Spotted lanternfly

713: 1718: 973: 1746: 1115: 1734: 432: 1762: 1216: 588: 31: 1694: 890: 1706: 56: 1465:), saving only male trees to use as "trap" trees, since the spotted lanternfly is attracted to its preferred hosts. The remaining male "trap trees" should be wrapped with sticky bands starting in early spring to catch any nymphs. Wildlife experts have warned to cover the sticky bands on trees with chicken wire or another similar wire after many reports of other animals (e.g. birds) becoming trapped on them, resulting in injury and/or death. 1778: 704:

extent adults, multiple opportunities to repeatedly attempt righting themselves following a jump. Additionally, the spotted lanternfly will also perform aerial reorientation and terrestrial righting as means to right itself, collectively giving the spotted lanternfly the ability to land on a variety of surfaces and spread rapidly through an environment.

969:, requiring two weeks of warm temperatures following winter before hatching is induced. Eggs overwintered for five months or longer demonstrate higher hatch rates and more synchronous hatching, suggesting that cold temperatures increase overall egg survival rates. The majority of adults die off by the end of December. 2946:

Murman, Kelly; Setliff, Gregory P; Pugh, Cathryn V; Toolan, Michael J; Canlas, Isaiah; Cannon, Stefani; Abreu, Leslie; Fetchen, Miranda; Zhang, Longwa; Warden, Melissa L; Wallace, Matthew; Wickham, Jacob; Spichiger, Sven-Erik; Swackhamer, Emelie; Carrillo, Daniel (14 December 2020). Morrison, William

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from seven municipalities on 1 November 2014. Considering the old egg masses that were found, it was estimated that the spotted lanternfly may have been in the United States since at least 2012 and had survived the unusually cold 2013–14 winter. By March 2021, lanternflies had been found in 34 of the

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Nymphs typically hatch from their egg cases starting in late April or early May, marking the beginning of the lanternfly's developmental stages. A nymph passes through several immature stages, all of which are wingless. In the first instar, it is black with white spots. Later instars have red patches

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affects the eggs of this species. The highest temperature that will still kill eggs was estimated by South Korean researchers to be between −12.7 and −3.4 °C (9.1 and 25.9 °F) on the basis of mean daily temperatures during their winter of 2009–2010. This estimate contrasts with eggs having

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can lower plants' photosynthetic potential and affect their health; this reduction is even more pronounced by the possible growth of molds over the sugary compound, which further limits light available to affected plants. The accumulation of sap and honeydew also attracts many species of ants, bees,

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The lanternfly is a planthopper, and uses its wings to assist these jumps rather than making sustained flights. The spotted lanternfly will perform a number of "successive collisions" upon jumping, employing both passive and active righting as it falls. This bouncing provides nymphs, and to a lesser

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of host plants, including the tree of heaven, an invasive tree. The damage to native hardwood forests which was feared at the outset did not come to pass. Unlike some insects, the spotted lanternfly does not pose direct danger to humans through biting or stinging. Spotted lanternflies lay egg masses

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to the adult form, with a black head and grayish wings with black spots. Nymphs hop or crawl to search for plants on which to feed. Young nymphs (first through third instars) appear to have a wider host range early on, which narrows as they grow older. Though lanternflies have been recorded feeding

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may have preferences for feeding on certain plants. Two examples of possible factors being investigated are the contribution of the overall sugar composition in the plant and the presence of toxic chemicals. The lanternfly lays eggs upon any smooth-trunked tree, stone, or vertical smooth surface,

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for feeding and laying eggs, but if it is not present, the insect can lay its eggs on any stationary object, natural or man-made, and feeds on a wide variety of plants. Eggs can easily be moved from place to place without being noticed, giving them an easy way to spread to new areas. Its initial

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eggs between the months of October and May by scraping them off surfaces, "double bag them and throw them in the garbage." People can scrape the eggs directly into plastic bags containing alcohol and/or hand sanitizer to kill them. The PDA has recommended removal of preferred spotted lanternfly

1195:. Researchers have estimated that the spotted lanternfly may have been entering the country sporadically since the 1930s, but only established a reproducing population around the beginning of the twenty-first century. Specimens found in Japan are genetically identical to populations in Beijing, 871:

also indirectly affects the health and productivity of their hosts and nearby plants through the production of large amounts of honeydew, the lanternfly's sugary secretions of excess waste and sap, as well as by leaving feeding scars in the host plant's branches that continue to drip sap. The

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Modeling for the life cycle of the spotted lanternfly in Australia has found that development and survival may vary throughout the country, should the spotted lanternfly be introduced. Australia's southwest coastal regions are estimated to have a high viability for the spotted lanternfly but

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has identified a risk of the SLF entering the country and has previously intercepted adult lanternflies traveling to Ontario in shipping crates. The primary concern is the potential damage the lanternflies could cause to the wine, fruit, and vegetable industries. The Canadian government and

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has been effective at capturing nymphs, though adults may be strong enough to escape the adhesive. The spotted lanternfly is known to avoid changes in tree texture and obstacles along the tree trunk, and inward facing traps have shown the greatest efficacy with minimized ecological impact.

3952: 4298: 1357:, although no live sightings have been reported from these states as of September 2023. The discoveries of dead specimens have increased concerns for possible accidental introduction of the insect to yet more states where they can potentially become established. 1682:, due to its high rates of parasitism of eggs. This wasp is under investigation in South Korea and in the United States, where it is being evaluated under quarantine until researchers are certain it will not become an invasive species and attack other insects. 1245:. Based on its host affinities, it presents a threat to the state's grape, fruit tree, and logging industries. The greatest risk of spread was seen in transportation of materials containing egg masses laid on smooth bark, stone, and other vertical surfaces. 4904: 2039: 4368: 1717: 2611: 354:

and wings as they mature. Early life instars have a large host range that narrows with maturation. Adult spotted lanternflies have a black head, grey wings, and red hind wings. Adults do not have any specialized feeding associations with

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has been documented as a pest to grapevines, leading to a decline in the total number and overall quality of the harvested grapes. No quantitative data has been published supporting this correlation, but it has been widely hypothesized.

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In 2006, the spotted lanternfly was introduced in Korea, and has been considered a pest since about 2007. It has since expanded its host range, attacking at least 65 plant species, uninhibited by natural predators. The distribution of

1264:, annually. Models of the spotted lanternfly's spread projected an annual loss of $ 554 million, with an additional loss of 4,987 jobs, should it continue to spread to the entirety of Pennsylvania. A national working group led by the 864:

feed on sap from the trunk or branches of their host plants; because they can appear in such large numbers on a single plant, they can directly cause substantial damage to, and effectively kill parts or the whole of the host.

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suggested that −25 °C (−13 °F) is about the temperature at which no eggs are hatched, while −15 °C (5 °F) still had limited hatching, depending upon how long they were chilled and where they were kept.

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pathogen between different individuals of the insects' preferred host. Researchers have not yet found this to be an effective method, and more research is needed to confirm if this is possible in field settings with

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on several herbaceous plants, this is most likely due to early-instar nymphs climbing or falling onto these plants; late-instar nymphs and adult lanternflies have no reliable association with herbaceous plants.

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agricultural industries are concerned about maintaining quarantine and have expressed hope that quarantine efforts in the United States are successful. Although most of Canada is an unsuitable habitat for

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Kim, Hyojoong; Kim, Minyoung; Kwon, Deok Ho; Park, Sangwook; Lee, Yerim; Huang, Junhao; Kai, Shi; Lee, Heung-Sik; Hong, Ki-Jeong; Jang, Yikweon; Lee, Seunghwan (December 2013). "Molecular comparison of

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When resting, the crimson hind wings are partially visible through the semi-translucent forewings, giving the lanternfly a red cast. Neatly spaced black rectangular markings color the tips of the

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In July 2022, live spotted lanternfly was found in Oakland County. Treatment of the infested area was completed and MDARD is working to contain and limit the spread of this small population.

1331:,. In late September 2023, the State of Tennessee confirmed the detection of a spotted lanternfly (SLF) in Davidson County, making Tennessee the 16th state to detect the spotted lanternfly. 4573: 2514: 4124: 4502: 4259: 4098: 3889: 2871: 1390: 1745: 656:(mated), the females' abdomens swell to the point where they have difficulty moving. Adult females, when measured from head to wing tip, have a body length of 20 to 27 millimetres ( 3912: 1405:

populations. Risk analysis of the spotted lanternfly to become established in Europe has been classified as a moderate to high risk, predominately due to the pervasiveness of

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Cooperband, Miriam F.; Wickham, Jacob; Cleary, Kaitlin; Spichiger, Sven-Erik; Zhang, Longwa; Baker, John; Canlas, Isaiah; Derstine, Nathan; Carrillo, Daniel (21 March 2019).

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scientists, was organized "to determine what is known about the lanternfly and what research is needed, including DNA analysis to pinpoint where the infestation originated."

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The Pennsylvania Department of Agriculture banned the transport of items that could harbor the egg masses, including firewood, lawn mowers, outdoor chairs, trucks, and

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Projected map by the US Department of Agriculture and Xinjiang Institute of Ecology and Geography on potentially suitable habitats for the spotted lanternfly globally.

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may thus be hinted at by unusual amounts of molds or stinging insects around specific plants. Bees often use spotted lanternfly honeydew to produce late-season honey.

2916: 1576:. It is thought that lanternflies, which feed on this host plant by piercing and sucking the sap from its vasculature, may be able to "aid" in the ongoing removal of 4072: 4471: 3440:

Lee, Jung-Su; Kim, Il-Kwon; Koh, Sang-Hyun; Cho, Sung Jong; Jang, Suk-Jun; Pyo, Seung-Hyeon; Choi, Won IL (January 2011). "Impact of minimum winter temperature on

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A large potential range exists for the spotted lanternfly to become established in almost all of the eastern part of the country, as well as critical wine- and

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is also referred to as the spot clothing wax cicada ("chu-ki" or "banyi-la-chan" in Chinese) and the Chinese blistering cicada ("ggot-mae-mi" 꽃매미 in Korean).

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Stål C. Beitrag zur Kenntnis der Fulgoriden. Entomologische Zeitung. Herausgegeben von dem entomologischen Vereine zu Stettin. Stettin, 24: 230-251. (1863).

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Liu, Houping; Lusk, Robert; Gallardy, Ross (1 March 2021). "Infrared thermography for insect detection: lighting up the spotted lanternfly in the field".

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Du, Zhenyong; Wu, Yunfei; Chen, Zhuo; Cao, Liangming; Ishikawa, Tadashi; Kamitani, Satoshi; Sota, Teiji; Song, Fan; Tian, Li; Cai, Wanzhi; Li, Hu (2021).

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introduction to the United States is believed to have been through this pathway, hitchhiking on an object imported from Asia into Pennsylvania. Adults of

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Francese, Joseph A; Cooperband, Miriam F; Murman, Kelly M; Cannon, Stefani L; Booth, Everett G; Devine, Sarah M; Wallace, Matthew S (14 April 2020).

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Barringer, Lawrence; Ciafré, Claire M. (2020). "Worldwide Feeding Host Plants of Spotted Lanternfly, With Significant Additions From North America".

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jump around the area to find new host plants. As nymphs, they feed on whatever host plant the egg was laid on before moving to another in the area.

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has also been reported in Taiwan, Vietnam, and India, but ongoing research has yet to conclude whether the species is native to these regions.

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and plays an important role in the lanternfly life cycle. This tree is the preferred host at all documented locations where the lanternfly and

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Essler, Jennifer L.; Kane, Sarah A.; Collins, Amanda; Ryder, Kaley; DeAngelo, Annemarie; Kaynaroglu, Patricia; Otto, Cynthia M. (3 May 2021).

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Dead lanternflies were found in Michigan as early as 2018 but its first live infestation was confirmed in August 2022, as was Rhode Island's.

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Other states began seeing spotted lanternflies as soon as 2018, and by 2021, they were also confirmed to be established in at least parts of

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Uyi, Osariyekemwen; Keller, Joseph A; Johnson, Anne; Long, David; Walsh, Brian; Hoover, Kelli (17 October 2020). Ranger, Christopher (ed.).

972: 3174: 2436:"Ecology and genetic structure of the invasive spotted lanternfly Lycorma delicatula in Japan where its distribution is slowly expanding" 2757:

Barringer, Lawrence E.; Donovall, Leo R.; Spichiger, Sven-Erik; Lynch, Daniel; Henry, David (June 2015). "The First New World Record of

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Ciolli, Marco; Anfora, Gianfranco; Hoelmer, Kim A.; Malek, Robert; Kaser, Joe M.; Broadley, Hannah J.; Gould, Juli (11 October 2019).

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in a pattern sometimes likened to brick and mortar. In flight, the spotted lanternfly displays red hind wings with black spots on the

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Choi, Man-Young; Yang, Zhong-Qi; Wang, Xiao-Yi; Tang, Yan-Long; Hou, Zhen-Rong; Kim, Jeong Hwan; Byeon, Young Woong (1 June 2014).

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Brooks, Rachel K., Ashley Toland, Andrew C. Dechaine, Thomas McAvoy, and Scott Salom. 2020. "The Inability of Spotted Lanternfly (

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from June to August. The PDA recommends tree injection and bark sprays, applied by professional applicators, and soil drench and

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has been theorized to be capable of maintaining a small population in the region although no specimens have yet been discovered.

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Lee, Jeong-Eun; Moon, Sang-Rae; Ahn, Hee-Geun; Cho, Sun-Ran; Yang, Jeong-Oh; Yoon, Chang-Mann; Kim, Gil-Hah (30 December 2009).

6013: 4525: 4178: 3475: 3075: 2106:"Descriptions of a new genus and some new species of homopterous insects from the East in the collection of the British Museum" 1365: 1339: 2162: 2127: 1673:

and was first documented to parasitize spotted lanternfly egg masses in 2017. Another biocontrol predator being tested is the

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Choi, Duck-Soo; Kim, Do-Ik; Ko, Suk-Ju; Kang, Beom-Ryong; Park, Jong-Dae; Kim, Seon-Gon; Choi, Kyeong-Ju (1 December 2012).

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estimated statewide costs of the spotted lanternfly to be $ 99.1 million in agricultural losses, and $ 236.3 million to the

2016: 4064: 3053: 1768: 1451: 1269: 1257: 229: 4455: 3301:"Spotted Lanternfly (Hemiptera: Fulgoridae) can complete development and reproduce without access to the preferred host, 910:

As early as July, adults can be seen, and they mate and lay eggs from late September through the onset of winter. During

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Xin, Bei; Wang, Xiao-yi; Cao, Liang-ming; Zhang, Yan-long; Gould, Juli; Broadley, Hannah; Hoelmer, Kim (February 2021).

2378: 1777: 1234: 1160: 1095: 649: 5031:"Discovery of three kairomones in relation to trap and lure development for spotted lanternfly (Hemiptera: Fulgoridae)" 3274: 5936: 5835: 3136: 2924: 802: 5262: 1401:

has found that the United Kingdom, France, Belgium, Switzerland, Spain, and Italy may be capable of supporting small

5668:"Biology and Management of the Spotted Lanternfly, Lycorma delicatula (Hemiptera: Fulgoridae), in the United States" 2708:"Many ways to land upright: novel righting strategies allow spotted lanternfly nymphs to land on diverse substrates" 2706:

Kane, Suzanne Amador; Bien, Theodore; Contreras-Orendain, Luis; Ochs, Michael F.; Tonia Hsieh, S. (11 August 2021).

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As of 2013, researchers are investigating another method of controlling lanternfly populations through reduction of

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A few natural predators have been identified in the lanternfly's native habitat in China, but are not yet used in

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and overall damage to grapevines. The tree is commonly found growing on the peripheries of Korean vineyards and

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Leach, Heather; Biddinger, David; Krawczyk, Greg; Swackhamer, Emelie; Korman, Amy; Walsh, Brian (8 May 2020).

3642:"Global phylogeography and invasion history of the spotted lanternfly revealed by mitochondrial phylogenomics" 2219: 1319:, with several of these states issuing quarantine orders. In 2022, the species was confirmed to be present in 431: 2676: 856:. The sugary waste fluid they produce can coat leaves and stems, which encourages mold growth and can impede 391:

was first recorded in the United States, and as of 2022, it is considered an invasive species in much of the

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Park, Ji-Doo; Kim, Min-Young; Lee, Sang-Gil; Shin, Sang-Chul; Kim, Jun-Heon; Park, Il-Kwon (30 March 2009).

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Pest control measures and guidelines have been issued by the Pennsylvania Department of Agriculture and its

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Mapping The Life-History, Development, And Survival of Spotted Lantern Fly In Occupied And Uninvaded Ranges

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efforts to cull the spotted lanternfly's population. However, this process indirectly harms other species.

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including man-made items such as vehicles, yard furniture, farm equipment, or other items stored outside.

808: 474: 187: 5840: 4873: 4147: 1923:"Invasive spotted lanternfly may not damage hardwood trees as previously thought | Penn State University" 4033: 1573: 1397:

as an A1 pest and anticipates it becoming invasive in Europe. CLIMEX modeling of potential habitats for

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in addition to the white spots. The final nymphal instar has red wing pads and a red upper body, before

800:. The lanternfly has also been recorded causing serious damage on at least 12 ornamental plants such as 735:

is a tree native to China but is invasive to many other areas worldwide. It is considered to be the key

494: 482: 331: 170: 5763: 4293: 4004:"State Agricultural Officials Urge Residents to Report Signs of Invasive Spotted Lanternfly | Mass.gov" 747:

co-occur. The spotted lanternfly has a host range of over 173 plant species worldwide, including grape

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as having a minimum of 20 black spots, with six spots located on the anterior margin of the forewings.

4897:"Wildlife experts warn spotted lanternfly traps are killing birds, bats, squirrels, and other animals" 1559:, indicating a means of early detection. Trained dogs are capable of detecting winterized egg masses. 957:

The egg masses contain 30–50 eggs covered in a yellowish-brown, waxy deposit, often referred to as an

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Rebbeck, J.; Malone, M. A.; Short, D. P. G.; Kasson, M. T.; O'Neal, E. S.; Davis, D. D. (July 2013).

5138: 3653: 3584: 3508: 3216: 2949:"Distribution, Survival, and Development of Spotted Lanternfly on Host Plants Found in North America" 2558: 2449: 1678: 1427: 1374: 1249: 653: 368: 276: 1948:"Spotted lanternfly lore: Penn State experts clear up falsehoods about pest | Penn State University" 1898:"Spotted lanternfly lore: Penn State experts clear up falsehoods about pest | Penn State University" 1536:

released by their host plants, and adults and second- to fourth-instar nymphs are also attracted to

6008: 5526:"Exploratory Survey of Spotted Lanternfly (Hemiptera: Fulgoridae) and Its Natural Enemies in China" 4815:

Maino, James Louis; Schouten, Rafael; Lye, Jessica C.; Umina, Paul A.; Reynolds, Olivia L. (2021),

4210: 1461: 1188: 915: 731: 470: 293: 5954: 5454: 4754:"Model-based prediction of potential distribution of the invasive insect pest, spotted lanternfly 3757:"Model-based prediction of potential distribution of the invasive insect pest, spotted lanternfly 1138:

is capable of surviving and completing its life cycle in environments without its preferred host,

5525: 5334: 5103: 4830: 3846: 3570:"The oldest known Lophopidae planthopper (Hemiptera: Fulgoromorpha) from the European Palaeocene" 3240: 2788: 2763: 2440: 2370: 1655: 1602: 1163:

approach, which showed that this pest has the potential to occur in the majority of South Korea.

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will perform a brief courtship and copulation which will last for up to 4 hours. In their native

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Han, Jung Min; Kim, Hyojoong; Lim, Eun Ji; Lee, Seunghwan; Kwon, Yong-Jung; Cho, Soowon (2008).

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Potential Economic Impact of the Spotted Lanternfly on Agriculture and Forestry in Pennsylvania

2610:. Cornell College of Agriculture and Life Sciences, New York State Integrated Pest Management. 2434:

Nakashita, Ayano; Wang, Yayun; Lu, Sihan; Shimada, Keisuke; Tsuchida, Tsutomu (February 2022).

5975: 5861: 5697: 5689: 5608: 5547: 5494: 5401: 5383: 5326: 5318: 5233: 5225: 5174: 5156: 5095: 5060: 5052: 5004: 4996: 4953: 4722: 4463: 4456:"A Kansas boy entered a unique insect at the state fair. It triggered a federal investigation" 4424: 4367:. State of Tennessee Department of Agriculture. State of Tennessee Department of Agriculture. 3851: 3822: 3782: 3723: 3687: 3669: 3535: 3391: 3334: 3326: 3232: 2976: 2968: 2833: 2780: 2739: 2586: 2539:

Wang, Rong-Rong; Liu, Jia-Jia; Li, Xin-Yu; Liang, Ai-Ping; Bourgoin, Thierry (27 March 2018).

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As of 2020, the PDA recommended several different pesticides to treat infestations, including

1261: 1184: 1143: 923: 641: 637: 575: 2541:"Relating antennal sensilla diversity and possible species behaviour in the planthopper pest 1377:

and southerly parts of other provinces are modeled to have low suitability for inhabitation.

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Kim, Jae Geun; Lee, Eun-Hyuk; Seo, Yeo-Min; Kim, Na-Yeon (8 June 2011). "Cyclic behavior of

2960: 2823: 2772: 2729: 2721: 2576: 2566: 2465: 2457: 2362: 2348:(Hemiptera: Auchenorrhyncha: Fulgoridae: Aphaeninae) finally, but suddenly arrived in Korea" 2310: 2119: 2065: 1871: 1835: 1300: 1192: 931: 911: 873: 841: 796:) – and common forest plants, as the nymphs are known to associate with other plants beyond 452: 364: 356: 283:

to Japan, South Korea, and the United States, where it is often referred to by the acronym "

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The spotted lanternfly has been found to be capable of reproducing using hosts other than

785: 700: in). Females also have longer legs as compared to those of their male counterparts. 384: 347: 324: 117: 818:. In the United States, high populations are seen infesting common forest trees, such as 5369: 5352:

Clifton, Eric H.; Castrillo, Louela A.; Gryganskyi, Andrii; Hajek, Ann E. (7 May 2019).

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Kime, Lynn F.; Harper, Jayson K.; Stone, William; Kelsey, Timothy W. (December 2019).

5997: 5338: 5107: 4834: 4790: 4289:"Michigan DNR advises residents to be on the lookout for invasive spotted lanternfly" 2366: 2215:(White 1845) review: biology, ecology and pest management with reference to kiwifruit 1481: 1316: 1288: 1036: 1024: 981: 934:

that was introduced to North America. This host choice is thought to have evolved as

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influence Host-Searching and -Acceptance of the Egg-Parasitoid Anastatus orientalis"

4566:"Oregon nursery finds destructive spotted lanternfly, first ever reported in Oregon" 3244: 3083: 2792: 2374: 5866: 4660:"Spotted lanternfly: A serious threat to Canada's wine and agricultural industries" 4629:"Spotted lanternfly: A serious threat to Canada's wine and agricultural industries" 4495:"Kansas boy's insect entry at state fair wins prize – and triggers federal inquiry" 4095:"New York State announces confirmed finding of Spotted Lanternfly on Staten Island" 3984:

U.S. Department of Agriculture, Animal and Plant Health Inspection Service (n.d.).

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was previously introduced into the United States in 1908 for population control of

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In 2009, Japan reported its first confirmed presence of the spotted lanternfly, in

894: 493:, another planthopper species native to Asia, and referenced prior descriptions by 420: 408: 3596: 1544:) may be used to lure them into sticky traps to augment this pest control method. 645: 501: 218: 207: 5811: 5151: 4826: 2677:"Entomologists hope vigilance, research stop newly discovered spotted lanternfly" 2571: 2248:"You Ain't From Around Here! Exotic Invasive of the Quarter: Spotted Lanternfly ( 5949: 5923: 5820: 5220: 5193: 3818: 3755:

Jung, Jae-Min; Jung, Sunghoon; Byeon, Dae-hyeon; Lee, Wang-Hee (December 2017).

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accumulation of this thick honeydew and tree sap on leaves below the host plant

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third, a white wedge in the middle of the wing, and a solid black wing tip. The

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Proceedings of the National Academy of Sciences of the United States of America

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waste they excrete are harmful to the health of host plants. They feed on the

137: 5693: 5612: 5421:"There's fungus among us, and it might kill off the spotted lanternfly in Pa" 5387: 5322: 5229: 5160: 5099: 5056: 5000: 4957: 4467: 3855: 3826: 3786: 3673: 3395: 3330: 3236: 2972: 2837: 2784: 2343: 5941: 5887: 5480: 5378: 4928:"Environmentally-friendly control methods and forecasting the hatching time 4097:. The New York State Department of Agriculture and Markets. 14 August 2020. 3262: 2123: 1650: 1597: 1556: 1537: 1533: 1513: 1477: 1071: 1007:

The spotted lanternfly is native to subtropical regions of Southeast Asia.

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has been found on more than 20 newly recorded host species of woody plants.

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populations seen as a risk factor for further infestation globally. Ongoing

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into South Korea in 2006 and Japan in 2009, and has since been considered a

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and the lack of consistent freezing seasons are considered risk factors in

5259: 3370:(Hemiptera: Fulgoridae) and response on feeding stimulants of some plants" 2315: 5853: 5848: 5805: 5542: 5281:

Clifton, Eric H.; Castrillo, Louela A.; Hajek, Ann E. (1 November 2021).

5047: 5030: 4419: 4065:"These three N.J. counties are now under a spotted lanternfly quarantine" 3531: 3321: 2964: 2776: 2549: 2210: 2192:

Lin, You-Sheng, et al. "Lanternflies (Hemiptera: Fulgoridae) of Taiwan."

1875: 1552: 1501: 1489: 1473: 1431: 1324: 1312: 1292: 1284: 1204: 1091: 1055: 1020: 903: 848:

tissue of foliage and stems with specialized mouthparts, and sucking the

756: 748: 680: in) while males are a smaller size, between 21 and 22 millimetres ( 625: 346:) of the spotted lanternfly are characterized by spotted black and white 4992: 4707: 2608:"Spotted Lanternfly Biology: Life Cycle, Identification, and Dispersion" 5915: 4752:

Jung, Jae-Min; Jung, Sunghoon; Byeon, Dae-hyeon; Lee, Wang-Hee (2017).

4717: 4183: 1308: 1200: 1196: 1083: 1079: 1067: 958: 789: 629: 486: 447: 351: 302: 147: 5928: 4010:. Massachusetts Department of Agricultural Resources. September 2020. 3665: 2866: 1628:) have been found to parasitize and kill the spotted lanternfly, with 1102:, and has been used since the 1100s for topical relief from swelling. 780:

The spotted lanternfly has feeding behavior associated with 103 plant

376:

containing 30–50 eggs, often covered with a grayish mud-like coating.

5283:"Discovery of two hypocrealean fungi infecting spotted lanternflies, 3842:"Lanternflies Eat Everything in Sight. The U.S. Is Looking Delicious" 1434:

were found to have the lowest suitability. Both the pervasiveness of

1350: 1346: 1099: 1087: 1051: 1031:

originally evolved in southwest China, eventually diverging into six

1008: 966: 845: 827: 556: 343: 97: 77: 5782: 4972: 3411:"The spotted lanternfly, a new insect pest detected in Pennsylvania" 1663:, have been found to parasitize spotted lanternfly eggs and nymphs. 1166:

A correlation seems to exist between the widespread distribution of

1035:

lineages. One of these lineages then moved northward, following the

893:

Spotted lanternfly's life cycle (instars) with size comparison to a

609: in) wide. Adult lanternflies have a black head and gray-brown 5892: 2291:

Dara, Surendra K.; Barringer, Lawrence; Steven P., Arthurs (2015).

1572:, has been found to effectively kill the invasive trees by causing 989:

temperatures in Pennsylvania, United States. Another study done at

2297:(Hemiptera: Fulgoridae): A New Invasive Pest in the United States" 1214: 1075: 1063: 1059: 971: 888: 823: 819: 781: 711: 599:

measure about 25 millimetres (1 in) long and 12 millimetres (

586: 443: 430: 411:

efforts have sought to limit population growth, due to the threat

315: 310: 306: 5628:"Vineyards Facing An Insect Invasion May Turn To Aliens For Help" 4397:

Wakie, T. T.; Neven, L. G.; Yee, W. L.; Lu, Z. (3 October 2019).

4326: 4216:

North Carolina Department of Agriculture & Consumer Services

3917: 2633:

Gillett-Kaufman, Jennifer L.; Griffith, Taryn B. (August 2018).

1335: 1304: 360: 298: 5879: 5786: 3610: 3509:"Effects of Temperature on Development and Survival of Nymphal 3417:. Penn State's College of Agricultural Sciences. Archived from 2405:"First Genome of Spotted Lanternfly Built from a Single Insect" 632:

is yellowish with black and white bands on the top and bottom.

591:

Spotted lanternfly in New York, where it is an invasive species

5635: 3805:(Hemiptera: Fulgoridae) isolates in Korea, China, and Japan". 3015: 1016: 613:

adorned with black spots. White's original account identified

372: 3137:"Finally, Something Spotted Lanternflies Are Good for: Honey" 2923:. Penton Agriculture Market. 10 November 2014. Archived from 1632:

having also shown an ability to kill spotted lanternflies in

1551:

have found that the spotted lanternfly emits long-wavelength

4537:

Oregon Department of Agriculture Fact Sheets and Pest Alerts

342:

can associate with more than 173 plants. Early life stages (

3011:"Invasive Bug Prompts Quarantine In Pennsylvania Townships" 2917:"Spotted lanternfly – a new threat to grapes, stone fruit?" 926:

habitat, they lay their eggs preferably on tree of heaven (

4403:(Hemiptera: Fulgoridae) in the United States and globally" 1596:

native to the United States have been identified to cause

1237:

first confirmed the presence of the spotted lanternfly in

5254:) to Vector a Plant Pathogen between its Preferred Host, 4179:"Invasive spotted lanternfly makes journey into Virginia" 3106:"Spotted Lanternfly in New Jersey: What you need to know" 1767:

Adult spotted lanternfly caught and eaten by a spider in

945:

but they generally fail to reach high populations unless

536:) as closely related to the spotted lanternfly. The name 497:

in his initial classification of the spotted lanternfly.

367:. The piercing wounds caused by their mouthparts and the 4211:"NCDA&CS finds spotted lanternfly in Forsyth County" 1391:

European and Mediterranean Plant Protection Organization

1345:

Dead spotted lanternfly specimens have been reported in

4322:"Spotted lanternfly: Should you be worried about them?" 640:

covered in needle-like tips. The spotted lanternfly is

415:

poses to global agricultural industries. Parts of the

4392: 4390: 4388: 4386: 1711:

Spotted lanternfly early-instar nymph on a grape leaf

1191:). In 2013, it was confirmed to have spread into the 5738:

PennState Extension spotted lantern information page

5455:"An Old Remedy for a New Problem? Identification of 5123:"Egg masses as training aids for spotted lanternfly 3913:"Spotted lanternfly spreading farther into our area" 1807:"Bourgoin T: Fulgoromorpha Lists On the Web (FLOW): 5795: 5666:Urban, Julie M.; Leach, Heather (23 January 2023). 5258:, in a Laboratory Setting" Insects 11, no. 8: 515. 4123:. Ohio Department of Agriculture. 27 October 2020. 3476:"New Invasive Pest Has Pennsylvania Towns on Alert" 1062:. Since then, it has expanded its range to include 489:, China. White described the species as similar to 16:

Species of planthopper indigenous to Southeast Asia

4596:"Spotted lanternfly: A colorful cause for concern" 3205:(Insecta: Hemiptera: Fulgoridae) on host plants". 1225:in Berks County, Pennsylvania, in mid-October 2019 759:with stone fruits. Depending on the life stage of 4361:"Spotted Lanternfly Detected in Middle Tennessee" 1847: 1845: 1843: 5731:, National Invasive Species Information Center, 5459:(Hymenoptera: Encyrtidae), an Egg Parasitoid of 1727:in Berks County, Pennsylvania, in late July 2018 949:is present. Many hypotheses are given as to why 1555:(8–14 μm) light during active feeding and 1050:was known to inhabit the Northern Provinces of 465:. Species within this genus are found in Asia. 4973:"Developing Traps for the Spotted Lanternfly, 4866:"Spotted Lanternfly Management for Homeowners" 4121:"Spotted Lanternfly Found in Jefferson County" 3082:. First Detection Program. n.d. Archived from 5453:Liu, Houping; Mottern, Jason (January 2017). 5194:"First Report of Verticillium Wilt Caused by 3568:Jacek Szwedo, Adam Stroin´ski (August 2012). 3409:Department of Entomology (10 November 2014). 2812:and the Control Effects of Some Insecticides" 1699:Spotted lanternfly eggs on the bark of a tree 279:to parts of China and Vietnam. It has spread 8: 4932:(Hemiptera: Fulgoridae) in Jeonnam Province" 4603:College of Agriculture and Natural Resources 3882:"Spotted Lanternfly – Profile and Resources" 1723:Late-instar nymphs of spotted lanternfly on 1454:Extension. These guidelines include killing 976:Closeup of "brick and mortar" wing structure 5733:United States National Agricultural Library 4493:Villarreal, Alexandra (15 September 2021). 3611:"Spotted Lanternfly - Encyclopedia of Life" 2861: 2859: 2857: 2855: 2853: 2851: 2849: 2847: 2040:"Unearthing a Better Way to Use Pesticides" 1566:populations; the use of a fungal pathogen, 1098:, the spotted lanternfly is believed to be 652:end of the abdomen, which males lack. When 5783: 5716:Hungry Pests Campaign – Spotted lanternfly 1532:has been found to be attracted to certain 1442:establishing populations in the country. 852:; it does not eat the fruit or the leaves 579:both being classified as close relatives. 29: 20: 5683: 5602: 5541: 5488: 5463:(Hemiptera: Fulgoridae) in North America" 5395: 5377: 5312: 5219: 5168: 5150: 5046: 4947: 4791:"Spotted Lanternfly (Lycorma delicatula)" 4773: 4758:(Hemiptera: Fulgoridae), by using CLIMEX" 4716: 4706: 4418: 3951:(Report). Pennsylvania State University. 3776: 3761:(Hemiptera: Fulgoridae), by using CLIMEX" 3681: 3385: 3320: 3135:Cunningham, Caroline (23 November 2020). 2827: 2733: 2580: 2570: 2469: 2314: 755:trees, including apple trees and several 524:classification places two other species ( 500:In 1863, the species was reclassified by 395:and is rapidly spreading south and west. 334:is often centered on its preferred host, 4255:Michigan Department of Natural Resources 4243:"Forest Pest Alert: Spotted Lanternfly ( 3444:(Hemiptera: Fulgoridae) egg mortality". 2099: 2097: 2095: 2064:Barringer, Lawrence (17 December 2021). 2038:Djajapranata, Cliff (20 November 2023). 1811:(White, 1845) (retrieved 26 April 2020)" 1523:In Pennsylvania and Korea, use of brown 1113: 5720:United States Department of Agriculture 5260:https://doi.org/10.3390/insects11080515 4859: 4857: 4855: 4454:Edwards, Jonathan (15 September 2021). 2670: 2668: 2666: 2664: 2545:(Hemiptera: Fulgoromorpha: Fulgoridae)" 1798: 1689: 980:Testing has been done to determine how 5725:Species Profile – Spotted lanternfly ( 5276: 5274: 4907:from the original on 11 September 2020 4666:from the original on 30 September 2020 4505:from the original on 15 September 2021 4474:from the original on 19 September 2021 4334:from the original on 28 September 2023 4301:from the original on 28 September 2023 4287:Roberts, Andy M. (19 September 2023). 4265:from the original on 28 September 2023 3892:from the original on 10 September 2021 3617:. National Museum of Natural History. 3469: 3467: 3256: 3254: 3177:from the original on 13 September 2023 3004: 3002: 2911: 2909: 2907: 2905: 2903: 2901: 2899: 2897: 2895: 2713:Journal of the Royal Society Interface 2602: 2600: 2534: 2532: 2511:Pennsylvania Department of Agriculture 2501: 2499: 2497: 2495: 2493: 2491: 2489: 2111:Annals and Magazine of Natural History 1520:, which can be applied by homeowners. 1231:Pennsylvania Department of Agriculture 5643:from the original on 20 February 2020 5558:from the original on 30 December 2021 5505:from the original on 30 December 2021 5448: 5446: 5431:from the original on 1 September 2019 4546:from the original on 25 December 2021 4371:from the original on 16 November 2023 3992:from the original on 20 January 2022. 3979: 3977: 3975: 3973: 3862:from the original on 15 December 2019 3703: 3701: 3621:from the original on 27 February 2022 3294: 3292: 3196: 3194: 3192: 3167:"Spotted Lanternflies and Beekeeping" 3040: 3038: 3023:from the original on 25 December 2014 2649:from the original on 29 November 2021 2302:Journal of Integrated Pest Management 2204: 2202: 1512:. Infested trees can be treated with 751:, fruit trees, ornamental trees, and 7: 6034:Taxa named by Adam White (zoologist) 5591:Korean Journal of Applied Entomology 4936:Korean Journal of Applied Entomology 4841:from the original on 17 January 2022 4762:Journal of Asia-Pacific Biodiversity 4733:from the original on 12 January 2022 4639:from the original on 2 December 2022 4576:from the original on 18 October 2020 4127:from the original on 1 November 2020 4101:from the original on 16 October 2020 4014:from the original on 20 October 2020 3911:Harrison, Courtney (22 March 2021). 3765:Journal of Asia-Pacific Biodiversity 3549:from the original on 22 January 2022 3488:from the original on 3 December 2014 3374:Korean Journal of Applied Entomology 3277:from the original on 11 January 2022 3261:CAB, International (November 2021). 3147:from the original on 19 October 2023 3116:from the original on 1 December 2021 3056:from the original on 17 October 2022 2880:from the original on 26 January 2022 2816:Korean Journal of Applied Entomology 2687:from the original on 1 February 2023 2614:from the original on 24 January 2022 2384:from the original on 22 January 2022 2272:from the original on 22 January 2022 2227:from the original on 28 January 2022 2165:from the original on 25 January 2022 2080:from the original on 11 January 2022 2019:from the original on 23 January 2023 1989:from the original on 23 January 2023 1739:Spotted lanternfly late-instar nymph 914:, spotted lanternflies will perform 784:, accounting for 33 families and 17 435:Spotted lanternfly showing underwing 5579:"Parasitism rate of egg parasitoid 4609:from the original on 1 October 2020 4191:from the original on 5 October 2018 4075:from the original on 5 October 2019 4063:Warren, Michael (30 January 2019). 4044:from the original on 1 October 2019 3925:from the original on 5 January 2022 2761:(Insecta: Hemiptera: Fulgoridae)". 2415:from the original on 9 January 2022 2209:Mauchline, C.; McKenna, C. (2019). 2130:from the original on 5 October 2021 1580:by functioning as a vector for the 1417:in Madrid, Spain on July 17, 2024. 767:it may prefer other hosts, such as 726:, United States, in early July 2018 5685:10.1146/annurev-ento-120220-111140 5626:Charles, Dan (16 September 2019). 4038:Delaware Department of Agriculture 3958:from the original on 10 April 2022 3807:Journal of Asia-Pacific Entomology 3446:Journal of Asia-Pacific Entomology 2987:from the original on 16 March 2023 14: 6024:Insect pests of temperate forests 5587:(Hemiptera: Fulgoridae) in China" 5300:Journal of Invertebrate Pathology 4895:Griffaton, Grace (15 June 2020). 4435:from the original on 23 July 2020 4223:from the original on 29 June 2022 4158:from the original on 27 July 2021 3345:from the original on 17 June 2022 3009:Chapell, Bill (3 November 2014). 2517:from the original on 20 June 2020 2323:from the original on 8 April 2022 1219:Adult spotted lanternflies on an 1159:has since been predicted using a 918:. During these migrations, adult 5419:Buffone, John (30 August 2019). 4570:Oregon Department of Agriculture 4526:"Pest Alert: Spotted lanternfly 4320:Bink, Addy (17 September 2023). 4177:McGrath, Mike (5 October 2018). 3736:from the original on 18 May 2023 3474:Maher, Kris (18 November 2014). 2675:Gill, Chuck (17 November 2014). 2367:10.1111/j.1748-5967.2008.00188.x 2246:Gagnon, Jennifer (Spring 2016). 1868:Entomological Society of America 1776: 1760: 1744: 1732: 1716: 1704: 1692: 54: 2808:"Biological Characteristics of 2403:Kaplan, Kim (16 October 2019). 1973:"Spotted Lanternfly Pest Alert" 1459:hosts, such as tree of heaven ( 1428:Australian Great Dividing Range 1366:Canadian Food Inspection Agency 5035:Journal of Economic Entomology 4407:Journal of Economic Entomology 4146:Entomology (26 January 2021). 3840:Montague, Zach (21 May 2018). 3708:Burne, Allan (17 March 2020). 2448:(1543 (2022)). Figure 5. 1751:Adult spotted lanternflies in 1266:U.S. Department of Agriculture 1142:, albeit with greatly reduced 359:but cause extensive damage to 1: 5764:"Spotted Lanternfly Spotters" 5583:(Hymenoptera: Eupelmidae) on 3597:10.1016/j.geobios.2011.10.007 1769:Delaware County, Pennsylvania 1258:Pennsylvania State University 1253:67 counties in Pennsylvania. 965:eggs undergo diapause during 716:Spotted lanternfly nymphs on 644:. Females have a set of red 506:Lycorma delicatula delicatula 323:populations are regulated by 5152:10.1371/journal.pone.0250945 2572:10.1371/journal.pone.0194995 2211:BS1847: Spotted lanternfly, 1235:Pennsylvania Game Commission 1096:traditional Chinese medicine 1015:evolved between 55 and 51.6 319:spp. In its native habitat, 5672:Annual Review of Entomology 5221:10.1094/PDIS-01-13-0062-PDN 4687:"Footprints and Ootheca of 4662:. Invasive Species Centre. 4524:Department of Agriculture. 4399:"The establishment risk of 3819:10.1016/j.aspen.2013.07.003 3458:10.1016/j.aspen.2010.09.004 1600:in the spotted lanternfly. 1557:rapid hemolymph circulation 877:and wasps; infestations of 803:Parthenocissus quinquefolia 399:egg masses are the primary 6050: 5604:10.5656/ksae.2014.01.1.075 5092:10.1007/s10340-021-01338-7 4949:10.5656/ksae.2012.09.0.022 4827:10.21203/rs.3.rs-400798/v1 4789:Christian (18 July 2024). 4775:10.1016/j.japb.2017.07.001 3778:10.1016/j.japb.2017.07.001 3387:10.5656/ksae.2009.48.4.467 3208:Journal of Insect Behavior 2829:10.5656/ksae.2009.48.1.053 2462:10.1038/s41598-022-05541-z 2149:Thierry, Bourgoin (2015). 1540:oil. Such chemicals (like 1229:On 29 September 2014, the 724:Berks County, Pennsylvania 518:Lycorma delicatula operosa 393:Northeastern United States 6029:Insects described in 1845 6004:Agricultural pest insects 5631:(Radio broadcast episode) 5468:Journal of Insect Science 5314:10.1016/j.jip.2021.107689 5293:Ophiocordyceps delicatula 4601:Michigan State University 3646:Evolutionary Applications 3229:10.1007/s10905-011-9266-8 2645:. University of Florida. 2259:Virginia Forest Landowner 2220:Plant & Food Research 1621:Ophiocordyceps delicatula 985:survived the much colder 936:a mechanism of protection 897:(24 mm, 1.0 in) 830:; in Pennsylvania alone, 636:displays orange, bulbous 330:The spotted lanternfly's 246: 239: 200: 195: 176: 169: 51:Scientific classification 49: 37: 28: 23: 5530:Environmental Entomology 5196:Verticillium nonalfalfae 4981:Environmental Entomology 4977:(Hemiptera: Fulgoridae)" 4695:Environmental Entomology 3520:Environmental Entomology 3513:(Hemiptera: Fulgoridae)" 3507:Kreitman, Devin (2021). 3309:Environmental Entomology 2953:Environmental Entomology 2009:"Spotted Lanternfly Map" 1855:Environmental Entomology 1836:CDFA: Spotted Lanternfly 1569:Verticillium nonalfalfae 1338:-growing valleys of the 1027:analysis has shown that 930:), which is a poisonous 548:, meaning "luxurious". 5379:10.1073/pnas.1903579116 5265:7 December 2023 at the 5080:Journal of Pest Science 4633:Invasive Species Centre 3886:Invasive Species Centre 3719:Biosecurity New Zealand 3710:"Pest risk assessment: 3481:The Wall Street Journal 2159:www.catalogueoflife.org 2153:Lycorma delicatula jole 2124:10.1080/037454809495244 1864:Oxford University Press 1753:Brooklyn Botanic Garden 1670:Lymantria dispar dispar 1592:Four species of fungal 1110:Accidental introduction 514:Lycorma delicatula jole 419:are undergoing massive 40:Brooklyn Botanic Garden 6014:Endemic fauna of China 2726:10.1098/rsif.2021.0367 2355:Entomological Research 1340:Pacific coastal states 1226: 1119: 977: 938:from natural enemies. 898: 809:Phellodendron amurense 727: 592: 508:, with two additional 483:scientific collections 436: 427:Taxonomy and discovery 5481:10.1093/jisesa/iew114 5427:. USA Today Network. 5291:and a novel species, 4658:Ehnes, Mandy (n.d.). 3714:(Spotted lanternfly)" 3366:"Feeding behavior of 3267:(spotted lanternfly)" 3141:Philadelphia Magazine 3110:Mendham Garden Center 2876:. 30 September 2016. 2683:. Science X network. 2635:"Spotted lanternfly, 2070:(spotted lanternfly)" 2044:Georgetown University 1549:infrared thermography 1446:Possible pest control 1250:recreational vehicles 1218: 1117: 975: 967:embryonic development 892: 844:plants, piercing the 715: 590: 544:, meaning "lamp" and 495:George Tradescant Lay 485:were made outside of 451:, in the planthopper 434: 387:. In September 2014, 5744:"Spotted Lanternfly" 5581:Anastatus orientalis 5289:Metarhizium pemphigi 4605:. 12 November 2020. 4148:"Spotted Lanternfly" 4034:"Spotted Lanternfly" 3986:"Spotted Lanternfly" 3415:Entomology News 2014 3076:"Spotted Lanternfly" 3050:Penn State Extension 3046:"Spotted Lanternfly" 2867:"Lycorma delicatula" 2777:10.3157/021.125.0105 2507:"Spotted Lanternfly" 2104:White, Adam (1845). 1783:4th instar nymph in 1679:Anastatus orientalis 1615:Metarhizium pemphigi 1589:or other pathogens. 1375:southwestern Ontario 1211:In the United States 1011:evidence indicates 442:is a species in the 397:L. delicatula's 5370:2019PNAS..116.9178C 5256:Ailanthus altissima 5200:Ailanthus altissima 5198:on Tree-of-Heaven ( 5143:2021PLoSO..1650945E 4870:PennState Extension 4460:The Washington Post 3658:2021EvApp..14..915D 3589:2012Geobi..45..413S 3303:Ailanthus altissima 3221:2011JIBeh..24..423K 3086:on 25 December 2021 2563:2018PLoSO..1394995W 2454:2022NatSR..12.1543N 2316:10.1093/jipm/pmv021 1771:, in September 2020 1587:V. nonalfalfae 1582:V. nonalfalfae 1564:Ailanthus altissima 1514:systemic pesticides 1462:Ailanthus altissima 1189:Ishikawa Prefecture 840:feeds on woody and 732:Ailanthus altissima 405:Ailanthus altissima 350:that develop a red 336:Ailanthus altissima 297:), but it feeds on 294:Ailanthus altissima 291:is tree of heaven ( 24:Spotted lanternfly 5827:Lycorma delicatula 5797:Lycorma delicatula 5750:. 19 November 2020 5727:Lycorma delicatula 5585:Lycorma delicatula 5543:10.1093/ee/nvaa137 5461:Lycorma delicatula 5457:Ooencyrtus kuvanae 5285:Lycorma delicatula 5252:Lycorma delicatula 5125:Lycorma delicatula 5048:10.1093/jee/toy412 4975:Lycorma delicatula 4930:Lycorma delicatula 4756:Lycorma delicatula 4689:Lycorma delicatula 4572:. 8 October 2020. 4528:Lycorma delicatula 4420:10.1093/jee/toz259 4401:Lycorma delicatula 4245:Lycorma delicatula 3847:The New York Times 3803:Lycorma delicatula 3759:Lycorma delicatula 3712:Lycorma delicatula 3532:10.1093/ee/nvaa155 3511:Lycorma delicatula 3442:Lycorma delicatula 3368:Lycorma delicatula 3322:10.1093/ee/nvaa083 3265:Lycorma delicatula 3203:Lycorma delicatula 2965:10.1093/ee/nvaa126 2921:Western Farm Press 2810:Lycorma delicatula 2764:Entomological News 2759:Lycorma delicatula 2643:entnemdept.ufl.edu 2637:Lycorma delicatula 2543:Lycorma delicatula 2441:Scientific Reports 2346:Lycorma delicatula 2295:Lycorma delicatula 2250:Lycorma delicatula 2213:Lycorma delicatula 2068:Lycorma delicatula 1876:10.1093/ee/nvaa093 1870:(ESA)): 999–1011. 1809:Lycorma delicatula 1665:Ooencyrtus kuvanae 1656:Ooencyrtus kuvanae 1630:Beauveria bassiana 1603:Beauveria bassiana 1547:Researchers using 1530:L. delicatula 1470:insecticidal soaps 1456:L. delicatula 1440:L. delicatula 1403:L. delicatula 1399:L. delicatula 1395:L. delicatula 1379:L. delicatula 1371:L. delicatula 1227: 1172:L. delicatula 1157:L. delicatula 1136:L. delicatula 1132:L. delicatula 1123:Lycorma delicatula 1120: 1048:L. delicatula 991:Rutgers University 978: 963:L. delicatula 951:L. delicatula 943:A. altissima, 920:L. delicatula 899: 879:L. delicatula 869:L. delicatula 862:Lycorma delicatula 838:L. delicatula 832:L. delicatula 776:A. altissima. 761:L. delicatula 741:L. delicatula 728: 642:sexually dimorphic 634:L. delicatula 615:L. delicatula 597:L. delicatula 593: 561:L. delicatula 550:L. delicatula 538:Lycorma delicatula 527:L. imperialis 479:Aphaena delicatula 467:L. delicatula 440:Lycorma delicatula 437: 413:L. delicatula 389:L. delicatula 340:L. delicatula 321:L. delicatula 269:Lycorma delicatula 264:spotted lanternfly 250:Aphaena delicatula 180:Lycorma delicatula 162:L. delicatula 6019:Hemiptera of Asia 5991: 5990: 5976:Open Tree of Life 5789:Taxon identifiers 5770:. 27 October 2020 5425:York Daily Record 5364:(19): 9178–9180. 4993:10.1093/ee/nvz166 4821:, pp. 1–18, 4708:10.1093/ee/nvz110 4294:White Lake Beacon 3729:978-1-99-001754-4 3666:10.1111/eva.13170 3421:on 5 January 2015 3171:extension.psu.edu 2927:on 30 August 2018 2720:(181): 20210367. 2155:Stål, 1863 | COL" 1755:in September 2021 1626:Entomophthoraceae 1578:A. altissima 1542:methyl salicylate 1510:zeta-cypermethrin 1436:A. altissima 1413:was observed via 1407:A. altissima 1262:forestry industry 1168:A. altissima 1140:A. altissima 1127:A. altissima 947:A. altissima 928:A. altissima 916:migratory flights 798:A. altissima 745:A. altissima 708:Host associations 576:Pyrops candelaria 491:Aphaena variegata 365:ornamental plants 357:herbaceous plants 287:". Its preferred 260: 259: 254: 233: 222: 211: 6041: 5984: 5983: 5971: 5970: 5958: 5957: 5945: 5944: 5932: 5931: 5919: 5918: 5906: 5905: 5896: 5895: 5883: 5882: 5870: 5869: 5857: 5856: 5844: 5843: 5831: 5830: 5829: 5816: 5815: 5814: 5784: 5779: 5777: 5775: 5759: 5757: 5755: 5705: 5687: 5653: 5652: 5650: 5648: 5632: 5623: 5617: 5616: 5606: 5574: 5568: 5567: 5565: 5563: 5545: 5521: 5515: 5514: 5512: 5510: 5492: 5450: 5441: 5440: 5438: 5436: 5416: 5410: 5409: 5399: 5381: 5349: 5343: 5342: 5316: 5278: 5269: 5248: 5242: 5241: 5223: 5189: 5183: 5182: 5172: 5154: 5118: 5112: 5111: 5075: 5069: 5068: 5050: 5026: 5020: 5019: 5017: 5015: 4968: 4962: 4961: 4951: 4923: 4917: 4916: 4914: 4912: 4892: 4886: 4885: 4883: 4881: 4876:on 25 April 2021 4872:. Archived from 4861: 4850: 4849: 4848: 4846: 4812: 4806: 4805: 4803: 4801: 4786: 4780: 4779: 4777: 4749: 4743: 4742: 4740: 4738: 4720: 4710: 4701:(6): 1270–1276. 4682: 4676: 4675: 4673: 4671: 4655: 4649: 4648: 4646: 4644: 4635:. 16 June 2020. 4625: 4619: 4618: 4616: 4614: 4592: 4586: 4585: 4583: 4581: 4562: 4556: 4555: 4553: 4551: 4545: 4534: 4521: 4515: 4514: 4512: 4510: 4490: 4484: 4483: 4481: 4479: 4451: 4445: 4444: 4442: 4440: 4422: 4394: 4381: 4380: 4378: 4376: 4357: 4351: 4350: 4341: 4339: 4317: 4311: 4310: 4308: 4306: 4284: 4278: 4277: 4272: 4270: 4264: 4251: 4239: 4233: 4232: 4230: 4228: 4219:. 29 June 2022. 4207: 4201: 4200: 4198: 4196: 4174: 4168: 4167: 4165: 4163: 4143: 4137: 4136: 4134: 4132: 4117: 4111: 4110: 4108: 4106: 4091: 4085: 4084: 4082: 4080: 4060: 4054: 4053: 4051: 4049: 4030: 4024: 4023: 4021: 4019: 4000: 3994: 3993: 3981: 3968: 3967: 3965: 3963: 3957: 3950: 3941: 3935: 3934: 3932: 3930: 3908: 3902: 3901: 3899: 3897: 3878: 3872: 3871: 3869: 3867: 3837: 3831: 3830: 3797: 3791: 3790: 3780: 3752: 3746: 3745: 3743: 3741: 3705: 3696: 3695: 3685: 3637: 3631: 3630: 3628: 3626: 3607: 3601: 3600: 3574: 3565: 3559: 3558: 3556: 3554: 3548: 3517: 3504: 3498: 3497: 3495: 3493: 3471: 3462: 3461: 3437: 3431: 3430: 3428: 3426: 3406: 3400: 3399: 3389: 3361: 3355: 3354: 3352: 3350: 3324: 3315:(5): 1185–1190. 3296: 3287: 3286: 3284: 3282: 3258: 3249: 3248: 3198: 3187: 3186: 3184: 3182: 3163: 3157: 3156: 3154: 3152: 3132: 3126: 3125: 3123: 3121: 3112:. 31 July 2020. 3102: 3096: 3095: 3093: 3091: 3072: 3066: 3065: 3063: 3061: 3042: 3033: 3032: 3030: 3028: 3006: 2997: 2996: 2994: 2992: 2959:(6): 1270–1281. 2943: 2937: 2936: 2934: 2932: 2913: 2890: 2889: 2887: 2885: 2863: 2842: 2841: 2831: 2803: 2797: 2796: 2754: 2748: 2747: 2737: 2703: 2697: 2696: 2694: 2692: 2672: 2659: 2658: 2656: 2654: 2630: 2624: 2623: 2621: 2619: 2604: 2595: 2594: 2584: 2574: 2536: 2527: 2526: 2524: 2522: 2503: 2484: 2483: 2473: 2431: 2425: 2424: 2422: 2420: 2409:www.ars.usda.gov 2400: 2394: 2393: 2391: 2389: 2383: 2352: 2339: 2333: 2332: 2330: 2328: 2318: 2288: 2282: 2281: 2279: 2277: 2271: 2256: 2243: 2237: 2236: 2234: 2232: 2206: 2197: 2190: 2184: 2181: 2175: 2174: 2172: 2170: 2146: 2140: 2139: 2137: 2135: 2101: 2090: 2089: 2087: 2085: 2061: 2055: 2054: 2052: 2050: 2035: 2029: 2028: 2026: 2024: 2005: 1999: 1998: 1996: 1994: 1988: 1977: 1969: 1963: 1962: 1960: 1958: 1944: 1938: 1937: 1935: 1933: 1919: 1913: 1912: 1910: 1908: 1894: 1888: 1887: 1849: 1838: 1833: 1827: 1826: 1824: 1822: 1817:on 1 August 2020 1813:. Archived from 1803: 1780: 1764: 1748: 1736: 1720: 1708: 1696: 1272:Researchers and 1268:, consisting of 1193:Fukui Prefecture 1039:during the late 932:invasive species 699: 698: 694: 689: 688: 684: 679: 678: 674: 671: 665: 664: 660: 608: 607: 603: 570:Aphaena amabilis 481:, and the first 403:of spread, with 379:The species was 252: 228: 217: 206: 204:L. d. delicatula 182: 59: 58: 33: 21: 6049: 6048: 6044: 6043: 6042: 6040: 6039: 6038: 5994: 5993: 5992: 5987: 5979: 5974: 5966: 5961: 5953: 5948: 5940: 5935: 5927: 5922: 5914: 5909: 5901: 5899: 5891: 5886: 5878: 5873: 5865: 5860: 5852: 5847: 5839: 5834: 5825: 5824: 5819: 5810: 5809: 5804: 5791: 5773: 5771: 5762: 5753: 5751: 5742: 5712: 5665: 5662: 5660:Further reading 5657: 5656: 5646: 5644: 5630: 5625: 5624: 5620: 5576: 5575: 5571: 5561: 5559: 5523: 5522: 5518: 5508: 5506: 5452: 5451: 5444: 5434: 5432: 5418: 5417: 5413: 5351: 5350: 5346: 5280: 5279: 5272: 5267:Wayback Machine 5249: 5245: 5191: 5190: 5186: 5137:(5): e0250945. 5127:detection dogs" 5120: 5119: 5115: 5077: 5076: 5072: 5028: 5027: 5023: 5013: 5011: 4970: 4969: 4965: 4925: 4924: 4920: 4910: 4908: 4894: 4893: 4889: 4879: 4877: 4863: 4862: 4853: 4844: 4842: 4814: 4813: 4809: 4799: 4797: 4788: 4787: 4783: 4751: 4750: 4746: 4736: 4734: 4684: 4683: 4679: 4669: 4667: 4657: 4656: 4652: 4642: 4640: 4627: 4626: 4622: 4612: 4610: 4594: 4593: 4589: 4579: 4577: 4564: 4563: 4559: 4549: 4547: 4543: 4532: 4523: 4522: 4518: 4508: 4506: 4492: 4491: 4487: 4477: 4475: 4453: 4452: 4448: 4438: 4436: 4396: 4395: 4384: 4374: 4372: 4359: 4358: 4354: 4337: 4335: 4319: 4318: 4314: 4304: 4302: 4286: 4285: 4281: 4268: 4266: 4262: 4249: 4241: 4240: 4236: 4226: 4224: 4209: 4208: 4204: 4194: 4192: 4176: 4175: 4171: 4161: 4159: 4145: 4144: 4140: 4130: 4128: 4119: 4118: 4114: 4104: 4102: 4093: 4092: 4088: 4078: 4076: 4062: 4061: 4057: 4047: 4045: 4032: 4031: 4027: 4017: 4015: 4002: 4001: 3997: 3983: 3982: 3971: 3961: 3959: 3955: 3948: 3943: 3942: 3938: 3928: 3926: 3910: 3909: 3905: 3895: 3893: 3880: 3879: 3875: 3865: 3863: 3839: 3838: 3834: 3799: 3798: 3794: 3754: 3753: 3749: 3739: 3737: 3730: 3707: 3706: 3699: 3639: 3638: 3634: 3624: 3622: 3609: 3608: 3604: 3572: 3567: 3566: 3562: 3552: 3550: 3546: 3515: 3506: 3505: 3501: 3491: 3489: 3473: 3472: 3465: 3439: 3438: 3434: 3424: 3422: 3408: 3407: 3403: 3363: 3362: 3358: 3348: 3346: 3298: 3297: 3290: 3280: 3278: 3260: 3259: 3252: 3200: 3199: 3190: 3180: 3178: 3165: 3164: 3160: 3150: 3148: 3134: 3133: 3129: 3119: 3117: 3104: 3103: 3099: 3089: 3087: 3074: 3073: 3069: 3059: 3057: 3044: 3043: 3036: 3026: 3024: 3008: 3007: 3000: 2990: 2988: 2945: 2944: 2940: 2930: 2928: 2915: 2914: 2893: 2883: 2881: 2865: 2864: 2845: 2805: 2804: 2800: 2756: 2755: 2751: 2705: 2704: 2700: 2690: 2688: 2674: 2673: 2662: 2652: 2650: 2632: 2631: 2627: 2617: 2615: 2606: 2605: 2598: 2557:(3): e0194995. 2538: 2537: 2530: 2520: 2518: 2505: 2504: 2487: 2433: 2432: 2428: 2418: 2416: 2402: 2401: 2397: 2387: 2385: 2381: 2350: 2341: 2340: 2336: 2326: 2324: 2290: 2289: 2285: 2275: 2273: 2269: 2254: 2245: 2244: 2240: 2230: 2228: 2208: 2207: 2200: 2191: 2187: 2182: 2178: 2168: 2166: 2148: 2147: 2143: 2133: 2131: 2103: 2102: 2093: 2083: 2081: 2063: 2062: 2058: 2048: 2046: 2037: 2036: 2032: 2022: 2020: 2007: 2006: 2002: 1992: 1990: 1986: 1975: 1971: 1970: 1966: 1956: 1954: 1946: 1945: 1941: 1931: 1929: 1921: 1920: 1916: 1906: 1904: 1896: 1895: 1891: 1851: 1850: 1841: 1834: 1830: 1820: 1818: 1805: 1804: 1800: 1795: 1788: 1781: 1772: 1765: 1756: 1749: 1740: 1737: 1728: 1721: 1712: 1709: 1700: 1697: 1688: 1646:Dryinus sinicus 1624:(in the family 1594:entomopathogens 1496:as bark spray, 1448: 1423: 1387: 1363: 1241:, northwest of 1213: 1181: 1152: 1112: 1033:phylogeographic 1005: 1000: 887: 710: 696: 692: 691: 686: 682: 681: 676: 672: 669: 667: 662: 658: 657: 605: 601: 600: 585: 565:fully sequenced 469:was originally 429: 325:parasitic wasps 191: 184: 178: 165: 118:Auchenorrhyncha 53: 17: 12: 11: 5: 6047: 6045: 6037: 6036: 6031: 6026: 6021: 6016: 6011: 6006: 5996: 5995: 5989: 5988: 5986: 5985: 5972: 5959: 5946: 5933: 5920: 5907: 5897: 5884: 5871: 5858: 5845: 5832: 5817: 5801: 5799: 5793: 5792: 5787: 5781: 5780: 5760: 5740: 5735: 5722: 5711: 5710:External links 5708: 5707: 5706: 5678:(1): 151–167. 5661: 5658: 5655: 5654: 5618: 5597:(2): 135–139. 5569: 5516: 5442: 5411: 5344: 5270: 5243: 5184: 5113: 5086:(2): 231–240. 5070: 5041:(2): 671–682. 5021: 4987:(2): 269–276. 4963: 4942:(4): 371–376. 4918: 4887: 4851: 4807: 4781: 4768:(4): 532–538. 4744: 4677: 4650: 4620: 4587: 4557: 4516: 4485: 4446: 4413:(1): 306–314. 4382: 4352: 4312: 4279: 4234: 4202: 4169: 4138: 4112: 4086: 4055: 4025: 3995: 3969: 3936: 3903: 3873: 3832: 3813:(4): 503–506. 3792: 3771:(4): 532–538. 3747: 3728: 3697: 3652:(4): 915–930. 3632: 3602: 3583:(4): 413–420. 3560: 3526:(1): 183–191. 3499: 3463: 3452:(1): 123–125. 3432: 3401: 3380:(4): 467–477. 3356: 3288: 3250: 3215:(6): 423–435. 3188: 3158: 3127: 3097: 3080:First Detector 3067: 3034: 2998: 2938: 2891: 2843: 2798: 2749: 2698: 2660: 2625: 2596: 2528: 2485: 2426: 2395: 2361:(4): 281–296. 2334: 2283: 2238: 2198: 2185: 2176: 2141: 2091: 2056: 2030: 2000: 1964: 1939: 1914: 1889: 1839: 1828: 1797: 1796: 1794: 1791: 1790: 1789: 1782: 1775: 1773: 1766: 1759: 1757: 1750: 1743: 1741: 1738: 1731: 1729: 1725:Vitis labrusca 1722: 1715: 1713: 1710: 1703: 1701: 1698: 1691: 1687: 1684: 1651:dryinidae wasp 1482:essential oils 1447: 1444: 1422: 1419: 1386: 1383: 1362: 1359: 1321:North Carolina 1233:(PDA) and the 1212: 1209: 1180: 1177: 1151: 1150:In South Korea 1148: 1125:normally uses 1111: 1108: 1046:In the 1930s, 1004: 1001: 999: 996: 987:winter 2013–14 912:mating periods 886: 883: 858:photosynthesis 815:Toona sinensis 719:Vitis labrusca 709: 706: 584: 581: 533:L. meliae 428: 425: 258: 257: 256: 255: 244: 243: 237: 236: 235: 234: 223: 212: 198: 197: 193: 192: 185: 174: 173: 167: 166: 159: 157: 153: 152: 145: 141: 140: 135: 131: 130: 125: 121: 120: 115: 111: 110: 105: 101: 100: 95: 91: 90: 85: 81: 80: 75: 71: 70: 65: 61: 60: 47: 46: 35: 34: 26: 25: 15: 13: 10: 9: 6: 4: 3: 2: 6046: 6035: 6032: 6030: 6027: 6025: 6022: 6020: 6017: 6015: 6012: 6010: 6007: 6005: 6002: 6001: 5999: 5982: 5977: 5973: 5969: 5964: 5960: 5956: 5951: 5947: 5943: 5938: 5934: 5930: 5925: 5921: 5917: 5912: 5908: 5904: 5898: 5894: 5889: 5885: 5881: 5876: 5872: 5868: 5863: 5859: 5855: 5850: 5846: 5842: 5837: 5833: 5828: 5822: 5818: 5813: 5807: 5803: 5802: 5800: 5798: 5794: 5790: 5785: 5769: 5765: 5761: 5749: 5745: 5741: 5739: 5736: 5734: 5730: 5728: 5723: 5721: 5717: 5714: 5713: 5709: 5703: 5699: 5695: 5691: 5686: 5681: 5677: 5673: 5669: 5664: 5663: 5659: 5642: 5638: 5637: 5629: 5622: 5619: 5614: 5610: 5605: 5600: 5596: 5592: 5588: 5586: 5582: 5573: 5570: 5557: 5553: 5549: 5544: 5539: 5535: 5531: 5527: 5520: 5517: 5504: 5500: 5496: 5491: 5486: 5482: 5478: 5474: 5470: 5469: 5464: 5462: 5458: 5449: 5447: 5443: 5430: 5426: 5422: 5415: 5412: 5407: 5403: 5398: 5393: 5389: 5385: 5380: 5375: 5371: 5367: 5363: 5359: 5355: 5348: 5345: 5340: 5336: 5332: 5328: 5324: 5320: 5315: 5310: 5306: 5302: 5301: 5296: 5294: 5290: 5286: 5277: 5275: 5271: 5268: 5264: 5261: 5257: 5253: 5247: 5244: 5239: 5235: 5231: 5227: 5222: 5217: 5213: 5209: 5208: 5207:Plant Disease 5203: 5201: 5197: 5188: 5185: 5180: 5176: 5171: 5166: 5162: 5158: 5153: 5148: 5144: 5140: 5136: 5132: 5128: 5126: 5117: 5114: 5109: 5105: 5101: 5097: 5093: 5089: 5085: 5081: 5074: 5071: 5066: 5062: 5058: 5054: 5049: 5044: 5040: 5036: 5032: 5025: 5022: 5010: 5006: 5002: 4998: 4994: 4990: 4986: 4982: 4978: 4976: 4967: 4964: 4959: 4955: 4950: 4945: 4941: 4937: 4933: 4931: 4922: 4919: 4906: 4902: 4898: 4891: 4888: 4875: 4871: 4867: 4860: 4858: 4856: 4852: 4840: 4836: 4832: 4828: 4824: 4820: 4819: 4811: 4808: 4796: 4792: 4785: 4782: 4776: 4771: 4767: 4763: 4759: 4757: 4748: 4745: 4732: 4728: 4724: 4719: 4714: 4709: 4704: 4700: 4696: 4692: 4690: 4681: 4678: 4665: 4661: 4654: 4651: 4638: 4634: 4630: 4624: 4621: 4608: 4604: 4602: 4597: 4591: 4588: 4575: 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3689: 3684: 3679: 3675: 3671: 3667: 3663: 3659: 3655: 3651: 3647: 3643: 3636: 3633: 3620: 3616: 3612: 3606: 3603: 3598: 3594: 3590: 3586: 3582: 3578: 3571: 3564: 3561: 3545: 3541: 3537: 3533: 3529: 3525: 3521: 3514: 3512: 3503: 3500: 3487: 3483: 3482: 3477: 3470: 3468: 3464: 3459: 3455: 3451: 3447: 3443: 3436: 3433: 3420: 3416: 3412: 3405: 3402: 3397: 3393: 3388: 3383: 3379: 3375: 3371: 3369: 3360: 3357: 3344: 3340: 3336: 3332: 3328: 3323: 3318: 3314: 3310: 3306: 3304: 3295: 3293: 3289: 3276: 3272: 3268: 3266: 3257: 3255: 3251: 3246: 3242: 3238: 3234: 3230: 3226: 3222: 3218: 3214: 3210: 3209: 3204: 3197: 3195: 3193: 3189: 3176: 3172: 3168: 3162: 3159: 3146: 3142: 3138: 3131: 3128: 3115: 3111: 3107: 3101: 3098: 3085: 3081: 3077: 3071: 3068: 3055: 3051: 3047: 3041: 3039: 3035: 3022: 3018: 3017: 3012: 3005: 3003: 2999: 2986: 2982: 2978: 2974: 2970: 2966: 2962: 2958: 2954: 2950: 2942: 2939: 2926: 2922: 2918: 2912: 2910: 2908: 2906: 2904: 2902: 2900: 2898: 2896: 2892: 2879: 2875: 2873: 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1420: 1418: 1416: 1412: 1411:L. delicatula 1408: 1404: 1400: 1396: 1392: 1384: 1382: 1380: 1376: 1372: 1367: 1360: 1358: 1356: 1352: 1348: 1343: 1341: 1337: 1332: 1330: 1326: 1322: 1318: 1317:West Virginia 1314: 1310: 1306: 1302: 1298: 1294: 1290: 1289:Massachusetts 1286: 1282: 1277: 1275: 1271: 1267: 1263: 1259: 1254: 1251: 1246: 1244: 1240: 1236: 1232: 1224: 1223: 1217: 1210: 1208: 1206: 1202: 1198: 1194: 1190: 1186: 1178: 1176: 1173: 1169: 1164: 1162: 1158: 1149: 1147: 1145: 1141: 1137: 1133: 1128: 1124: 1116: 1109: 1107: 1105: 1104:L. delicatula 1101: 1097: 1093: 1089: 1085: 1081: 1077: 1073: 1069: 1065: 1061: 1057: 1053: 1049: 1044: 1042: 1038: 1037:Yangtze River 1034: 1030: 1029:L. delicatula 1026: 1022: 1019:, during the 1018: 1014: 1013:L. delicatula 1010: 1002: 997: 995: 992: 988: 983: 982:overwintering 974: 970: 968: 964: 960: 955: 952: 948: 944: 939: 937: 933: 929: 925: 921: 917: 913: 908: 905: 896: 891: 884: 882: 880: 875: 870: 866: 863: 859: 855: 851: 847: 843: 839: 835: 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Oregon. 4344:Yahoo News 4152:Entomology 4131:28 October 4018:19 October 3866:13 January 3740:11 January 3625:24 January 3349:7 December 3281:26 January 3181:7 December 3151:7 December 3060:19 January 2884:26 January 2653:26 January 2618:26 January 2388:17 January 2327:17 January 2309:(1): 1–6. 2276:19 January 2223:(Report). 2169:25 January 2023:23 January 1993:23 January 1980:USDA-APHIS 1793:References 1641:biocontrol 1534:kairomones 1486:bifenthrin 1478:pyrethrins 1452:Penn State 1355:California 1297:New Jersey 1270:Penn State 885:Life cycle 546:delicatula 510:subspecies 475:Adam White 463:Aphaeninae 456:Fulgoridae 381:introduced 332:life cycle 301:including 281:invasively 277:indigenous 215:L. d. jole 138:Fulgoridae 114:Suborder: 88:Arthropoda 5812:Q10571341 5694:0066-4170 5613:1225-0171 5475:(1): 18. 5388:0027-8424 5339:244070660 5323:0022-2011 5230:0191-2917 5161:1932-6203 5108:234011992 5100:1612-4766 5057:0022-0493 5001:0046-225X 4958:1225-0171 4911:27 August 4901:fox43.com 4835:236227250 4670:8 October 4580:9 October 4468:0190-8286 4439:8 October 4375:2 October 4195:5 October 4105:9 October 4079:5 October 4048:5 October 3929:5 January 3856:0362-4331 3827:1226-8615 3787:2287-884X 3674:1752-4571 3492:5 October 3396:1225-0171 3331:0046-225X 3237:0892-7553 2973:0046-225X 2838:1225-0171 2785:0013-872X 2419:9 January 2231:9 January 2194:Zool Stud 2084:9 January 1538:spearmint 1385:In Europe 1361:In Canada 1256:In 2019, 1207:, China. 1100:poisonous 1072:Guangdong 646:valvifers 622:forewings 611:forewings 522:Taxonomic 502:Carl Stål 471:described 460:subfamily 156:Species: 108:Hemiptera 74:Kingdom: 68:Eukaryota 5955:2.943390 5849:BugGuide 5806:Wikidata 5702:36206772 5641:Archived 5556:Archived 5552:33301027 5503:Archived 5499:28069731 5429:Archived 5406:31010927 5331:34774856 5263:Archived 5238:30722582 5179:33939739 5131:PLOS ONE 5065:30753676 5009:31990325 4905:Archived 4839:archived 4731:Archived 4727:31603497 4664:Archived 4637:Archived 4607:Archived 4574:Archived 4541:Archived 4503:Archived 4472:Archived 4433:Archived 4429:31579914 4369:Archived 4332:Archived 4299:Archived 4260:Archived 4221:Archived 4189:Archived 4156:Archived 4125:Archived 4099:Archived 4073:Archived 4042:Archived 4012:Archived 3990:Archived 3953:Archived 3923:Archived 3890:Archived 3860:Archived 3734:Archived 3692:33897812 3619:Archived 3544:Archived 3540:33269378 3486:Archived 3343:Archived 3339:32725170 3275:Archived 3245:44841539 3175:Archived 3145:Archived 3114:Archived 3054:Archived 3021:Archived 2991:16 March 2985:Archived 2981:33128562 2878:Archived 2793:86233694 2744:34376093 2685:Archived 2681:phys.org 2647:Archived 2639:(White)" 2612:Archived 2591:29584780 2550:PLOS One 2515:Archived 2480:35105894 2413:Archived 2379:Archived 2375:86593102 2321:Archived 2267:Archived 2225:Archived 2163:Archived 2128:Archived 2078:Archived 2017:Archived 2015:. 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