3244:
136:
3250:
3255:
3276:
3262:
392:(inside branch) has seven segments, the first six of which are roughly even in size and shape, with small spines on the underside, and the last of which terminates with a claw and appears to have been used for walking and eating, as the claw could be useful in grabbing hold of and tearing up food. The exopod (outside branch) was not segmented, but flap-like, with
115:
368:, a piece of exoskeleton that covers part of the underside of the animal. Its doublure is, on average, about twice as long as the length of its cephalon. It has been hypothesized that this large doublure could have aided the animal in digging through sediment to find food, as it was thought to have been both a predator of small animals and a scavenger.
414:, which show that it was narrow annulated tube running along the midline of the animal, with three or possibly four pairs of midgut glands within the cephalic shield, with other gland pairs extending along the trunk.
374:
has one pair of antennae attached to its cephalon, but no other appendages or auditory or visual structures. No mouth is present in the specimens, but the gut is present, so
2192:
2382:
Berks, Harry O.; Lunde
Nielsen, Morten; Flannery‐Sutherland, Joseph; Thorshøj Nielsen, Arne; Park, Tae‐Yoon S.; Vinther, Jakob (May 2023).
410:
though the differences between the two species are relatively minor. The species also preserves elements of the gut tract not preserved in
2441:
2168:
3243:
2136:
2283:"Redescription of the Chengjiang arthropod Squamacula clypeata Hou and Bergström, from the Lower Cambrian, south-west China"
2214:
2384:"A possibly deep branching artiopodan arthropod from the lower Cambrian Sirius Passet Lagerstätte (North Greenland)"
135:
2196:
349:
3254:
2434:
607:
2746:
2215:"New artiopodan arthropods from the early Cambrian Emu Bay Shale Konservat-Lagerstätte of South Australia"
1513:
378:
must have had a mouth, which is thought to have been between the cephalon and doublure on the underside.
3249:
3261:
2753:
2346:
2294:
2226:
2798:
2579:
1633:
515:
3219:
2826:
2777:
1568:
3153:
2989:
2881:
2833:
2769:
2700:
1938:
1921:
1897:
1814:
1616:
1592:
931:
746:
3300:
3279:
3162:
2895:
2874:
2805:
2686:
2671:
2563:
2427:
2250:
2186:
1873:
1544:
760:
476:
130:
2909:
2853:
2761:
2693:
1766:
2978:
2819:
2623:
1742:
1486:
3190:
2916:
2902:
2888:
2405:
2364:
2312:
2242:
2174:
2164:
2142:
2132:
1790:
1725:
858:
319:
276:
3033:
2707:
1403:
2789:
2679:
2395:
2354:
2302:
2234:
1420:
284:
2936:
2739:
2572:
1701:
620:
565:
548:
2335:"A new small soft-bodied non-trilobite artiopod from the Cambrian Stage 4 Guanshan Biota"
2350:
2298:
2230:
406:
in that it has 11 thoracic tergites, which appear to exhibit less overlap that those of
3012:
2964:
2602:
2523:
1313:
1051:
396:(bristles) near the tip, which may have been used for both locomotion and respiration.
2333:
Jiao, De-Guang; Du, Kun-Sheng; Zhang, Xi-Guang; Yang, Jie; Eggink, Daniel (May 2022).
3294:
3005:
2655:
2477:
2307:
2282:
2104:
1365:
986:
969:
296:
72:
2254:
426:
has consistently been placed as the earliest diverging and most primitive member of
337:, meaning shield-shaped. It was named this in reference to its shield-like outline.
287:
of Yunnan, China. At the time of description there were only two known specimens of
3053:
2957:
2844:
2664:
2161:
The
Cambrian fossils of Chengjiang, China : the flowering of early animal life
1379:
673:
206:
17:
2213:
Paterson, John R.; García-Bellido, Diego C.; Edgecombe, Gregory D. (March 2012).
2950:
2867:
2731:
2718:
2616:
1499:
1352:
715:
47:
3089:
3080:
3069:
3044:
3026:
3019:
2998:
2971:
2812:
2359:
2334:
1684:
1246:
1100:
1034:
1010:
945:
635:
92:
57:
2409:
2368:
2316:
2246:
2178:
3208:
3197:
3138:
3131:
3116:
2644:
2630:
2544:
2516:
2498:
2483:
2468:
2450:
2163:. Hou, Xianguang. (Second ed.). Chichester, West Sussex. 8 March 2017.
2146:
2099:
1462:
1155:
1138:
1114:
841:
698:
431:
427:
388:
appendages (an appendage which branches into two) per thoracic tergite. The
365:
272:
269:
180:
167:
147:
97:
2281:
Zhang, Xingliang; Han, Jian; Zhang, Zhifei; Liu, Huqin; Shu, Degan (2004).
291:, but now there are at least six known specimens. In 2012 a second species
3183:
3169:
3107:
3062:
2860:
2637:
2586:
2551:
1831:
1328:
875:
808:
498:
385:
361:
87:
82:
67:
62:
52:
114:
3176:
3096:
2530:
1229:
1205:
791:
389:
353:
327:
102:
77:
2400:
2383:
2462:
2238:
357:
157:
3124:
2129:
Arthropods of the Lower
Cambrian Chengjiang fauna, southwest China
1172:
308:
393:
348:
is flattened (dorsoventrally). It has 11 segments in total: the
2423:
434:
and a variety of morphologically similar soft-bodied forms.
2419:
2097:
A close relationship has been proposed with the genus
326:, indicating that the animal is relatively small. The
247:
236:
213:
3207:
3152:
3106:
3079:
3052:
3043:
2988:
2934:
2843:
2788:
2717:
2654:
2600:
2562:
2496:
2131:. Bergström. Oslo: Scandinavian University Press.
2435:
8:
3049:
2942:
2723:
2608:
2508:
2504:
2493:
2442:
2428:
2420:
2191:: CS1 maint: location missing publisher (
113:
31:
2399:
2358:
2306:
2127:Hou, Xianguang; Bergström, Jan (1997).
2116:
2328:
2326:
2184:
7:
2276:
2274:
2272:
2270:
2268:
2266:
2264:
2208:
2206:
2122:
2120:
1913:
1889:
1865:
1806:
1782:
1758:
1717:
1676:
1608:
1584:
1560:
1536:
1529:
1505:
1478:
1454:
1395:
1371:
1344:
1320:
1221:
1197:
1130:
1106:
1026:
1002:
961:
937:
927:
833:
783:
776:
752:
690:
665:
658:
651:
627:
599:
540:
490:
483:
472:
448:
440:
437:Cladogram after Jiao et al. 2021.
25:
3275:
3274:
3260:
3253:
3248:
3242:
2308:10.1111/j.0031-0239.2004.00363.x
134:
333:is derived from the Latin word
314:is derived from the Latin word
283:was described in 1997 from the
1:
1922:Glypharthrus trispinicaudatus
674:Kwanyinaspis maotiashanensis
27:Extinct genus of arthropods
3317:
1702:Australaglaspis stonyensis
566:Acanthomeridion anacanthus
422:In phylogenetic analyses,
352:(the head), nine thoracic
3270:
3240:
2945:
2726:
2611:
2511:
2507:
2492:
2457:
2360:10.1017/S0016756821001254
1935:
1918:
1911:
1894:
1887:
1870:
1863:
1828:
1811:
1804:
1787:
1780:
1763:
1756:
1739:
1722:
1715:
1698:
1681:
1674:
1634:Brachyaglaspis singularis
1630:
1613:
1606:
1589:
1582:
1565:
1558:
1541:
1534:
1527:
1510:
1503:
1483:
1476:
1459:
1452:
1417:
1400:
1393:
1376:
1369:
1349:
1342:
1325:
1318:
1243:
1226:
1219:
1202:
1195:
1169:
1152:
1135:
1128:
1111:
1104:
1048:
1031:
1024:
1007:
1000:
983:
966:
959:
942:
935:
925:
872:
855:
838:
831:
805:
788:
781:
774:
757:
750:
712:
695:
688:
670:
663:
656:
649:
632:
625:
604:
597:
562:
545:
538:
512:
495:
488:
481:
470:
453:
446:
318:, meaning scale, and the
233:
228:
212:
205:
131:Scientific classification
129:
121:
112:
34:
1052:Phytophilaspis pergamena
549:Acanthomeridion serratum
430:, a clade that includes
356:(each of which covers a
2388:Papers in Palaeontology
2219:Journal of Paleontology
1939:Glypharthrus magnoculus
1247:Misszhouia longicaudata
295:was described from the
243:Hou and Bergström, 1997
223:Hou and Bergström, 1997
200:Hou and Bergström, 1997
1767:Aglaspella granulifera
1569:Quasimodaspis brentsae
1035:Xandarella spectaculum
970:Luohuilinella deletres
716:Eoredlichia intermedia
608:Bailongia longicaudata
516:Australimicola spriggi
2195:) CS1 maint: others (
636:Retifacies abnormalis
1898:Glypharthrus thomasi
1815:Glypharthrus simplex
1617:Tremaglaspis vanroyi
1404:Triopus draboviensis
1380:Cheloniellon calmani
1329:Sidneyia inexpectans
1139:Tariccoia arrusensis
254:Paterson et al. 2012
2351:2022GeoM..159..730J
2339:Geological Magazine
2299:2004Palgy..47..605Z
2231:2012JPal...86..340P
2107:site of Greenland.
1874:Chraspedops modesta
1726:Uarthrus instabilis
1685:Cyclopites vulgaris
1156:Soomaspis splendida
987:Luohuilinella rarus
457:Squamacula clypeata
382:Squamacula clypeata
372:Squamacula clypeata
346:Squamacula clypeata
279:. The type species
251:Squamacula buckorum
240:Squamacula clypeata
218:Squamacula clypeata
124:Squamacula clypeata
18:Squamacula clypeata
1593:Tremaglaspis unite
1487:Eozetetes gemmelli
1353:Emeraldella brocki
1011:Sinoburius lunaris
946:Cindarella eucalla
876:Saperion glumaceum
761:Haifengella corona
699:Olenoides serratus
122:Reconstruction of
3288:
3287:
3238:
3237:
3234:
3233:
3230:
3229:
3148:
3147:
2930:
2929:
2926:
2925:
2596:
2595:
2401:10.1002/spp2.1495
2094:
2093:
2085:
2084:
2076:
2075:
2067:
2066:
2058:
2057:
2049:
2048:
2040:
2039:
2031:
2030:
2022:
2021:
2013:
2012:
2004:
2003:
1995:
1994:
1986:
1985:
1977:
1976:
1968:
1967:
1959:
1958:
1950:
1949:
1852:
1851:
1843:
1842:
1832:Aglaspis spinifer
1663:
1662:
1654:
1653:
1645:
1644:
1545:Chlupacaris dubia
1441:
1440:
1432:
1431:
1303:
1302:
1294:
1293:
1285:
1284:
1276:
1275:
1267:
1266:
1258:
1257:
1184:
1183:
1115:Buenaspis forteyi
1090:
1089:
1081:
1080:
1072:
1071:
1063:
1062:
914:
913:
905:
904:
896:
895:
887:
886:
820:
819:
736:
735:
727:
726:
586:
585:
577:
576:
527:
526:
499:Zhiwenia coronata
320:diminutive suffix
277:Cambrian Series 2
261:
260:
255:
244:
201:
43:Cambrian Series 2
16:(Redirected from
3308:
3278:
3277:
3265:
3264:
3257:
3252:
3246:
3050:
2943:
2790:Tremaglaspididae
2724:
2609:
2509:
2505:
2494:
2444:
2437:
2430:
2421:
2414:
2413:
2403:
2379:
2373:
2372:
2362:
2330:
2321:
2320:
2310:
2278:
2259:
2258:
2239:10.1666/11-077.1
2210:
2201:
2200:
2190:
2182:
2157:
2151:
2150:
2124:
1914:
1890:
1866:
1807:
1791:Gogglops ensifer
1783:
1759:
1718:
1677:
1609:
1585:
1561:
1537:
1530:
1506:
1479:
1463:Kodymirus vagans
1455:
1396:
1372:
1345:
1321:
1230:Naraoia compacta
1222:
1198:
1131:
1107:
1027:
1003:
962:
938:
928:
834:
809:Helmetia expansa
784:
777:
753:
691:
666:
659:
652:
628:
600:
541:
491:
484:
473:
449:
444:Artiopoda
441:
384:has one pair of
285:Chengjiang biota
253:
249:
242:
238:
215:
199:
192:
179:
139:
138:
117:
107:
44:
40:Temporal range:
32:
21:
3316:
3315:
3311:
3310:
3309:
3307:
3306:
3305:
3291:
3290:
3289:
3284:
3266:
3259:
3226:
3203:
3144:
3102:
3075:
3039:
2984:
2939:
2937:Trilobitomorpha
2922:
2839:
2784:
2754:Chacharejocaris
2740:Australaglaspis
2713:
2650:
2605:
2592:
2573:Acanthomeridion
2558:
2501:
2488:
2453:
2448:
2418:
2417:
2381:
2380:
2376:
2332:
2331:
2324:
2280:
2279:
2262:
2212:
2211:
2204:
2183:
2171:
2159:
2158:
2154:
2139:
2126:
2125:
2118:
2113:
2095:
2086:
2077:
2068:
2059:
2050:
2041:
2032:
2023:
2014:
2005:
1996:
1987:
1978:
1969:
1960:
1951:
1853:
1844:
1743:Flobertia kochi
1664:
1655:
1646:
1514:Beckwithia typa
1442:
1433:
1421:Duslia insignis
1304:
1295:
1286:
1277:
1268:
1259:
1206:Naraoia spinosa
1185:
1091:
1082:
1073:
1064:
915:
906:
897:
888:
842:Tegopelte gigas
821:
737:
728:
621:Trilobitomorpha
587:
578:
528:
420:
343:
305:
224:
221:
198:
190:
177:
133:
108:
106:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
42:
41:
38:
28:
23:
22:
15:
12:
11:
5:
3314:
3312:
3304:
3303:
3293:
3292:
3286:
3285:
3283:
3282:
3271:
3268:
3267:
3241:
3239:
3236:
3235:
3232:
3231:
3228:
3227:
3225:
3224:
3223:
3222:
3213:
3211:
3205:
3204:
3202:
3201:
3194:
3187:
3180:
3173:
3166:
3158:
3156:
3150:
3149:
3146:
3145:
3143:
3142:
3135:
3128:
3121:
3112:
3110:
3104:
3103:
3101:
3100:
3093:
3085:
3083:
3077:
3076:
3074:
3073:
3066:
3058:
3056:
3047:
3041:
3040:
3038:
3037:
3030:
3023:
3016:
3013:Phytophilaspis
3009:
3002:
2994:
2992:
2986:
2985:
2983:
2982:
2975:
2968:
2965:Pygmaclypeatus
2961:
2954:
2946:
2940:
2935:
2932:
2931:
2928:
2927:
2924:
2923:
2921:
2920:
2913:
2906:
2899:
2892:
2885:
2878:
2871:
2864:
2857:
2849:
2847:
2841:
2840:
2838:
2837:
2830:
2823:
2816:
2809:
2802:
2799:Brachyaglaspis
2794:
2792:
2786:
2785:
2783:
2782:
2774:
2766:
2758:
2750:
2743:
2736:
2727:
2721:
2715:
2714:
2712:
2711:
2704:
2697:
2690:
2683:
2676:
2668:
2660:
2658:
2652:
2651:
2649:
2648:
2641:
2634:
2627:
2620:
2612:
2606:
2603:Vicissicaudata
2601:
2598:
2597:
2594:
2593:
2591:
2590:
2583:
2580:Australimicola
2576:
2568:
2566:
2560:
2559:
2557:
2556:
2548:
2541:
2534:
2527:
2524:Falcatamacaris
2520:
2512:
2502:
2497:
2490:
2489:
2487:
2486:
2480:
2471:
2465:
2458:
2455:
2454:
2449:
2447:
2446:
2439:
2432:
2424:
2416:
2415:
2374:
2345:(5): 730–734.
2322:
2293:(3): 605–617.
2260:
2225:(2): 340–357.
2202:
2169:
2152:
2137:
2115:
2114:
2112:
2109:
2092:
2091:
2088:
2087:
2083:
2082:
2079:
2078:
2074:
2073:
2070:
2069:
2065:
2064:
2061:
2060:
2056:
2055:
2052:
2051:
2047:
2046:
2043:
2042:
2038:
2037:
2034:
2033:
2029:
2028:
2025:
2024:
2020:
2019:
2016:
2015:
2011:
2010:
2007:
2006:
2002:
2001:
1998:
1997:
1993:
1992:
1989:
1988:
1984:
1983:
1980:
1979:
1975:
1974:
1971:
1970:
1966:
1965:
1962:
1961:
1957:
1956:
1953:
1952:
1948:
1947:
1944:
1943:
1934:
1931:
1930:
1927:
1926:
1917:
1912:
1910:
1907:
1906:
1903:
1902:
1893:
1888:
1886:
1883:
1882:
1879:
1878:
1869:
1864:
1862:
1859:
1858:
1855:
1854:
1850:
1849:
1846:
1845:
1841:
1840:
1837:
1836:
1827:
1824:
1823:
1820:
1819:
1810:
1805:
1803:
1800:
1799:
1796:
1795:
1786:
1781:
1779:
1776:
1775:
1772:
1771:
1762:
1757:
1755:
1752:
1751:
1748:
1747:
1738:
1735:
1734:
1731:
1730:
1721:
1716:
1714:
1711:
1710:
1707:
1706:
1697:
1694:
1693:
1690:
1689:
1680:
1675:
1673:
1670:
1669:
1666:
1665:
1661:
1660:
1657:
1656:
1652:
1651:
1648:
1647:
1643:
1642:
1639:
1638:
1629:
1626:
1625:
1622:
1621:
1612:
1607:
1605:
1602:
1601:
1598:
1597:
1588:
1583:
1581:
1578:
1577:
1574:
1573:
1564:
1559:
1557:
1554:
1553:
1550:
1549:
1540:
1535:
1533:
1528:
1526:
1523:
1522:
1519:
1518:
1509:
1504:
1502:
1496:
1495:
1492:
1491:
1482:
1477:
1475:
1472:
1471:
1468:
1467:
1458:
1453:
1451:
1448:
1447:
1444:
1443:
1439:
1438:
1435:
1434:
1430:
1429:
1426:
1425:
1416:
1413:
1412:
1409:
1408:
1399:
1394:
1392:
1389:
1388:
1385:
1384:
1375:
1370:
1368:
1362:
1361:
1358:
1357:
1348:
1343:
1341:
1338:
1337:
1334:
1333:
1324:
1319:
1317:
1314:Vicissicaudata
1310:
1309:
1306:
1305:
1301:
1300:
1297:
1296:
1292:
1291:
1288:
1287:
1283:
1282:
1279:
1278:
1274:
1273:
1270:
1269:
1265:
1264:
1261:
1260:
1256:
1255:
1252:
1251:
1242:
1239:
1238:
1235:
1234:
1225:
1220:
1218:
1215:
1214:
1211:
1210:
1201:
1196:
1194:
1191:
1190:
1187:
1186:
1182:
1181:
1178:
1177:
1168:
1165:
1164:
1161:
1160:
1151:
1148:
1147:
1144:
1143:
1134:
1129:
1127:
1124:
1123:
1120:
1119:
1110:
1105:
1103:
1097:
1096:
1093:
1092:
1088:
1087:
1084:
1083:
1079:
1078:
1075:
1074:
1070:
1069:
1066:
1065:
1061:
1060:
1057:
1056:
1047:
1044:
1043:
1040:
1039:
1030:
1025:
1023:
1020:
1019:
1016:
1015:
1006:
1001:
999:
996:
995:
992:
991:
982:
979:
978:
975:
974:
965:
960:
958:
955:
954:
951:
950:
941:
936:
934:
926:
924:
921:
920:
917:
916:
912:
911:
908:
907:
903:
902:
899:
898:
894:
893:
890:
889:
885:
884:
881:
880:
871:
868:
867:
864:
863:
859:Skioldia aldna
854:
851:
850:
847:
846:
837:
832:
830:
827:
826:
823:
822:
818:
817:
814:
813:
804:
801:
800:
797:
796:
787:
782:
780:
775:
773:
770:
769:
766:
765:
756:
751:
749:
743:
742:
739:
738:
734:
733:
730:
729:
725:
724:
721:
720:
711:
708:
707:
704:
703:
694:
689:
687:
683:
682:
679:
678:
669:
664:
662:
657:
655:
650:
648:
645:
644:
641:
640:
631:
626:
624:
617:
616:
613:
612:
603:
598:
596:
593:
592:
589:
588:
584:
583:
580:
579:
575:
574:
571:
570:
561:
558:
557:
554:
553:
544:
539:
537:
534:
533:
530:
529:
525:
524:
521:
520:
511:
508:
507:
504:
503:
494:
489:
487:
482:
480:
471:
469:
466:
465:
462:
461:
452:
447:
445:
439:
419:
416:
342:
339:
304:
301:
299:of Australia.
268:is an extinct
259:
258:
257:
256:
245:
231:
230:
226:
225:
222:
210:
209:
203:
202:
188:
184:
183:
175:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
127:
126:
119:
118:
110:
109:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
45:
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
3313:
3302:
3299:
3298:
3296:
3281:
3273:
3272:
3269:
3263:
3258:
3256:
3251:
3245:
3221:
3217:
3216:
3215:
3214:
3212:
3210:
3206:
3200:
3199:
3195:
3193:
3192:
3188:
3186:
3185:
3181:
3179:
3178:
3174:
3172:
3171:
3167:
3165:
3164:
3160:
3159:
3157:
3155:
3151:
3141:
3140:
3136:
3134:
3133:
3129:
3127:
3126:
3122:
3119:
3118:
3114:
3113:
3111:
3109:
3105:
3099:
3098:
3094:
3092:
3091:
3087:
3086:
3084:
3082:
3078:
3072:
3071:
3067:
3065:
3064:
3060:
3059:
3057:
3055:
3051:
3048:
3046:
3042:
3036:
3035:
3031:
3029:
3028:
3024:
3022:
3021:
3017:
3015:
3014:
3010:
3008:
3007:
3006:Luohuilinella
3003:
3001:
3000:
2996:
2995:
2993:
2991:
2987:
2981:
2980:
2976:
2974:
2973:
2969:
2967:
2966:
2962:
2960:
2959:
2955:
2953:
2952:
2948:
2947:
2944:
2941:
2938:
2933:
2919:
2918:
2914:
2912:
2911:
2907:
2905:
2904:
2900:
2898:
2897:
2893:
2891:
2890:
2886:
2884:
2883:
2879:
2877:
2876:
2872:
2870:
2869:
2865:
2863:
2862:
2858:
2856:
2855:
2851:
2850:
2848:
2846:
2842:
2836:
2835:
2831:
2829:
2828:
2827:Quasimodaspis
2824:
2822:
2821:
2817:
2815:
2814:
2810:
2808:
2807:
2803:
2801:
2800:
2796:
2795:
2793:
2791:
2787:
2780:
2779:
2778:Rozhkovocaris
2775:
2772:
2771:
2767:
2764:
2763:
2759:
2756:
2755:
2751:
2749:
2748:
2744:
2742:
2741:
2737:
2734:
2733:
2729:
2728:
2725:
2722:
2720:
2716:
2710:
2709:
2705:
2703:
2702:
2698:
2696:
2695:
2691:
2689:
2688:
2684:
2682:
2681:
2677:
2674:
2673:
2669:
2667:
2666:
2662:
2661:
2659:
2657:
2656:Cheloniellida
2653:
2647:
2646:
2642:
2640:
2639:
2635:
2633:
2632:
2628:
2626:
2625:
2621:
2619:
2618:
2614:
2613:
2610:
2607:
2604:
2599:
2589:
2588:
2584:
2582:
2581:
2577:
2575:
2574:
2570:
2569:
2567:
2565:
2561:
2554:
2553:
2549:
2547:
2546:
2542:
2540:
2539:
2535:
2533:
2532:
2528:
2526:
2525:
2521:
2519:
2518:
2514:
2513:
2510:
2506:
2503:
2500:
2495:
2491:
2485:
2481:
2479:
2478:Panarthropoda
2475:
2472:
2470:
2467:Superphylum:
2466:
2464:
2460:
2459:
2456:
2452:
2445:
2440:
2438:
2433:
2431:
2426:
2425:
2422:
2411:
2407:
2402:
2397:
2393:
2389:
2385:
2378:
2375:
2370:
2366:
2361:
2356:
2352:
2348:
2344:
2340:
2336:
2329:
2327:
2323:
2318:
2314:
2309:
2304:
2300:
2296:
2292:
2288:
2287:Palaeontology
2284:
2277:
2275:
2273:
2271:
2269:
2267:
2265:
2261:
2256:
2252:
2248:
2244:
2240:
2236:
2232:
2228:
2224:
2220:
2216:
2209:
2207:
2203:
2198:
2194:
2188:
2180:
2176:
2172:
2170:9781118896310
2166:
2162:
2156:
2153:
2148:
2144:
2140:
2134:
2130:
2123:
2121:
2117:
2110:
2108:
2106:
2105:Sirius Passet
2102:
2101:
2090:
2089:
2081:
2080:
2072:
2071:
2063:
2062:
2054:
2053:
2045:
2044:
2036:
2035:
2027:
2026:
2018:
2017:
2009:
2008:
2000:
1999:
1991:
1990:
1982:
1981:
1973:
1972:
1964:
1963:
1955:
1954:
1946:
1945:
1942:
1941:
1940:
1933:
1932:
1929:
1928:
1925:
1924:
1923:
1916:
1915:
1909:
1908:
1905:
1904:
1901:
1900:
1899:
1892:
1891:
1885:
1884:
1881:
1880:
1877:
1876:
1875:
1868:
1867:
1861:
1860:
1857:
1856:
1848:
1847:
1839:
1838:
1835:
1834:
1833:
1826:
1825:
1822:
1821:
1818:
1817:
1816:
1809:
1808:
1802:
1801:
1798:
1797:
1794:
1793:
1792:
1785:
1784:
1778:
1777:
1774:
1773:
1770:
1769:
1768:
1761:
1760:
1754:
1753:
1750:
1749:
1746:
1745:
1744:
1737:
1736:
1733:
1732:
1729:
1728:
1727:
1720:
1719:
1713:
1712:
1709:
1708:
1705:
1704:
1703:
1696:
1695:
1692:
1691:
1688:
1687:
1686:
1679:
1678:
1672:
1671:
1668:
1667:
1659:
1658:
1650:
1649:
1641:
1640:
1637:
1636:
1635:
1628:
1627:
1624:
1623:
1620:
1619:
1618:
1611:
1610:
1604:
1603:
1600:
1599:
1596:
1595:
1594:
1587:
1586:
1580:
1579:
1576:
1575:
1572:
1571:
1570:
1563:
1562:
1556:
1555:
1552:
1551:
1548:
1547:
1546:
1539:
1538:
1532:
1531:
1525:
1524:
1521:
1520:
1517:
1516:
1515:
1508:
1507:
1501:
1498:
1497:
1494:
1493:
1490:
1489:
1488:
1481:
1480:
1474:
1473:
1470:
1469:
1466:
1465:
1464:
1457:
1456:
1450:
1449:
1446:
1445:
1437:
1436:
1428:
1427:
1424:
1423:
1422:
1415:
1414:
1411:
1410:
1407:
1406:
1405:
1398:
1397:
1391:
1390:
1387:
1386:
1383:
1382:
1381:
1374:
1373:
1367:
1366:Cheloniellida
1364:
1363:
1360:
1359:
1356:
1355:
1354:
1347:
1346:
1340:
1339:
1336:
1335:
1332:
1331:
1330:
1323:
1322:
1315:
1312:
1311:
1308:
1307:
1299:
1298:
1290:
1289:
1281:
1280:
1272:
1271:
1263:
1262:
1254:
1253:
1250:
1249:
1248:
1241:
1240:
1237:
1236:
1233:
1232:
1231:
1224:
1223:
1217:
1216:
1213:
1212:
1209:
1208:
1207:
1200:
1199:
1193:
1192:
1189:
1188:
1180:
1179:
1176:
1175:
1174:
1173:Liwia convexa
1167:
1166:
1163:
1162:
1159:
1158:
1157:
1150:
1149:
1146:
1145:
1142:
1141:
1140:
1133:
1132:
1126:
1125:
1122:
1121:
1118:
1117:
1116:
1109:
1108:
1102:
1099:
1098:
1095:
1094:
1086:
1085:
1077:
1076:
1068:
1067:
1059:
1058:
1055:
1054:
1053:
1046:
1045:
1042:
1041:
1038:
1037:
1036:
1029:
1028:
1022:
1021:
1018:
1017:
1014:
1013:
1012:
1005:
1004:
998:
997:
994:
993:
990:
989:
988:
981:
980:
977:
976:
973:
972:
971:
964:
963:
957:
956:
953:
952:
949:
948:
947:
940:
939:
933:
930:
929:
923:
922:
919:
918:
910:
909:
901:
900:
892:
891:
883:
882:
879:
878:
877:
870:
869:
866:
865:
862:
861:
860:
853:
852:
849:
848:
845:
844:
843:
836:
835:
829:
828:
825:
824:
816:
815:
812:
811:
810:
803:
802:
799:
798:
795:
794:
793:
786:
785:
779:
778:
772:
771:
768:
767:
764:
763:
762:
755:
754:
748:
745:
744:
741:
740:
732:
731:
723:
722:
719:
718:
717:
710:
709:
706:
705:
702:
701:
700:
693:
692:
685:
684:
681:
680:
677:
676:
675:
668:
667:
661:
660:
654:
653:
647:
646:
643:
642:
639:
638:
637:
630:
629:
622:
619:
618:
615:
614:
611:
610:
609:
602:
601:
595:
594:
591:
590:
582:
581:
573:
572:
569:
568:
567:
560:
559:
556:
555:
552:
551:
550:
543:
542:
536:
535:
532:
531:
523:
522:
519:
518:
517:
510:
509:
506:
505:
502:
501:
500:
493:
492:
486:
485:
478:
475:
474:
468:
467:
464:
463:
460:
459:
458:
451:
450:
443:
442:
438:
435:
433:
429:
425:
417:
415:
413:
409:
405:
402:differs from
401:
397:
395:
391:
387:
383:
379:
377:
373:
369:
367:
363:
359:
355:
351:
347:
340:
338:
336:
332:
329:
325:
321:
317:
313:
310:
302:
300:
298:
297:Emu Bay Shale
294:
290:
286:
282:
278:
274:
271:
267:
266:
252:
246:
241:
235:
234:
232:
227:
220:
219:
211:
208:
204:
197:
196:
189:
186:
185:
182:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
137:
132:
128:
125:
120:
116:
111:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
37:
33:
30:
19:
3247:
3196:
3189:
3182:
3175:
3168:
3161:
3154:Conciliterga
3137:
3130:
3123:
3115:
3095:
3088:
3068:
3061:
3054:Emucarididae
3032:
3025:
3018:
3011:
3004:
2997:
2990:Xandarellida
2977:
2970:
2963:
2958:Kwanyinaspis
2956:
2949:
2915:
2908:
2901:
2894:
2887:
2882:Glypharthrus
2880:
2873:
2866:
2859:
2852:
2845:Aglaspididae
2834:Tremaglaspis
2832:
2825:
2818:
2811:
2804:
2797:
2776:
2770:Obrutschewia
2768:
2760:
2752:
2745:
2738:
2730:
2706:
2701:Pseudarthron
2699:
2692:
2685:
2678:
2670:
2665:Cheloniellon
2663:
2643:
2636:
2629:
2622:
2615:
2585:
2578:
2571:
2550:
2543:
2537:
2536:
2529:
2522:
2515:
2473:
2391:
2387:
2377:
2342:
2338:
2290:
2286:
2222:
2218:
2160:
2155:
2128:
2098:
2096:
1937:
1936:
1920:
1919:
1896:
1895:
1872:
1871:
1830:
1829:
1813:
1812:
1789:
1788:
1765:
1764:
1741:
1740:
1724:
1723:
1700:
1699:
1683:
1682:
1632:
1631:
1615:
1614:
1591:
1590:
1567:
1566:
1543:
1542:
1512:
1511:
1485:
1484:
1461:
1460:
1419:
1418:
1402:
1401:
1378:
1377:
1351:
1350:
1327:
1326:
1245:
1244:
1228:
1227:
1204:
1203:
1171:
1170:
1154:
1153:
1137:
1136:
1113:
1112:
1050:
1049:
1033:
1032:
1009:
1008:
985:
984:
968:
967:
944:
943:
932:Xandarellida
874:
873:
857:
856:
840:
839:
807:
806:
792:Kuamaia lata
790:
789:
759:
758:
747:Conciliterga
714:
713:
697:
696:
672:
671:
634:
633:
606:
605:
564:
563:
547:
546:
514:
513:
497:
496:
456:
455:
454:
436:
423:
421:
411:
408:S. clypeata,
407:
403:
399:
398:
381:
380:
375:
371:
370:
345:
344:
334:
330:
323:
315:
311:
306:
292:
288:
280:
264:
263:
262:
250:
239:
217:
216:
207:Type species
194:
193:
174:(unranked):
123:
35:
29:
3163:Haifengella
2951:Arthroaspis
2896:Hesselbonia
2875:Chraspedops
2868:Aglaspoides
2806:Chlupacaris
2732:Angarocaris
2719:Aglaspidida
2687:Neostrabops
2672:Drabovaspis
2617:Emeraldella
2564:Protosutura
1500:Aglaspidida
477:Protosutura
412:S. clypeata
404:S. clypeata
400:S. buckorum
376:S. clypeata
364:. It has a
360:), and one
341:Description
293:S. buckorum
289:S. clypeata
281:S. clypeata
3090:Misszhouia
3081:Naraoiidae
3070:Kangacaris
3045:Nektaspida
3027:Xandarella
3020:Sinoburius
2999:Cindarella
2972:Retifacies
2910:Tuboculops
2854:Aglaspella
2813:Cyclopites
2762:Girardevia
2747:Beckwithia
2694:Paraduslia
2538:Squamacula
2484:Arthropoda
2138:8200376931
2111:References
1101:Nektaspida
686:Trilobita
432:trilobites
424:Squamacula
312:Squamacula
270:artiopodan
265:Squamacula
195:Squamacula
168:Arthropoda
36:Squamacula
3301:Artiopoda
3220:Trilobita
3209:Trilobita
3198:Tegopelte
3139:Tariccoia
3132:Soomaspis
3117:Buenaspis
2979:Tonglaiia
2820:Flobertia
2645:Carimersa
2631:Kodymirus
2624:Eozetetes
2545:Thulaspis
2517:Bailongia
2499:Artiopoda
2469:Ecdysozoa
2461:Kingdom:
2451:Artiopoda
2410:2056-2799
2369:0016-7568
2317:1475-4983
2247:0022-3360
2187:cite book
2179:970396735
2103:from the
2100:Thulaspis
428:Artiopoda
335:clypeatus
303:Etymology
275:from the
273:arthropod
181:Artiopoda
154:Kingdom:
148:Eukaryota
3295:Category
3280:Category
3191:Skioldia
3184:Saperion
3170:Helmetia
3108:Liwiidae
3063:Emucaris
2917:Uarthrus
2903:Setaspis
2889:Gogglops
2861:Aglaspis
2638:Sidneyia
2587:Zhiwenia
2552:Urokodia
2482:Phylum:
2463:Animalia
2255:87520649
2147:38305908
418:Taxonomy
386:biramous
366:doublure
362:pygidium
354:tergites
350:cephalon
331:clypeata
229:Species
164:Phylum:
158:Animalia
144:Domain:
3177:Kuamaia
3097:Naraoia
3034:Zhugeia
2708:Triopus
2531:Molaria
2347:Bibcode
2295:Bibcode
2227:Bibcode
1316:
623:
479:
390:endopod
328:species
187:Genus:
2680:Duslia
2408:
2367:
2315:
2253:
2245:
2177:
2167:
2145:
2135:
358:somite
316:squama
3125:Liwia
2474:Clade
2394:(3).
2251:S2CID
394:setae
324:culus
309:genus
3218:see
2406:ISSN
2365:ISSN
2313:ISSN
2243:ISSN
2197:link
2193:link
2175:OCLC
2165:ISBN
2143:OCLC
2133:ISBN
307:The
48:PreꞒ
2396:doi
2355:doi
2343:159
2303:doi
2235:doi
3297::
2476::
2404:.
2390:.
2386:.
2363:.
2353:.
2341:.
2337:.
2325:^
2311:.
2301:.
2291:47
2289:.
2285:.
2263:^
2249:.
2241:.
2233:.
2223:86
2221:.
2217:.
2205:^
2189:}}
2185:{{
2173:.
2141:.
2119:^
98:Pg
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2781:?
2773:?
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2757:?
2735:?
2675:?
2555:?
2443:e
2436:t
2429:v
2412:.
2398::
2392:9
2371:.
2357::
2349::
2319:.
2305::
2297::
2257:.
2237::
2229::
2199:)
2181:.
2149:.
322:-
248:†
237:†
214:†
191:†
178:†
103:N
93:K
88:J
83:T
78:P
73:C
68:D
63:S
58:O
53:Ꞓ
20:)
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