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Taxonomy of Banksia

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occur superficially or in shallow pits. These taxa are typically tall shrubs and trees that occur in moist areas; they have unbeaked follicles and soft, short-lived leaves that are in many cases needle-like. The clade includes all eastern taxa of the series Salicinae and Spicigerae (that is, all taxa
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Most aspects of Thiele and Ladiges' arrangement were not accepted by George in his 1999 revision. He stated that "the infrageneric classification and systematic sequence presented here are modified from that of George (1981) and take into account new data revealed in the work of Thiele & Ladiges
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The change has been adopted by a number of Australian herbaria including the Western Australian Herbarium, now headed by Thiele (this has the largest holdings of specimens), and by the Australian Plant Census, a project of the Council of Heads of Australasian Herbaria. This is in accordance with
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were defined as those taxa in which the leaf stomata occur in crypts with constricted entrances. These are usually small shrubs that occur on dry, infertile sandplains. They have beaked follicles and thick, tough, long-lived serrated leaves. It includes all other western taxa, plus the eastern
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in recognition of "the great quantity of plants Mr Banks and Dr Solander found in this place". Over the next seven weeks, Banks and Solander collected thousands of plant specimens, including the first specimens of a new genus that would later be named
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and seed characters, with the criterion that a taxon was considered a distinct species only if it exhibited a "significant and consistent difference in the morphology of flowers and/or fruit". It was the first thorough revision of the taxonomy of
97:, along with secondary characteristics such as leaf structure and growth habit. Increasingly, molecular evidence from DNA is providing important new insights into relationships within the genus and between this and other genera in the Proteaceae. 2181:) are adjusted to achieve a holophyletic naming system. Alex George maintains a firm position against the change, arguing variously that the phylogenetic analyses are flawed and that paraphyly does not necessitate taxonomic changes. 729: 686:, was erected to contain the species that did not belong in the other sections, and was therefore highly heterogeneous. Despite these shortcomings, this arrangement would stand for over 100 years. 2040:(1996)", but none of the four promotions to species rank was accepted, and none of the thirteen infrageneric taxa introduced by Thiele and Ladiges was retained. However, a number of Australian 2735:
Mast, Austin R. (1998) Molecular systematics of subtribe Banksiinae (Banksia and Dryandra; Proteaceae) based on cpDNA and nrDNA sequence data: implications for taxonomy and biogeography.
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differed substantially from the current taxonomic arrangement, they published a revised arrangement that accorded better with their results. Four varieties were promoted to species rank:
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Mast, Austin R., Eric H. Jones and Shawn P. Havery (2005). "An Assessment of Old and New DNA sequence evidence for the Paraphyly of Banksia with respect to Dryandra (Proteaceae)".
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for species with hooked styles. These two sections were then divided into nine and three series respectively. The arrangement into series largely followed Bentham, with series
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is shown to be nested inside, and hence should be included within, Banksia. This interpretation has been controversial in some circles, and is not universally accepted.
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Although the taxonomic legitimacy of tribe Banksiinae is universally recognised, there has been some debate about the legitimacy of the tribe's resolution into genera
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book series; this recognises two subgenera, three sections, 13 series, 77 species, 6 subspecies and 18 varieties, and treats the traditionally related genus
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History in the service of systematics : papers from the Conference to celebrate the centenary of the British Museum (Natural History) 13-16 April, 1981
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industries. However they are threatened by a number of processes, including land clearing, frequent burning, and disease; and a number of species are
2474:. Vol. 17B: Proteaceae 3: Hakea to Dryandra. Collingwood, Victoria: CSIRO Publishing / Australian Biological Resources Study. pp. 175–251. 2225:
Captain Cook's Journal during his First Voyage Round the World made in H. M. Bark "Endeavour" 1768–71: A Literal Transcription of the Original MSS
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species that the Forsters had collected during Cook's second voyage. However Linnaeus incorrectly attributed the Forsters' specimens to the genus
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Supplementum Plantarum Systema Vegetabilium Editionis Decima Tertia, Generum Plantarum Editionis Fexta, Et Specierum Plantarum Editionis Secunda
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L.f.". Linnaeus placed the genus in class Tetrandra, order Monogynia of his father's classification, and named it in honour of Banks. The name
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on a number of grounds of which the most obvious and easily recognised is the occurrence of flowers in condensed heads. The placement of
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Salkin, A. I. (1981). "A Short History of the Discovery and Naming of Banksias in Eastern Australia: Part I, Banks & Solander".
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majority current taxonomic practice, in which only holophyletic groups are named as taxa, and groups found to be paraphyletic (e.g.
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characters, taking the number of sections to five. Three of these sections were fairly well-defined and homogeneous, while another,
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sectional rank, and divided the latter into four series based on leaf properties; these series were all highly heterogeneous.
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have continued to follow Thiele and Ladiges on some points, for example by recognising the four species that they promoted.
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species was reduced to 46, and Meissner's four heterogeneous series were replaced by four sections based on leaf, style and
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species were formally published. By this time, a sixth species had been collected; in 1776, during Cook's third voyage,
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because the individual flowers in its inflorescence occur in pairs. On the basis of certain characters of the
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Mast, Austin R.; Thiele, Kevin R. (2007). "The transfer of Dryandra R.Br. to Banksia L.f. (Proteaceae)".
3024: 3012: 3002: 2876: 2862: 2653: 2011: 1341: 1132: 1124: 1079: 1065: 410: 370:", but for various reasons the project was never completed, and it would be ten years before any of the 338: 987:. This has been put into effect in a 2007 paper by Mast & Thiele, in which all existing species of 367: 3121: 1756: 1714: 1676: 1658: 1402: 1396: 1261: 1238: 1197: 1168: 733: 300: 283:. Cook landed on Australian soil for the first time on 29 April 1770, at a place that he later named 3079: 2930: 2416: 1791: 1564: 1467: 1324: 1162: 666: 231: 77: 3034: 2867: 2855: 2832: 2808: 1871: 1785: 1670: 1641: 1520: 1503: 1479: 1449: 1414: 1361: 821: 589: 385: 306: 219: 2342: 732:". Their arrangement has been refined somewhat over the ensuing three decades, most notably by 3073: 2888: 2716: 2693: 2641: 2603: 2595: 2569: 2543: 2475: 2435: 2295: 2142: 1877: 1838: 1803: 1762: 1699: 1624: 1570: 1491: 1485: 1455: 1437: 1367: 1355: 1349: 1318: 1215: 1191: 841: 725: 513: 438: 380: 271: 2685: 2633: 2585: 2508: 2497:"A new suprageneric classification of the Proteaceae, with an annotated checklist of genera" 2447: 2393: 1844: 1832: 1820: 1664: 1589: 1532: 1390: 1330: 1312: 1252: 1227: 1209: 1203: 1035: 675: 341: 48: 2997: 2923: 2381: 1919: 1883: 1682: 1497: 1306: 1277: 1246: 1156: 1003: 907:. For a number of years this debate centred on similarities between the inflorescences of 809: 447: 326: 321: 294: 266: 60: 25: 3039: 2849: 2451: 2411: 2068:
species, in 2002 and 2005. Their results indicated the presence of two large clades of
2032: 1826: 1797: 1768: 1739: 1547: 1526: 1461: 1408: 1384: 1289: 1283: 834: 815: 762: 745: 653: 195: 105: 2420:. Vol. 5: Myoporineae to Proteaceae. London: L. Reeve & Co. pp. 541–562. 3135: 1473: 1443: 1431: 1221: 1174: 631: 600: 551: 509: 429: 312: 280: 2438:(1975). "On the Proteaceae: the evolution and classification of a southern family". 2223: 2104:) and also the western Spicigerae, Quercinae, Grandes, Abietinae and Dryandroideae. 2377: 1915: 855: 781: 262: 215: 203: 44: 30: 959:
are not especially closely related, but have also provided powerful evidence that
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from Parkinson's sketches. Banks had plans to publish his entire collection as "
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published his 1856 classification of the Proteaceae, there were 58 described
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Koenig was published in 1783, but subsequently determined to be a synonym of
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Flora Australiensis: A Description of the Plants of the Australian Territory
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L.f. has since been challenged a number of times. The later near-homonym
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On the Proteaceae: the evolution and classification of a southern family
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in Banks' honour. Four species were present in this first collection:
2632:(1). CSIRO Publishing / Australian Systematic Botany Society: 75–88. 1047:, which remains the standard text on the genus, and the treatment of 1031: 939:
were matters of superficial appearance, whereas similarities between
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on the basis of its hooked styles, which can be seen clearly here.
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analyses led by Austin Mast have confirmed George's position that
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and co-authors published cladistic analyses of genetic data from
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George, Alex S. (1999). "Banksia". In Wilson, Annette (ed.).
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to trees up to 35 metres tall, and occur in all but the most
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species (Proteaceae: Grevilleoideae) from Western Australia"
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for over a century, and formed the basis for George's 1984
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in 2006. Proteaceae is divided into five subfamilies, with
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for species with straight or slightly curved styles, and
159:(in the traditional sense, not including Dryandra), is a 116:. The most recent complete revision is that published in 2384:(1996). "A Cladistic Analysis of Banksia (Proteaceae)". 51:, who was present when the genus was first collected at 2000:. Two new series and eleven subseries were introduced; 947:
species were far more important diagnostically. Recent
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Prodromus Florae Novae Hollandiae et Insulae Van Diemen
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was published, but this was later determined by be a
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Thiele and Ladiges' taxonomic arrangement of Banksia
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species in his 1830 supplement, placing them all in
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Australian Native Plants Society (Australia) 2528: 2526: 2524: 2052:Following on from an earlier molecular study, 530:proposed to enforce the right of precedent of 495:Bruce, but the name was rejected in favour of 421:had in fact already been published in 1775 as 2793: 2465: 2463: 2461: 2372: 2370: 2368: 2366: 2364: 2362: 2290:L.f. — a case history in Australian botany". 2239: 2237: 2087:were defined as those taxa in which the leaf 2030:was largely redefined. Six species were left 1105:, as provided by George and published in the 206:, and they are of economic importance to the 8: 2658:: CS1 maint: multiple names: authors list ( 2619: 2617: 2560: 2558: 1922:published a cladistic analysis of the genus 796:in Proteaceae can be summarised as follows: 615:in accordance with his 1810 classification. 603:. All other species were placed in subgenus 599:, in recognition of its unusual dome-shaped 2495:Weston, Peter H.; Barker, Nigel P. (2006). 2313: 2311: 1101:This conventional taxonomic arrangement of 136:have provided an alternative view in which 3062: 2838: 2820: 2800: 2786: 2778: 682:, was somewhat heterogeneous. The fourth, 194:areas of Australia. As heavy producers of 2589: 2512: 16:Classification of the plant genus Banksia 2762:Australian Plants Online: ANPSA website 2210: 43:species, it was produced by one of Sir 2651: 2318:Olde, Peter; Neil R. Marriott (2002). 1109:series, may be summarised as follows: 1094:remaining somewhat heterogeneous, and 316:(Swamp Banksia). In June the ship was 2221:(1893). William J. L. Wharton (ed.). 1022:published his classic 1981 monograph 931:argued that the similarities between 728:in their influential 1975 monograph " 362:, and artists were employed to paint 7: 2149:Thus, the results presented by Mast 558:L.f. was formally conserved against 442:, and therefore considered the name 333:Every specimen collected during the 550:L.f. This challenge failed, as did 399:was finally described and named by 330:(Tropical Banksia) were collected. 2452:10.1111/j.1095-8339.1975.tb01644.x 1060:George followed Brown in dividing 772:, both of which occur only in the 534:J.R.Forst & G.Forst, renaming 275:during Lieutenant (later Captain) 14: 3103:Commonly fossilised invertebrates 2060:samples of almost all species of 664:in his landmark 1870 publication 39:. One of the first drawings of a 2157:is paraphyletic with respect to 1098:remaining highly heterogeneous. 963:is paraphyletic with respect to 450:corrected Banks' error in 1788, 72:has traditionally been based on 446:available for use. By the time 2124:, and also appears to include 709:Banksia ser. dryandra sessilis 403:in his April 1782 publication 337:voyage was sketched by Banks' 100:The genus is placed in family 1: 2272:. Brunsvigae: Orphanotrophei. 999:Classical taxonomic treatment 554:'s 1905 challenge. In 1940, 2737:Australian Systematic Botany 2678:Australian Systematic Botany 2626:Australian Systematic Botany 2386:Australian Systematic Botany 2264:Carolus Linnaeus the Younger 1929:Australian Systematic Botany 562:J.R.Forst. & G.Forst by 401:Carolus Linnaeus the Younger 245:Carolus Linnaeus the Younger 2076:" ("hidden stomates") and " 1074:. He then divided subgenus 670:. The number of recognised 577:of the 31 known species of 573:published descriptions and 520:that referred to the genus 461:J.R.Forst & G.Forst to 310:(Heath-leaved Banksia) and 3158: 2578:American Journal of Botany 2341:(1): 85–99. Archived from 2128:. It is worth noting that 1907: 491:Bruce, later corrected to 247:'s April 1782 publication 229: 149: 120:'s 1999 monograph for the 2533:George, Alex S. (1981). " 2191:Cultivated plant taxonomy 780:, are placed in subtribe 542:, and proposing the name 2754:"You don't have to call 2080:" ("visible stomates"). 261:were first collected by 2514:10.7751/telopea20065733 575:a taxonomic arrangement 497:Johann Friedrich Gmelin 378:collected specimens of 279:'s first voyage to the 243:was first described in 179:wildflower and popular 23:A 1770s watercolour of 2153:strongly suggest that 1015: 991:were transferred into 923:to be moved across to 713: 640:Meissner's arrangement 406:Supplementum Plantarum 384:(Silver Banksia) from 254: 250:Supplementum Plantarum 56: 2752:George, Alex (2008). 2711:George, Alex (1996). 2591:10.3732/ajb.89.8.1311 1010:is placed in section 1006: 915:species and those of 702:was mereged into the 697: 339:botanical illustrator 324:, where specimens of 269:, naturalists on the 239: 22: 2246:Victorian Naturalist 2072:, which they named " 2020:were discarded; and 1998:B. dolichostyla 1989:B. sphaerocarpa 1970:B. brevidentata 1757:B. sphaerocarpa 1715:B. dryandroides 1677:B. occidentalis 1659:B. verticillata 1403:B. blechnifolia 1397:B. chamaephyton 1198:B. integrifolia 1169:B. oblongifolia 1064:into two subgenera, 1034:, pollen-presenter, 760:. Two small genera, 744:placed in subfamily 505:. In 1820 the name 432:, referring to some 301:B. integrifolia 3142:Taxonomy of Banksia 2414:(1870). "Banksia". 2382:Ladiges, Pauline Y. 2286:(1981). "The genus 2230:. London: E. Stock. 1946:B. penicillata 1792:B. leptophylla 1565:B. quercifolia 1468:B. benthamiana 1325:B. candolleana 1163:B. plagiocarpa 1078:into two sections: 788:is placed subtribe 667:Flora Australiensis 546:Kuntze in place of 232:Timeline of Banksia 2472:Flora of Australia 2064:, along with five 1904:Thiele and Ladiges 1899:Cladistic analyses 1872:B. ilicifolia 1786:B. telmatiaea 1671:B. littoralis 1642:B. ericifolia 1521:B. lemanniana 1504:B. lindleyana 1480:B. lullfitzii 1450:B. pilostylis 1415:B. petiolaris 1362:B. hookeriana 1107:Flora of Australia 1054:Flora of Australia 1016: 1008:B. ericifolia 971:is a sub-group of 822:Symphionematoideae 714: 593:alone in subgenus 590:B. ilicifolia 413:for the genus is " 386:South Bruny Island 368:Banks' Florilegium 307:B. ericifolia 267:Dr Daniel Solander 255: 134:cladistic analyses 123:Flora of Australia 91:fruiting structure 80:properties of the 57: 3129: 3128: 3111: 3110: 3093:Cultivated plants 3088: 3087: 3052: 3051: 2570:Thomas J. Givnish 2539:L.f. (Proteaceae) 2432:Johnson, L. A. S. 2398:10.1071/SB9960661 2143:B. attenuata 1878:B. oligantha 1839:B. pulchella 1804:B. scabrella 1763:B. micrantha 1700:B. tricuspis 1625:B. spinulosa 1571:B. oreophila 1492:B. laevigata 1486:B. elderiana 1456:B. attenuata 1438:B. praemorsa 1368:B. victoriae 1356:B. burdettii 1350:B. prionotes 1319:B. menziesii 1216:B. marginata 1192:B. aquilonia 1028:L.f. (Proteaceae) 979:would be to sink 698:The former genus 625:Stephan Endlicher 381:B. marginata 304:(Coast Banksia), 226:Taxonomic history 3149: 3063: 2839: 2821: 2802: 2795: 2788: 2779: 2773: 2772: 2770: 2769: 2756:Dryandra Banksia 2749: 2743: 2733: 2727: 2726: 2713:The Banksia Book 2708: 2702: 2701: 2673: 2664: 2663: 2657: 2649: 2621: 2612: 2611: 2593: 2584:(8): 1311–1323. 2562: 2553: 2552: 2530: 2519: 2518: 2516: 2492: 2486: 2485: 2467: 2456: 2455: 2428: 2422: 2421: 2408: 2402: 2401: 2374: 2357: 2356: 2354: 2353: 2347: 2332: 2315: 2306: 2305: 2280: 2274: 2273: 2260: 2254: 2253: 2241: 2232: 2231: 2229: 2215: 1984:B. hiemalis 1975:B. gardneri 1961:B. gardneri 1951:B. conferta 1938:B. conferta 1845:B. meisneri 1833:B. laricina 1821:B. violacea 1665:B. seminuda 1590:B. coccinea 1533:B. aculeata 1391:B. gardneri 1331:B. sceptrum 1313:B. speciosa 1253:B. solandri 1228:B. saxicola 1210:B. paludosa 1204:B. conferta 1045:The Banksia Book 722:L. A. S. Johnson 676:pollen-presenter 350: 342:Sydney Parkinson 263:Sir Joseph Banks 61:flowering plants 49:Sydney Parkinson 3157: 3156: 3152: 3151: 3150: 3148: 3147: 3146: 3132: 3131: 3130: 3125: 3107: 3084: 3048: 2944: 2911:B. integrifolia 2837: 2812: 2806: 2776: 2767: 2765: 2751: 2750: 2746: 2734: 2730: 2723: 2710: 2709: 2705: 2690:10.1071/SB06016 2675: 2674: 2667: 2650: 2638:10.1071/SB04015 2623: 2622: 2615: 2572:(August 2002). 2566:Mast, Austin R. 2564: 2563: 2556: 2532: 2531: 2522: 2494: 2493: 2489: 2482: 2469: 2468: 2459: 2430: 2429: 2425: 2412:Bentham, George 2410: 2409: 2405: 2376: 2375: 2360: 2351: 2349: 2345: 2330: 2317: 2316: 2309: 2302: 2282: 2281: 2277: 2262: 2261: 2257: 2243: 2242: 2235: 2217: 2216: 2212: 2208: 2187: 2138:B. elegans 2114:B. serrata 2094:B. serrata 2085:/Phanerostomata 2078:/Phanerostomata 2050: 1926:in the journal 1920:Pauline Ladiges 1912: 1906: 1901: 1884:B. cuneata 1683:B. brownii 1498:B. elegans 1307:B. baxteri 1278:B. serrata 1247:B. grandis 1157:B. dentata 1001: 810:Bellendenoideae 734:Peter H. Weston 692: 658:his arrangement 448:Joseph Gaertner 348: 327:B. dentata 322:Endeavour River 298:(Saw Banksia), 295:B. serrata 234: 228: 154: 148: 26:Banksia serrata 17: 12: 11: 5: 3155: 3153: 3145: 3144: 3134: 3133: 3127: 3126: 3124: 3119: 3117: 3113: 3112: 3109: 3108: 3106: 3105: 3100: 3095: 3089: 3086: 3085: 3083: 3082: 3077: 3069: 3067: 3060: 3054: 3053: 3050: 3049: 3047: 3046: 3045: 3044: 3037: 3027: 3022: 3017: 3010: 3005: 3000: 2995: 2990: 2985: 2980: 2979: 2978: 2973: 2968: 2960: 2952: 2950: 2946: 2945: 2943: 2942: 2935: 2928: 2921: 2916: 2915: 2914: 2900: 2893: 2886: 2881: 2874: 2873: 2872: 2860: 2853: 2845: 2843: 2836: 2835: 2829: 2827: 2818: 2814: 2813: 2807: 2805: 2804: 2797: 2790: 2782: 2775: 2774: 2744: 2728: 2721: 2703: 2665: 2613: 2554: 2520: 2507:(3): 314–344. 2487: 2480: 2457: 2436:Barbara Briggs 2423: 2403: 2392:(5): 661–733. 2358: 2307: 2300: 2275: 2255: 2233: 2209: 2207: 2204: 2203: 2202: 2193: 2186: 2183: 2177:not including 2122:B. ornata 2118:B. aemula 2109:/Cryptostomata 2102:B. ornata 2098:B. aemula 2074:/Cryptostomata 2049: 2046: 2033:incertae sedis 1908:Main article: 1905: 1902: 1900: 1897: 1896: 1895: 1894: 1893: 1892: 1891: 1890: 1889: 1888: 1887: 1856: 1855: 1854: 1853: 1852: 1851: 1850: 1849: 1848: 1827:B. incana 1809: 1808: 1807: 1798:B. lanata 1774: 1773: 1772: 1769:B. grossa 1745: 1744: 1743: 1740:B. nutans 1720: 1719: 1718: 1705: 1704: 1703: 1690: 1689: 1688: 1687: 1686: 1647: 1646: 1645: 1630: 1629: 1628: 1597: 1596: 1595: 1594: 1593: 1578: 1577: 1576: 1575: 1574: 1553: 1552: 1551: 1548:B. baueri 1538: 1537: 1536: 1527:B. caleyi 1509: 1508: 1507: 1462:B. ashbyi 1420: 1419: 1418: 1409:B. repens 1385:B. goodii 1373: 1372: 1371: 1338: 1337: 1336: 1335: 1334: 1295: 1294: 1293: 1290:B. ornata 1284:B. aemula 1258: 1257: 1256: 1235: 1234: 1233: 1232: 1231: 1180: 1179: 1178: 1000: 997: 897: 896: 895: 894: 893: 892: 886: 880: 879: 878: 877: 876: 858: 844: 835:Grevilleoideae 830: 824: 818: 816:Persoonioideae 812: 763:Austromuellera 746:Grevilleoideae 726:Barbara Briggs 691: 688: 654:George Bentham 564:Thomas Sprague 227: 224: 147: 144: 106:Grevilleoideae 59:As with other 15: 13: 10: 9: 6: 4: 3: 2: 3154: 3143: 3140: 3139: 3137: 3123: 3120: 3118: 3114: 3104: 3101: 3099: 3098:Invertebrates 3096: 3094: 3091: 3090: 3081: 3078: 3076: 3075: 3071: 3070: 3068: 3064: 3061: 3059: 3055: 3043: 3042: 3038: 3036: 3033: 3032: 3031: 3028: 3026: 3023: 3021: 3018: 3016: 3015: 3011: 3009: 3006: 3004: 3001: 2999: 2996: 2994: 2991: 2989: 2986: 2984: 2981: 2977: 2974: 2972: 2969: 2967: 2964: 2963: 2961: 2959: 2958: 2954: 2953: 2951: 2947: 2941: 2940: 2936: 2934: 2933: 2929: 2927: 2926: 2922: 2920: 2917: 2913: 2912: 2908: 2907: 2906: 2905: 2901: 2899: 2898: 2894: 2892: 2891: 2887: 2885: 2882: 2880: 2879: 2875: 2871: 2870: 2866: 2865: 2864: 2861: 2859: 2858: 2854: 2852: 2851: 2847: 2846: 2844: 2840: 2834: 2831: 2830: 2828: 2826: 2822: 2819: 2815: 2810: 2803: 2798: 2796: 2791: 2789: 2784: 2783: 2780: 2763: 2759: 2757: 2748: 2745: 2741: 2738: 2732: 2729: 2724: 2722:0-86417-818-2 2718: 2714: 2707: 2704: 2699: 2695: 2691: 2687: 2683: 2679: 2672: 2670: 2666: 2661: 2655: 2647: 2643: 2639: 2635: 2631: 2627: 2620: 2618: 2614: 2609: 2605: 2601: 2597: 2592: 2587: 2583: 2579: 2575: 2571: 2567: 2561: 2559: 2555: 2551:(3): 239–473. 2550: 2546: 2545: 2540: 2538: 2529: 2527: 2525: 2521: 2515: 2510: 2506: 2502: 2498: 2491: 2488: 2483: 2481:0-643-06454-0 2477: 2473: 2466: 2464: 2462: 2458: 2453: 2449: 2446:(2): 83–182. 2445: 2441: 2437: 2433: 2427: 2424: 2419: 2418: 2413: 2407: 2404: 2399: 2395: 2391: 2387: 2383: 2379: 2378:Thiele, Kevin 2373: 2371: 2369: 2367: 2365: 2363: 2359: 2348:on 2006-09-23 2344: 2340: 2336: 2329: 2327: 2323: 2314: 2312: 2308: 2303: 2301:0-901843-05-9 2297: 2293: 2289: 2285: 2284:George, A. S. 2279: 2276: 2271: 2270: 2265: 2259: 2256: 2251: 2247: 2240: 2238: 2234: 2228: 2226: 2220: 2214: 2211: 2205: 2201: 2199: 2194: 2192: 2189: 2188: 2184: 2182: 2180: 2176: 2170: 2168: 2164: 2160: 2156: 2152: 2147: 2145: 2144: 2139: 2135: 2131: 2127: 2123: 2119: 2115: 2110: 2105: 2103: 2099: 2095: 2090: 2086: 2081: 2079: 2075: 2071: 2067: 2063: 2059: 2055: 2047: 2045: 2043: 2037: 2035: 2034: 2029: 2028: 2024: 2019: 2018: 2014: 2009: 2008: 2004: 1999: 1995: 1994: 1990: 1985: 1981: 1980: 1976: 1971: 1967: 1966: 1962: 1957: 1956: 1952: 1947: 1943: 1939: 1935: 1931: 1930: 1925: 1921: 1917: 1911: 1903: 1898: 1886: 1885: 1880: 1879: 1874: 1873: 1869: 1868: 1867: 1866: 1865: 1864: 1863: 1862: 1857: 1847: 1846: 1841: 1840: 1835: 1834: 1829: 1828: 1823: 1822: 1818: 1817: 1816: 1815: 1810: 1806: 1805: 1800: 1799: 1794: 1793: 1788: 1787: 1783: 1782: 1781: 1780: 1775: 1771: 1770: 1765: 1764: 1759: 1758: 1754: 1753: 1752: 1751: 1750:Sphaerocarpae 1746: 1742: 1741: 1737: 1736: 1735: 1734: 1729: 1728: 1727: 1726: 1721: 1717: 1716: 1712: 1711: 1710: 1706: 1702: 1701: 1697: 1696: 1695: 1691: 1685: 1684: 1679: 1678: 1673: 1672: 1667: 1666: 1661: 1660: 1656: 1655: 1654: 1653: 1648: 1644: 1643: 1639: 1638: 1637: 1636: 1631: 1627: 1626: 1622: 1621: 1620: 1619: 1614: 1613: 1612: 1611: 1606: 1605: 1604: 1603: 1598: 1592: 1591: 1587: 1586: 1585: 1584: 1583: 1579: 1573: 1572: 1567: 1566: 1562: 1561: 1560: 1559: 1554: 1550: 1549: 1545: 1544: 1543: 1539: 1535: 1534: 1529: 1528: 1523: 1522: 1518: 1517: 1516: 1515: 1510: 1506: 1505: 1500: 1499: 1494: 1493: 1488: 1487: 1482: 1481: 1476: 1475: 1474:B. audax 1470: 1469: 1464: 1463: 1458: 1457: 1452: 1451: 1446: 1445: 1444:B. epica 1440: 1439: 1434: 1433: 1432:B. media 1429: 1428: 1427: 1426: 1421: 1417: 1416: 1411: 1410: 1405: 1404: 1399: 1398: 1393: 1392: 1387: 1386: 1382: 1381: 1380: 1379: 1374: 1370: 1369: 1364: 1363: 1358: 1357: 1352: 1351: 1347: 1346: 1345: 1344: 1339: 1333: 1332: 1327: 1326: 1321: 1320: 1315: 1314: 1309: 1308: 1304: 1303: 1302: 1301: 1296: 1292: 1291: 1286: 1285: 1280: 1279: 1275: 1274: 1273: 1272: 1267: 1266: 1265: 1264: 1259: 1255: 1254: 1249: 1248: 1244: 1243: 1242: 1241: 1236: 1230: 1229: 1224: 1223: 1222:B. canei 1218: 1217: 1212: 1211: 1206: 1205: 1200: 1199: 1194: 1193: 1189: 1188: 1187: 1186: 1185:Integrifoliae 1181: 1177: 1176: 1175:B. robur 1171: 1170: 1165: 1164: 1159: 1158: 1154: 1153: 1152: 1151: 1146: 1145: 1144: 1143: 1138: 1137: 1136: 1135: 1130: 1129: 1128: 1127: 1122: 1121: 1120: 1119: 1118: 1112: 1111: 1110: 1108: 1104: 1099: 1097: 1093: 1089: 1088: 1083: 1082: 1077: 1073: 1069: 1068: 1063: 1058: 1056: 1055: 1050: 1046: 1042: 1037: 1033: 1029: 1025: 1021: 1013: 1009: 1005: 998: 996: 994: 990: 986: 982: 978: 974: 970: 966: 962: 958: 954: 950: 946: 942: 938: 934: 930: 926: 922: 918: 914: 910: 906: 902: 891: 887: 885: 881: 875: 874: 873: 867: 866: 865: 864: 859: 857: 853: 852: 851: 850: 845: 843: 839: 838: 837: 836: 831: 829: 825: 823: 819: 817: 813: 811: 807: 806: 805: 804: 799: 798: 797: 795: 791: 787: 783: 779: 775: 771: 770: 765: 764: 759: 755: 751: 750:leaf venation 747: 743: 739: 735: 731: 727: 723: 719: 711: 710: 705: 701: 696: 689: 687: 685: 681: 677: 673: 669: 668: 663: 659: 655: 651: 649: 645: 641: 637: 633: 632:Carl Meissner 628: 626: 622: 618: 617:Banksia verae 614: 613:Banksia verae 610: 606: 605:Banksia verae 602: 601:inflorescence 598: 597: 592: 591: 586: 585: 580: 576: 572: 567: 565: 561: 557: 553: 552:James Britten 549: 545: 541: 537: 533: 529: 525: 524: 519: 515: 511: 508: 504: 503: 498: 494: 490: 486: 482: 481: 476: 472: 468: 466: 465: 460: 457: 453: 449: 445: 441: 440: 435: 431: 427: 424: 420: 416: 412: 408: 407: 402: 398: 393: 391: 387: 383: 382: 377: 373: 369: 365: 361: 358: 354: 347: 343: 340: 336: 331: 329: 328: 323: 319: 315: 314: 313:B. robur 309: 308: 303: 302: 297: 296: 291: 286: 282: 281:Pacific Ocean 278: 274: 273: 268: 264: 260: 257:Specimens of 252: 251: 246: 242: 238: 233: 225: 223: 221: 217: 213: 209: 205: 204:honey possums 201: 197: 193: 189: 185: 182: 178: 174: 170: 166: 163:of around 80 162: 158: 153: 145: 143: 141: 140: 135: 131: 130: 125: 124: 119: 115: 112:and subtribe 111: 107: 103: 98: 96: 92: 88: 85: 84: 79: 78:morphological 75: 71: 70: 69: 62: 54: 50: 46: 42: 38: 37: 32: 28: 27: 21: 3072: 3058:Polyphyletic 3041:Homo sapiens 3040: 3013: 2955: 2938: 2931: 2924: 2910: 2903: 2902: 2896: 2889: 2877: 2868: 2856: 2848: 2825:Monophyletic 2766:. Retrieved 2761: 2755: 2747: 2739: 2736: 2731: 2712: 2706: 2684:(1): 63–71. 2681: 2677: 2654:cite journal 2629: 2625: 2581: 2577: 2548: 2542: 2536: 2504: 2500: 2490: 2471: 2443: 2439: 2426: 2415: 2406: 2389: 2385: 2350:. Retrieved 2343:the original 2338: 2334: 2325: 2324:and two new 2321: 2291: 2287: 2278: 2268: 2258: 2249: 2245: 2224: 2213: 2197: 2178: 2174: 2171: 2166: 2162: 2158: 2154: 2150: 2148: 2141: 2137: 2133: 2129: 2125: 2121: 2117: 2113: 2108: 2106: 2101: 2097: 2093: 2084: 2082: 2077: 2073: 2069: 2065: 2061: 2051: 2048:DNA analysis 2038: 2031: 2026: 2022: 2016: 2012: 2006: 2002: 1997: 1993:dolichostyla 1992: 1988: 1983: 1978: 1974: 1969: 1965:brevidentata 1964: 1960: 1954: 1950: 1945: 1941: 1937: 1927: 1923: 1916:Kevin Thiele 1913: 1882: 1876: 1870: 1860: 1843: 1837: 1831: 1825: 1819: 1813: 1802: 1796: 1790: 1784: 1779:Leptophyllae 1778: 1767: 1761: 1755: 1749: 1738: 1732: 1724: 1713: 1709:Dryandroidae 1708: 1698: 1693: 1681: 1675: 1669: 1663: 1657: 1652:Occidentales 1651: 1640: 1634: 1623: 1617: 1609: 1601: 1588: 1581: 1569: 1563: 1557: 1546: 1541: 1531: 1525: 1519: 1513: 1502: 1496: 1490: 1484: 1478: 1472: 1466: 1460: 1454: 1448: 1442: 1436: 1430: 1424: 1413: 1407: 1401: 1395: 1389: 1383: 1377: 1366: 1360: 1354: 1348: 1342: 1329: 1323: 1317: 1311: 1305: 1299: 1288: 1282: 1276: 1270: 1262: 1251: 1245: 1239: 1226: 1220: 1214: 1208: 1202: 1196: 1190: 1184: 1173: 1167: 1161: 1155: 1149: 1141: 1133: 1125: 1115: 1113: 1106: 1102: 1100: 1095: 1091: 1086: 1080: 1075: 1071: 1066: 1061: 1059: 1052: 1048: 1044: 1040: 1027: 1023: 1017: 1011: 1007: 992: 988: 984: 980: 972: 968: 964: 960: 956: 952: 944: 940: 936: 932: 928: 924: 920: 916: 912: 908: 904: 900: 898: 870: 868: 860: 856:Musgraveinae 846: 832: 800: 793: 785: 782:Musgraveinae 767: 761: 752:, hairs and 741: 738:Nigel Barker 720:was laid by 715: 707: 703: 683: 679: 671: 665: 661: 652: 647: 643: 635: 630:By the time 629: 620: 619:was renamed 616: 612: 608: 604: 595: 588: 587:. He placed 582: 578: 571:Robert Brown 568: 559: 555: 547: 543: 539: 535: 531: 526:, In 1891, 521: 506: 500: 492: 488: 478: 474: 470: 469: 462: 458: 451: 443: 437: 422: 418: 414: 409:; hence the 404: 396: 394: 379: 376:David Nelson 371: 364:watercolours 351:s return to 345: 334: 332: 325: 311: 305: 299: 293: 289: 270: 258: 256: 248: 240: 175:. An iconic 156: 155: 137: 127: 121: 104:, subfamily 99: 81: 66: 65:taxonomy of 64: 58: 45:Joseph Banks 40: 34: 31:type species 24: 3030:Vertebrates 3020:Lepidoptera 2993:Brachiopoda 2962:Gastropoda 2939:Pachypodium 2884:Orchidaceae 2219:Cook, James 2054:Austin Mast 2027:Cyrtostylis 1955:penicillata 1942:penicillata 1932:. As their 1814:Longistyles 1694:Tricuspidae 1635:Ericifoliae 1425:Cyrtostylis 1300:Cratistylis 1096:Cyrtostylis 1092:Orthostylis 1020:Alex George 890:Macadamieae 884:Embothrieae 828:Proteoideae 774:rainforests 684:Cyrtostylis 680:Orthostylis 544:Sirmuellera 528:Otto Kuntze 518:nomen nudum 485:James Bruce 483:L. In 1790 434:New Zealand 200:honeyeaters 118:Alex George 3008:Dermaptera 3003:Scorpiones 2897:Adenanthos 2768:2009-12-16 2535:The Genus 2352:2007-01-10 2206:References 2007:Oncostylis 1811:Subseries 1776:Subseries 1747:Subseries 1730:Subseries 1649:Subseries 1632:Subseries 1618:Spinulosae 1615:Subseries 1610:Spicigerae 1602:Oncostylis 1514:Tetragonae 1378:Prostratae 1297:Subseries 1268:Subseries 1182:Subseries 1147:Subseries 1087:Oncostylis 1024:The Genus 1012:Oncostylis 967:(that is, 943:and other 929:status quo 863:Banksiinae 833:Subfamily 826:Subfamily 820:Subfamily 814:Subfamily 808:Subfamily 803:Proteaceae 790:Banksiinae 778:Queensland 718:Proteaceae 712:pictured). 706:in 2007. ( 656:published 487:published 395:The genus 285:Botany Bay 277:James Cook 230:See also: 220:endangered 212:cut flower 177:Australian 173:Proteaceae 150:See also: 146:Background 114:Banksiinae 102:Proteaceae 74:anatomical 53:Botany Bay 3035:Lemuridae 3025:Lucanidae 3014:Anopheles 2919:Cactaceae 2878:Narcissus 2863:Liliaceae 2742:: 321–342 2698:1030-1887 2646:1030-1887 2600:0002-9122 2326:Grevillea 2320:"One new 2140:and then 2134:Isostylis 1934:cladogram 1914:In 1996, 1861:Isostylis 1858:Subgenus 1725:Abietinae 1558:Quercinae 1542:Bauerinae 1142:Salicinae 1123:Subgenus 1072:Isostylis 1018:In 1981, 977:monophyly 957:Isostylis 941:Isostylis 933:Isostylis 921:Isostylis 913:Isostylis 861:Subtribe 854:Subtribe 849:Banksieae 769:Musgravea 758:Banksieae 648:Eubanksia 644:Isostylis 638:species. 627:in 1847. 621:Eubanksia 596:Isostylis 569:In 1810, 439:Passerina 426:J.R.Forst 411:full name 360:herbarium 346:Endeavour 344:. On the 335:Endeavour 272:Endeavour 110:Banksieae 3136:Category 3080:Diatomea 3074:Protista 3066:Protists 2988:Bivalvia 2983:Conoidea 2957:Animalia 2890:Triticum 2833:Bacteria 2809:Taxonomy 2608:21665734 2266:(1782). 2196:List of 2185:See also 2179:Dryandra 2163:Dryandra 2159:Dryandra 2130:Dryandra 2126:Dryandra 2112:species 2066:Dryandra 2042:herbaria 2017:Crocinae 1979:hiemalis 1881: – 1875: – 1842: – 1836: – 1830: – 1824: – 1801: – 1795: – 1789: – 1766: – 1760: – 1733:Nutantes 1680: – 1674: – 1668: – 1662: – 1599:Section 1582:Coccinea 1580:Section 1568: – 1530: – 1524: – 1501: – 1495: – 1489: – 1483: – 1477: – 1471: – 1465: – 1459: – 1453: – 1447: – 1441: – 1435: – 1412: – 1406: – 1400: – 1394: – 1388: – 1365: – 1359: – 1353: – 1343:Crocinae 1328: – 1322: – 1316: – 1310: – 1287: – 1281: – 1250: – 1225: – 1219: – 1213: – 1207: – 1201: – 1195: – 1172: – 1166: – 1160: – 1150:Acclives 1131:Section 1057:series. 1036:follicle 989:Dryandra 981:Dryandra 969:Dryandra 965:Dryandra 953:Dryandra 937:Dryandra 925:Dryandra 917:Dryandra 905:Dryandra 842:Roupalae 700:Dryandra 499:'s name 489:Bankesia 390:Tasmania 318:careened 139:Dryandra 129:Dryandra 108:, tribe 3122:Viruses 2998:Araneae 2949:Animals 2932:Drosera 2904:Banksia 2850:Plantae 2544:Nuytsia 2537:Banksia 2501:Telopea 2335:Nuytsia 2322:Banksia 2288:Banksia 2200:species 2198:Banksia 2175:Banksia 2167:Banksia 2155:Banksia 2092:except 2089:stomata 2070:Banksia 2062:Banksia 2023:Banksia 2013:Banksia 2003:Banksia 1953:subsp. 1924:Banksia 1722:Series 1707:Series 1692:Series 1607:Series 1555:Series 1540:Series 1511:Series 1422:Series 1375:Series 1340:Series 1271:Banksia 1263:Banksia 1260:Series 1240:Grandes 1237:Series 1139:Series 1134:Banksia 1126:Banksia 1117:Banksia 1103:Banksia 1081:Banksia 1076:Banksia 1067:Banksia 1062:Banksia 1051:in the 1049:Banksia 1041:Banksia 1026:Banksia 993:Banksia 985:Banksia 973:Banksia 961:Banksia 945:Banksia 909:Banksia 901:Banksia 872:Banksia 801:Family 794:Banksia 786:Banksia 742:Banksia 704:Banksia 672:Banksia 662:Banksia 660:of the 636:Banksia 609:Banksia 581:in his 579:Banksia 560:Banksia 556:Banksia 548:Banksia 540:Banksia 536:Pimelea 532:Banksia 507:Banksia 502:Hagenia 493:Banksia 475:Banksea 471:Banksia 464:Pimelea 459:Banksia 456:renamed 452:Banksia 444:Banksia 430:G.Forst 423:Banksia 419:Banksia 415:Banksia 397:Banksia 372:Banksia 353:England 290:Banksia 259:Banksia 241:Banksia 208:nursery 171:family 167:in the 165:species 157:Banksia 152:Banksia 83:Banksia 68:Banksia 41:Banksia 36:Banksia 2925:Citrus 2869:Tulipa 2857:Allium 2842:Plants 2811:of ... 2719:  2696:  2644:  2606:  2598:  2478:  2298:  2227:  2151:et al. 2005:sect. 1986:; and 1114:Genus 911:subg. 888:Tribe 882:Tribe 869:Genus 847:Tribe 840:Tribe 754:pollen 523:Cuphea 510:Dombey 480:Costus 428:& 357:London 196:nectar 188:shrubs 181:garden 87:flower 63:, the 29:, the 3116:Other 2346:(PDF) 2331:(PDF) 2165:into 2025:ser. 2015:ser. 1991:var. 1977:var. 1963:var. 1948:(now 1940:var. 1032:style 983:into 642:gave 349:' 184:plant 169:plant 161:genus 2976:1997 2971:2005 2966:2017 2817:Life 2717:ISBN 2694:ISSN 2660:link 2642:ISSN 2604:PMID 2596:ISSN 2476:ISBN 2296:ISBN 2252:(2). 2120:and 2107:The 2100:and 2083:The 2010:and 1918:and 1070:and 955:and 935:and 903:and 766:and 736:and 724:and 646:and 265:and 218:and 216:rare 210:and 202:and 192:arid 95:seed 93:and 76:and 2686:doi 2634:doi 2586:doi 2541:". 2509:doi 2448:doi 2394:doi 2058:DNA 1996:to 1982:to 1968:to 1958:); 1944:to 949:DNA 776:of 623:by 538:to 514:DC. 512:ex 320:at 33:of 3138:: 2760:. 2740:11 2692:. 2682:20 2680:. 2668:^ 2656:}} 2652:{{ 2640:. 2630:18 2628:. 2616:^ 2602:. 2594:. 2582:89 2580:. 2576:. 2568:; 2557:^ 2547:. 2523:^ 2505:11 2503:. 2499:. 2460:^ 2444:70 2442:. 2434:; 2388:. 2380:; 2361:^ 2339:15 2337:. 2333:. 2310:^ 2250:98 2248:. 2236:^ 2146:. 2116:, 2096:, 2036:. 1972:; 995:. 784:. 566:. 392:. 388:, 222:. 89:, 2801:e 2794:t 2787:v 2771:. 2758:" 2725:. 2700:. 2688:: 2662:) 2648:. 2636:: 2610:. 2588:: 2549:3 2517:. 2511:: 2484:. 2454:. 2450:: 2400:. 2396:: 2390:9 2355:. 2304:. 253:. 55:.

Index


Banksia serrata
type species
Banksia
Joseph Banks
Sydney Parkinson
Botany Bay
flowering plants
Banksia
anatomical
morphological
Banksia
flower
fruiting structure
seed
Proteaceae
Grevilleoideae
Banksieae
Banksiinae
Alex George
Flora of Australia
Dryandra
cladistic analyses
Dryandra
Banksia
genus
species
plant
Proteaceae
Australian

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