451:IUCN Red List of Threatened Species
554:. American Society of Mammalogists
14:
854:Environment of São Paulo (state)
844:Environment of Rio Grande do Sul
809:IUCN Red List vulnerable species
609:(Linnaeus) (Carnivora, Felidae)"
605:Cabrera (Primates, Atelidae) by
80:
626:10.1590/S0101-81752005000300043
291:Aparados da Serra National Park
613:Revista Brasileira de Zoologia
16:Subspecies of New World monkey
1:
839:Environment of Paraná (state)
575:The New Neotropical Companion
829:Fauna of the Atlantic Forest
246:) is a monkey subspecies of
864:Taxa named by Ángel Cabrera
834:Environment of Minas Gerais
885:
681:Alouatta guariba clamitans
651:Alouatta guariba clamitans
603:Alouatta guariba clamitans
525:Alouatta guariba clamitans
243:Alouatta guariba clamitans
220:Alouatta guariba clamitans
39:Alouatta guariba clamitans
869:Mammals described in 1940
381:), as well as the larger
216:
209:
77:Scientific classification
75:
53:
44:
37:
28:
23:
278:Distribution and habitat
262:) and far northeastern
250:native to southeastern
572:Kricher, John (2017).
294:
24:Southern brown howler
357:of these monkeys are
285:
238:southern brown howler
819:Mammals of Argentina
500:10.1896/052.025.0103
487:Primate Conservation
202:A. g. clamitans
458:: e.T39918A17979180
339:Jequitinhonha River
47:Conservation status
824:Primates of Brazil
607:Leopardus pardalis
481:Alouatta clamitans
363:Leopardus pardalis
295:
272:Alouatta clamitans
859:Misiones Province
796:
795:
643:Taxon identifiers
585:978-0-691-11525-2
440:Alouatta guariba
407:Rio Grande do Sul
331:Misiones Province
323:Rio Grande do Sul
260:Rio Grande do Sul
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329:and eastward to
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781:Observation.org
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188:A. guariba
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814:Howler monkeys
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619:(3): 793–795.
601:"Predation on
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319:Santa Catarina
303:Rio de Janeiro
299:Espírito Santo
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211:Trinomial name
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493:(25): 11–19.
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383:birds of prey
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379:Panthera onca
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372:
371:Puma concolor
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344:
342:
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327:Camaquã River
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567:
556:. Retrieved
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542:
531:. Retrieved
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486:
480:
472:
460:. Retrieved
455:
449:
443:
439:
403:yellow fever
392:
378:
370:
362:
352:
348:
307:Minas Gerais
296:
271:
256:Minas Gerais
248:brown howler
242:
241:
237:
235:
219:
217:
201:
200:
196:Subspecies:
186:
174:
150:Infraorder:
38:
18:
716:iNaturalist
675:Wikispecies
154:Simiiformes
803:Categories
558:2020-04-18
533:2020-04-18
414:References
399:vulnerable
335:Doce River
289:jungle in
144:Haplorhini
140:Suborder:
62:Vulnerable
444:clamitans
355:predators
311:São Paulo
287:Araucaria
264:Argentina
182:Species:
100:Kingdom:
94:Eukaryota
760:12100382
666:Q2583127
660:Wikidata
550:Alouatta
509:86045499
268:Misiones
175:Alouatta
164:Atelidae
160:Family:
134:Primates
124:Mammalia
114:Chordata
110:Phylum:
104:Animalia
90:Domain:
67:IUCN 3.1
708:4267229
462:3 March
375:jaguars
373:), and
367:cougars
359:ocelots
345:Ecology
337:or the
227:Cabrera
170:Genus:
130:Order:
120:Class:
65: (
786:206616
773:183328
734:945165
721:147529
582:
529:. ITIS
507:
389:Status
315:Paraná
252:Brazil
229:, 1940
747:39918
695:5FJ8D
505:S2CID
768:NCBI
742:IUCN
729:ITIS
703:GBIF
580:ISBN
491:2010
464:2021
456:2020
442:ssp.
393:The
321:and
236:The
755:MSW
690:CoL
621:doi
495:doi
365:),
258:to
805::
783::
770::
757::
744::
731::
718::
705::
692::
677::
662::
617:22
615:.
611:.
503:.
489:.
485:.
454:.
448:.
422:^
385:.
317:,
313:,
309:,
305:,
301:,
629:.
623::
588:.
561:.
552:"
548:"
536:.
527:"
523:"
511:.
497::
466:.
446:"
438:"
377:(
369:(
361:(
266:(
254:(
240:(
69:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
