131:
1856:
1873:
1850:
118:
477:, another closely related trematopid, has been described as having five distal carpals. In the pes of the holotype specimen, there are only four digits preserved. However, since the third preserved digit is the longest, it likely represents the fourth digit, with the first digit absent due to imperfect preservation. The terminal
455:
into anterior and posterior portions is evidence of the presence of salt glands. The anterior portion is sub-circular and is thought to be the true narial opening. The posterior portion likely held the salt gland. The expansion of the posterior portion may not be an accommodation for the salt gland,
437:, or eye sockets, are centrally placed so that the preorbital length of the skull is equal to that of the postorbital length. They make up around 30% of the skull length. The skull table seems to extend posteriorly past the level of the jaw articulation in
463:, given the completeness of the holotype specimen. The relatively short tail, preserved only as a faint impression in MNG 7722, is 8.5 centimetres (3.3 in) long. The limbs are short, with four digits on the
432:
The skull of MNG 7722 is severely crushed dorsoventrally, making it hard to determine the shape of the skull in lateral view. It is 6.9 centimetres (2.7 in) in length, which is small for a trematopid. The
938:"A new trematopid amphibian (Temnospondyli: Dissorophoidea) from the Upper Pennsylvanian of western Pennsylvania: earliest record of terrestrial vertebrates responding to a warmer, drier climate"
937:
326:
of central
Germany. It consists of a skull and much of the postcranial skeleton. The only major portion of the skeleton that is missing is the presacral
1989:
1925:
1054:
986:
899:
441:, which is unique among trematopids. However, this is likely to have been the result of crushing of the skull, which displaced the skull table.
1999:
2004:
1021:
1047:
130:
1855:
968:
1994:
1849:
1133:
322:, known as MNG 7722, has been found from an outcrop of the Tambach Formation at the Bromacker locality in the
1684:
1040:
398:
1893:
1876:
1160:
512:
366:
Although it is difficult to determine the maturity of MNG 7722, it is thought to be an early adult. Poor
1979:
1951:
1828:
1027:
1001:
914:
1984:
1253:
520:
357:
213:
1784:
1290:
1269:
1805:
1763:
1320:
960:
718:
margin. In other trematopids, the margin is biconcave. Below is a cladogram modified from Berman
125:
1956:
1496:
1938:
1791:
375:
323:
279:
1943:
1740:
952:
424:
that was frequently eaten in the
Bromacker Quarry by the authors who described the species.
287:
269:
1798:
1594:
1448:
1283:
1118:
1096:
464:
434:
1756:
1664:
1262:
1224:
1202:
1172:
1063:
715:
262:
1973:
1777:
1675:
1657:
1622:
1615:
1480:
1434:
1414:
1145:
1102:
532:
524:
516:
468:
406:
303:
276:
175:
68:
30:
964:
1816:
1770:
1729:
1709:
1650:
1560:
1469:
1441:
1331:
1306:
1276:
691:
554:
527:, and a contribution to the ventral orbital rim (the rim of the eye socket) by the
379:
367:
272:
265:
221:
188:
386:
suggest that MNG 7722 represents an early stage of development, while the pitted
1636:
1524:
1515:
1504:
1489:
1455:
1427:
1389:
1375:
626:
43:
1916:
1722:
1691:
1574:
1538:
1354:
1347:
593:
511:. Characteristics that supported this relationship included the absence of an
503:
448:
387:
283:
88:
53:
1698:
1643:
1629:
1567:
1462:
1361:
536:
478:
421:
390:, with tightly closed sutures between bones, indicates a mature individual.
383:
345:
331:
291:
142:
93:
37:
1910:
1608:
1552:
1545:
1396:
1340:
1087:
956:
643:
473:
360:
338:
327:
315:
162:
83:
78:
63:
58:
48:
1930:
1749:
1368:
1242:
748:
678:
528:
452:
417:
353:
349:
341:
295:
255:
98:
73:
900:"A new trematopid amphibian from the Lower Permian of central Germany"
471:, or foot. It is possible that the manus possessed five digits, since
117:
1531:
1382:
1081:
371:
334:
152:
1887:
987:"Biology and evolution of the nasal region in trematopid amphibians"
936:
Berman, D.S.; Henrici, A.C.; Brezinski, D.K.; Kollar, A.D. (2010).
456:
but rather a means of coping with cranial stresses during feeding.
258:
1032:
451:
similar to those seen in modern reptiles. The subdivision of the
1891:
1074:
1036:
485:
were narrow, and would have been the core supports of claws.
344:
and articulated skeletons of terrestrial and semiterrestrial
302:
is the first trematopid to have been discovered outside the
523:
trait), the reduction of the suborbital process of the
228:
1900:
1815:
1739:
1708:
1674:
1592:
1514:
1479:
1413:
1330:
1304:
1252:
1222:
1171:
1144:
1117:
898:Sumida, S.S; Berman, D.S.; Martens, T. (1998).
1048:
543:(1998) showing the proposed relationships of
447:, like other trematopids, likely possessed a
8:
1888:
1600:
1419:
1312:
1234:
1230:
1219:
1150:
1123:
1114:
1071:
1055:
1041:
1033:
714:all possess a shallow and broadly concave
459:The postcranial skeleton is well known in
116:
20:
931:
929:
927:
893:
891:
889:
887:
885:
881:
397:refers to the Tambach Formation. The
7:
420:. The name refers to the Thuringian
330:column. The Bromacker locality is a
821:
764:
740:
733:
725:
618:
568:
561:
550:
467:, or hand, and five digits on the
14:
1872:
1871:
1854:
1848:
352:. MNG 7722 was found in red-bed
129:
1990:Permian temnospondyls of Europe
493:In the original description of
412:, meaning munch or nibble, and
363:in flat-bedded channel fills.
1:
2000:Fossil taxa described in 1998
2005:Prehistoric amphibian genera
1000:(4): 839β853. Archived from
913:(4): 605β629. Archived from
507:was the closest relative of
356:deposits consisting of well
519:, but was interpreted as a
282:(the lowermost unit of the
16:Extinct genus of amphibians
2021:
1867:
1845:
1603:
1422:
1315:
1237:
1233:
1218:
1199:
1153:
1126:
1113:
1070:
945:Annals of Carnegie Museum
841:
826:
819:
786:
769:
762:
745:
738:
731:
640:
623:
616:
590:
573:
566:
559:
227:
220:
126:Scientific classification
124:
115:
23:
694:of the clade containing
676:In their description of
547:and other trematopids:
515:(which is also seen in
337:that is well known for
275:. It is known from the
501:(1998) concluded that
1952:Paleobiology Database
1028:Paleobiology Database
985:Dilkes, D.W. (1993).
539:modified from Sumida
1254:Micromelerpetontidae
957:10.2992/007.078.0401
513:internarial fenestra
405:is derived from the
1902:Tambachia trogallas
729:Trematopidae
320:Tambachia trogallas
286:) near the town of
233:Tambachia trogallas
1967:
1966:
1939:Open Tree of Life
1894:Taxon identifiers
1885:
1884:
1863:
1862:
1843:
1842:
1839:
1838:
1792:Schoenfelderpeton
1588:
1587:
1584:
1583:
1300:
1299:
1214:
1213:
1210:
1209:
1195:
1194:
1191:
1190:
872:
871:
863:
862:
854:
853:
808:
807:
799:
798:
673:
672:
664:
663:
655:
654:
605:
604:
324:Thuringian Forest
280:Tambach Formation
247:
246:
216:
2012:
1960:
1959:
1947:
1946:
1934:
1933:
1921:
1920:
1919:
1889:
1875:
1874:
1858:
1853:
1852:
1741:Branchiosauridae
1601:
1420:
1313:
1235:
1231:
1220:
1151:
1124:
1115:
1108:
1107:
1072:
1057:
1050:
1043:
1034:
1009:
1008:
1006:
991:
982:
976:
975:
973:
967:. Archived from
942:
933:
922:
921:
919:
904:
895:
822:
765:
741:
734:
726:
619:
569:
562:
551:
531:rather than the
288:Tambach-Dietharz
284:Upper Rotliegend
230:
207:
200:
187:
174:
134:
133:
120:
110:
40:
29:Temporal range:
21:
2020:
2019:
2015:
2014:
2013:
2011:
2010:
2009:
1995:Permian Germany
1970:
1969:
1968:
1963:
1955:
1950:
1942:
1937:
1929:
1924:
1915:
1914:
1909:
1896:
1886:
1881:
1859:
1847:
1835:
1811:
1799:Tungussogyrinus
1735:
1712:
1704:
1670:
1597:
1595:Amphibamiformes
1580:
1510:
1475:
1449:Astreptorhachis
1409:
1326:
1309:
1296:
1284:Micromelerpeton
1248:
1227:
1206:
1187:
1167:
1140:
1134:Tetrapodomorpha
1119:Tetrapodomorpha
1109:
1097:Tetrapodomorpha
1077:
1076:
1066:
1061:
1018:
1013:
1012:
1004:
989:
984:
983:
979:
971:
940:
935:
934:
925:
917:
902:
897:
896:
883:
878:
873:
864:
855:
809:
800:
674:
665:
656:
606:
491:
430:
382:portion of the
312:
243:
236:
206:
198:
185:
172:
128:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
36:284β279.5
35:
34:
27:
17:
12:
11:
5:
2018:
2016:
2008:
2007:
2002:
1997:
1992:
1987:
1982:
1972:
1971:
1965:
1964:
1962:
1961:
1948:
1935:
1922:
1906:
1904:
1898:
1897:
1892:
1883:
1882:
1880:
1879:
1868:
1865:
1864:
1861:
1860:
1846:
1844:
1841:
1840:
1837:
1836:
1834:
1833:
1832:
1831:
1822:
1820:
1813:
1812:
1810:
1809:
1802:
1795:
1788:
1781:
1774:
1767:
1760:
1757:Branchiosaurus
1753:
1745:
1743:
1737:
1736:
1734:
1733:
1726:
1718:
1716:
1706:
1705:
1703:
1702:
1695:
1688:
1680:
1678:
1672:
1671:
1669:
1668:
1665:Rubeostratilia
1661:
1654:
1647:
1640:
1633:
1626:
1619:
1612:
1604:
1598:
1593:
1590:
1589:
1586:
1585:
1582:
1581:
1579:
1578:
1571:
1564:
1557:
1549:
1542:
1535:
1528:
1520:
1518:
1512:
1511:
1509:
1508:
1501:
1493:
1485:
1483:
1477:
1476:
1474:
1473:
1466:
1459:
1452:
1445:
1438:
1431:
1423:
1417:
1411:
1410:
1408:
1407:
1400:
1393:
1386:
1379:
1372:
1365:
1358:
1351:
1344:
1336:
1334:
1328:
1327:
1325:
1324:
1316:
1310:
1305:
1302:
1301:
1298:
1297:
1295:
1294:
1287:
1280:
1273:
1266:
1263:Branchierpeton
1258:
1256:
1250:
1249:
1247:
1246:
1238:
1228:
1225:Dissorophoidea
1223:
1216:
1215:
1212:
1211:
1208:
1207:
1203:Dissorophoidea
1200:
1197:
1196:
1193:
1192:
1189:
1188:
1186:
1185:
1184:
1183:
1177:
1175:
1173:Dissorophoidea
1169:
1168:
1166:
1165:
1164:
1163:
1154:
1148:
1142:
1141:
1139:
1138:
1137:
1136:
1127:
1121:
1111:
1110:
1106:
1105:
1099:
1090:
1084:
1075:
1068:
1067:
1064:Dissorophoidea
1062:
1060:
1059:
1052:
1045:
1037:
1031:
1030:
1017:
1016:External links
1014:
1011:
1010:
1007:on 2011-09-28.
977:
974:on 2010-06-15.
951:(4): 289β318.
923:
920:on 2012-02-24.
880:
879:
877:
874:
870:
869:
866:
865:
861:
860:
857:
856:
852:
851:
848:
847:
840:
837:
836:
833:
832:
825:
820:
818:
815:
814:
811:
810:
806:
805:
802:
801:
797:
796:
793:
792:
785:
782:
781:
778:
777:
768:
763:
761:
758:
757:
754:
753:
744:
739:
737:
732:
730:
724:
686:(2010) placed
671:
670:
667:
666:
662:
661:
658:
657:
653:
652:
649:
648:
639:
636:
635:
632:
631:
622:
617:
615:
612:
611:
608:
607:
603:
602:
599:
598:
589:
586:
585:
582:
581:
572:
567:
565:
560:
558:
549:
490:
489:Classification
487:
453:external naris
429:
426:
311:
308:
245:
244:
237:
225:
224:
218:
217:
196:
192:
191:
183:
179:
178:
170:
166:
165:
160:
156:
155:
150:
146:
145:
140:
136:
135:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
2017:
2006:
2003:
2001:
1998:
1996:
1993:
1991:
1988:
1986:
1983:
1981:
1978:
1977:
1975:
1958:
1953:
1949:
1945:
1940:
1936:
1932:
1927:
1923:
1918:
1912:
1908:
1907:
1905:
1903:
1899:
1895:
1890:
1878:
1870:
1869:
1866:
1857:
1851:
1830:
1826:
1825:
1824:
1823:
1821:
1818:
1814:
1808:
1807:
1803:
1801:
1800:
1796:
1794:
1793:
1789:
1787:
1786:
1785:Piasimotriton
1782:
1780:
1779:
1778:Milnererpeton
1775:
1773:
1772:
1768:
1766:
1765:
1761:
1759:
1758:
1754:
1752:
1751:
1747:
1746:
1744:
1742:
1738:
1732:
1731:
1727:
1725:
1724:
1720:
1719:
1717:
1715:
1714:sensu stricto
1711:
1707:
1701:
1700:
1696:
1694:
1693:
1689:
1687:
1686:
1682:
1681:
1679:
1677:
1676:Micropholidae
1673:
1667:
1666:
1662:
1660:
1659:
1658:Plemmyradytes
1655:
1653:
1652:
1648:
1646:
1645:
1641:
1639:
1638:
1634:
1632:
1631:
1627:
1625:
1624:
1623:Gerobatrachus
1620:
1618:
1617:
1616:Georgenthalia
1613:
1611:
1610:
1606:
1605:
1602:
1599:
1596:
1591:
1577:
1576:
1572:
1570:
1569:
1565:
1563:
1562:
1558:
1555:
1554:
1550:
1548:
1547:
1543:
1541:
1540:
1536:
1534:
1533:
1529:
1527:
1526:
1522:
1521:
1519:
1517:
1513:
1507:
1506:
1502:
1499:
1498:
1494:
1492:
1491:
1487:
1486:
1484:
1482:
1481:Dissorophinae
1478:
1472:
1471:
1467:
1465:
1464:
1460:
1458:
1457:
1453:
1451:
1450:
1446:
1444:
1443:
1439:
1437:
1436:
1435:Arkanserpeton
1432:
1430:
1429:
1425:
1424:
1421:
1418:
1416:
1415:Dissorophidae
1412:
1406:
1405:
1401:
1399:
1398:
1394:
1392:
1391:
1387:
1385:
1384:
1380:
1378:
1377:
1373:
1371:
1370:
1366:
1364:
1363:
1359:
1357:
1356:
1352:
1350:
1349:
1345:
1343:
1342:
1338:
1337:
1335:
1333:
1329:
1323:
1322:
1318:
1317:
1314:
1311:
1308:
1303:
1293:
1292:
1288:
1286:
1285:
1281:
1279:
1278:
1274:
1272:
1271:
1267:
1265:
1264:
1260:
1259:
1257:
1255:
1251:
1245:
1244:
1240:
1239:
1236:
1232:
1229:
1226:
1221:
1217:
1205:
1204:
1198:
1181:
1180:
1179:
1178:
1176:
1174:
1170:
1162:
1161:Temnospondyli
1158:
1157:
1156:
1155:
1152:
1149:
1147:
1146:Temnospondyli
1143:
1135:
1131:
1130:
1129:
1128:
1125:
1122:
1120:
1116:
1112:
1104:
1103:Temnospondyli
1100:
1098:
1094:
1091:
1089:
1085:
1083:
1079:
1078:
1073:
1069:
1065:
1058:
1053:
1051:
1046:
1044:
1039:
1038:
1035:
1029:
1025:
1024:
1020:
1019:
1015:
1003:
999:
995:
994:Palaeontology
988:
981:
978:
970:
966:
962:
958:
954:
950:
946:
939:
932:
930:
928:
924:
916:
912:
908:
907:Palaeontology
901:
894:
892:
890:
888:
886:
882:
875:
868:
867:
859:
858:
850:
849:
846:
845:
839:
838:
835:
834:
831:
830:
824:
823:
817:
816:
813:
812:
804:
803:
795:
794:
791:
790:
784:
783:
780:
779:
776:
775:
774:
767:
766:
760:
759:
756:
755:
752:
751:
750:
743:
742:
736:
735:
728:
727:
723:
721:
717:
713:
709:
705:
701:
697:
693:
689:
685:
681:
680:
669:
668:
660:
659:
651:
650:
647:
646:
645:
638:
637:
634:
633:
630:
629:
628:
621:
620:
614:
613:
610:
609:
601:
600:
597:
596:
595:
588:
587:
584:
583:
580:
579:
578:
571:
570:
564:
563:
556:
553:
552:
548:
546:
542:
538:
535:. Below is a
534:
530:
526:
522:
518:
514:
510:
506:
505:
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454:
450:
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440:
436:
427:
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423:
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415:
411:
408:
404:
400:
399:specific name
396:
391:
389:
385:
381:
377:
373:
369:
364:
362:
359:
355:
351:
347:
343:
340:
336:
333:
329:
325:
321:
317:
309:
307:
305:
304:United States
301:
297:
293:
289:
285:
281:
278:
277:Early Permian
274:
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267:
264:
260:
257:
253:
252:
241:
235:
234:
226:
223:
219:
215:
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197:
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184:
181:
180:
177:
176:Temnospondyli
171:
168:
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138:
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114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
31:Early Permian
26:
22:
19:
1980:Dissorophids
1901:
1829:Lissamphibia
1817:Lissamphibia
1804:
1797:
1790:
1783:
1776:
1771:Melanerpeton
1769:
1762:
1755:
1748:
1730:Doleserpeton
1728:
1721:
1713:
1710:Amphibamidae
1697:
1690:
1683:
1663:
1656:
1651:Platyrhinops
1649:
1642:
1635:
1628:
1621:
1614:
1607:
1573:
1566:
1561:Reiszerpeton
1559:
1551:
1544:
1537:
1530:
1523:
1503:
1495:
1488:
1470:Platyhystrix
1468:
1461:
1454:
1447:
1442:Aspidosaurus
1440:
1433:
1426:
1403:
1402:
1395:
1388:
1381:
1374:
1367:
1360:
1353:
1346:
1339:
1332:Trematopidae
1319:
1307:Olsoniformes
1291:Nyranerpeton
1289:
1282:
1277:Limnogyrinus
1275:
1270:Eimerisaurus
1268:
1261:
1241:
1201:
1092:
1022:
1002:the original
997:
993:
980:
969:the original
948:
944:
915:the original
910:
906:
843:
842:
828:
827:
788:
787:
772:
771:
770:
747:
746:
719:
711:
707:
703:
699:
695:
692:sister taxon
687:
683:
677:
675:
642:
641:
625:
624:
592:
591:
576:
575:
574:
555:Trematopidae
544:
540:
517:dissorophids
508:
502:
498:
494:
492:
482:
472:
460:
458:
444:
443:
438:
431:
413:
409:
403:T. trogallas
402:
394:
392:
380:endochondral
368:ossification
365:
358:consolidated
319:
318:specimen of
313:
299:
273:Trematopidae
266:temnospondyl
263:dissorophoid
250:
249:
248:
239:
232:
231:
222:Type species
209:
202:
201:
189:Trematopidae
24:
18:
1985:Trematopids
1806:Xerodromeus
1764:Leptorophus
1685:Micropholis
1637:Limnerpeton
1525:Brevidorsum
1516:Eucacopinae
1505:Dissorophus
1490:Broiliellus
1456:Iratusaurus
1428:Anakamacops
1390:Phonerpeton
1376:Mattauschia
1321:Palodromeus
829:Phonerpeton
627:Phonerpeton
428:Description
268:within the
1974:Categories
1723:Amphibamus
1692:Pasawioops
1575:Zygosaurus
1539:Conjunctio
1497:Diploseira
1355:Anconastes
1348:Actiobates
1182:see belowβ
876:References
789:Anconastes
708:Anconastes
700:Anconastes
594:Anconastes
521:convergent
504:Anconastes
449:salt gland
416:, meaning
388:skull roof
346:amphibians
1699:Tersomius
1644:Nanobamus
1630:Kermitops
1568:Scapanops
1463:Nooxobeia
1404:Tambachia
1362:Ecolsonia
1080:Kingdom:
1023:Tambachia
773:Tambachia
716:occipital
704:Tambachia
696:Tambachia
682:, Berman
577:Tambachia
545:Tambachia
537:cladogram
509:Tambachia
497:, Sumida
495:Tambachia
483:Tambachia
479:phalanges
461:Tambachia
445:Tambachia
439:Tambachia
422:bratwurst
395:Tambachia
393:The name
384:braincase
361:mudstones
342:trackways
332:sandstone
328:vertebral
310:Discovery
300:Tambachia
292:Thuringia
251:Tambachia
203:Tambachia
149:Kingdom:
143:Eukaryota
25:Tambachia
1917:Q7681002
1911:Wikidata
1877:Category
1609:Eoscopus
1553:Parioxys
1546:Kamacops
1397:Rotaryus
1341:Acheloma
1088:Chordata
1086:Phylum:
1082:Animalia
965:85974488
844:Acheloma
722:(2010):
644:Acheloma
533:palatine
525:lacrimal
474:Acheloma
350:reptiles
339:tetrapod
316:holotype
182:Family:
163:Chordata
159:Phylum:
153:Animalia
139:Domain:
1944:6159410
1931:8555050
1750:Apateon
1369:Fedexia
1243:Stegops
1101:Order:
1026:in the
749:Fedexia
712:Fedexia
690:as the
688:Fedexia
679:Fedexia
557:
529:maxilla
418:sausage
376:tarsals
372:carpals
370:of the
354:fluvial
296:Germany
256:extinct
238:Sumida
208:Sumida
195:Genus:
169:Order:
105:↓
1957:152960
1532:Cacops
1383:Mordex
963:
720:et al.
710:, and
684:et al.
541:et al.
499:et al.
435:orbits
378:, and
335:quarry
270:family
254:is an
242:, 1998
240:et al.
210:et al.
1093:Clade
1005:(PDF)
990:(PDF)
972:(PDF)
961:S2CID
941:(PDF)
918:(PDF)
903:(PDF)
465:manus
414:allas
410:trogo
407:Greek
259:genus
1926:GBIF
1827:see
1159:see
1132:see
698:and
348:and
314:The
214:1998
44:Preκ
953:doi
481:of
469:pes
401:of
290:in
261:of
1976::
1954::
1941::
1928::
1913::
1095::
998:36
996:.
992:.
959:.
949:78
947:.
943:.
926:^
911:41
909:.
905:.
884:^
706:,
702:.
374:,
306:.
298:.
294:,
212:,
94:Pg
38:Ma
33:,
1819:?
1556:?
1500:?
1056:e
1049:t
1042:v
955::
229:β
199:β
186:β
173:β
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:κ
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