677:
2532:
2555:
2547:
1951:
2540:
3415:
1336:
144:
3401:
123:
808:
3464:
1936:
1944:
3408:
799:(outer shin bone), a continuous facet between this condyle and the facet for the fourth distal tarsal (a minor ankle bone), and a rear projection known as a calcaneal tuber which was wider than tall and had flared edges. Some of these characteristics were not considered to be present until a re-evalutation of the specimen in 2011.
555:. A similar situation is visible in other gracilisuchids, which have a broad contact between the jugal and the lower branch of the squamosal. However, they did not retain the upper portion of the lower temporal fenestra, leaving only a small, triangular remnant of the lower temporal fenestra under the squamosal-jugal contact.
718:(upper plate of the hip) with large, fan-shaped sacral ribs. No intercentra were preserved, and the vertebrae lacked bevelled edges, but since the vertebrae were disarticulated and some euparkeriids retain intercentra while lacking bevelled edges, their absence in
740:(bony plates) were initially reported as being absent, but later preparation revealed one. This osteoderm was broad, with a ridge on its upper surface culminating in a pointed front prong. A corresponding groove-like depression was present on the lower surface.
480:
These posterodorsal processes are very characteristic, either by bisecting the nasal (in the case of the premaxilla's) or existing in the first place (in the case of the maxilla's). As such, they were considered by Wu & Russell (2001) to be
516:
The premaxilla probably had five teeth, while the maxilla had at least 13. Preserved teeth were curved and finely serrated. The nasal bones project over the side of the snout and form the upper border of the antorbital fossa. The
852:
533:. The portion of the jugal which rises behind the eye socket has a very wide base, and its surface is set inwards relative to the portion below the eye socket. As with other diapsids,
469:(the main toothed bone of the snout) also had a posterodorsal process, a much more rare trait compared to the posterodorsal process of the premaxilla (which was present in most
965:, in the newly created family Gracilisuchidae. Most of the similarities justifying this classification were present in the skull, and some were originally considered unique to
652:
The lower jaw was slender, with an elliptical hole known as a mandibular fenestra. The toothed dentary bone had two rear prongs surrounding the mandibular fenestra. Unique to
946:) had internal carotid arteries which entered the braincase from below. In addition, the calcaneum was re-evaluated and Parrish's observations of the ankle were reconfirmed.
668:
is how the outer surface of the surangular was highly concave. There were at least 16 teeth in the lower jaw, and they were similar in shape to those of the upper jaw.
3529:
499:, so they likely diagnose the entire family Gracilisuchidae, or possibly even larger subsets of Archosauria. For example, the premaxillary posterodorsal process of
473:). The maxilla's posterodorsal process was a small, triangular peak of bone which formed the lower rear corner of the antorbital fossa and connected to the
521:(which form the portion of the skull above the eye sockets) wedge into the nasals as a large, V-shaped suture. Both of these traits are also present in
3623:
3593:
3516:
819:
has gone through much revision in the past due to its combination of features from both suchian archosaurs and earlier archosaur relatives such as
3588:
922:. This reassignment was based on the structure of the calcaneum (heel bone), which is similar to that of other early pseudosuchians, especially
3613:
1369:
3618:
901:
which entered the braincase from below, rather than from the side. However, they also ruled out the possibility of a close relationship with
1172:"New clade of enigmatic early archosaurs yields insights into early pseudosuchian phylogeny and the biogeography of the archosaur radiation"
461:(which formed the upper surface of the snout). As with other archosaurs and their relatives, the side of the snout has a hole (known as an
676:
416:, which contains all archosaurs more closely related to crocodilians than to birds. Some skull features previously considered unique to
664:, which forms the upper rear portion of the jaw (and the upper border of the mandibular fenestra), was large. Another autapomorphy of
2554:
2531:
1950:
3414:
2546:
1026:
833:. The fossil, however, was not fully prepared. Subsequent analysis by Parrish in 1993 indicated that the fossils represented a
685:
859:) redescribed the fossil. Wu and Russell prepared the fossil, and noted the limb bones (femur and humerus) resembled those of
537:
had a pair of openings at the rear portion of the skull known as temporal fenestra. The bone separating the two holes, the
985:
841:, was described in 1982. However, due to its large intercentra, this species is not considered to be a close relative of
2539:
143:
656:
among gracilisuchids, the lower prong was much longer than the upper prong. This had the added effect of excluding the
359:
had a peculiar combination of features which has made it difficult to classify in the past. It possessed teeth on the
3400:
1362:
906:
748:
3608:
1475:
1049:
614:
does not preclude its classification as a crown-archosaur. Another seemingly non-archosaurian trait reported in
1935:
1170:
Butler, Richard J; Sullivan, Corwin; Ezcurra, Martín D; Liu, Jun; Lecuona, Agustina; Sookias, Roland B (2014).
930:'s comprehensive analysis on archosaurs and archosauriforms. This analysis noted that some archosaurs (such as
3603:
3583:
3467:
2838:
898:
660:(which formed the rear lower edge of the jaw) from forming the lower border of the mandibular fenestra. The
619:
545:
into two separate holes, leaving three holes at the back of the skull in total. This trait is also known is
364:
3484:
2025:
1943:
732:(hip socket). The pubis (lower front plate of the hip) was moderate in length and also similar to that of
542:
3598:
3555:
1355:
856:
622:
enter the braincase from below, rather than from the sides as in almost all other archosaurs (including
346:
345:
in 1973, based on a partially complete but disarticulated fossil skeleton (IVPP V.32237) found in the
3332:
1301:
1183:
457:(nostril holes). This prong, known as a posterodorsal process, bisects a corresponding branch of the
261:
3031:
1981:
1968:
1902:
1834:
1448:
1341:
462:
3407:
3361:
2698:
1317:
1135:
1061:
138:
3560:
384:, and some features of the skull have only been observed in true archosaurs, particularly early
3542:
3534:
3212:
3176:
2688:
1270:
1239:"Systematics of putative euparkeriids (Diapsida: Archosauriformes) from the Triassic of China"
1219:
1201:
1053:
792:
764:
3547:
3257:
3126:
3109:
2886:
2058:
1460:
1309:
1260:
1250:
1209:
1191:
1127:
1117:
1045:
927:
916:
to be a pseudosuchian once again during the study of the recently discovered archosauriform
583:
470:
409:
367:
which entered the braincase from below. These two traits were rare among
Archosauria (true,
122:
3232:
2987:
2855:
2776:
2753:
2728:
2586:
2573:
2498:
2486:
2478:
2130:
1853:
1785:
1597:
912:
In 2010, paleontologists Martín
Ezcurra, Agustina Lecuona, and Augustín Martinelli found
807:
772:
768:
315:
237:
1305:
1187:
586:. However, more recently pterygoid teeth have been reported in crown-archosaurs such as
17:
3437:
3294:
3239:
3197:
3156:
3009:
2980:
2959:
2827:
2708:
2617:
2293:
2095:
2088:
2081:
1812:
1725:
1635:
1562:
1265:
1238:
1214:
1171:
847:
710:, with concavities on their sides and neural spines with expanded tops. There were two
567:
538:
501:
342:
308:
257:
505:
also bisects its nasal, and a posterodorsal process of the maxilla is present in some
441:
was proportionally similar to that of other small generalized archosauriforms such as
3577:
3273:
3187:
3166:
3062:
2950:
2926:
2916:
2876:
2813:
2769:
2762:
2668:
2658:
2637:
2627:
2441:
2426:
2355:
2335:
2242:
2222:
2148:
2067:
2039:
2012:
1862:
1805:
1679:
1621:
955:
943:
918:
830:
632:
592:
510:
474:
398:
80:
1321:
1139:
1069:
1065:
747:(upper arm bone) had a deltopectoral crest which was poorly differentiated from the
3447:
3287:
3225:
3085:
3071:
2973:
2936:
2906:
2799:
2790:
2718:
2456:
2448:
2412:
2396:
2388:
2379:
2365:
2325:
2304:
2262:
2212:
2121:
2102:
2074:
1891:
1798:
1764:
1757:
1750:
1711:
1614:
1606:
1541:
1528:
1487:
1378:
1105:
961:
882:
861:
760:
728:, with a short forward projection, a much larger rear projection, and a large bony
724:
715:
707:
657:
546:
518:
491:
482:
413:
404:
350:
334:
286:
211:
1313:
1106:"The early evolution of archosaurs: relationships and the origin of major clades"
3452:
3321:
3266:
3136:
3093:
3078:
3047:
3040:
3017:
3001:
2994:
2966:
2647:
2419:
2345:
2314:
2282:
2272:
2202:
2171:
2164:
2110:
2047:
1998:
1877:
1734:
1700:
1690:
1668:
1628:
1572:
991:
784:
661:
638:
571:
551:
380:. On the other hand, the ankle was much more advanced than that of animals like
368:
55:
3507:
541:, had a lower branch which curved forwards to contact the jugal and divide the
3457:
3442:
3432:
3367:
3338:
3309:
3146:
2739:
2463:
2434:
2404:
2252:
2232:
2137:
2032:
1433:
1408:
1331:
938:
893:
in the group). This conclusion was supported by features such as teeth on the
886:
825:
780:
729:
702:
644:
602:
596:
526:
458:
450:
389:
376:
100:
65:
1205:
1196:
1057:
909:("advanced") features, such as a calcaneal tuber directed further backwards.
630:). However, this trait is also known to occur in other crown-archosaurs like
3350:
3280:
2678:
1582:
1417:
890:
874:
870:
823:. Young originally believed that the fossils came from an animal similar to
788:
776:
737:
575:
465:) surrounded by a lowered basin of bone (known as an antorbital fossa). The
371:
318:
198:
155:
105:
49:
1274:
1223:
453:(a toothed bone at the tip of the snout) projects a small prong behind the
3501:
3246:
2896:
2806:
2607:
2186:
1823:
1771:
1402:
1290:"A new basal archosauriform diapsid from the Lower Triassic of Argentina"
1031:(Archosauriformes) and new information on its phylogenetic relationships"
932:
923:
693:
588:
506:
322:
175:
95:
90:
75:
70:
60:
3521:
2471:
2157:
1869:
1742:
744:
466:
454:
326:
311:
185:
110:
85:
41:
1255:
1651:
1396:
1131:
894:
834:
796:
711:
559:
385:
360:
224:
165:
3478:
885:(archosaurs closer to crocodilians than to dinosaurs, also known as
1289:
649:
Otherwise, the braincase is generally similar to other archosaurs.
1122:
806:
752:
675:
330:
304:
570:. Traditionally, pterygoid teeth are considered to be absent in
374:) and are more similar to non-archosaurian archosauriforms like
3482:
2571:
1966:
1526:
1389:
1351:
1347:
791:
and suchians). These include a hemicylindrical (barrel-shaped)
477:(which lies between the antorbital fossa and the eye socket).
1237:
Sookias, R. B.; Sullivan, C.; Liu, J.; Butler, R. J. (2014).
566:. One particular similarity is the presence of teeth on the
1050:
10.1671/0272-4634(2001)021[0040:ROTDAA]2.0.CO;2
578:. Wu & Russell (2001) used this to justify classifying
424:
retains a few unique features of the jaw and cheek region.
853:
Institute of
Vertebrate Paleontology and Paleoanthropology
845:, and was likely an indeterminate euparkeriid related to
420:
953:
was found to be closely related to the small suchians
408:, and a 2014 study grouped these three genera in the
3318:
3209:
3193:
3183:
3172:
3162:
3152:
3142:
3132:
3122:
3059:
3028:
2947:
2932:
2922:
2912:
2902:
2892:
2882:
2872:
2824:
2787:
2750:
2735:
2724:
2714:
2704:
2694:
2684:
2674:
2664:
2654:
2643:
2633:
2623:
2613:
2603:
2376:
2361:
2351:
2341:
2331:
2321:
2310:
2300:
2289:
2278:
2268:
2258:
2248:
2238:
2228:
2218:
2208:
2198:
2009:
1994:
1850:
1782:
1722:
1707:
1696:
1686:
1675:
1664:
1594:
1578:
1568:
1558:
529:(cheek bone), on the other hand, is truly unique to
276:
3491:
3349:
3317:
3308:
3256:
3208:
3107:
3058:
3027:
2946:
2853:
2823:
2786:
2749:
2584:
2497:
2375:
2185:
2147:
2120:
2057:
2008:
1979:
1901:
1849:
1822:
1781:
1721:
1649:
1593:
1539:
1486:
1459:
1432:
1288:Ezcurra, M.D.; Lecuona, A.; Martinelli, A. (2010).
722:cannot be proven. The ilium was similar to that of
1110:Bulletin of the American Museum of Natural History
759:and most pseudosuchians, but it lacked the offset
489:However, they are now both known to be present in
926:. This conclusion was also supported in 2011 by
873:did not. Further preparation revealed a partial
779:(heel bone) had many similarities with that of "
562:(roof of the mouth) were generally similar to
1363:
936:) also had palatal teeth, and other (such as
412:Gracilisuchidae. Gracilisuchidae lies within
8:
881:was not a suchian nor even a member of the
3479:
3314:
3217:
3115:
2865:
2861:
2596:
2592:
2581:
2568:
2191:
2017:
1987:
1976:
1963:
1790:
1657:
1551:
1547:
1536:
1523:
1465:
1438:
1429:
1386:
1370:
1356:
1348:
121:
31:
1264:
1254:
1213:
1195:
1121:
1025:Wu, Xiao-Chun; Russell, Anthony (2001).
855:in Beijing) and Anthony Russell (of the
714:(hip) vertebrae, which connected to the
130:Multiple views of the holotype skull of
1099:
1097:
1095:
1093:
1091:
1089:
978:
1020:
7:
1165:
1163:
1161:
1159:
1157:
1155:
1153:
1151:
1149:
1018:
1016:
1014:
1012:
1010:
1008:
1006:
1004:
1002:
1000:
783:" archosauriforms (i.e. ones with a
755:(thigh bone) was similar to that of
1294:Journal of Vertebrate Paleontology
1038:Journal of Vertebrate Paleontology
485:(unique distinguishing traits) of
396:shares the most similarities with
25:
3624:Prehistoric pseudosuchian genera
3594:Middle Triassic reptiles of Asia
3463:
3462:
3413:
3406:
3399:
2553:
2545:
2538:
2530:
1949:
1942:
1934:
1334:
877:. Wu and Russell concluded that
851:. In 2001, Xiao-Chun Wu (of the
829:, and assigned it to the family
321:, which lived during the Middle
142:
686:Paleozoological Museum of China
610:. Therefore, their presence in
3589:Middle Triassic pseudosuchians
889:in publications which include
1:
3614:Fossil taxa described in 1973
837:. A putative second species,
388:(distant ancestors of modern
3619:Taxa named by Yang Zhongjian
1314:10.1080/02724634.2010.501446
839:"Turfanosuchus" shageduensis
3640:
558:Preserved portions of the
3427:
3396:
3220:
3118:
2868:
2864:
2599:
2595:
2580:
2567:
2527:
2194:
2020:
1990:
1975:
1962:
1931:
1793:
1660:
1554:
1550:
1535:
1522:
1468:
1441:
1428:
1385:
899:internal carotid arteries
700:were similar to those of
620:internal carotid arteries
495:and (to a lesser extent)
365:internal carotid arteries
273:
268:
139:Scientific classification
137:
129:
120:
34:
27:Extinct genus of reptiles
1197:10.1186/1471-2148-14-128
1176:BMC Evolutionary Biology
1029:Turfanosuchus dabanensis
682:Turfanosuchus dabanensis
132:Turfanosuchus dabanensis
18:Turfanosuchus dabanensis
543:lower temporal fenestra
2026:Benggwigwishingasuchus
1104:Nesbitt, S.J. (2011).
815:The classification of
812:
689:
392:). Out of this group,
3556:Paleobiology Database
857:University of Calgary
810:
679:
618:is the fact that its
684:, on display at the
672:Postcranial skeleton
1982:Paracrocodylomorpha
1969:Paracrocodylomorpha
1903:Paracrocodylomorpha
1342:Paleontology portal
1306:2010JVPal..30.1433E
1188:2014BMCEE..14..128B
582:as a non-archosaur
525:. The shape of the
463:antorbital fenestra
341:, was described by
2699:Pseudhesperosuchus
1027:"Redescription of
813:
690:
347:Kelamayi Formation
329:) of northwestern
3571:
3570:
3543:Open Tree of Life
3485:Taxon identifiers
3476:
3475:
3423:
3422:
3394:
3393:
3390:
3389:
3386:
3385:
3382:
3381:
3378:
3377:
3304:
3303:
3213:Shartegosuchoidea
3177:Pachycheilosuchus
3103:
3102:
2849:
2848:
2689:Phyllodontosuchus
2563:
2562:
2525:
2524:
2521:
2520:
2517:
2516:
2181:
2180:
1958:
1957:
1929:
1928:
1925:
1924:
1921:
1920:
1845:
1844:
1645:
1644:
1518:
1517:
1514:
1513:
1510:
1509:
1506:
1505:
1256:10.7717/peerj.658
905:, noting several
769:avemetatarsalians
765:fourth trochanter
296:
295:
290:
264:
16:(Redirected from
3631:
3609:Fossils of China
3564:
3563:
3551:
3550:
3538:
3537:
3525:
3524:
3512:
3511:
3510:
3480:
3466:
3465:
3418:
3417:
3410:
3404:
3403:
3320:
3315:
3258:Shartegosuchidae
3218:
3211:
3195:
3185:
3174:
3164:
3154:
3144:
3134:
3127:Dianchungosaurus
3124:
3116:
3110:Mesoeucrocodylia
3061:
3030:
2949:
2934:
2924:
2914:
2904:
2894:
2887:Eopneumatosuchus
2884:
2874:
2866:
2862:
2826:
2789:
2752:
2737:
2726:
2716:
2706:
2696:
2686:
2676:
2666:
2656:
2645:
2635:
2625:
2615:
2605:
2597:
2593:
2582:
2569:
2558:
2557:
2550:
2549:
2543:
2542:
2535:
2534:
2378:
2363:
2353:
2343:
2333:
2323:
2312:
2302:
2291:
2280:
2270:
2260:
2250:
2240:
2230:
2220:
2210:
2200:
2192:
2059:Ctenosauriscidae
2018:
2011:
1996:
1988:
1977:
1964:
1953:
1947:
1946:
1939:
1938:
1852:
1791:
1784:
1724:
1709:
1698:
1688:
1677:
1666:
1658:
1596:
1580:
1570:
1560:
1552:
1548:
1537:
1524:
1476:Archosauromorpha
1466:
1461:Archosauromorpha
1439:
1430:
1423:
1422:
1387:
1372:
1365:
1358:
1349:
1344:
1339:
1338:
1337:
1326:
1325:
1300:(5): 1433–1450.
1285:
1279:
1278:
1268:
1258:
1234:
1228:
1227:
1217:
1199:
1167:
1144:
1143:
1125:
1101:
1084:
1083:
1081:
1080:
1074:
1068:. Archived from
1035:
1022:
995:
983:
928:Sterling Nesbitt
471:archosauromorphs
284:
278:
256:
249:
236:
147:
146:
125:
115:
52:
40:Temporal range:
32:
21:
3639:
3638:
3634:
3633:
3632:
3630:
3629:
3628:
3574:
3573:
3572:
3567:
3559:
3554:
3546:
3541:
3533:
3528:
3520:
3515:
3506:
3505:
3500:
3487:
3477:
3472:
3419:
3412:
3398:
3374:
3345:
3300:
3252:
3233:Shantungosuchus
3204:
3112:
3099:
3054:
3023:
2988:Hemiprotosuchus
2942:
2858:
2856:Crocodyliformes
2845:
2819:
2782:
2777:Terrestrisuchus
2754:Saltoposuchidae
2745:
2729:Syntomiprosopus
2589:
2587:Crocodylomorpha
2576:
2574:Crocodylomorpha
2559:
2552:
2544:
2537:
2529:
2513:
2499:Crocodylomorpha
2493:
2487:Vytshegdosuchus
2479:Vjushkovisaurus
2371:
2177:
2143:
2131:Dolichobrachium
2116:
2053:
2004:
1984:
1971:
1954:
1941:
1933:
1917:
1897:
1854:Gracilisuchidae
1841:
1818:
1786:Aetosauriformes
1777:
1717:
1654:
1641:
1598:Ornithosuchidae
1589:
1544:
1531:
1502:
1482:
1455:
1424:
1392:
1391:
1381:
1376:
1340:
1335:
1333:
1330:
1329:
1287:
1286:
1282:
1236:
1235:
1231:
1169:
1168:
1147:
1103:
1102:
1087:
1078:
1076:
1072:
1033:
1024:
1023:
998:
984:
980:
975:
805:
773:crocodylomorphs
680:A sculpture of
674:
435:
430:
255:
247:
238:Gracilisuchidae
234:
141:
116:
114:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
48:247.2–242
47:
46:
44:
38:
28:
23:
22:
15:
12:
11:
5:
3637:
3635:
3627:
3626:
3621:
3616:
3611:
3606:
3604:Triassic China
3601:
3596:
3591:
3586:
3584:Gracilisuchids
3576:
3575:
3569:
3568:
3566:
3565:
3552:
3539:
3526:
3513:
3497:
3495:
3489:
3488:
3483:
3474:
3473:
3471:
3470:
3460:
3455:
3450:
3445:
3440:
3438:Prestosuchidae
3435:
3428:
3425:
3424:
3421:
3420:
3397:
3395:
3392:
3391:
3388:
3387:
3384:
3383:
3380:
3379:
3376:
3375:
3373:
3372:
3371:
3370:
3364:
3355:
3353:
3347:
3346:
3344:
3343:
3342:
3341:
3335:
3326:
3324:
3312:
3306:
3305:
3302:
3301:
3299:
3298:
3295:Shartegosuchus
3291:
3284:
3277:
3270:
3262:
3260:
3254:
3253:
3251:
3250:
3243:
3240:Sichuanosuchus
3236:
3229:
3221:
3215:
3206:
3205:
3203:
3202:
3198:Thalattosuchia
3191:
3181:
3170:
3160:
3157:Lusitanisuchus
3150:
3140:
3130:
3119:
3113:
3108:
3105:
3104:
3101:
3100:
3098:
3097:
3090:
3082:
3075:
3067:
3065:
3056:
3055:
3053:
3052:
3044:
3036:
3034:
3032:Notochampsidae
3025:
3024:
3022:
3021:
3014:
3010:Pedeticosaurus
3006:
2998:
2991:
2984:
2981:Erythrochampsa
2977:
2970:
2963:
2960:Coloradisuchus
2955:
2953:
2944:
2943:
2941:
2940:
2930:
2920:
2910:
2900:
2890:
2880:
2869:
2859:
2854:
2851:
2850:
2847:
2846:
2844:
2843:
2842:
2841:
2839:Thalattosuchia
2832:
2830:
2828:Thalattosuchia
2821:
2820:
2818:
2817:
2810:
2803:
2795:
2793:
2784:
2783:
2781:
2780:
2773:
2766:
2758:
2756:
2747:
2746:
2744:
2743:
2733:
2722:
2712:
2709:Redondavenator
2702:
2692:
2682:
2672:
2662:
2652:
2641:
2631:
2621:
2618:Dibothrosuchus
2611:
2600:
2590:
2585:
2578:
2577:
2572:
2565:
2564:
2561:
2560:
2528:
2526:
2523:
2522:
2519:
2518:
2515:
2514:
2512:
2511:
2510:
2509:
2503:
2501:
2495:
2494:
2492:
2491:
2483:
2475:
2468:
2460:
2453:
2445:
2438:
2431:
2423:
2416:
2409:
2401:
2393:
2384:
2382:
2373:
2372:
2370:
2369:
2359:
2349:
2339:
2329:
2319:
2308:
2298:
2294:Incertovenator
2287:
2276:
2266:
2256:
2246:
2236:
2226:
2216:
2206:
2195:
2189:
2183:
2182:
2179:
2178:
2176:
2175:
2168:
2161:
2153:
2151:
2145:
2144:
2142:
2141:
2134:
2126:
2124:
2118:
2117:
2115:
2114:
2107:
2099:
2096:Hypselorhachis
2092:
2089:Ctenosauriscus
2085:
2082:Bystrowisuchus
2078:
2071:
2063:
2061:
2055:
2054:
2052:
2051:
2044:
2036:
2029:
2021:
2015:
2006:
2005:
2003:
2002:
1991:
1985:
1980:
1973:
1972:
1967:
1960:
1959:
1956:
1955:
1932:
1930:
1927:
1926:
1923:
1922:
1919:
1918:
1916:
1915:
1914:
1913:
1907:
1905:
1899:
1898:
1896:
1895:
1888:
1881:
1874:
1866:
1858:
1856:
1847:
1846:
1843:
1842:
1840:
1839:
1838:
1837:
1828:
1826:
1820:
1819:
1817:
1816:
1813:Revueltosaurus
1809:
1802:
1794:
1788:
1779:
1778:
1776:
1775:
1768:
1761:
1754:
1747:
1739:
1730:
1728:
1726:Erpetosuchidae
1719:
1718:
1716:
1715:
1705:
1694:
1684:
1673:
1661:
1655:
1650:
1647:
1646:
1643:
1642:
1640:
1639:
1636:Venaticosuchus
1632:
1625:
1618:
1611:
1602:
1600:
1591:
1590:
1588:
1587:
1576:
1566:
1563:Dasygnathoides
1555:
1545:
1540:
1533:
1532:
1527:
1520:
1519:
1516:
1515:
1512:
1511:
1508:
1507:
1504:
1503:
1501:
1500:
1499:
1498:
1492:
1490:
1484:
1483:
1481:
1480:
1479:
1478:
1469:
1463:
1457:
1456:
1454:
1453:
1452:
1451:
1442:
1436:
1426:
1425:
1421:
1420:
1411:
1405:
1399:
1390:
1383:
1382:
1377:
1375:
1374:
1367:
1360:
1352:
1346:
1345:
1328:
1327:
1280:
1229:
1145:
1085:
996:
977:
976:
974:
971:
848:Halazhaisuchus
804:
803:Classification
801:
708:pseudosuchians
673:
670:
584:archosauriform
568:pterygoid bone
502:Revueltosaurus
487:Turfanosuchus.
483:autapomorphies
434:
431:
429:
426:
309:archosauriform
294:
293:
292:
291:
271:
270:
266:
265:
245:
241:
240:
232:
228:
227:
222:
215:
214:
209:
202:
201:
196:
189:
188:
183:
179:
178:
173:
169:
168:
163:
159:
158:
153:
149:
148:
135:
134:
127:
126:
118:
117:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
53:
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
3636:
3625:
3622:
3620:
3617:
3615:
3612:
3610:
3607:
3605:
3602:
3600:
3597:
3595:
3592:
3590:
3587:
3585:
3582:
3581:
3579:
3562:
3557:
3553:
3549:
3544:
3540:
3536:
3531:
3527:
3523:
3518:
3514:
3509:
3503:
3499:
3498:
3496:
3494:
3493:Turfanosuchus
3490:
3486:
3481:
3469:
3461:
3459:
3456:
3454:
3451:
3449:
3446:
3444:
3441:
3439:
3436:
3434:
3430:
3429:
3426:
3416:
3411:
3409:
3402:
3369:
3365:
3363:
3359:
3358:
3357:
3356:
3354:
3352:
3348:
3340:
3336:
3334:
3330:
3329:
3328:
3327:
3325:
3323:
3316:
3313:
3311:
3307:
3297:
3296:
3292:
3290:
3289:
3285:
3283:
3282:
3278:
3276:
3275:
3274:Fruitachampsa
3271:
3269:
3268:
3264:
3263:
3261:
3259:
3255:
3249:
3248:
3244:
3242:
3241:
3237:
3235:
3234:
3230:
3228:
3227:
3223:
3222:
3219:
3216:
3214:
3207:
3200:
3199:
3192:
3190:
3189:
3188:Titanochampsa
3182:
3179:
3178:
3171:
3169:
3168:
3167:Neuquensuchus
3161:
3159:
3158:
3151:
3149:
3148:
3141:
3139:
3138:
3131:
3129:
3128:
3121:
3120:
3117:
3114:
3111:
3106:
3096:
3095:
3091:
3088:
3087:
3083:
3081:
3080:
3076:
3074:
3073:
3069:
3068:
3066:
3064:
3063:Gobiosuchidae
3057:
3050:
3049:
3045:
3043:
3042:
3038:
3037:
3035:
3033:
3026:
3020:
3019:
3015:
3012:
3011:
3007:
3004:
3003:
2999:
2997:
2996:
2992:
2990:
2989:
2985:
2983:
2982:
2978:
2976:
2975:
2971:
2969:
2968:
2964:
2962:
2961:
2957:
2956:
2954:
2952:
2951:Protosuchidae
2945:
2939:
2938:
2931:
2929:
2928:
2927:Stegomosuchus
2921:
2919:
2918:
2917:Platyognathus
2911:
2909:
2908:
2901:
2899:
2898:
2891:
2889:
2888:
2881:
2879:
2878:
2877:Entradasuchus
2871:
2870:
2867:
2863:
2860:
2857:
2852:
2840:
2836:
2835:
2834:
2833:
2831:
2829:
2822:
2816:
2815:
2814:Macelognathus
2811:
2809:
2808:
2804:
2802:
2801:
2797:
2796:
2794:
2792:
2785:
2779:
2778:
2774:
2772:
2771:
2770:Saltoposuchus
2767:
2765:
2764:
2763:Litargosuchus
2760:
2759:
2757:
2755:
2748:
2742:
2741:
2734:
2731:
2730:
2723:
2721:
2720:
2713:
2711:
2710:
2703:
2701:
2700:
2693:
2691:
2690:
2683:
2681:
2680:
2673:
2671:
2670:
2669:Kayentasuchus
2663:
2661:
2660:
2659:Junggarsuchus
2653:
2650:
2649:
2642:
2640:
2639:
2638:Hesperosuchus
2632:
2630:
2629:
2628:Dromicosuchus
2622:
2620:
2619:
2612:
2610:
2609:
2602:
2601:
2598:
2594:
2591:
2588:
2583:
2579:
2575:
2570:
2566:
2556:
2551:
2548:
2541:
2533:
2507:
2506:
2505:
2504:
2502:
2500:
2496:
2489:
2488:
2484:
2481:
2480:
2476:
2474:
2473:
2469:
2466:
2465:
2461:
2459:
2458:
2454:
2451:
2450:
2446:
2444:
2443:
2442:Scolotosuchus
2439:
2437:
2436:
2432:
2429:
2428:
2427:Procerosuchus
2424:
2422:
2421:
2417:
2415:
2414:
2410:
2407:
2406:
2402:
2399:
2398:
2394:
2391:
2390:
2386:
2385:
2383:
2381:
2374:
2368:
2367:
2360:
2358:
2357:
2356:Stagonosuchus
2350:
2348:
2347:
2340:
2338:
2337:
2336:Procerosuchus
2330:
2328:
2327:
2320:
2317:
2316:
2309:
2307:
2306:
2299:
2296:
2295:
2288:
2285:
2284:
2277:
2275:
2274:
2267:
2265:
2264:
2257:
2255:
2254:
2247:
2245:
2244:
2243:Decuriasuchus
2237:
2235:
2234:
2227:
2225:
2224:
2223:Batrachotomus
2217:
2215:
2214:
2207:
2205:
2204:
2197:
2196:
2193:
2190:
2188:
2184:
2174:
2173:
2169:
2167:
2166:
2162:
2160:
2159:
2155:
2154:
2152:
2150:
2149:Shuvosauridae
2146:
2140:
2139:
2135:
2133:
2132:
2128:
2127:
2125:
2123:
2119:
2113:
2112:
2108:
2105:
2104:
2100:
2098:
2097:
2093:
2091:
2090:
2086:
2084:
2083:
2079:
2077:
2076:
2072:
2070:
2069:
2068:Arizonasaurus
2065:
2064:
2062:
2060:
2056:
2050:
2049:
2045:
2042:
2041:
2040:Schultzsuchus
2037:
2035:
2034:
2030:
2028:
2027:
2023:
2022:
2019:
2016:
2014:
2013:Poposauroidea
2007:
2001:
2000:
1993:
1992:
1989:
1986:
1983:
1978:
1974:
1970:
1965:
1961:
1952:
1948:
1945:
1937:
1911:
1910:
1909:
1908:
1906:
1904:
1900:
1894:
1893:
1889:
1887:
1886:
1885:Turfanosuchus
1882:
1880:
1879:
1875:
1872:
1871:
1867:
1865:
1864:
1863:Gracilisuchus
1860:
1859:
1857:
1855:
1848:
1836:
1832:
1831:
1830:
1829:
1827:
1825:
1821:
1815:
1814:
1810:
1808:
1807:
1806:Euscolosuchus
1803:
1801:
1800:
1796:
1795:
1792:
1789:
1787:
1780:
1774:
1773:
1769:
1767:
1766:
1762:
1760:
1759:
1755:
1753:
1752:
1748:
1745:
1744:
1740:
1737:
1736:
1732:
1731:
1729:
1727:
1720:
1714:
1713:
1706:
1703:
1702:
1695:
1693:
1692:
1685:
1682:
1681:
1680:Koilamasuchus
1674:
1671:
1670:
1663:
1662:
1659:
1656:
1653:
1648:
1638:
1637:
1633:
1631:
1630:
1626:
1624:
1623:
1622:Ornithosuchus
1619:
1617:
1616:
1612:
1609:
1608:
1604:
1603:
1601:
1599:
1592:
1585:
1584:
1577:
1575:
1574:
1567:
1565:
1564:
1557:
1556:
1553:
1549:
1546:
1543:
1538:
1534:
1530:
1525:
1521:
1496:
1495:
1494:
1493:
1491:
1489:
1485:
1477:
1473:
1472:
1471:
1470:
1467:
1464:
1462:
1458:
1450:
1446:
1445:
1444:
1443:
1440:
1437:
1435:
1431:
1427:
1419:
1415:
1412:
1410:
1406:
1404:
1400:
1398:
1394:
1393:
1388:
1384:
1380:
1373:
1368:
1366:
1361:
1359:
1354:
1353:
1350:
1343:
1332:
1323:
1319:
1315:
1311:
1307:
1303:
1299:
1295:
1291:
1284:
1281:
1276:
1272:
1267:
1262:
1257:
1252:
1248:
1244:
1240:
1233:
1230:
1225:
1221:
1216:
1211:
1207:
1203:
1198:
1193:
1189:
1185:
1181:
1177:
1173:
1166:
1164:
1162:
1160:
1158:
1156:
1154:
1152:
1150:
1146:
1141:
1137:
1133:
1129:
1124:
1123:10.1206/352.1
1119:
1115:
1111:
1107:
1100:
1098:
1096:
1094:
1092:
1090:
1086:
1075:on 2011-07-16
1071:
1067:
1063:
1059:
1055:
1051:
1047:
1043:
1039:
1032:
1030:
1021:
1019:
1017:
1015:
1013:
1011:
1009:
1007:
1005:
1003:
1001:
997:
993:
989:
988:
987:Turfanosuchus
982:
979:
972:
970:
968:
967:Turfanosuchus
964:
963:
958:
957:
956:Gracilisuchus
952:
951:Turfanosuchus
947:
945:
941:
940:
935:
934:
929:
925:
921:
920:
919:Koilamasuchus
915:
914:Turfanosuchus
910:
908:
904:
900:
896:
892:
888:
884:
880:
879:Turfanosuchus
876:
872:
869:, though the
868:
864:
863:
858:
854:
850:
849:
844:
843:Turfanosuchus
840:
836:
832:
831:Euparkeriidae
828:
827:
822:
818:
817:Turfanosuchus
809:
802:
800:
798:
794:
790:
786:
782:
778:
774:
770:
766:
762:
758:
754:
750:
746:
741:
739:
735:
731:
727:
726:
721:
720:Turfanosuchus
717:
713:
709:
705:
704:
699:
698:Turfanosuchus
695:
687:
683:
678:
671:
669:
667:
666:Turfanosuchus
663:
659:
655:
654:Turfanosuchus
650:
648:
646:
641:
640:
635:
634:
633:Arizonasaurus
629:
625:
624:Gracilisuchus
621:
617:
616:Turfanosuchus
613:
612:Turfanosuchus
609:
605:
604:
600:
598:
594:
593:Eudimorphodon
590:
585:
581:
580:Turfanosuchus
577:
573:
569:
565:
561:
556:
554:
553:
548:
544:
540:
536:
535:Turfanosuchus
532:
531:Turfanosuchus
528:
524:
523:Gracilisuchus
520:
519:frontal bones
514:
512:
508:
504:
503:
498:
497:Gracilisuchus
494:
493:
488:
484:
478:
476:
475:lacrimal bone
472:
468:
464:
460:
456:
452:
448:
447:Gracilisuchus
444:
440:
439:Turfanosuchus
437:The skull of
432:
427:
425:
423:
422:Turfanosuchus
419:
418:Turfanosuchus
415:
411:
407:
406:
401:
400:
399:Gracilisuchus
395:
394:Turfanosuchus
391:
387:
383:
379:
378:
373:
370:
366:
362:
358:
357:Turfanosuchus
354:
352:
348:
344:
340:
339:T. dabanensis
336:
332:
328:
324:
320:
317:
316:gracilisuchid
313:
310:
306:
302:
301:
300:Turfanosuchus
288:
285:Young, 1973 (
283:
282:
281:T. dabanensis
275:
274:
272:
267:
263:
259:
254:
253:
252:Turfanosuchus
246:
243:
242:
239:
233:
230:
229:
226:
223:
220:
217:
216:
213:
210:
207:
204:
203:
200:
197:
194:
191:
190:
187:
184:
181:
180:
177:
174:
171:
170:
167:
164:
161:
160:
157:
154:
151:
150:
145:
140:
136:
133:
128:
124:
119:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
51:
43:
37:
36:Turfanosuchus
33:
30:
19:
3599:Anisian life
3492:
3448:Sphenosuchia
3405:
3368:crocodilians
3339:notosuchians
3293:
3288:Nominosuchus
3286:
3279:
3272:
3265:
3245:
3238:
3231:
3226:Hadongsuchus
3224:
3196:
3186:
3175:
3165:
3155:
3145:
3135:
3125:
3092:
3086:Lisboasaurus
3084:
3077:
3072:Cassissuchus
3070:
3046:
3039:
3016:
3008:
3000:
2993:
2986:
2979:
2974:Edentosuchus
2972:
2965:
2958:
2937:Tagarosuchus
2935:
2925:
2915:
2907:Microchampsa
2905:
2895:
2885:
2875:
2812:
2805:
2800:Almadasuchus
2798:
2791:Hallopodidae
2775:
2768:
2761:
2738:
2727:
2719:Sphenosuchus
2717:
2707:
2697:
2687:
2677:
2667:
2657:
2646:
2636:
2626:
2616:
2606:
2536:
2485:
2477:
2470:
2462:
2457:Teratosaurus
2455:
2449:Scythosuchus
2447:
2440:
2433:
2425:
2418:
2413:Polonosuchus
2411:
2403:
2397:Energosuchus
2395:
2389:Arganasuchus
2387:
2380:Rauisuchidae
2366:Youngosuchus
2364:
2354:
2344:
2334:
2326:Prestosuchus
2324:
2313:
2305:Luperosuchus
2303:
2292:
2281:
2271:
2263:Fasolasuchus
2261:
2251:
2241:
2231:
2221:
2213:Arganasuchus
2211:
2201:
2170:
2163:
2156:
2136:
2129:
2122:Poposauridae
2109:
2103:Tsylmosuchus
2101:
2094:
2087:
2080:
2075:Bromsgroveia
2073:
2066:
2046:
2038:
2031:
2024:
1997:
1940:
1892:Yonghesuchus
1890:
1884:
1883:
1876:
1868:
1861:
1811:
1804:
1799:Acaenasuchus
1797:
1770:
1765:Parringtonia
1763:
1758:Pagosvenator
1756:
1751:Erpetosuchus
1749:
1741:
1733:
1712:Ticinosuchus
1710:
1699:
1689:
1678:
1667:
1634:
1627:
1620:
1615:Dynamosuchus
1613:
1607:Aenigmaspina
1605:
1581:
1571:
1561:
1542:Pseudosuchia
1529:Pseudosuchia
1488:Pseudosuchia
1413:
1379:Pseudosuchia
1297:
1293:
1283:
1246:
1242:
1232:
1179:
1175:
1113:
1109:
1077:. Retrieved
1070:the original
1044:(1): 40–50.
1041:
1037:
1028:
986:
981:
966:
962:Yonghesuchus
960:
954:
950:
948:
944:poposauroids
937:
931:
917:
913:
911:
902:
883:Pseudosuchia
878:
866:
862:Ticinosuchus
860:
846:
842:
838:
824:
820:
816:
814:
761:femoral head
756:
749:humeral head
742:
734:Ticinosuchus
733:
725:Ticinosuchus
723:
719:
701:
697:
691:
681:
665:
658:angular bone
653:
651:
643:
637:
631:
628:Yonghesuchus
627:
623:
615:
611:
608:Yonghesuchus
607:
601:
587:
579:
563:
557:
550:
534:
530:
522:
515:
511:poposauroids
500:
496:
492:Yonghesuchus
490:
486:
479:
446:
442:
438:
436:
421:
417:
414:Pseudosuchia
405:Yonghesuchus
403:
397:
393:
390:crocodilians
381:
375:
356:
355:
351:Turfan Basin
338:
335:type species
299:
298:
297:
280:
279:
251:
250:
218:
212:Pseudosuchia
205:
192:
131:
35:
29:
3453:Protosuchia
3333:Ziphosuchia
3322:Ziphosuchia
3267:Adzhosuchus
3137:Hsisosuchus
3094:Zaraasuchus
3079:Gobiosuchus
3048:Orthosuchus
3041:Notochampsa
3018:Protosuchus
3002:Orthosuchus
2995:Hoplosuchus
2967:Dianosuchus
2648:Heteropelta
2420:Postosuchus
2346:Saurosuchus
2315:Mandasuchus
2283:Heteropelta
2273:Heptasuchus
2203:Apatosuchus
2172:Sillosuchus
2165:Shuvosaurus
2111:Xilousuchus
2048:Qianosuchus
1999:Mambawakale
1878:Parvosuchus
1735:Archeopelta
1701:Nundasuchus
1691:Mandasuchus
1669:Heteropelta
1629:Riojasuchus
1583:Phytosauria
1573:Nundasuchus
1418:Archosauria
992:Fossilworks
811:Restoration
787:ankle like
785:crurotarsal
781:crurotarsan
763:and raised
639:Qianosuchus
552:Postosuchus
547:rauisuchids
459:nasal bones
428:Description
369:crown-group
314:, likely a
199:Archosauria
3578:Categories
3458:Mesosuchia
3443:Bathyotica
3433:Rauisuchia
3431:See also:
3310:Metasuchia
3147:Induszalim
2740:Trialestes
2508:see below↓
2464:Tikisuchus
2435:Rauisuchus
2405:Jushatyria
2253:Etjosuchus
2233:Dagasuchus
2138:Poposaurus
2033:Lotosaurus
1912:see below↓
1835:Aetosauria
1824:Aetosauria
1497:see below↓
1449:Sauropsida
1434:Sauropsida
1409:Sauropsida
1182:(1): 128.
1079:2008-06-08
973:References
942:and early
939:Silesaurus
907:apomorphic
903:Euparkeria
891:phytosaurs
887:Crurotarsi
867:Euparkeria
826:Euparkeria
821:Euparkeria
789:phytosaurs
757:Euparkeria
738:Osteoderms
730:acetabulum
703:Euparkeria
662:surangular
645:Silesaurus
603:Eodromaeus
597:Lewisuchus
576:archosaurs
564:Euparkeria
451:premaxilla
443:Euparkeria
382:Euparkeria
377:Euparkeria
372:archosaurs
343:C.C. Young
3366:includes
3362:Neosuchia
3351:Neosuchia
3337:includes
3281:Kyasuchus
2679:Parrishia
1395:Kingdom:
1206:1471-2148
1132:2246/6112
1116:: 1–292.
1058:0272-4634
949:In 2014,
924:aetosaurs
875:osteoderm
871:calcaneum
777:calcaneum
771:or early
694:vertebrae
539:squamosal
507:aetosaurs
319:archosaur
162:Kingdom:
156:Eukaryota
3508:Q1838372
3502:Wikidata
3468:Category
3247:Zosuchus
2897:Kemkemia
2807:Hallopus
2608:Carnufex
2187:Loricata
1772:Tarjadia
1403:Chordata
1401:Phylum:
1397:Animalia
1322:83826840
1275:25469319
1249:: e658.
1224:24916124
1140:83493714
1066:56213383
933:Eoraptor
795:for the
589:Eoraptor
549:such as
386:suchians
323:Triassic
269:Species
231:Family:
186:Reptilia
176:Chordata
172:Phylum:
166:Animalia
152:Domain:
3548:4127073
3535:1161419
3522:4818074
2472:Vivaron
2158:Effigia
1870:Maehary
1743:Dyoplax
1407:Class:
1302:Bibcode
1266:4250070
1215:4061117
1184:Bibcode
835:suchian
793:condyle
745:humerus
467:maxilla
349:of the
327:Anisian
312:reptile
244:Genus:
182:Class:
42:Anisian
3561:100513
1652:Suchia
1320:
1273:
1263:
1222:
1212:
1204:
1138:
1064:
1056:
895:palate
797:fibula
775:. The
751:. The
712:sacral
642:, and
606:, and
560:palate
449:. The
410:family
363:, and
361:palate
333:. The
225:Suchia
3530:IRMNG
1414:Clade
1318:S2CID
1243:PeerJ
1136:S2CID
1073:(PDF)
1062:S2CID
1034:(PDF)
753:femur
716:ilium
572:crown
527:jugal
455:nares
433:Skull
331:China
305:genus
303:is a
258:Young
219:Clade
206:Clade
193:Clade
3517:GBIF
3360:see
3331:see
2837:see
1833:see
1474:see
1447:see
1271:PMID
1220:PMID
1202:ISSN
1054:ISSN
994:.org
959:and
897:and
865:and
743:The
706:and
692:The
626:and
509:and
445:and
402:and
287:type
262:1973
56:PreꞒ
1310:doi
1261:PMC
1251:doi
1210:PMC
1192:doi
1128:hdl
1118:doi
1114:352
1046:doi
990:at
767:of
696:of
307:of
3580::
3558::
3545::
3532::
3519::
3504::
1416::
1316:.
1308:.
1298:30
1296:.
1292:.
1269:.
1259:.
1245:.
1241:.
1218:.
1208:.
1200:.
1190:.
1180:14
1178:.
1174:.
1148:^
1134:.
1126:.
1112:.
1108:.
1088:^
1060:.
1052:.
1042:21
1040:.
1036:.
999:^
969:.
736:.
636:,
595:,
591:,
513:.
353:.
337:,
260:,
221::
208::
195::
106:Pg
50:Ma
3319:†
3210:†
3201:?
3194:†
3184:†
3180:?
3173:†
3163:†
3153:†
3143:†
3133:†
3123:†
3089:?
3060:†
3051:?
3029:†
3013:?
3005:?
2948:†
2933:†
2923:†
2913:†
2903:†
2893:†
2883:†
2873:†
2825:†
2788:†
2751:†
2736:†
2732:?
2725:†
2715:†
2705:†
2695:†
2685:†
2675:†
2665:†
2655:†
2651:?
2644:†
2634:†
2624:†
2614:†
2604:†
2490:?
2482:?
2467:?
2452:?
2430:?
2408:?
2400:?
2392:?
2377:†
2362:†
2352:†
2342:†
2332:†
2322:†
2318:?
2311:†
2301:†
2297:?
2290:†
2286:?
2279:†
2269:†
2259:†
2249:†
2239:†
2229:†
2219:†
2209:†
2199:†
2106:?
2043:?
2010:†
1995:†
1873:?
1851:†
1783:†
1746:?
1738:?
1723:†
1708:†
1704:?
1697:†
1687:†
1683:?
1676:†
1672:?
1665:†
1610:?
1595:†
1586:?
1579:†
1569:†
1559:†
1371:e
1364:t
1357:v
1324:.
1312::
1304::
1277:.
1253::
1247:2
1226:.
1194::
1186::
1142:.
1130::
1120::
1082:.
1048::
688:.
647:.
599:,
574:-
325:(
289:)
277:†
248:†
235:†
111:N
101:K
96:J
91:T
86:P
81:C
76:D
71:S
66:O
61:Ꞓ
45:~
20:)
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