641:
793:
612:
626:
781:
752:
769:
516:
598:, females matured and stopped growing before males, which is often seen in extant polygynous mammals. Males may have fought for mating rights; healed wounds on skulls have been observed, and healed broken ribs are not uncommon (although not all have their sexes determined). This is further supported by the breeding age female-to-male ratio in the Ashfall Fossil Beds being 4.25:1. There is also a rarity of young adult males preserved at Ashfall, which may be accounted for if they formed bachelor herds away from females and dominant bulls.
127:
657:
495:
170:
144:
1231:
716:
Most of the skeletons are adult females and young, the breeding age female-to-male ratio being 4.25:1. There is also a rarity of young adult males. If the rhinos at
Ashfall represent a herd, this may be accounted for if young adult males formed bachelor herds away from females and dominant bulls. The
780:
640:
792:
611:
552:
suggested in 1898 that it was semi-aquatic and hippo-like in habits. This idea persisted for about a century, but has recently been discounted by isotopic evidence.
1319:
1127:"The geology and paleontology of Ashfall Fossil Beds, a late Miocene (Clarendonian) mass-death assemblage, Antelope County and adjacent Knox County, Nebraska, USA"
625:
594:
was sexually dimorphic. Males were larger, with larger tusks (lower incisors), a more massive head and neck, and significantly larger forelimbs. As a result of
583:
or mixed feeding (both browsing and grazing) diet. Carbon and oxygen isotope analysis of tooth enamel suggests hippo-like grazing habits, but not aquatic.
1380:
1405:
1370:
1306:
846:
1400:
1146:
710:
728:
skeletons at the site, of which most were preserved in a nearly complete state. One extraordinary specimen includes the remains of a
1218:
1204:
1131:
Geologic Field Trips along the
Boundary between the Central Lowlands and Great Plains: 2014 Meeting of the GSA North-Central Section
705:
was buried above the rest, being the last of the animals to succumb (small animals died faster), several weeks or months after the
888:(Rhinocerotidae) through an interval of changing climate and vegetation in the Neogene of the Great Plains, central United States"
695:
751:
1365:
1087:
1027:
1375:
1360:
768:
1167:
724:
The greatest concentration of
Ashfall fossils is housed in a building called the "Rhino Barn", due to the prevalence of
135:
515:
169:
927:
1103:
126:
1026:
Mihlbachler, Matthew C.; Campbell, Daniel; Chen, Charlotte; Ayoub, Michael; Kaur, Pawandeep (February 2018).
1246:
698:
549:
1235:
988:
850:
1390:
1332:
960:
156:
721:
herds, as amongst the skeletons, 54% are immature, 30% are young adults, and 16% are older adults.
1395:
1293:
1039:
899:
706:
580:
1385:
1126:
742:
683:
544:
had much shorter legs than modern rhinos, and a barrel chest, making its build more like that of a
429:
1063:
457:
368:
164:
1337:
989:"A new species of Teleoceras (Mammalia, Rhinocerotidae) from the late Hemphillian of Tennessee"
953:"A New Species of Teleoceras (Mammalia, Rhinocerotidae) from the Late Hemphillian of Tennessee"
656:
494:
440:
from around 17.5 to 4.9 million years ago. It grew up to lengths of 13 feet (4.0 meters) long.
1324:
1214:
1200:
1142:
1107:
1055:
1008:
595:
264:
1134:
1099:
1047:
1000:
907:
572:
468:
1284:
973:
1088:"Sexual dimorphism and paleoecology in Teleoceras , a North American Miocene rhinoceros"
1043:
903:
571:) molar teeth, which has historically led to suggestions that the species were grazers.
1028:"Microwear–mesowear congruence and mortality bias in rhinoceros mass-death assemblages"
759:
244:
221:
1354:
952:
526:
433:
418:
84:
1067:
718:
545:
456:, most likely due to rapid climate cooling, increased seasonality and expansion of
414:
273:
143:
17:
912:
879:
475:, dated to 4.5-5 million years ago, hosts one of the latest-known populations of
1278:
559:
have a small nasal horn, but this appears to be absent in other species such as
453:
437:
59:
45:
1138:
231:
104:
69:
41:
1269:
1171:
1111:
1059:
1012:
1004:
568:
472:
181:
109:
53:
1230:
1263:
687:
576:
448:
426:
201:
99:
94:
79:
74:
64:
1051:
757:(1) 8-9 year old young adult male "Tusker". Wisdom teeth emerging. (2)
1311:
422:
114:
89:
1125:
Tucker, S.T.; Otto, R.E.; Joeckel, R.M.; Voorhies, M.R. (April 2014),
211:
191:
1240:
531:
410:
1298:
786:
10-12 year old adult male "Dr. Marie", with all permanent teeth.
713:(HPOD), as a result of lung failure from the fine volcanic ash.
504:
1244:
987:
Short, Rachel; Wallace, Steven; Emmert, Laura (2019-04-27).
409:(Greek: "perfect" (teleos), "horn" (keratos)) is an extinct
1104:
10.1666/0094-8373(2000)026<0689:SDAPIT>2.0.CO;2
463:
grasses, as isotopic evidence suggests that the uptake of C
467:
plants was far less than that in contemporary horses. The
1213:
709:. Their skeletons show evidence of bone disease, ie
1253:
1199:Columbia University Press, New York, 631 pp.
1197:Classification of Mammals Above the Species Level.
717:age demographic is very similar to that of modern
993:Bulletin of the Florida Museum of Natural History
892:Palaeogeography, Palaeoclimatology, Palaeoecology
1133:, Geological Society of America, pp. 1–22,
548:than a modern rhino. Based on this description,
1195:McKenna, Malcolm C., and Bell, Susan K. 1997.
847:"Glossary. American Museum of Natural History"
1211:The Evolution of North American Rhinoceroses.
8:
774:(3) Adult female "Sandy" (4) Calf "Justin".
1241:
873:
871:
869:
142:
125:
31:
911:
694:skeletons are preserved in ash from the
446:went extinct in North America alongside
811:
798:Fetus within birth canal of the pelvis.
732:calf trying to suckle from its mother.
969:
958:
1081:
1079:
1077:
711:hypertrophic pulmonary osteodystrophy
7:
951:Short, Rachel A; Wallace, Steven C.
25:
827:McKenna & Bell, 1997, p. 483.
682:is the most common fossil in the
579:analysis alternatively suggest a
1381:Neogene mammals of North America
1229:
932:geology.teacherfriendlyguide.org
791:
779:
767:
750:
655:
639:
624:
610:
514:
493:
168:
1406:Ringold Formation Miocene Fauna
1371:Miocene genus first appearances
1:
1401:Fossil taxa described in 1894
878:Wang, B.; Secord, R. (2020).
928:"Region 4: The Great Plains"
913:10.1016/j.palaeo.2019.109411
136:Natural History Museum of LA
701:. Of the 20+ taxa present,
1422:
1209:Prothero, Donald R. 2005.
760:Cormohipparion occidentale
707:pyroclastic airfall event
374:
367:
295:
290:
279:
272:
165:Scientific classification
163:
150:
141:
133:
124:
34:
1086:Mead, Alfred J. (2000).
27:Extinct genus of mammals
1005:10.58782/flmnh.kpcf8483
836:Prothero, 2005, p. 122.
968:Cite journal requires
818:Prothero, 2005, p. 94.
699:supervolcanic eruption
550:Henry Fairfield Osborn
527:Barbourofelis loveorum
1366:Pliocene rhinoceroses
1333:Paleobiology Database
1238:at Wikimedia Commons
1168:"Ashfall Fossil Beds"
1139:10.1130/2014.0036(01)
157:Robert Bruce Horsfall
1376:Zanclean extinctions
1361:Miocene rhinoceroses
853:on 20 November 2021.
480:Teleoceras aepysoma.
1052:10.1017/pab.2017.13
1044:2018Pbio...44..131M
904:2020PPP...54209411W
743:Ashfall Fossil Beds
684:Ashfall Fossil Beds
675:Ashfall Fossil Beds
18:Teleoceras fossiger
690:. Over 100 intact
567:has high crowned (
452:at the end of the
1348:
1347:
1247:Taxon identifiers
1234:Media related to
1148:978-0-8137-0036-6
880:"Paleoecology of
587:Sexual dimorphism
466:
461:
402:
401:
396:
385:
268:
16:(Redirected from
1413:
1341:
1340:
1328:
1327:
1315:
1314:
1302:
1301:
1289:
1288:
1287:
1274:
1273:
1272:
1242:
1233:
1183:
1182:
1180:
1179:
1170:. Archived from
1164:
1158:
1157:
1156:
1155:
1122:
1116:
1115:
1083:
1072:
1071:
1023:
1017:
1016:
984:
978:
977:
971:
966:
964:
956:
948:
942:
941:
939:
938:
924:
918:
917:
915:
875:
864:
863:(Prothero, 2005)
861:
855:
854:
849:. Archived from
843:
837:
834:
828:
825:
819:
816:
795:
783:
771:
754:
696:Bruneau-Jarbidge
680:Teleoceras major
659:
643:
628:
614:
573:Dental microwear
555:Some species of
518:
497:
469:Gray Fossil Site
464:
459:
394:
383:
283:Teleoceras major
263:
256:
243:
173:
172:
155:illustration by
146:
134:Specimen at the
129:
119:
56:
40:Temporal range:
32:
21:
1421:
1420:
1416:
1415:
1414:
1412:
1411:
1410:
1351:
1350:
1349:
1344:
1336:
1331:
1323:
1318:
1310:
1305:
1297:
1292:
1283:
1282:
1277:
1268:
1267:
1262:
1249:
1227:
1192:
1187:
1186:
1177:
1175:
1166:
1165:
1161:
1153:
1151:
1149:
1124:
1123:
1119:
1085:
1084:
1075:
1025:
1024:
1020:
986:
985:
981:
967:
957:
950:
949:
945:
936:
934:
926:
925:
921:
877:
876:
867:
862:
858:
845:
844:
840:
835:
831:
826:
822:
817:
813:
808:
803:
802:
801:
800:
799:
796:
788:
787:
784:
776:
775:
772:
764:
763:
755:
746:
745:
677:
672:
667:
666:
665:
664:
663:
660:
652:
651:
644:
636:
635:
629:
621:
620:
615:
606:
605:
589:
539:
538:
537:
536:
535:
519:
510:
509:
508:
498:
487:
348:T. medicornutum
318:T. brachyrhinum
286:
262:
254:
241:
167:
120:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
52:15.97–4.9
51:
50:
48:
38:
28:
23:
22:
15:
12:
11:
5:
1419:
1417:
1409:
1408:
1403:
1398:
1393:
1388:
1383:
1378:
1373:
1368:
1363:
1353:
1352:
1346:
1345:
1343:
1342:
1329:
1316:
1303:
1290:
1275:
1259:
1257:
1251:
1250:
1245:
1226:
1225:External links
1223:
1222:
1221:
1207:
1191:
1188:
1185:
1184:
1159:
1147:
1117:
1098:(4): 689–706.
1073:
1038:(1): 131–154.
1018:
999:(5): 183–260.
979:
970:|journal=
943:
919:
865:
856:
838:
829:
820:
810:
809:
807:
804:
797:
790:
789:
785:
778:
777:
773:
766:
765:
756:
749:
748:
747:
737:
736:
735:
734:
676:
673:
671:
668:
661:
654:
653:
645:
638:
637:
630:
623:
622:
616:
609:
608:
607:
603:
602:
601:
600:
588:
585:
520:
513:
512:
511:
499:
492:
491:
490:
489:
488:
486:
483:
436:to the end of
417:. It lived in
400:
399:
398:
397:
386:
372:
371:
365:
364:
363:
362:
356:
350:
344:
338:
332:
326:
320:
314:
308:
302:
293:
292:
288:
287:
277:
276:
270:
269:
252:
248:
247:
245:Aceratheriinae
239:
235:
234:
232:Rhinocerotidae
229:
225:
224:
222:Perissodactyla
219:
215:
214:
209:
205:
204:
199:
195:
194:
189:
185:
184:
179:
175:
174:
161:
160:
148:
147:
139:
138:
131:
130:
122:
121:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
57:
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
1418:
1407:
1404:
1402:
1399:
1397:
1394:
1392:
1389:
1387:
1384:
1382:
1379:
1377:
1374:
1372:
1369:
1367:
1364:
1362:
1359:
1358:
1356:
1339:
1334:
1330:
1326:
1321:
1317:
1313:
1308:
1304:
1300:
1295:
1291:
1286:
1280:
1276:
1271:
1265:
1261:
1260:
1258:
1256:
1252:
1248:
1243:
1239:
1237:
1232:
1224:
1220:
1219:0-521-83240-3
1216:
1212:
1208:
1206:
1205:0-231-11013-8
1202:
1198:
1194:
1193:
1189:
1174:on 2011-05-18
1173:
1169:
1163:
1160:
1150:
1144:
1140:
1136:
1132:
1128:
1121:
1118:
1113:
1109:
1105:
1101:
1097:
1093:
1089:
1082:
1080:
1078:
1074:
1069:
1065:
1061:
1057:
1053:
1049:
1045:
1041:
1037:
1033:
1029:
1022:
1019:
1014:
1010:
1006:
1002:
998:
994:
990:
983:
980:
975:
962:
954:
947:
944:
933:
929:
923:
920:
914:
909:
905:
901:
897:
893:
889:
887:
883:
874:
872:
870:
866:
860:
857:
852:
848:
842:
839:
833:
830:
824:
821:
815:
812:
805:
794:
782:
770:
762:
761:
753:
744:
740:
733:
731:
727:
722:
720:
714:
712:
708:
704:
700:
697:
693:
689:
685:
681:
674:
669:
658:
649:
646:Young female
642:
633:
627:
619:
613:
599:
597:
593:
586:
584:
582:
578:
574:
570:
566:
562:
558:
553:
551:
547:
543:
533:
529:
528:
523:
517:
506:
502:
496:
484:
482:
481:
478:
474:
471:in northeast
470:
462:
455:
451:
450:
445:
441:
439:
435:
434:Hemingfordian
431:
428:
424:
420:
419:North America
416:
412:
408:
407:
393:
392:
387:
382:
381:
376:
375:
373:
370:
366:
361:
357:
355:
354:T. meridianum
351:
349:
345:
343:
339:
337:
336:T. guymonense
333:
331:
327:
325:
321:
319:
315:
313:
312:T. americanum
309:
307:
303:
301:
297:
296:
294:
289:
285:
284:
278:
275:
271:
266:
261:
260:
253:
250:
249:
246:
240:
237:
236:
233:
230:
227:
226:
223:
220:
217:
216:
213:
210:
207:
206:
203:
200:
197:
196:
193:
190:
187:
186:
183:
180:
177:
176:
171:
166:
162:
158:
154:
149:
145:
140:
137:
132:
128:
123:
116:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
55:
47:
43:
37:
33:
30:
19:
1391:Clarendonian
1254:
1228:
1210:
1196:
1190:Bibliography
1176:. Retrieved
1172:the original
1162:
1152:, retrieved
1130:
1120:
1095:
1092:Paleobiology
1091:
1035:
1032:Paleobiology
1031:
1021:
996:
992:
982:
961:cite journal
946:
935:. Retrieved
931:
922:
895:
891:
885:
881:
859:
851:the original
841:
832:
823:
814:
758:
738:
729:
725:
723:
715:
702:
691:
679:
678:
647:
631:
617:
591:
590:
564:
560:
556:
554:
546:hippopotamus
541:
540:
525:
521:
500:
479:
476:
447:
443:
442:
415:rhinocerotid
405:
404:
403:
390:
389:
384:(Cook, 1930)
379:
378:
359:
353:
347:
341:
335:
329:
323:
317:
311:
305:
299:
282:
281:
274:Type species
258:
257:
152:
35:
29:
1396:Hemphillian
1279:Wikispecies
650:, Nebraska.
632:T. fossiger
618:T. aginense
561:T. aepysoma
522:T. proterum
501:T. fossiger
485:Description
477:Teleoceras,
454:Hemphillian
438:Hemphillian
432:during the
421:during the
391:Paraphelops
360:T. proterum
330:T. fossiger
306:T. aginense
300:T. aepysoma
238:Subfamily:
153:T. fossiger
46:Hemphillian
1386:Barstovian
1355:Categories
1285:Teleoceras
1255:Teleoceras
1236:Teleoceras
1178:2005-12-13
1154:2024-08-13
937:2021-06-26
898:: 109411.
886:Teleoceras
806:References
730:Teleoceras
596:bimaturism
592:Teleoceras
565:Teleoceras
557:Teleoceras
542:Teleoceras
444:Teleoceras
406:Teleoceras
395:Lane, 1927
259:Teleoceras
42:Barstovian
36:Teleoceras
1112:0094-8373
1060:0094-8373
1013:2373-9991
741:from the
670:Discovery
634:, Kansas.
569:hypsodont
473:Tennessee
380:Mesoceras
324:T. hicksi
188:Kingdom:
182:Eukaryota
1264:Wikidata
1068:90987376
882:Aphelops
739:T. major
726:T. major
703:T. major
692:T. major
688:Nebraska
648:T. major
581:browsing
577:mesowear
449:Aphelops
427:Pliocene
369:Synonyms
342:T. major
291:Species
228:Family:
212:Mammalia
202:Chordata
198:Phylum:
192:Animalia
178:Domain:
1325:1374709
1312:3239442
1299:4452636
1270:Q133358
1040:Bibcode
900:Bibcode
423:Miocene
265:Hatcher
251:Genus:
218:Order:
208:Class:
1217:
1203:
1145:
1110:
1066:
1058:
1011:
604:Skulls
430:epochs
267:, 1894
1338:43226
1320:IRMNG
1064:S2CID
719:hippo
662:Baby.
532:FLMNH
411:genus
151:1913
1307:GBIF
1215:ISBN
1201:ISBN
1143:ISBN
1108:ISSN
1056:ISSN
1009:ISSN
974:help
884:and
575:and
524:and
505:NMNH
425:and
60:PreꞒ
1294:EoL
1135:doi
1100:doi
1048:doi
1001:doi
908:doi
896:542
686:of
413:of
1357::
1335::
1322::
1309::
1296::
1281::
1266::
1141:,
1129:,
1106:.
1096:26
1094:.
1090:.
1076:^
1062:.
1054:.
1046:.
1036:44
1034:.
1030:.
1007:.
997:56
995:.
991:.
965::
963:}}
959:{{
930:.
906:.
894:.
890:.
868:^
563:.
530:,
503:,
159:.
110:Pg
54:Ma
1181:.
1137::
1114:.
1102::
1070:.
1050::
1042::
1015:.
1003::
976:)
972:(
955:.
940:.
916:.
910::
902::
534:.
507:.
465:4
460:4
458:C
388:†
377:†
358:†
352:†
346:†
340:†
334:†
328:†
322:†
316:†
310:†
304:†
298:†
280:†
255:†
242:†
115:N
105:K
100:J
95:T
90:P
85:C
80:D
75:S
70:O
65:Ꞓ
49:~
44:-
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.