492:
125:
104:
563:
sides, extending back to partly cover the seventh segment, with a subrectangular shape when viewed from above and subtriangular shape when seen from the side. Gastric ridges are present on either side of the carapace, extending from the base of the
1812:"Archaeostomatopodea (Malacostraca, Hoplocarida) aus dem Namur B (höheres Marsdenium, Karbon) von Hagen-Vorhalle (NRW, Deutschland) und eine Neudefinition einiger Arten und Famile Tyrannophontidaae"
1550:
on these appendages would have prevented the animal from lifting the front of its body while standing, and its short walking appendages make a wide stance impossible. This further suggests
398:, and thus was given the specific name meaning "gigantic" in Greek. As the tail fan is not preserved, this species may actually belong in a separate genus, but is prudently placed in
579:(walking appendages), and a back part of three larger segments. The back part of the thorax is not covered by the carapace. The entire thorax is slightly shorter than the
2341:
534:
has a carapace length of 27 mm (1.1 in) and the entire living animal likely measured 135 mm (5.3 in) long, comparable in size to extant
1558:
hunt primarily while swimming, and it has been suggested that research into mantis shrimp larvae is needed to better understand to lifestyle of early
2287:
1874:
Smith, C.P.A.; Aubier, P.; Charbonnier, S.; Laville, T.; Olivier, N.; Escarguel, G.; Jenks, J.F.; Bylund, K.G.; Fara, E.; Brayard, A. (2023-03-31).
1590:
period, around 309 million years ago. At the time of deposition, this locality would have been located at a latitude between 4 and 10° south of the
591:
is ovate and has a pair of moveable spines a bit over halfway along its length, as well as a differentset of spines nearer the base of the telson.
2356:
2351:
1546:
animal, swimming above the seabed while it hunted, and perhaps grabbing prey from above using its raptorial appendages. The presence of large
2220:
2186:
511:
587:
is slightly longer than the one immediately in front of it, with the first segment being the shortest and the sixth being the longest. The
2361:
461:
1835:"Palaeo- and archaeostomatopods (Hoplocarida, Crustacea) from the Bear Gulch Limestone, Mississippian (Namurian), of central Montana"
360:
313:, was also named by Schram in 2007. Two other species were formerly assigned to the genus, but have since been reclassified.
386:, was described in 2007 by Frederick Schram. It is known from only the holotype (PE 36987), which preserves the head and
124:
394:
of the Mazon Creek fossil beds in
Illinois. This species is significantly larger than even the largest specimens of
556:
387:
1583:
379:
have also been collected from the Mazon Creek fossil beds, though Schram noted that the species was quite rare.
555:
are large and oval in shape, attached to the head by stalks. The smooth carapace envelops the front part of the
491:
2239:
Pennsylvanian invertebrates of the Mazon Creek Area, Illinois: the morphology and affinities of
Tullimonstrum
1579:
615:. Schram used the same classification again in 2007 in a study revising the morphology and relationships of
352:
286:
603:, Frederick Schram also erected the family Tyrannophontidae, to which the genus is assigned. The family is
2255:
2166:
612:
2313:
1633:, nicknamed the Tully monster, is also present and one of the most famous species from the assemblage.
623:
2100:"The Mazon Creek Lagerstätte: a diverse late Paleozoic ecosystem entombed within siderite concretions"
2346:
2111:
2044:
1995:
1698:
503:
453:
1195:
536:
429:
391:
2031:
Haug, Joachim T.; Haug, Carolin; Maas, Andreas; Kutschera, Verena; Waloszek, Dieter (2010-09-21).
1168:
2147:
1901:
1876:"Closing a major gap in mantis shrimp evolution - first fossils of Stomatopoda from the Triassic"
1792:
1730:
1722:
328:
119:
575:
The thorax can be divided into a front part made up of small, unfused segments which bear short
2318:
2300:
2216:
2182:
2139:
2080:
2062:
2033:"Evolution of mantis shrimps (Stomatopoda, Malacostraca) in the light of new Mesozoic fossils"
2013:
1960:
1942:
1893:
1856:
1784:
1714:
565:
560:
2305:
2208:
1606:
preserving fossils of an extremely wide array of organisms, which would have lived alongside
2174:
2129:
2119:
2070:
2052:
2003:
1950:
1932:
1883:
1846:
1706:
364:
304:
604:
499:
368:
2115:
2048:
1999:
1702:
1658:
323:, swimming above the seabed while it hunted, perhaps grabbing prey from above using its
2178:
2075:
2032:
1955:
1920:
1603:
1356:
976:
959:
943:
569:
442:
1921:"A new fossil mantis shrimp and the convergent evolution of a lobster-like morphotype"
530:
length of 4.4–13.6 mm (0.17–0.54 in). Meanwhile, the only known specimen of
2335:
2237:
1905:
1811:
1660:
Some Middle
Pennsylvanian Hoplocarida (Crustacea) and their phylogenetic significance
1629:
1587:
1543:
1141:
1114:
627:
317:
282:
278:
176:
61:
2151:
1749:
1734:
2278:
2242:. University of Illinois Urbana-Champaign. : Field Museum of Natural History.
2008:
1979:
1750:"Quelques eumalacostracés nouveaux du Westphalien inférieur d'Argenteau près Liége"
1615:
1312:
790:
773:
757:
696:
679:
663:
552:
423:
219:
166:
1875:
1619:
1595:
1021:
648:
619:
36:
1851:
1559:
1339:
1097:
576:
544:
81:
46:
2143:
2066:
2057:
2017:
1946:
1897:
1888:
1860:
1788:
1718:
1623:
1531:
1523:
1285:
631:
616:
324:
156:
136:
103:
86:
2084:
1964:
2272:
1710:
828:
739:
548:
527:
433:
356:
344:
320:
290:
76:
71:
56:
51:
41:
2124:
2099:
1937:
2292:
1796:
1773:"Upper Pennsylvanian Arthropods from Black Shales of Iowa and Nebraska"
1772:
1726:
1685:
1591:
1547:
1535:
580:
507:
457:
332:
91:
66:
2134:
1594:. This crustacean is part of a diverse fossil assemblage known as the
1554:
was not a bottom-dwelling animal. In modern mantis shrimps, only the
1249:
1229:
916:
896:
855:
588:
478:
473:
146:
2249:
1834:
452:
was named in 1998 and is known from several specimens found in the
1555:
584:
490:
274:
1833:
Jenner, Ronald A.; Hof, Cees H. J.; Schram, Frederick R. (1998).
1627:
being the most abundant animal from this location. The enigmatic
1611:
2253:
572:
flattened, appearing almost triangular when seen from the top.
1599:
611:
being its type and only genus, and is placed in the suborder
1534:
animal. However, while modern mantis shrimps are primarily
2236:
Johnson, Ralph Gordon; Richardson, Eugene
Stanley (1969).
2215:(second ed.). Manson Publishing Ltd. pp. 94–96.
440:
in 1984 by
Frederick Schram, and then reassigned again to
410:
The following species were formerly assigned to the genus
2165:
Bardack, David (1979-01-01), Nitecki, Matthew H. (ed.),
2098:
Clements, Thomas; Purnell, Mark; Gabbott, Sarah (2019).
1980:"Larval development and behaviour of the mantis shrimp,
1526:
appendages and the lifestyles of other mantis shrimps,
476:. In 2007, this species was moved to a separate genus,
1335:
1308:
1281:
1245:
1225:
1191:
1164:
1137:
1110:
1080:
1053:
1017:
939:
912:
892:
851:
824:
753:
735:
659:
472:(meaning "tail"), in reference to the spines on the
464:
34453. The specific name is combined from the Greek
375:. Aside from the holotype, a few other specimens of
249:
226:
2262:
1684:
1602:. The Mazon Creek fossil beds are an important
568:to the corners of the carapace. The rostrum is
1919:Haug, Carolin; Haug, Joachim T. (2021-04-16).
498:. 325 million years old, Early Carboniferous,
1984:Milne Edwards (Crustacea : Stomatopoda)"
8:
1754:Annales de la Société Géologique de Belgique
630:, the results of which are displayed in the
1988:Journal of the Royal Society of New Zealand
1663:. Chicago: Field Museum of Natural History.
2250:
626:to determine the relations between fossil
436:, Belgium. It was renamed as a species of
359:, USA by Calvin George and donated to the
102:
20:
2133:
2123:
2074:
2056:
2007:
1954:
1936:
1887:
1850:
1686:"Paleozoic Proto-Mantis Shrimp Revisited"
331:which would have been just south of the
432:in 1922 based on fossils collected near
1816:Geologie und Paläontologie in Westfalen
1642:
2342:Carboniferous animals of North America
367:, who at the time considered it to be
1598:, which would have lived in a marine
512:Royal Tyrrell Museum of Palaeontology
293:. It is the only genus in the family
7:
2173:, Academic Press, pp. 501–528,
1678:
1676:
1674:
1672:
1670:
1652:
1650:
1648:
1646:
1582:of Illinois, which date back to the
1610:. These include over 25 species of
1328:
1301:
1274:
1218:
1211:
1184:
1157:
1130:
1103:
1073:
1046:
1010:
951:
885:
878:
871:
844:
765:
728:
721:
671:
654:
644:
551:of approximately equal length. The
363:. The genus was erected in 1969 by
2207:Selden, Paul; Nudds, John (2012).
2179:10.1016/b978-0-12-519650-5.50024-2
622:. Smith et al. (2023) conducted a
482:, of which it is the type species.
414:, but have since been reassigned:
14:
2167:"Fishes of the Mazon Creek Fauna"
2104:Journal of the Geological Society
351:(PE12098) was collected from the
316:This animal is theorized to be a
307:. A second, much larger species,
402:until more specimens are found.
123:
371:and include the single species
361:Field Museum of Natural History
2213:Evolution of Fossil Ecosystems
2009:10.1080/03036758.1972.10429369
599:In his initial description of
16:Extinct genus of mantis shrimp
1:
2357:Prehistoric crustacean genera
2352:Fossil taxa described in 1998
1771:Schram, Frederick R. (1984).
1748:van Straelen, Victor (1922).
1683:Schram, Frederick R. (2007).
1657:Schram, Frederick R. (1969).
547:each possess three annulated
390:, and was collected from the
446:in 2004 by Lothar Schöllman.
1810:Schöllmann, Lothar (2004).
1542:has been theorized to be a
639:
303:, was described in 1969 by
281:that lived during the late
2378:
2362:Prehistoric mantis shrimps
460:, with the holotype being
327:appendages. It lived in a
285:period in what is now the
1852:10.1163/18759866-06703001
1353:
1333:
1326:
1306:
1299:
1279:
1272:
1243:
1223:
1216:
1209:
1189:
1182:
1162:
1155:
1135:
1128:
1108:
1101:
1078:
1071:
1051:
1044:
1015:
1008:
973:
956:
949:
910:
890:
883:
876:
869:
849:
842:
822:
787:
770:
763:
733:
726:
719:
693:
676:
669:
652:
246:
241:
225:
218:
120:Scientific classification
118:
110:
101:
23:
2058:10.1186/1471-2148-10-290
2037:BMC Evolutionary Biology
1889:10.3140/bull.geosci.1864
1839:Contributions to Zoology
1313:Lysiosquilla nkporoensis
1085:Tyrannophontes gigantion
1058:Tyrannophontes theridion
960:Gorgonophontes fraiponti
680:Archaeocaris vermiformis
522:differ greatly in size.
349:Tyrannophontes theridion
231:Tyrannophontes theridion
1880:Bulletin of Geosciences
1777:Journal of Paleontology
1691:Journal of Paleontology
1580:Mazon Creek fossil beds
1196:Archaeosculda phoenicia
829:Bairdops beargulchensis
421:was initially named as
382:An additional species,
353:Mazon Creek fossil beds
287:Mazon Creek fossil beds
1614:and numerous types of
977:Gorgonophontes peleron
697:Archaeocaris graffhami
583:. Each segment of the
515:
468:(meaning "thorn") and
2314:Paleobiology Database
1978:Pyne, Rex R. (1972).
1340:Nodosculda fisherorum
1115:Triassosculda ahyongi
624:phylogenetic analysis
494:
111:3D reconstruction of
1711:10.1666/pleo05-075.1
1142:Tyrannosculda laurae
917:Daidal acanthocercus
454:Bear Gulch Limestone
339:Discovery and naming
297:. The type species,
2171:Mazon Creek Fossils
2125:10.1144/jgs2018-088
2116:2019JGSoc.176....1C
2049:2010BMCEE..10..290H
2000:1972JRSNZ...2..121P
1938:10.7717/peerj.11124
1703:2007JPal...81..895S
1578:originate from the
1169:Pseudosculda laevis
1022:Chabardella spinosa
897:Daidal schoellmanni
613:Archaeostomatopodea
518:The two species of
430:Victor van Straelen
392:Francis Creek Shale
1530:would have been a
1500:Archaeostomatopods
1286:Ursquilla yehoachi
791:Perimecturus rapax
774:Perimecturus parki
516:
406:Reassigned species
329:marine environment
2329:
2328:
2301:Open Tree of Life
2256:Taxon identifiers
2222:978-1-84076-623-3
2188:978-0-12-519650-5
1570:Palaeoenvironment
1522:Judging from its
1512:
1511:
1503:Palaeostomatopods
1494:
1493:
1485:
1484:
1476:
1475:
1467:
1466:
1458:
1457:
1449:
1448:
1440:
1439:
1431:
1430:
1422:
1421:
1413:
1412:
1404:
1403:
1395:
1394:
1386:
1385:
1377:
1376:
1368:
1367:
1261:
1260:
1033:
1032:
997:
996:
988:
987:
928:
927:
811:
810:
802:
801:
708:
707:
559:on the upper and
266:
265:
260:
214:
195:
2369:
2322:
2321:
2309:
2308:
2296:
2295:
2283:
2282:
2281:
2251:
2244:
2243:
2233:
2227:
2226:
2204:
2198:
2197:
2196:
2195:
2162:
2156:
2155:
2137:
2127:
2095:
2089:
2088:
2078:
2060:
2028:
2022:
2021:
2011:
1975:
1969:
1968:
1958:
1940:
1916:
1910:
1909:
1891:
1871:
1865:
1864:
1854:
1830:
1824:
1823:
1807:
1801:
1800:
1768:
1762:
1761:
1745:
1739:
1738:
1688:
1680:
1665:
1664:
1654:
1586:age of the late
1574:Both species of
1337:
1329:
1310:
1302:
1283:
1275:
1247:
1227:
1219:
1212:
1193:
1185:
1166:
1158:
1139:
1131:
1112:
1104:
1082:
1074:
1055:
1047:
1019:
1011:
952:
941:
914:
894:
886:
879:
872:
853:
845:
826:
766:
755:
740:Bairdops elegans
737:
729:
722:
672:
661:
655:
645:
640:
450:T. acanthocercus
365:Frederick Schram
305:Frederick Schram
295:Tyrannophontidae
258:
251:
228:
212:
205:
193:
190:Tyrannophontidae
188:
128:
127:
106:
96:
33:
29:Temporal range:
21:
2377:
2376:
2372:
2371:
2370:
2368:
2367:
2366:
2332:
2331:
2330:
2325:
2317:
2312:
2304:
2299:
2291:
2286:
2277:
2276:
2271:
2258:
2248:
2247:
2235:
2234:
2230:
2223:
2206:
2205:
2201:
2193:
2191:
2189:
2164:
2163:
2159:
2097:
2096:
2092:
2030:
2029:
2025:
1977:
1976:
1972:
1918:
1917:
1913:
1873:
1872:
1868:
1832:
1831:
1827:
1809:
1808:
1804:
1770:
1769:
1765:
1747:
1746:
1742:
1682:
1681:
1668:
1656:
1655:
1644:
1639:
1572:
1520:
1514:
1508:
1507:
1504:
1501:
1495:
1486:
1477:
1468:
1459:
1450:
1441:
1432:
1423:
1414:
1405:
1396:
1387:
1378:
1369:
1262:
1034:
998:
989:
947:
929:
812:
803:
761:
709:
667:
637:
597:
500:Heath Formation
489:
408:
341:
257:
237:
234:
211:
203:
192:
186:
122:
97:
95:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
31:
30:
27:
17:
12:
11:
5:
2375:
2373:
2365:
2364:
2359:
2354:
2349:
2344:
2334:
2333:
2327:
2326:
2324:
2323:
2310:
2297:
2284:
2268:
2266:
2264:Tyrannophontes
2260:
2259:
2254:
2246:
2245:
2228:
2221:
2199:
2187:
2157:
2090:
2023:
1994:(2): 121–146.
1982:Squilla armata
1970:
1911:
1866:
1825:
1802:
1783:(1): 197–209.
1763:
1740:
1697:(5): 895–916.
1666:
1641:
1640:
1638:
1635:
1608:Tyrannophontes
1576:Tyrannophontes
1571:
1568:
1564:Tyrannophontes
1552:Tyrannophontes
1540:Tyrannophontes
1528:Tyrannophontes
1519:
1516:
1510:
1509:
1506:Pseudosculdids
1505:
1502:
1499:
1498:
1496:
1492:
1491:
1488:
1487:
1483:
1482:
1479:
1478:
1474:
1473:
1470:
1469:
1465:
1464:
1461:
1460:
1456:
1455:
1452:
1451:
1447:
1446:
1443:
1442:
1438:
1437:
1434:
1433:
1429:
1428:
1425:
1424:
1420:
1419:
1416:
1415:
1411:
1410:
1407:
1406:
1402:
1401:
1398:
1397:
1393:
1392:
1389:
1388:
1384:
1383:
1380:
1379:
1375:
1374:
1371:
1370:
1366:
1365:
1362:
1361:
1357:Squilla mantis
1352:
1349:
1348:
1345:
1344:
1332:
1327:
1325:
1322:
1321:
1318:
1317:
1305:
1300:
1298:
1295:
1294:
1291:
1290:
1278:
1273:
1271:
1268:
1267:
1264:
1263:
1259:
1258:
1255:
1254:
1250:Sculda syriaca
1242:
1239:
1238:
1235:
1234:
1230:Sculda pennata
1222:
1217:
1215:
1210:
1208:
1205:
1204:
1201:
1200:
1188:
1183:
1181:
1178:
1177:
1174:
1173:
1161:
1156:
1154:
1151:
1150:
1147:
1146:
1134:
1129:
1127:
1124:
1123:
1120:
1119:
1107:
1102:
1100:
1094:
1093:
1090:
1089:
1077:
1072:
1070:
1067:
1066:
1063:
1062:
1050:
1045:
1043:
1040:
1039:
1036:
1035:
1031:
1030:
1027:
1026:
1014:
1009:
1007:
1004:
1003:
1000:
999:
995:
994:
991:
990:
986:
985:
982:
981:
972:
969:
968:
965:
964:
955:
950:
948:
944:Gorgonophontes
938:
935:
934:
931:
930:
926:
925:
922:
921:
909:
906:
905:
902:
901:
889:
884:
882:
877:
875:
870:
868:
865:
864:
861:
860:
856:Daidal pattoni
848:
843:
841:
838:
837:
834:
833:
821:
818:
817:
814:
813:
809:
808:
805:
804:
800:
799:
796:
795:
786:
783:
782:
779:
778:
769:
764:
762:
752:
749:
748:
745:
744:
732:
727:
725:
720:
718:
715:
714:
711:
710:
706:
705:
702:
701:
692:
689:
688:
685:
684:
675:
670:
668:
658:
653:
651:
643:
628:mantis shrimps
609:Tyrannophontes
601:Tyrannophontes
596:
595:Classification
593:
570:dorsoventrally
520:Tyrannophontes
488:
485:
484:
483:
447:
443:Gorgonophontes
438:Tyrannophontes
412:Tyrannophontes
407:
404:
400:Tyrannophontes
340:
337:
273:is an extinct
270:Tyrannophontes
264:
263:
262:
261:
244:
243:
242:Other species
239:
238:
235:
223:
222:
216:
215:
208:Tyrannophontes
201:
197:
196:
184:
180:
179:
174:
170:
169:
164:
160:
159:
154:
150:
149:
144:
140:
139:
134:
130:
129:
116:
115:
108:
107:
99:
98:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
35:
34:
28:
25:Tyrannophontes
15:
13:
10:
9:
6:
4:
3:
2:
2374:
2363:
2360:
2358:
2355:
2353:
2350:
2348:
2345:
2343:
2340:
2339:
2337:
2320:
2315:
2311:
2307:
2302:
2298:
2294:
2289:
2285:
2280:
2274:
2270:
2269:
2267:
2265:
2261:
2257:
2252:
2241:
2240:
2232:
2229:
2224:
2218:
2214:
2210:
2209:"Mazon Creek"
2203:
2200:
2190:
2184:
2180:
2176:
2172:
2168:
2161:
2158:
2153:
2149:
2145:
2141:
2136:
2131:
2126:
2121:
2117:
2113:
2109:
2105:
2101:
2094:
2091:
2086:
2082:
2077:
2072:
2068:
2064:
2059:
2054:
2050:
2046:
2042:
2038:
2034:
2027:
2024:
2019:
2015:
2010:
2005:
2001:
1997:
1993:
1989:
1985:
1983:
1974:
1971:
1966:
1962:
1957:
1952:
1948:
1944:
1939:
1934:
1930:
1926:
1922:
1915:
1912:
1907:
1903:
1899:
1895:
1890:
1885:
1881:
1877:
1870:
1867:
1862:
1858:
1853:
1848:
1844:
1840:
1836:
1829:
1826:
1821:
1817:
1813:
1806:
1803:
1798:
1794:
1790:
1786:
1782:
1778:
1774:
1767:
1764:
1759:
1756:(in French).
1755:
1751:
1744:
1741:
1736:
1732:
1728:
1724:
1720:
1716:
1712:
1708:
1704:
1700:
1696:
1692:
1687:
1679:
1677:
1675:
1673:
1671:
1667:
1662:
1661:
1653:
1651:
1649:
1647:
1643:
1636:
1634:
1632:
1631:
1630:Tullimonstrum
1626:
1625:
1621:
1617:
1616:invertebrates
1613:
1609:
1605:
1601:
1597:
1593:
1589:
1588:Carboniferous
1585:
1581:
1577:
1569:
1567:
1565:
1561:
1557:
1553:
1549:
1545:
1544:benthopelagic
1541:
1537:
1533:
1529:
1525:
1518:Palaeobiology
1517:
1515:
1497:
1490:
1489:
1481:
1480:
1472:
1471:
1463:
1462:
1454:
1453:
1445:
1444:
1436:
1435:
1427:
1426:
1418:
1417:
1409:
1408:
1400:
1399:
1391:
1390:
1382:
1381:
1373:
1372:
1364:
1363:
1360:
1359:
1358:
1351:
1350:
1347:
1346:
1343:
1342:
1341:
1331:
1330:
1324:
1323:
1320:
1319:
1316:
1315:
1314:
1304:
1303:
1297:
1296:
1293:
1292:
1289:
1288:
1287:
1277:
1276:
1270:
1269:
1266:
1265:
1257:
1256:
1253:
1252:
1251:
1241:
1240:
1237:
1236:
1233:
1232:
1231:
1221:
1220:
1214:
1213:
1207:
1206:
1203:
1202:
1199:
1198:
1197:
1187:
1186:
1180:
1179:
1176:
1175:
1172:
1171:
1170:
1160:
1159:
1153:
1152:
1149:
1148:
1145:
1144:
1143:
1133:
1132:
1126:
1125:
1122:
1121:
1118:
1117:
1116:
1106:
1105:
1099:
1096:
1095:
1092:
1091:
1088:
1087:
1086:
1076:
1075:
1069:
1068:
1065:
1064:
1061:
1060:
1059:
1049:
1048:
1042:
1041:
1038:
1037:
1029:
1028:
1025:
1024:
1023:
1013:
1012:
1006:
1005:
1002:
1001:
993:
992:
984:
983:
980:
979:
978:
971:
970:
967:
966:
963:
962:
961:
954:
953:
946:
945:
937:
936:
933:
932:
924:
923:
920:
919:
918:
908:
907:
904:
903:
900:
899:
898:
888:
887:
881:
880:
874:
873:
867:
866:
863:
862:
859:
858:
857:
847:
846:
840:
839:
836:
835:
832:
831:
830:
820:
819:
816:
815:
807:
806:
798:
797:
794:
793:
792:
785:
784:
781:
780:
777:
776:
775:
768:
767:
760:
759:
751:
750:
747:
746:
743:
742:
741:
731:
730:
724:
723:
717:
716:
713:
712:
704:
703:
700:
699:
698:
691:
690:
687:
686:
683:
682:
681:
674:
673:
666:
665:
657:
656:
650:
647:
646:
642:
641:
638:
635:
633:
629:
625:
621:
618:
614:
610:
606:
602:
594:
592:
590:
586:
582:
578:
573:
571:
567:
562:
558:
554:
553:compound eyes
550:
546:
541:
539:
538:
537:Harpiosquilla
533:
529:
525:
521:
513:
509:
505:
501:
497:
493:
486:
481:
480:
475:
471:
467:
463:
459:
455:
451:
448:
445:
444:
439:
435:
431:
427:
425:
420:
417:
416:
415:
413:
405:
403:
401:
397:
393:
389:
385:
380:
378:
374:
370:
366:
362:
358:
354:
350:
346:
338:
336:
334:
330:
326:
322:
319:
318:benthopelagic
314:
312:
311:
306:
302:
301:
296:
292:
288:
284:
283:Carboniferous
280:
279:mantis shrimp
276:
272:
271:
256:
255:
248:
247:
245:
240:
233:
232:
224:
221:
217:
210:
209:
202:
199:
198:
191:
185:
182:
181:
178:
175:
172:
171:
168:
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
138:
135:
132:
131:
126:
121:
117:
114:
109:
105:
100:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
26:
22:
19:
2263:
2238:
2231:
2212:
2202:
2192:, retrieved
2170:
2160:
2107:
2103:
2093:
2040:
2036:
2026:
1991:
1987:
1981:
1973:
1928:
1924:
1914:
1879:
1869:
1842:
1838:
1828:
1819:
1815:
1805:
1780:
1776:
1766:
1757:
1753:
1743:
1694:
1690:
1659:
1628:
1622:
1607:
1575:
1573:
1563:
1551:
1539:
1527:
1521:
1513:
1355:
1354:
1338:
1334:
1311:
1307:
1284:
1280:
1248:
1244:
1228:
1224:
1194:
1190:
1167:
1163:
1140:
1136:
1113:
1109:
1084:
1083:
1079:
1057:
1056:
1052:
1020:
1016:
975:
974:
958:
957:
942:
915:
911:
895:
891:
854:
850:
827:
823:
789:
788:
772:
771:
758:Perimecturus
756:
738:
734:
695:
694:
678:
677:
664:Archaeocaris
662:
636:
608:
600:
598:
574:
542:
535:
532:T. gigantion
531:
524:T. theridion
523:
519:
517:
496:T. theridion
495:
477:
469:
465:
449:
441:
437:
424:Perimecturus
422:
419:T. fraiponti
418:
411:
409:
399:
396:T. theridion
395:
384:T. gigantion
383:
381:
377:T. theridion
376:
373:T. theridion
372:
348:
342:
315:
310:T. gigantion
309:
308:
300:T. theridion
299:
298:
294:
269:
268:
267:
259:Schram, 2007
254:T. gigantion
253:
252:
236:Schram, 1969
230:
229:
220:Type species
213:Schram, 1969
207:
206:
194:Schram, 1969
189:
167:Malacostraca
113:T. theridion
112:
24:
18:
2347:Stomatopoda
2110:(1): 1–11.
1620:Sea anemone
1618:, with the
1604:lagerstätte
1596:Essex biota
1560:stomatopods
649:Stomatopoda
620:holpocarids
577:thoracopods
487:Description
177:Stomatopoda
2336:Categories
2279:Q123370868
2194:2023-11-08
2135:2381/44761
2043:(1): 290.
1931:: e11124.
1882:: 95–110.
1845:(3): 155.
1822:: 111–141.
1637:References
1098:Unipeltata
545:antennules
526:reached a
504:Bear Gulch
157:Arthropoda
2144:0016-7649
2067:1471-2148
2018:0303-6758
1947:2167-8359
1906:258089800
1898:1802-8225
1861:1383-4517
1789:0022-3360
1719:0022-3360
1624:Essexella
1584:Moscovian
1532:predatory
1524:raptorial
632:cladogram
617:Paleozoic
605:monotypic
540:species.
510:. At the
426:fraiponti
369:monotypic
325:raptorial
143:Kingdom:
137:Eukaryota
32:Moscovian
2273:Wikidata
2152:84178980
2085:20858249
1965:33959413
1760:: 35–40.
1735:85606671
1548:tergites
549:flagella
528:carapace
466:akanthos
357:Illinois
345:holotype
321:predator
291:Illinois
183:Family:
153:Phylum:
147:Animalia
133:Domain:
2306:3005037
2293:8480732
2112:Bibcode
2076:2955030
2045:Bibcode
1996:Bibcode
1956:8054755
1797:1304745
1727:4498847
1699:Bibcode
1592:equator
1536:benthic
634:below:
607:, with
581:abdomen
566:rostrum
561:lateral
508:Montana
458:Montana
333:equator
200:Genus:
173:Order:
163:Class:
2219:
2185:
2150:
2142:
2083:
2073:
2065:
2016:
1963:
1953:
1945:
1904:
1896:
1859:
1795:
1787:
1733:
1725:
1717:
1556:larvae
589:telson
557:thorax
479:Daidal
474:telson
470:kerkos
388:thorax
2319:22282
2148:S2CID
1925:PeerJ
1902:S2CID
1793:JSTOR
1731:S2CID
1723:JSTOR
1562:like
585:pleon
434:Liège
275:genus
2288:GBIF
2217:ISBN
2183:ISBN
2140:ISSN
2081:PMID
2063:ISSN
2014:ISSN
1961:PMID
1943:ISSN
1894:ISSN
1857:ISSN
1785:ISSN
1715:ISSN
1612:fish
543:The
343:The
37:Preęž’
2175:doi
2130:hdl
2120:doi
2108:176
2071:PMC
2053:doi
2004:doi
1951:PMC
1933:doi
1884:doi
1847:doi
1707:doi
1600:bay
456:of
428:by
355:in
347:of
289:of
277:of
2338::
2316::
2303::
2290::
2275::
2211:.
2181:,
2169:,
2146:.
2138:.
2128:.
2118:.
2106:.
2102:.
2079:.
2069:.
2061:.
2051:.
2041:10
2039:.
2035:.
2012:.
2002:.
1990:.
1986:.
1959:.
1949:.
1941:.
1927:.
1923:.
1900:.
1892:.
1878:.
1855:.
1843:67
1841:.
1837:.
1820:62
1818:.
1814:.
1791:.
1781:58
1779:.
1775:.
1758:45
1752:.
1729:.
1721:.
1713:.
1705:.
1695:81
1693:.
1689:.
1669:^
1645:^
1566:.
1538:,
506:,
502:,
462:CM
335:.
87:Pg
2225:.
2177::
2154:.
2132::
2122::
2114::
2087:.
2055::
2047::
2020:.
2006::
1998::
1992:2
1967:.
1935::
1929:9
1908:.
1886::
1863:.
1849::
1799:.
1737:.
1709::
1701::
1336:â€
1309:â€
1282:â€
1246:â€
1226:â€
1192:â€
1165:â€
1138:â€
1111:â€
1081:â€
1054:â€
1018:â€
940:â€
913:â€
893:â€
852:â€
825:â€
754:â€
736:â€
660:â€
514:.
250:â€
227:â€
204:â€
187:â€
92:N
82:K
77:J
72:T
67:P
62:C
57:D
52:S
47:O
42:ęž’
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