3233:
125:
3239:
3244:
3265:
3251:
381:(inside branch) has seven segments, the first six of which are roughly even in size and shape, with small spines on the underside, and the last of which terminates with a claw and appears to have been used for walking and eating, as the claw could be useful in grabbing hold of and tearing up food. The exopod (outside branch) was not segmented, but flap-like, with
104:
357:, a piece of exoskeleton that covers part of the underside of the animal. Its doublure is, on average, about twice as long as the length of its cephalon. It has been hypothesized that this large doublure could have aided the animal in digging through sediment to find food, as it was thought to have been both a predator of small animals and a scavenger.
403:, which show that it was narrow annulated tube running along the midline of the animal, with three or possibly four pairs of midgut glands within the cephalic shield, with other gland pairs extending along the trunk.
363:
has one pair of antennae attached to its cephalon, but no other appendages or auditory or visual structures. No mouth is present in the specimens, but the gut is present, so
2181:
2371:
Berks, Harry O.; Lunde
Nielsen, Morten; Flannery‐Sutherland, Joseph; Thorshøj Nielsen, Arne; Park, Tae‐Yoon S.; Vinther, Jakob (May 2023).
399:
though the differences between the two species are relatively minor. The species also preserves elements of the gut tract not preserved in
2430:
2157:
3232:
2125:
2272:"Redescription of the Chengjiang arthropod Squamacula clypeata Hou and Bergström, from the Lower Cambrian, south-west China"
2203:
2373:"A possibly deep branching artiopodan arthropod from the lower Cambrian Sirius Passet Lagerstätte (North Greenland)"
124:
2185:
338:
3243:
2423:
596:
2735:
2204:"New artiopodan arthropods from the early Cambrian Emu Bay Shale Konservat-Lagerstätte of South Australia"
1502:
367:
must have had a mouth, which is thought to have been between the cephalon and doublure on the underside.
3238:
3250:
2742:
2335:
2283:
2215:
2787:
2568:
1622:
504:
3208:
2815:
2766:
1557:
3142:
2978:
2870:
2822:
2758:
2689:
1927:
1910:
1886:
1803:
1605:
1581:
920:
735:
3289:
3268:
3151:
2884:
2863:
2794:
2675:
2660:
2552:
2416:
2239:
2175:
1862:
1533:
749:
465:
119:
2898:
2842:
2750:
2682:
1755:
2967:
2808:
2612:
1731:
1475:
3179:
2905:
2891:
2877:
2394:
2353:
2301:
2231:
2163:
2153:
2131:
2121:
1779:
1714:
847:
308:
265:
3022:
2696:
1392:
2778:
2668:
2384:
2343:
2291:
2223:
1409:
273:
2925:
2728:
2561:
1690:
609:
554:
537:
2324:"A new small soft-bodied non-trilobite artiopod from the Cambrian Stage 4 Guanshan Biota"
2339:
2287:
2219:
395:
in that it has 11 thoracic tergites, which appear to exhibit less overlap that those of
3001:
2953:
2591:
2512:
1302:
1040:
385:(bristles) near the tip, which may have been used for both locomotion and respiration.
2322:
Jiao, De-Guang; Du, Kun-Sheng; Zhang, Xi-Guang; Yang, Jie; Eggink, Daniel (May 2022).
3283:
2994:
2644:
2466:
2296:
2271:
2093:
1354:
975:
958:
285:
61:
2243:
415:
has consistently been placed as the earliest diverging and most primitive member of
326:, meaning shield-shaped. It was named this in reference to its shield-like outline.
276:
of Yunnan, China. At the time of description there were only two known specimens of
3042:
2946:
2833:
2653:
2150:
The
Cambrian fossils of Chengjiang, China : the flowering of early animal life
1368:
662:
195:
2202:
Paterson, John R.; García-Bellido, Diego C.; Edgecombe, Gregory D. (March 2012).
2939:
2856:
2720:
2707:
2605:
1488:
1341:
704:
36:
3078:
3069:
3058:
3033:
3015:
3008:
2987:
2960:
2801:
2348:
2323:
1673:
1235:
1089:
1023:
999:
934:
624:
81:
46:
2398:
2357:
2305:
2235:
2167:
3197:
3186:
3127:
3120:
3105:
2633:
2619:
2533:
2505:
2487:
2472:
2457:
2439:
2152:. Hou, Xianguang. (Second ed.). Chichester, West Sussex. 8 March 2017.
2135:
2088:
1451:
1144:
1127:
1103:
830:
687:
420:
416:
377:
appendages (an appendage which branches into two) per thoracic tergite. The
354:
261:
258:
169:
156:
136:
86:
2270:
Zhang, Xingliang; Han, Jian; Zhang, Zhifei; Liu, Huqin; Shu, Degan (2004).
280:, but now there are at least six known specimens. In 2012 a second species
3172:
3158:
3096:
3051:
2849:
2626:
2575:
2540:
1820:
1317:
864:
797:
487:
374:
350:
76:
71:
56:
51:
41:
103:
3165:
3085:
2519:
1218:
1194:
780:
378:
342:
316:
91:
66:
2389:
2372:
2451:
2227:
346:
146:
3113:
2118:
Arthropods of the Lower
Cambrian Chengjiang fauna, southwest China
1161:
297:
382:
337:
is flattened (dorsoventrally). It has 11 segments in total: the
2412:
423:
and a variety of morphologically similar soft-bodied forms.
2408:
2086:
A close relationship has been proposed with the genus
315:, indicating that the animal is relatively small. The
236:
225:
202:
3196:
3141:
3095:
3068:
3041:
3032:
2977:
2923:
2832:
2777:
2706:
2643:
2589:
2551:
2485:
2120:. Bergström. Oslo: Scandinavian University Press.
2424:
8:
3038:
2931:
2712:
2597:
2497:
2493:
2482:
2431:
2417:
2409:
2180:: CS1 maint: location missing publisher (
102:
20:
2388:
2347:
2295:
2116:Hou, Xianguang; Bergström, Jan (1997).
2105:
2317:
2315:
2173:
7:
2265:
2263:
2261:
2259:
2257:
2255:
2253:
2197:
2195:
2111:
2109:
1902:
1878:
1854:
1795:
1771:
1747:
1706:
1665:
1597:
1573:
1549:
1525:
1518:
1494:
1467:
1443:
1384:
1360:
1333:
1309:
1210:
1186:
1119:
1095:
1015:
991:
950:
926:
916:
822:
772:
765:
741:
679:
654:
647:
640:
616:
588:
529:
479:
472:
461:
437:
429:
426:Cladogram after Jiao et al. 2021.
14:
3264:
3263:
3249:
3242:
3237:
3231:
2297:10.1111/j.0031-0239.2004.00363.x
123:
322:is derived from the Latin word
303:is derived from the Latin word
272:was described in 1997 from the
1:
1911:Glypharthrus trispinicaudatus
663:Kwanyinaspis maotiashanensis
16:Extinct genus of arthropods
3306:
1691:Australaglaspis stonyensis
555:Acanthomeridion anacanthus
411:In phylogenetic analyses,
341:(the head), nine thoracic
3259:
3229:
2934:
2715:
2600:
2500:
2496:
2481:
2446:
2349:10.1017/S0016756821001254
1924:
1907:
1900:
1883:
1876:
1859:
1852:
1817:
1800:
1793:
1776:
1769:
1752:
1745:
1728:
1711:
1704:
1687:
1670:
1663:
1623:Brachyaglaspis singularis
1619:
1602:
1595:
1578:
1571:
1554:
1547:
1530:
1523:
1516:
1499:
1492:
1472:
1465:
1448:
1441:
1406:
1389:
1382:
1365:
1358:
1338:
1331:
1314:
1307:
1232:
1215:
1208:
1191:
1184:
1158:
1141:
1124:
1117:
1100:
1093:
1037:
1020:
1013:
996:
989:
972:
955:
948:
931:
924:
914:
861:
844:
827:
820:
794:
777:
770:
763:
746:
739:
701:
684:
677:
659:
652:
645:
638:
621:
614:
593:
586:
551:
534:
527:
501:
484:
477:
470:
459:
442:
435:
307:, meaning scale, and the
222:
217:
201:
194:
120:Scientific classification
118:
110:
101:
23:
1041:Phytophilaspis pergamena
538:Acanthomeridion serratum
419:, a clade that includes
345:(each of which covers a
2377:Papers in Palaeontology
2208:Journal of Paleontology
1928:Glypharthrus magnoculus
1236:Misszhouia longicaudata
284:was described from the
232:Hou and Bergström, 1997
212:Hou and Bergström, 1997
189:Hou and Bergström, 1997
1756:Aglaspella granulifera
1558:Quasimodaspis brentsae
1024:Xandarella spectaculum
959:Luohuilinella deletres
705:Eoredlichia intermedia
597:Bailongia longicaudata
505:Australimicola spriggi
2184:) CS1 maint: others (
625:Retifacies abnormalis
1887:Glypharthrus thomasi
1804:Glypharthrus simplex
1606:Tremaglaspis vanroyi
1393:Triopus draboviensis
1369:Cheloniellon calmani
1318:Sidneyia inexpectans
1128:Tariccoia arrusensis
243:Paterson et al. 2012
2340:2022GeoM..159..730J
2328:Geological Magazine
2288:2004Palgy..47..605Z
2220:2012JPal...86..340P
2096:site of Greenland.
1863:Chraspedops modesta
1715:Uarthrus instabilis
1674:Cyclopites vulgaris
1145:Soomaspis splendida
976:Luohuilinella rarus
446:Squamacula clypeata
371:Squamacula clypeata
361:Squamacula clypeata
335:Squamacula clypeata
268:. The type species
240:Squamacula buckorum
229:Squamacula clypeata
207:Squamacula clypeata
113:Squamacula clypeata
1582:Tremaglaspis unite
1476:Eozetetes gemmelli
1342:Emeraldella brocki
1000:Sinoburius lunaris
935:Cindarella eucalla
865:Saperion glumaceum
750:Haifengella corona
688:Olenoides serratus
111:Reconstruction of
3277:
3276:
3227:
3226:
3223:
3222:
3219:
3218:
3137:
3136:
2919:
2918:
2915:
2914:
2585:
2584:
2390:10.1002/spp2.1495
2083:
2082:
2074:
2073:
2065:
2064:
2056:
2055:
2047:
2046:
2038:
2037:
2029:
2028:
2020:
2019:
2011:
2010:
2002:
2001:
1993:
1992:
1984:
1983:
1975:
1974:
1966:
1965:
1957:
1956:
1948:
1947:
1939:
1938:
1841:
1840:
1832:
1831:
1821:Aglaspis spinifer
1652:
1651:
1643:
1642:
1634:
1633:
1534:Chlupacaris dubia
1430:
1429:
1421:
1420:
1292:
1291:
1283:
1282:
1274:
1273:
1265:
1264:
1256:
1255:
1247:
1246:
1173:
1172:
1104:Buenaspis forteyi
1079:
1078:
1070:
1069:
1061:
1060:
1052:
1051:
903:
902:
894:
893:
885:
884:
876:
875:
809:
808:
725:
724:
716:
715:
575:
574:
566:
565:
516:
515:
488:Zhiwenia coronata
309:diminutive suffix
266:Cambrian Series 2
250:
249:
244:
233:
190:
32:Cambrian Series 2
3297:
3267:
3266:
3254:
3253:
3246:
3241:
3235:
3039:
2932:
2779:Tremaglaspididae
2713:
2598:
2498:
2494:
2483:
2433:
2426:
2419:
2410:
2403:
2402:
2392:
2368:
2362:
2361:
2351:
2319:
2310:
2309:
2299:
2267:
2248:
2247:
2228:10.1666/11-077.1
2199:
2190:
2189:
2179:
2171:
2146:
2140:
2139:
2113:
1903:
1879:
1855:
1796:
1780:Gogglops ensifer
1772:
1748:
1707:
1666:
1598:
1574:
1550:
1526:
1519:
1495:
1468:
1452:Kodymirus vagans
1444:
1385:
1361:
1334:
1310:
1219:Naraoia compacta
1211:
1187:
1120:
1096:
1016:
992:
951:
927:
917:
823:
798:Helmetia expansa
773:
766:
742:
680:
655:
648:
641:
617:
589:
530:
480:
473:
462:
438:
433:Artiopoda
430:
373:has one pair of
274:Chengjiang biota
242:
238:
231:
227:
204:
188:
181:
168:
128:
127:
106:
96:
33:
29:Temporal range:
21:
3305:
3304:
3300:
3299:
3298:
3296:
3295:
3294:
3280:
3279:
3278:
3273:
3255:
3248:
3215:
3192:
3133:
3091:
3064:
3028:
2973:
2928:
2926:Trilobitomorpha
2911:
2828:
2773:
2743:Chacharejocaris
2729:Australaglaspis
2702:
2639:
2594:
2581:
2562:Acanthomeridion
2547:
2490:
2477:
2442:
2437:
2407:
2406:
2370:
2369:
2365:
2321:
2320:
2313:
2269:
2268:
2251:
2201:
2200:
2193:
2172:
2160:
2148:
2147:
2143:
2128:
2115:
2114:
2107:
2102:
2084:
2075:
2066:
2057:
2048:
2039:
2030:
2021:
2012:
2003:
1994:
1985:
1976:
1967:
1958:
1949:
1940:
1842:
1833:
1732:Flobertia kochi
1653:
1644:
1635:
1503:Beckwithia typa
1431:
1422:
1410:Duslia insignis
1293:
1284:
1275:
1266:
1257:
1248:
1195:Naraoia spinosa
1174:
1080:
1071:
1062:
1053:
904:
895:
886:
877:
831:Tegopelte gigas
810:
726:
717:
610:Trilobitomorpha
576:
567:
517:
409:
332:
294:
213:
210:
187:
179:
166:
122:
97:
95:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
31:
30:
27:
17:
12:
11:
5:
3303:
3301:
3293:
3292:
3282:
3281:
3275:
3274:
3272:
3271:
3260:
3257:
3256:
3230:
3228:
3225:
3224:
3221:
3220:
3217:
3216:
3214:
3213:
3212:
3211:
3202:
3200:
3194:
3193:
3191:
3190:
3183:
3176:
3169:
3162:
3155:
3147:
3145:
3139:
3138:
3135:
3134:
3132:
3131:
3124:
3117:
3110:
3101:
3099:
3093:
3092:
3090:
3089:
3082:
3074:
3072:
3066:
3065:
3063:
3062:
3055:
3047:
3045:
3036:
3030:
3029:
3027:
3026:
3019:
3012:
3005:
3002:Phytophilaspis
2998:
2991:
2983:
2981:
2975:
2974:
2972:
2971:
2964:
2957:
2954:Pygmaclypeatus
2950:
2943:
2935:
2929:
2924:
2921:
2920:
2917:
2916:
2913:
2912:
2910:
2909:
2902:
2895:
2888:
2881:
2874:
2867:
2860:
2853:
2846:
2838:
2836:
2830:
2829:
2827:
2826:
2819:
2812:
2805:
2798:
2791:
2788:Brachyaglaspis
2783:
2781:
2775:
2774:
2772:
2771:
2763:
2755:
2747:
2739:
2732:
2725:
2716:
2710:
2704:
2703:
2701:
2700:
2693:
2686:
2679:
2672:
2665:
2657:
2649:
2647:
2641:
2640:
2638:
2637:
2630:
2623:
2616:
2609:
2601:
2595:
2592:Vicissicaudata
2590:
2587:
2586:
2583:
2582:
2580:
2579:
2572:
2569:Australimicola
2565:
2557:
2555:
2549:
2548:
2546:
2545:
2537:
2530:
2523:
2516:
2513:Falcatamacaris
2509:
2501:
2491:
2486:
2479:
2478:
2476:
2475:
2469:
2460:
2454:
2447:
2444:
2443:
2438:
2436:
2435:
2428:
2421:
2413:
2405:
2404:
2363:
2334:(5): 730–734.
2311:
2282:(3): 605–617.
2249:
2214:(2): 340–357.
2191:
2158:
2141:
2126:
2104:
2103:
2101:
2098:
2081:
2080:
2077:
2076:
2072:
2071:
2068:
2067:
2063:
2062:
2059:
2058:
2054:
2053:
2050:
2049:
2045:
2044:
2041:
2040:
2036:
2035:
2032:
2031:
2027:
2026:
2023:
2022:
2018:
2017:
2014:
2013:
2009:
2008:
2005:
2004:
2000:
1999:
1996:
1995:
1991:
1990:
1987:
1986:
1982:
1981:
1978:
1977:
1973:
1972:
1969:
1968:
1964:
1963:
1960:
1959:
1955:
1954:
1951:
1950:
1946:
1945:
1942:
1941:
1937:
1936:
1933:
1932:
1923:
1920:
1919:
1916:
1915:
1906:
1901:
1899:
1896:
1895:
1892:
1891:
1882:
1877:
1875:
1872:
1871:
1868:
1867:
1858:
1853:
1851:
1848:
1847:
1844:
1843:
1839:
1838:
1835:
1834:
1830:
1829:
1826:
1825:
1816:
1813:
1812:
1809:
1808:
1799:
1794:
1792:
1789:
1788:
1785:
1784:
1775:
1770:
1768:
1765:
1764:
1761:
1760:
1751:
1746:
1744:
1741:
1740:
1737:
1736:
1727:
1724:
1723:
1720:
1719:
1710:
1705:
1703:
1700:
1699:
1696:
1695:
1686:
1683:
1682:
1679:
1678:
1669:
1664:
1662:
1659:
1658:
1655:
1654:
1650:
1649:
1646:
1645:
1641:
1640:
1637:
1636:
1632:
1631:
1628:
1627:
1618:
1615:
1614:
1611:
1610:
1601:
1596:
1594:
1591:
1590:
1587:
1586:
1577:
1572:
1570:
1567:
1566:
1563:
1562:
1553:
1548:
1546:
1543:
1542:
1539:
1538:
1529:
1524:
1522:
1517:
1515:
1512:
1511:
1508:
1507:
1498:
1493:
1491:
1485:
1484:
1481:
1480:
1471:
1466:
1464:
1461:
1460:
1457:
1456:
1447:
1442:
1440:
1437:
1436:
1433:
1432:
1428:
1427:
1424:
1423:
1419:
1418:
1415:
1414:
1405:
1402:
1401:
1398:
1397:
1388:
1383:
1381:
1378:
1377:
1374:
1373:
1364:
1359:
1357:
1351:
1350:
1347:
1346:
1337:
1332:
1330:
1327:
1326:
1323:
1322:
1313:
1308:
1306:
1303:Vicissicaudata
1299:
1298:
1295:
1294:
1290:
1289:
1286:
1285:
1281:
1280:
1277:
1276:
1272:
1271:
1268:
1267:
1263:
1262:
1259:
1258:
1254:
1253:
1250:
1249:
1245:
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1240:
1231:
1228:
1227:
1224:
1223:
1214:
1209:
1207:
1204:
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1200:
1199:
1190:
1185:
1183:
1180:
1179:
1176:
1175:
1171:
1170:
1167:
1166:
1157:
1154:
1153:
1150:
1149:
1140:
1137:
1136:
1133:
1132:
1123:
1118:
1116:
1113:
1112:
1109:
1108:
1099:
1094:
1092:
1086:
1085:
1082:
1081:
1077:
1076:
1073:
1072:
1068:
1067:
1064:
1063:
1059:
1058:
1055:
1054:
1050:
1049:
1046:
1045:
1036:
1033:
1032:
1029:
1028:
1019:
1014:
1012:
1009:
1008:
1005:
1004:
995:
990:
988:
985:
984:
981:
980:
971:
968:
967:
964:
963:
954:
949:
947:
944:
943:
940:
939:
930:
925:
923:
915:
913:
910:
909:
906:
905:
901:
900:
897:
896:
892:
891:
888:
887:
883:
882:
879:
878:
874:
873:
870:
869:
860:
857:
856:
853:
852:
848:Skioldia aldna
843:
840:
839:
836:
835:
826:
821:
819:
816:
815:
812:
811:
807:
806:
803:
802:
793:
790:
789:
786:
785:
776:
771:
769:
764:
762:
759:
758:
755:
754:
745:
740:
738:
732:
731:
728:
727:
723:
722:
719:
718:
714:
713:
710:
709:
700:
697:
696:
693:
692:
683:
678:
676:
672:
671:
668:
667:
658:
653:
651:
646:
644:
639:
637:
634:
633:
630:
629:
620:
615:
613:
606:
605:
602:
601:
592:
587:
585:
582:
581:
578:
577:
573:
572:
569:
568:
564:
563:
560:
559:
550:
547:
546:
543:
542:
533:
528:
526:
523:
522:
519:
518:
514:
513:
510:
509:
500:
497:
496:
493:
492:
483:
478:
476:
471:
469:
460:
458:
455:
454:
451:
450:
441:
436:
434:
428:
408:
405:
331:
328:
293:
290:
288:of Australia.
257:is an extinct
248:
247:
246:
245:
234:
220:
219:
215:
214:
211:
199:
198:
192:
191:
177:
173:
172:
164:
160:
159:
154:
150:
149:
144:
140:
139:
134:
130:
129:
116:
115:
108:
107:
99:
98:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
35:
34:
28:
15:
13:
10:
9:
6:
4:
3:
2:
3302:
3291:
3288:
3287:
3285:
3270:
3262:
3261:
3258:
3252:
3247:
3245:
3240:
3234:
3210:
3206:
3205:
3204:
3203:
3201:
3199:
3195:
3189:
3188:
3184:
3182:
3181:
3177:
3175:
3174:
3170:
3168:
3167:
3163:
3161:
3160:
3156:
3154:
3153:
3149:
3148:
3146:
3144:
3140:
3130:
3129:
3125:
3123:
3122:
3118:
3116:
3115:
3111:
3108:
3107:
3103:
3102:
3100:
3098:
3094:
3088:
3087:
3083:
3081:
3080:
3076:
3075:
3073:
3071:
3067:
3061:
3060:
3056:
3054:
3053:
3049:
3048:
3046:
3044:
3040:
3037:
3035:
3031:
3025:
3024:
3020:
3018:
3017:
3013:
3011:
3010:
3006:
3004:
3003:
2999:
2997:
2996:
2995:Luohuilinella
2992:
2990:
2989:
2985:
2984:
2982:
2980:
2976:
2970:
2969:
2965:
2963:
2962:
2958:
2956:
2955:
2951:
2949:
2948:
2944:
2942:
2941:
2937:
2936:
2933:
2930:
2927:
2922:
2908:
2907:
2903:
2901:
2900:
2896:
2894:
2893:
2889:
2887:
2886:
2882:
2880:
2879:
2875:
2873:
2872:
2868:
2866:
2865:
2861:
2859:
2858:
2854:
2852:
2851:
2847:
2845:
2844:
2840:
2839:
2837:
2835:
2831:
2825:
2824:
2820:
2818:
2817:
2816:Quasimodaspis
2813:
2811:
2810:
2806:
2804:
2803:
2799:
2797:
2796:
2792:
2790:
2789:
2785:
2784:
2782:
2780:
2776:
2769:
2768:
2767:Rozhkovocaris
2764:
2761:
2760:
2756:
2753:
2752:
2748:
2745:
2744:
2740:
2738:
2737:
2733:
2731:
2730:
2726:
2723:
2722:
2718:
2717:
2714:
2711:
2709:
2705:
2699:
2698:
2694:
2692:
2691:
2687:
2685:
2684:
2680:
2678:
2677:
2673:
2671:
2670:
2666:
2663:
2662:
2658:
2656:
2655:
2651:
2650:
2648:
2646:
2645:Cheloniellida
2642:
2636:
2635:
2631:
2629:
2628:
2624:
2622:
2621:
2617:
2615:
2614:
2610:
2608:
2607:
2603:
2602:
2599:
2596:
2593:
2588:
2578:
2577:
2573:
2571:
2570:
2566:
2564:
2563:
2559:
2558:
2556:
2554:
2550:
2543:
2542:
2538:
2536:
2535:
2531:
2529:
2528:
2524:
2522:
2521:
2517:
2515:
2514:
2510:
2508:
2507:
2503:
2502:
2499:
2495:
2492:
2489:
2484:
2480:
2474:
2470:
2468:
2467:Panarthropoda
2464:
2461:
2459:
2456:Superphylum:
2455:
2453:
2449:
2448:
2445:
2441:
2434:
2429:
2427:
2422:
2420:
2415:
2414:
2411:
2400:
2396:
2391:
2386:
2382:
2378:
2374:
2367:
2364:
2359:
2355:
2350:
2345:
2341:
2337:
2333:
2329:
2325:
2318:
2316:
2312:
2307:
2303:
2298:
2293:
2289:
2285:
2281:
2277:
2276:Palaeontology
2273:
2266:
2264:
2262:
2260:
2258:
2256:
2254:
2250:
2245:
2241:
2237:
2233:
2229:
2225:
2221:
2217:
2213:
2209:
2205:
2198:
2196:
2192:
2187:
2183:
2177:
2169:
2165:
2161:
2159:9781118896310
2155:
2151:
2145:
2142:
2137:
2133:
2129:
2123:
2119:
2112:
2110:
2106:
2099:
2097:
2095:
2094:Sirius Passet
2091:
2090:
2079:
2078:
2070:
2069:
2061:
2060:
2052:
2051:
2043:
2042:
2034:
2033:
2025:
2024:
2016:
2015:
2007:
2006:
1998:
1997:
1989:
1988:
1980:
1979:
1971:
1970:
1962:
1961:
1953:
1952:
1944:
1943:
1935:
1934:
1931:
1930:
1929:
1922:
1921:
1918:
1917:
1914:
1913:
1912:
1905:
1904:
1898:
1897:
1894:
1893:
1890:
1889:
1888:
1881:
1880:
1874:
1873:
1870:
1869:
1866:
1865:
1864:
1857:
1856:
1850:
1849:
1846:
1845:
1837:
1836:
1828:
1827:
1824:
1823:
1822:
1815:
1814:
1811:
1810:
1807:
1806:
1805:
1798:
1797:
1791:
1790:
1787:
1786:
1783:
1782:
1781:
1774:
1773:
1767:
1766:
1763:
1762:
1759:
1758:
1757:
1750:
1749:
1743:
1742:
1739:
1738:
1735:
1734:
1733:
1726:
1725:
1722:
1721:
1718:
1717:
1716:
1709:
1708:
1702:
1701:
1698:
1697:
1694:
1693:
1692:
1685:
1684:
1681:
1680:
1677:
1676:
1675:
1668:
1667:
1661:
1660:
1657:
1656:
1648:
1647:
1639:
1638:
1630:
1629:
1626:
1625:
1624:
1617:
1616:
1613:
1612:
1609:
1608:
1607:
1600:
1599:
1593:
1592:
1589:
1588:
1585:
1584:
1583:
1576:
1575:
1569:
1568:
1565:
1564:
1561:
1560:
1559:
1552:
1551:
1545:
1544:
1541:
1540:
1537:
1536:
1535:
1528:
1527:
1521:
1520:
1514:
1513:
1510:
1509:
1506:
1505:
1504:
1497:
1496:
1490:
1487:
1486:
1483:
1482:
1479:
1478:
1477:
1470:
1469:
1463:
1462:
1459:
1458:
1455:
1454:
1453:
1446:
1445:
1439:
1438:
1435:
1434:
1426:
1425:
1417:
1416:
1413:
1412:
1411:
1404:
1403:
1400:
1399:
1396:
1395:
1394:
1387:
1386:
1380:
1379:
1376:
1375:
1372:
1371:
1370:
1363:
1362:
1356:
1355:Cheloniellida
1353:
1352:
1349:
1348:
1345:
1344:
1343:
1336:
1335:
1329:
1328:
1325:
1324:
1321:
1320:
1319:
1312:
1311:
1304:
1301:
1300:
1297:
1296:
1288:
1287:
1279:
1278:
1270:
1269:
1261:
1260:
1252:
1251:
1243:
1242:
1239:
1238:
1237:
1230:
1229:
1226:
1225:
1222:
1221:
1220:
1213:
1212:
1206:
1205:
1202:
1201:
1198:
1197:
1196:
1189:
1188:
1182:
1181:
1178:
1177:
1169:
1168:
1165:
1164:
1163:
1162:Liwia convexa
1156:
1155:
1152:
1151:
1148:
1147:
1146:
1139:
1138:
1135:
1134:
1131:
1130:
1129:
1122:
1121:
1115:
1114:
1111:
1110:
1107:
1106:
1105:
1098:
1097:
1091:
1088:
1087:
1084:
1083:
1075:
1074:
1066:
1065:
1057:
1056:
1048:
1047:
1044:
1043:
1042:
1035:
1034:
1031:
1030:
1027:
1026:
1025:
1018:
1017:
1011:
1010:
1007:
1006:
1003:
1002:
1001:
994:
993:
987:
986:
983:
982:
979:
978:
977:
970:
969:
966:
965:
962:
961:
960:
953:
952:
946:
945:
942:
941:
938:
937:
936:
929:
928:
922:
919:
918:
912:
911:
908:
907:
899:
898:
890:
889:
881:
880:
872:
871:
868:
867:
866:
859:
858:
855:
854:
851:
850:
849:
842:
841:
838:
837:
834:
833:
832:
825:
824:
818:
817:
814:
813:
805:
804:
801:
800:
799:
792:
791:
788:
787:
784:
783:
782:
775:
774:
768:
767:
761:
760:
757:
756:
753:
752:
751:
744:
743:
737:
734:
733:
730:
729:
721:
720:
712:
711:
708:
707:
706:
699:
698:
695:
694:
691:
690:
689:
682:
681:
674:
673:
670:
669:
666:
665:
664:
657:
656:
650:
649:
643:
642:
636:
635:
632:
631:
628:
627:
626:
619:
618:
611:
608:
607:
604:
603:
600:
599:
598:
591:
590:
584:
583:
580:
579:
571:
570:
562:
561:
558:
557:
556:
549:
548:
545:
544:
541:
540:
539:
532:
531:
525:
524:
521:
520:
512:
511:
508:
507:
506:
499:
498:
495:
494:
491:
490:
489:
482:
481:
475:
474:
467:
464:
463:
457:
456:
453:
452:
449:
448:
447:
440:
439:
432:
431:
427:
424:
422:
418:
414:
406:
404:
402:
398:
394:
391:differs from
390:
386:
384:
380:
376:
372:
368:
366:
362:
358:
356:
352:
348:
344:
340:
336:
329:
327:
325:
321:
318:
314:
310:
306:
302:
299:
291:
289:
287:
286:Emu Bay Shale
283:
279:
275:
271:
267:
263:
260:
256:
255:
241:
235:
230:
224:
223:
221:
216:
209:
208:
200:
197:
193:
186:
185:
178:
175:
174:
171:
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
138:
135:
132:
131:
126:
121:
117:
114:
109:
105:
100:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
26:
22:
19:
3236:
3185:
3178:
3171:
3164:
3157:
3150:
3143:Conciliterga
3126:
3119:
3112:
3104:
3084:
3077:
3057:
3050:
3043:Emucarididae
3021:
3014:
3007:
3000:
2993:
2986:
2979:Xandarellida
2966:
2959:
2952:
2947:Kwanyinaspis
2945:
2938:
2904:
2897:
2890:
2883:
2876:
2871:Glypharthrus
2869:
2862:
2855:
2848:
2841:
2834:Aglaspididae
2823:Tremaglaspis
2821:
2814:
2807:
2800:
2793:
2786:
2765:
2759:Obrutschewia
2757:
2749:
2741:
2734:
2727:
2719:
2695:
2690:Pseudarthron
2688:
2681:
2674:
2667:
2659:
2654:Cheloniellon
2652:
2632:
2625:
2618:
2611:
2604:
2574:
2567:
2560:
2539:
2532:
2526:
2525:
2518:
2511:
2504:
2462:
2380:
2376:
2366:
2331:
2327:
2279:
2275:
2211:
2207:
2149:
2144:
2117:
2087:
2085:
1926:
1925:
1909:
1908:
1885:
1884:
1861:
1860:
1819:
1818:
1802:
1801:
1778:
1777:
1754:
1753:
1730:
1729:
1713:
1712:
1689:
1688:
1672:
1671:
1621:
1620:
1604:
1603:
1580:
1579:
1556:
1555:
1532:
1531:
1501:
1500:
1474:
1473:
1450:
1449:
1408:
1407:
1391:
1390:
1367:
1366:
1340:
1339:
1316:
1315:
1234:
1233:
1217:
1216:
1193:
1192:
1160:
1159:
1143:
1142:
1126:
1125:
1102:
1101:
1039:
1038:
1022:
1021:
998:
997:
974:
973:
957:
956:
933:
932:
921:Xandarellida
863:
862:
846:
845:
829:
828:
796:
795:
781:Kuamaia lata
779:
778:
748:
747:
736:Conciliterga
703:
702:
686:
685:
661:
660:
623:
622:
595:
594:
553:
552:
536:
535:
503:
502:
486:
485:
445:
444:
443:
425:
412:
410:
400:
397:S. clypeata,
396:
392:
388:
387:
370:
369:
364:
360:
359:
334:
333:
323:
319:
312:
304:
300:
295:
281:
277:
269:
253:
252:
251:
239:
228:
206:
205:
196:Type species
183:
182:
163:(unranked):
112:
24:
18:
3152:Haifengella
2940:Arthroaspis
2885:Hesselbonia
2864:Chraspedops
2857:Aglaspoides
2795:Chlupacaris
2721:Angarocaris
2708:Aglaspidida
2676:Neostrabops
2661:Drabovaspis
2606:Emeraldella
2553:Protosutura
1489:Aglaspidida
466:Protosutura
401:S. clypeata
393:S. clypeata
389:S. buckorum
365:S. clypeata
353:. It has a
349:), and one
330:Description
282:S. buckorum
278:S. clypeata
270:S. clypeata
3079:Misszhouia
3070:Naraoiidae
3059:Kangacaris
3034:Nektaspida
3016:Xandarella
3009:Sinoburius
2988:Cindarella
2961:Retifacies
2899:Tuboculops
2843:Aglaspella
2802:Cyclopites
2751:Girardevia
2736:Beckwithia
2683:Paraduslia
2527:Squamacula
2473:Arthropoda
2127:8200376931
2100:References
1090:Nektaspida
675:Trilobita
421:trilobites
413:Squamacula
301:Squamacula
259:artiopodan
254:Squamacula
184:Squamacula
157:Arthropoda
25:Squamacula
3290:Artiopoda
3209:Trilobita
3198:Trilobita
3187:Tegopelte
3128:Tariccoia
3121:Soomaspis
3106:Buenaspis
2968:Tonglaiia
2809:Flobertia
2634:Carimersa
2620:Kodymirus
2613:Eozetetes
2534:Thulaspis
2506:Bailongia
2488:Artiopoda
2458:Ecdysozoa
2450:Kingdom:
2440:Artiopoda
2399:2056-2799
2358:0016-7568
2306:1475-4983
2236:0022-3360
2176:cite book
2168:970396735
2092:from the
2089:Thulaspis
417:Artiopoda
324:clypeatus
292:Etymology
264:from the
262:arthropod
170:Artiopoda
143:Kingdom:
137:Eukaryota
3284:Category
3269:Category
3180:Skioldia
3173:Saperion
3159:Helmetia
3097:Liwiidae
3052:Emucaris
2906:Uarthrus
2892:Setaspis
2878:Gogglops
2850:Aglaspis
2627:Sidneyia
2576:Zhiwenia
2541:Urokodia
2471:Phylum:
2452:Animalia
2244:87520649
2136:38305908
407:Taxonomy
375:biramous
355:doublure
351:pygidium
343:tergites
339:cephalon
320:clypeata
218:Species
153:Phylum:
147:Animalia
133:Domain:
3166:Kuamaia
3086:Naraoia
3023:Zhugeia
2697:Triopus
2520:Molaria
2336:Bibcode
2284:Bibcode
2216:Bibcode
1305:
612:
468:
379:endopod
317:species
176:Genus:
2669:Duslia
2397:
2356:
2304:
2242:
2234:
2166:
2156:
2134:
2124:
347:somite
305:squama
3114:Liwia
2463:Clade
2383:(3).
2240:S2CID
383:setae
313:culus
298:genus
3207:see
2395:ISSN
2354:ISSN
2302:ISSN
2232:ISSN
2186:link
2182:link
2164:OCLC
2154:ISBN
2132:OCLC
2122:ISBN
296:The
37:PreꞒ
2385:doi
2344:doi
2332:159
2292:doi
2224:doi
3286::
2465::
2393:.
2379:.
2375:.
2352:.
2342:.
2330:.
2326:.
2314:^
2300:.
2290:.
2280:47
2278:.
2274:.
2252:^
2238:.
2230:.
2222:.
2212:86
2210:.
2206:.
2194:^
2178:}}
2174:{{
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2108:^
87:Pg
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2770:?
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2746:?
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2432:e
2425:t
2418:v
2401:.
2387::
2381:9
2360:.
2346::
2338::
2308:.
2294::
2286::
2246:.
2226::
2218::
2188:)
2170:.
2138:.
311:-
237:†
226:†
203:†
180:†
167:†
92:N
82:K
77:J
72:T
67:P
62:C
57:D
52:S
47:O
42:Ꞓ
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