1904:
376:
972:
603:
1910:
2460:
136:
1193:
457:
2490:
2466:
1015:
embayment, flattening and expansion of the parietal bar, reduction of size of the parietal fenestrae, the change of its shape from (sub)angular to round, and development of flanges on the medial bar. Geometric morphometric analysis supported the sequence, finding them to plot in the expected order. The phylogenetic analysis (see above), however, complicated matters, with the genera
1181:
506:
118:
1014:
would, under this model, represent a single population of organisms changing in form over time, rather than as a diverse assemblage of close relatives. This is evidenced by their parietal morphology, which show consistent trends of change. These are: the gradual closure of the initially large medial
515:
Like most other chasmosaurs, its paired fused parietals combined bear six epiparietals (small horns along the parietal), symmetrically arranged with three on each side. The first pair, small and triangular, project from the top edge of the median embayment, and in life would have touched each other.
520:
had a very large frill similar to that of its relatives. A singular, fused episquamosal (small horns along the squamosal) is also known from the holotype; it is rugose and indistinct from that of other ceratopsids. The left epijugal horn is known as well, fused to the jugal and quadratojugal bones;
399:
in New Mexico. It is the only diagnostic chasmosaurine specimen discovered from the middle or upper part of the Hunter Wash Member. The age of the specimen is undetermined; its frill texture indicates it is a young subadult, but its large size and epiparietal fusion would indicate it represents an
516:
The second pair are a larger set of triangles, whereas the third epiparietals have a rounded, "D" shape; both project upwards, angled in line with the rest of the parietal. The preserved right squamosal (bone which forms the right side of the frill) itself is long, indicating adult
480:
by the anatomy of its parietal (the upper portion of its frill), which forms a heart shape. The prominent median embayment (a large notch in the middle of the top of the frill) of earlier relatives is heavily reduced, being very narrow as opposed to wide and U-shaped.
490:. The parietal fenestrae (the holes in the frill) have a more rounded shape than the ancestral angular state, and are smaller due to the expanded parietal and median bars. Overall, the anatomy is intermediate between that of more primitive genera like
485:
parietal bars (the top edges of the frill) are thin and extremely broad compared to earlier relatives; they are more plate-shaped than bar-shaped. Its median bar (the middle strut) has also expanded, bearing more pronounced flanges than its ancestor
594:
data may requires revision, as it likely contains specimens of more than one species; it was noted this could be impacting the results negatively. Several taxa had been named too recently to be included in the study describing
387:
P-27468, collected in 1997, consists of a parietal (or fused paired parietals), other skull fragments, a partial sacrum, and vertebral fragments. It was discovered in grey siltstone deposits from the
367:, which it was closely related to, but had a distinctive heart-shaped upper frill with very narrow notch. It has been hypothesized to form an anagenetic series with several other chasmosaur species.
411:
specimens. It was informally named as a distinct genus in 2016. Later, in 2020, it was formally named and described by Denver W. Fowler and
Elizabeth A. Freedman Fowler. The name
2575:
415:
means "coming to the end of the cavity", referring to the parietal embayment being nearly closed off before being lost completely in more derived taxa. The specific name,
1027:
plotting within the supposed anagenetic lineage. It was noted, however, that these are fragmentary and undiagnostic respectively, and so they may not be of significance.
1126:"Redescription Of A Specimen Of Pentaceratops (Ornithischia: Ceratopsidae) And Phylogenetic Evaluation Of FiveReferred Specimens From The Upper Cretaceous Of New Mexico"
1125:
407:, though noted as aberrant for the genus. Joshua Fry questioned the referral in a 2015 masters thesis, with a phylogenetic analysis failing to group it alongside other
2595:
2580:
448:
due to its deep, narrow median embayment and broad parietal bar. The specimen is too distorted to allow confident referral to either, however.
1903:
599:, and their inclusion in an analysis could also shift its position. One analysis from Fowler and Freedman Fowler (2020) is reproduced below:
2600:
1225:
434:" in 1930, but by future authors have considered its distinctiveness to be a result of pathology. Though sometimes considered a synonym of
2590:
426:
The nearly complete, but highly distorted skull PMU 24923 may belong to the species as well. Also hailing from the
Kirtland Formation,
1909:
2483:
1036:
275:
2605:
1171:
2459:
375:
135:
971:
602:
1075:"Transitional evolutionary forms in chasmosaurine ceratopsid dinosaurs: evidence from the Campanian of New Mexico"
1316:
1218:
2585:
2570:
1434:
1149:"Dinosaurs and time: chronostratigraphic frameworks and their utility in analysis of dinosaur paleobiology"
2510:
419:, refers to the discoverer of the holotype specimen, Paul Sealey. Naming the specimen as a new species of
2542:
2493:
1343:
2465:
574:
belongs to the latter group. Phylogenetic analysis found it to be relatively derived, more basal than
1197:
427:
1599:
1211:
2319:
2269:
1701:
392:
348:
130:
2547:
379:
Geological map of the southeast San Juan Basin; B (upper middle) is where the holotype was found
2402:
2340:
2303:
1777:
1664:
1106:
2432:
2230:
2055:
1757:
1452:
1185:
1096:
1086:
1017:
732:
586:-line to collapse into an unresolved polytomy. Additionally, it was noted the coding of the
509:
327:
2326:
2278:
2237:
2173:
2076:
2027:
1729:
1653:
1608:
1482:
1468:
1427:
1301:
1285:
887:
582:. The tree, however, is unstable; removing some taxa from the analysis caused much of the
30:
355:
in 1997, and was later described and named in a 2020 study. It was similar in anatomy to
1148:
456:
2333:
2292:
2223:
2210:
2181:
2166:
2136:
2097:
2083:
2000:
1961:
1840:
1750:
1715:
1708:
1588:
1538:
1461:
1441:
1101:
1074:
903:
815:
621:
553:
499:
440:
396:
254:
212:
2564:
2411:
2390:
2360:
2351:
2251:
2145:
2090:
2069:
2041:
2034:
2020:
2013:
1819:
1808:
1801:
1743:
1736:
1722:
1680:
1615:
1581:
1552:
1505:
1023:
975:
863:
756:
683:
363:
357:
331:
68:
2425:
2376:
2369:
2285:
2258:
2244:
2152:
2125:
2118:
2062:
2048:
1940:
1927:
1858:
1847:
1832:
1794:
1786:
1768:
1673:
1639:
1622:
1574:
1567:
1523:
1328:
1276:
773:
666:
642:
558:
533:
335:
317:
283:
241:
196:
2533:
2418:
2312:
2195:
2159:
1982:
1694:
1559:
978:
diagram demonstrating the proposed anagenetic lineages of chasmosaurs; frill of
43:
2188:
2104:
1954:
1687:
1475:
1409:
1396:
1355:
1234:
991:
556:. Fowler and Freedman Folwler (2020) divide chasmosaurs into two lineages; a "
352:
324:
228:
88:
53:
388:
147:
93:
37:
1110:
438:, Fowler and Freedman Fowler considered it more likely to belong to either
505:
117:
2527:
1989:
1975:
1968:
1531:
1267:
1258:
344:
320:
180:
167:
83:
78:
63:
58:
48:
2383:
1545:
1489:
1091:
98:
73:
1252:
157:
2504:
970:
601:
504:
455:
384:
374:
313:
343:, known from a parietal and some other associated fragments. The
423:
was decided against to prevent the genus becoming paraphyletic.
2508:
1925:
1394:
1245:
1207:
982:
is "I", upper right, shown next to that of its close relatives
430:
discovered it in 1921. It was later named as the new species "
1203:
541:, indicating it was an animal of roughly similar adult size.
1169:
290:
2517:
2401:
2359:
2350:
2302:
2268:
2209:
2135:
1999:
1938:
1857:
1818:
1776:
1767:
1663:
1637:
1598:
1503:
1451:
1407:
1354:
1327:
1300:
403:A 2005 abstract referred the specimen to the genus
994:lineage of chasmosaurs. The genera (in sequence)
531:known remains are slightly smaller than those of
464:(H, lower right) compared to its close relatives
472:is distinguished from close relatives such as
1219:
8:
1073:Fowler, D.W.; Freedman Fowler, E.A. (2020).
1068:
1066:
1064:
1062:
1060:
1058:
1056:
1054:
1052:
334:. The genus contains a single species, the
125:Holotype parietals from the front and back
2576:Late Cretaceous dinosaurs of North America
2505:
2356:
2215:
2005:
1946:
1935:
1922:
1824:
1773:
1645:
1515:
1511:
1419:
1415:
1404:
1391:
1333:
1306:
1297:
1242:
1226:
1212:
1204:
116:
20:
1100:
1090:
552:was a member of the ceratopsid subfamily
521:it robust and large, but unlike that of
1176:
1048:
16:Genus of ceratopsid dinosaurs (fossil)
570:-line" leading to more derived taxa.
7:
1147:Denver Warwick Fowler (April 2016).
879:
855:
831:
807:
748:
724:
717:
658:
634:
627:
617:
494:and that of more derived ones like
300:Fowler & Freedman Fowler, 2020
14:
2596:Campanian genus first appearances
2489:
2488:
2464:
2458:
1908:
1902:
1191:
1179:
1037:Timeline of ceratopsian research
134:
347:specimen was discovered in the
460:Preserved parietal anatomy of
274:Fowler & Freedman Fowler,
1:
2581:Fossil taxa described in 2020
468:Known from limited material,
2601:Campanian genus extinctions
2622:
2591:Paleontology in New Mexico
391:Hunter Wash Member of the
2478:
2456:
2218:
2008:
1949:
1934:
1921:
1899:
1827:
1648:
1518:
1514:
1422:
1418:
1403:
1390:
1336:
1309:
1296:
1241:
901:
884:
877:
860:
853:
836:
829:
812:
805:
770:
753:
746:
729:
722:
715:
680:
663:
656:
639:
632:
625:
432:Pentaceratops fenestratus
289:
282:
131:Scientific classification
129:
124:
115:
23:
1153:scholarworks.montana.edu
525:is not especially long.
436:Pentaceratops sternbergi
1435:Micropachycephalosaurus
1124:Fry, Joshua J. (2015).
610:, proposed ancestor of
983:
614:
578:but more derived than
512:
465:
383:The holotype specimen
380:
2543:Paleobiology Database
986:It was proposed that
974:
605:
508:
459:
378:
2606:Ornithischian genera
990:is a part of a long
428:Charles H. Sternberg
371:Discovery and naming
340:Terminocavus sealeyi
295:Terminocavus sealeyi
1600:Archaeoceratopsidae
2320:Crittendenceratops
2270:Albertaceratopsini
1092:10.7717/peerj.9251
984:
615:
562:-line" leading to
513:
466:
393:Kirtland Formation
381:
349:Kirtland Formation
2558:
2557:
2511:Taxon identifiers
2502:
2501:
2474:
2473:
2454:
2453:
2450:
2449:
2446:
2445:
2442:
2441:
2403:Pachyrhinosaurini
2341:Yehuecauhceratops
2304:Nasutoceratopsini
2205:
2204:
1917:
1916:
1897:
1896:
1893:
1892:
1889:
1888:
1885:
1884:
1881:
1880:
1877:
1876:
1778:Protoceratopsidae
1665:Leptoceratopsidae
1633:
1632:
1499:
1498:
1386:
1385:
1382:
1381:
1378:
1377:
1374:
1373:
968:
967:
959:
958:
950:
949:
941:
940:
932:
931:
923:
922:
914:
913:
794:
793:
785:
784:
704:
703:
695:
694:
305:
304:
278:
2613:
2551:
2550:
2538:
2537:
2536:
2506:
2492:
2491:
2469:
2468:
2462:
2433:Pachyrhinosaurus
2357:
2231:Machairoceratops
2216:
2056:Coahuilaceratops
2006:
1947:
1936:
1923:
1912:
1907:
1906:
1825:
1774:
1758:Zhuchengceratops
1646:
1516:
1512:
1453:Chaoyangsauridae
1420:
1416:
1405:
1392:
1334:
1307:
1298:
1291:
1290:
1243:
1228:
1221:
1214:
1205:
1196:
1195:
1194:
1184:
1183:
1182:
1175:
1164:
1163:
1161:
1159:
1144:
1138:
1137:
1121:
1115:
1114:
1104:
1094:
1070:
1018:Coahuilaceratops
880:
856:
832:
808:
749:
733:Coahuilaceratops
725:
718:
659:
635:
628:
618:
510:Life restoration
292:
273:
266:
253:
240:
227:
211:
195:
139:
138:
120:
110:
40:
29:Temporal range:
21:
2621:
2620:
2616:
2615:
2614:
2612:
2611:
2610:
2561:
2560:
2559:
2554:
2546:
2541:
2532:
2531:
2526:
2513:
2503:
2498:
2470:
2463:
2438:
2397:
2346:
2327:Furcatoceratops
2298:
2279:Albertaceratops
2264:
2238:Menefeeceratops
2201:
2174:Tatankaceratops
2131:
2077:Mercuriceratops
2028:Arrhinoceratops
1995:
1943:
1930:
1913:
1901:
1873:
1853:
1814:
1763:
1730:Montanoceratops
1659:
1654:Graciliceratops
1642:
1629:
1609:Archaeoceratops
1594:
1508:
1495:
1483:Xuanhuaceratops
1469:Hualianceratops
1447:
1428:Albalophosaurus
1412:
1399:
1370:
1350:
1323:
1317:Avemetatarsalia
1302:Avemetatarsalia
1292:
1286:Marginocephalia
1248:
1247:
1237:
1232:
1202:
1192:
1190:
1180:
1178:
1170:
1168:
1167:
1157:
1155:
1146:
1145:
1141:
1123:
1122:
1118:
1072:
1071:
1050:
1045:
1033:
969:
960:
951:
942:
933:
924:
915:
888:Arrhinoceratops
795:
786:
705:
696:
547:
454:
373:
330:of what is now
301:
298:
272:
264:
251:
238:
225:
209:
193:
133:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
36:75.0–74.6
35:
34:
31:Late Cretaceous
27:
17:
12:
11:
5:
2619:
2617:
2609:
2608:
2603:
2598:
2593:
2588:
2586:Campanian life
2583:
2578:
2573:
2571:Chasmosaurines
2563:
2562:
2556:
2555:
2553:
2552:
2539:
2523:
2521:
2515:
2514:
2509:
2500:
2499:
2497:
2496:
2486:
2479:
2476:
2475:
2472:
2471:
2457:
2455:
2452:
2451:
2448:
2447:
2444:
2443:
2440:
2439:
2437:
2436:
2429:
2422:
2415:
2407:
2405:
2399:
2398:
2396:
2395:
2387:
2380:
2373:
2365:
2363:
2354:
2348:
2347:
2345:
2344:
2337:
2334:Nasutoceratops
2330:
2323:
2316:
2308:
2306:
2300:
2299:
2297:
2296:
2293:Medusaceratops
2289:
2282:
2274:
2272:
2266:
2265:
2263:
2262:
2255:
2248:
2241:
2234:
2227:
2224:Diabloceratops
2219:
2213:
2211:Centrosaurinae
2207:
2206:
2203:
2202:
2200:
2199:
2192:
2185:
2182:Titanoceratops
2178:
2170:
2167:Regaliceratops
2163:
2156:
2149:
2141:
2139:
2137:Triceratopsini
2133:
2132:
2130:
2129:
2122:
2115:
2108:
2101:
2098:Sierraceratops
2094:
2087:
2084:Navajoceratops
2080:
2073:
2066:
2059:
2052:
2045:
2038:
2031:
2024:
2017:
2009:
2003:
2001:Chasmosaurinae
1997:
1996:
1994:
1993:
1986:
1979:
1972:
1965:
1962:Brachyceratops
1958:
1950:
1944:
1939:
1932:
1931:
1926:
1919:
1918:
1915:
1914:
1900:
1898:
1895:
1894:
1891:
1890:
1887:
1886:
1883:
1882:
1879:
1878:
1875:
1874:
1872:
1871:
1870:
1869:
1863:
1861:
1855:
1854:
1852:
1851:
1844:
1841:Turanoceratops
1837:
1828:
1822:
1816:
1815:
1813:
1812:
1805:
1798:
1791:
1782:
1780:
1771:
1765:
1764:
1762:
1761:
1754:
1751:Unescoceratops
1747:
1740:
1733:
1726:
1719:
1716:Ischioceratops
1712:
1709:Gryphoceratops
1705:
1698:
1691:
1684:
1677:
1669:
1667:
1661:
1660:
1658:
1657:
1649:
1643:
1638:
1635:
1634:
1631:
1630:
1628:
1627:
1619:
1612:
1604:
1602:
1596:
1595:
1593:
1592:
1589:Sasayamagnomus
1585:
1578:
1571:
1564:
1556:
1549:
1542:
1539:Auroraceratops
1535:
1528:
1519:
1509:
1504:
1501:
1500:
1497:
1496:
1494:
1493:
1486:
1479:
1472:
1465:
1462:Chaoyangsaurus
1457:
1455:
1449:
1448:
1446:
1445:
1442:Psittacosaurus
1438:
1431:
1423:
1413:
1408:
1401:
1400:
1395:
1388:
1387:
1384:
1383:
1380:
1379:
1376:
1375:
1372:
1371:
1369:
1368:
1367:
1366:
1360:
1358:
1352:
1351:
1349:
1348:
1347:
1346:
1337:
1331:
1325:
1324:
1322:
1321:
1320:
1319:
1310:
1304:
1294:
1293:
1289:
1288:
1279:
1270:
1261:
1255:
1246:
1239:
1238:
1233:
1231:
1230:
1223:
1216:
1208:
1201:
1200:
1188:
1166:
1165:
1139:
1130:Masters Theses
1116:
1047:
1046:
1044:
1041:
1040:
1039:
1032:
1029:
1004:Navajoceratops
966:
965:
962:
961:
957:
956:
953:
952:
948:
947:
944:
943:
939:
938:
935:
934:
930:
929:
926:
925:
921:
920:
917:
916:
912:
911:
908:
907:
904:Triceratopsini
900:
897:
896:
893:
892:
883:
878:
876:
873:
872:
869:
868:
859:
854:
852:
849:
848:
845:
844:
835:
830:
828:
825:
824:
821:
820:
816:Navajoceratops
811:
806:
804:
801:
800:
797:
796:
792:
791:
788:
787:
783:
782:
779:
778:
769:
766:
765:
762:
761:
752:
747:
745:
742:
741:
738:
737:
728:
723:
721:
716:
714:
711:
710:
707:
706:
702:
701:
698:
697:
693:
692:
689:
688:
679:
676:
675:
672:
671:
662:
657:
655:
652:
651:
648:
647:
638:
633:
631:
626:
624:
622:Chasmosaurinae
616:
554:Chasmosaurinae
546:
545:Classification
543:
500:triceratopsins
488:Navajoceratops
453:
450:
441:Navajoceratops
397:San Juan Basin
372:
369:
323:from the late
303:
302:
299:
287:
286:
280:
279:
262:
258:
257:
255:Chasmosaurinae
249:
245:
244:
236:
232:
231:
223:
216:
215:
213:Neornithischia
207:
200:
199:
191:
184:
183:
178:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
127:
126:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
2618:
2607:
2604:
2602:
2599:
2597:
2594:
2592:
2589:
2587:
2584:
2582:
2579:
2577:
2574:
2572:
2569:
2568:
2566:
2549:
2544:
2540:
2535:
2529:
2525:
2524:
2522:
2520:
2516:
2512:
2507:
2495:
2487:
2485:
2481:
2480:
2477:
2467:
2461:
2435:
2434:
2430:
2428:
2427:
2423:
2421:
2420:
2416:
2414:
2413:
2412:Achelousaurus
2409:
2408:
2406:
2404:
2400:
2393:
2392:
2391:Styracosaurus
2388:
2386:
2385:
2381:
2379:
2378:
2374:
2372:
2371:
2367:
2366:
2364:
2362:
2361:Centrosaurini
2358:
2355:
2353:
2352:Eucentrosaura
2349:
2343:
2342:
2338:
2336:
2335:
2331:
2329:
2328:
2324:
2322:
2321:
2317:
2315:
2314:
2310:
2309:
2307:
2305:
2301:
2295:
2294:
2290:
2288:
2287:
2283:
2281:
2280:
2276:
2275:
2273:
2271:
2267:
2261:
2260:
2256:
2254:
2253:
2252:Wendiceratops
2249:
2247:
2246:
2242:
2240:
2239:
2235:
2233:
2232:
2228:
2226:
2225:
2221:
2220:
2217:
2214:
2212:
2208:
2198:
2197:
2193:
2191:
2190:
2186:
2184:
2183:
2179:
2176:
2175:
2171:
2169:
2168:
2164:
2162:
2161:
2157:
2155:
2154:
2150:
2148:
2147:
2146:Eotriceratops
2143:
2142:
2140:
2138:
2134:
2128:
2127:
2123:
2121:
2120:
2116:
2114:
2113:
2109:
2107:
2106:
2102:
2100:
2099:
2095:
2093:
2092:
2091:Pentaceratops
2088:
2086:
2085:
2081:
2079:
2078:
2074:
2072:
2071:
2070:Kosmoceratops
2067:
2065:
2064:
2060:
2058:
2057:
2053:
2051:
2050:
2046:
2044:
2043:
2042:Bravoceratops
2039:
2037:
2036:
2035:Bisticeratops
2032:
2030:
2029:
2025:
2023:
2022:
2021:Anchiceratops
2018:
2016:
2015:
2014:Agujaceratops
2011:
2010:
2007:
2004:
2002:
1998:
1992:
1991:
1987:
1985:
1984:
1980:
1978:
1977:
1973:
1971:
1970:
1966:
1964:
1963:
1959:
1957:
1956:
1952:
1951:
1948:
1945:
1942:
1937:
1933:
1929:
1924:
1920:
1911:
1905:
1867:
1866:
1865:
1864:
1862:
1860:
1856:
1850:
1849:
1845:
1843:
1842:
1838:
1835:
1834:
1830:
1829:
1826:
1823:
1821:
1820:Ceratopsoidea
1817:
1811:
1810:
1809:Protoceratops
1806:
1804:
1803:
1802:Breviceratops
1799:
1797:
1796:
1792:
1789:
1788:
1784:
1783:
1781:
1779:
1775:
1772:
1770:
1766:
1760:
1759:
1755:
1753:
1752:
1748:
1746:
1745:
1744:Udanoceratops
1741:
1739:
1738:
1737:Prenoceratops
1734:
1732:
1731:
1727:
1725:
1724:
1723:Leptoceratops
1720:
1718:
1717:
1713:
1711:
1710:
1706:
1704:
1703:
1699:
1697:
1696:
1692:
1690:
1689:
1685:
1683:
1682:
1681:Bainoceratops
1678:
1676:
1675:
1671:
1670:
1668:
1666:
1662:
1656:
1655:
1651:
1650:
1647:
1644:
1641:
1636:
1625:
1624:
1620:
1618:
1617:
1616:Helioceratops
1613:
1611:
1610:
1606:
1605:
1603:
1601:
1597:
1591:
1590:
1586:
1584:
1583:
1582:Mosaiceratops
1579:
1577:
1576:
1572:
1570:
1569:
1565:
1562:
1561:
1557:
1555:
1554:
1553:Koreaceratops
1550:
1548:
1547:
1543:
1541:
1540:
1536:
1534:
1533:
1529:
1526:
1525:
1521:
1520:
1517:
1513:
1510:
1507:
1506:Neoceratopsia
1502:
1492:
1491:
1487:
1485:
1484:
1480:
1478:
1477:
1473:
1471:
1470:
1466:
1464:
1463:
1459:
1458:
1456:
1454:
1450:
1444:
1443:
1439:
1437:
1436:
1432:
1430:
1429:
1425:
1424:
1421:
1417:
1414:
1411:
1406:
1402:
1398:
1393:
1389:
1364:
1363:
1362:
1361:
1359:
1357:
1353:
1345:
1341:
1340:
1339:
1338:
1335:
1332:
1330:
1326:
1318:
1314:
1313:
1312:
1311:
1308:
1305:
1303:
1299:
1295:
1287:
1283:
1280:
1278:
1274:
1271:
1269:
1265:
1262:
1260:
1256:
1254:
1250:
1249:
1244:
1240:
1236:
1229:
1224:
1222:
1217:
1215:
1210:
1209:
1206:
1199:
1198:United States
1189:
1187:
1177:
1173:
1154:
1150:
1143:
1140:
1135:
1131:
1127:
1120:
1117:
1112:
1108:
1103:
1098:
1093:
1088:
1084:
1080:
1076:
1069:
1067:
1065:
1063:
1061:
1059:
1057:
1055:
1053:
1049:
1042:
1038:
1035:
1034:
1030:
1028:
1026:
1025:
1024:Bravoceratops
1020:
1019:
1013:
1012:Anchiceratops
1009:
1005:
1001:
1000:Pentaceratops
997:
993:
989:
981:
977:
976:Stratigraphic
973:
964:
963:
955:
954:
946:
945:
937:
936:
928:
927:
919:
918:
910:
909:
906:
905:
899:
898:
895:
894:
891:
890:
889:
882:
881:
875:
874:
871:
870:
867:
866:
865:
864:Anchiceratops
858:
857:
851:
850:
847:
846:
843:
842:
841:
834:
833:
827:
826:
823:
822:
819:
818:
817:
810:
809:
803:
802:
799:
798:
790:
789:
781:
780:
777:
776:
775:
768:
767:
764:
763:
760:
759:
758:
757:Pentaceratops
751:
750:
744:
743:
740:
739:
736:
735:
734:
727:
726:
720:
719:
713:
712:
709:
708:
700:
699:
691:
690:
687:
686:
685:
684:Kosmoceratops
678:
677:
674:
673:
670:
669:
668:
661:
660:
654:
653:
650:
649:
646:
645:
644:
637:
636:
630:
629:
623:
620:
619:
613:
609:
608:Pentaceratops
604:
600:
598:
593:
589:
588:Pentaceratops
585:
584:Pentaceratops
581:
580:Navajocertops
577:
576:Anchiceratops
573:
569:
568:Pentaceratops
565:
564:Kosmoceratops
561:
560:
555:
551:
544:
542:
540:
539:Pentaceratops
536:
535:
530:
526:
524:
523:Pentaceratops
519:
511:
507:
503:
501:
497:
496:Anchiceratops
493:
492:Pentaceratops
489:
484:
483:Terminocavus'
479:
478:Anchiceratops
475:
474:Pentaceratops
471:
463:
458:
451:
449:
447:
443:
442:
437:
433:
429:
424:
422:
421:Pentaceratops
418:
414:
410:
409:Pentaceratops
406:
405:Pentaceratops
401:
398:
394:
390:
386:
377:
370:
368:
366:
365:
364:Anchiceratops
360:
359:
358:Pentaceratops
354:
350:
346:
342:
341:
337:
333:
332:North America
329:
326:
322:
319:
315:
311:
310:
297:
296:
288:
285:
281:
277:
271:
270:
263:
260:
259:
256:
250:
247:
246:
243:
237:
234:
233:
230:
224:
221:
218:
217:
214:
208:
205:
202:
201:
198:
192:
189:
186:
185:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
137:
132:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
2519:Terminocavus
2518:
2431:
2426:Stellasaurus
2424:
2417:
2410:
2389:
2382:
2377:Coronosaurus
2375:
2370:Centrosaurus
2368:
2339:
2332:
2325:
2318:
2311:
2291:
2286:Lokiceratops
2284:
2277:
2259:Xenoceratops
2257:
2250:
2245:Sinoceratops
2243:
2236:
2229:
2222:
2194:
2187:
2180:
2172:
2165:
2158:
2153:Nedoceratops
2151:
2144:
2126:Vagaceratops
2124:
2119:Utahceratops
2117:
2112:Terminocavus
2111:
2110:
2103:
2096:
2089:
2082:
2075:
2068:
2063:Judiceratops
2061:
2054:
2049:Chasmosaurus
2047:
2040:
2033:
2026:
2019:
2012:
1988:
1981:
1974:
1967:
1960:
1953:
1941:Ceratopsidae
1928:Ceratopsidae
1859:Ceratopsidae
1848:Zuniceratops
1846:
1839:
1833:Ajkaceratops
1831:
1807:
1800:
1795:Bagaceratops
1793:
1787:Ajkaceratops
1785:
1769:Coronosauria
1756:
1749:
1742:
1735:
1728:
1721:
1714:
1707:
1700:
1693:
1686:
1679:
1674:Asiaceratops
1672:
1652:
1640:Euceratopsia
1623:Yamaceratops
1621:
1614:
1607:
1587:
1580:
1575:Microceratus
1573:
1568:Liaoceratops
1566:
1558:
1551:
1544:
1537:
1530:
1524:Ajkaceratops
1522:
1488:
1481:
1474:
1467:
1460:
1440:
1433:
1426:
1344:Ornithischia
1329:Ornithischia
1281:
1277:Ornithischia
1272:
1263:
1156:. Retrieved
1152:
1142:
1133:
1129:
1119:
1082:
1078:
1022:
1016:
1011:
1008:Terminocavus
1007:
1003:
999:
996:Utahceratops
995:
988:Terminocavus
987:
985:
980:Terminocavus
979:
902:
886:
885:
862:
861:
840:Terminocavus
839:
838:
837:
814:
813:
774:Utahceratops
772:
771:
755:
754:
731:
730:
682:
681:
667:Vagaceratops
665:
664:
643:Chasmosaurus
641:
640:
612:Terminocavus
611:
607:
597:Terminocavus
596:
592:Chasmosaurus
591:
587:
583:
579:
575:
572:Terminocavus
571:
567:
563:
559:Chasmosaurus
557:
550:Terminocavus
549:
548:
538:
534:Utahceratops
532:
529:Terminocavus
528:
527:
522:
518:Terminocavus
517:
514:
495:
491:
487:
482:
477:
473:
470:Terminocavus
469:
467:
462:Terminocavus
461:
446:Terminocavus
445:
439:
435:
431:
425:
420:
416:
413:Terminocavus
412:
408:
404:
402:
382:
362:
356:
339:
338:
336:type species
309:Terminocavus
308:
307:
306:
294:
293:
284:Type species
269:Terminocavus
268:
267:
242:Ceratopsidae
219:
203:
197:Ornithischia
187:
174:
25:Terminocavus
24:
18:
2419:Einiosaurus
2313:Avaceratops
2196:Triceratops
2160:Ojoceratops
1983:Monoclonius
1695:Ferrisaurus
1560:Kulceratops
1158:17 February
452:Description
248:Subfamily:
2565:Categories
2482:See also:
2189:Torosaurus
2105:Spiclypeus
1955:Agathaumas
1868:see below↓
1688:Cerasinops
1476:Stenopelix
1410:Ceratopsia
1397:Ceratopsia
1365:see below↓
1356:Ceratopsia
1268:Dinosauria
1235:Ceratopsia
1043:References
992:anagenetic
353:New Mexico
325:Cretaceous
318:ceratopsid
229:Ceratopsia
181:Dinosauria
2534:Q96077702
1251:Kingdom:
1186:Dinosaurs
1085:: e9251.
606:Skull of
389:Campanian
154:Kingdom:
148:Eukaryota
2528:Wikidata
2494:Category
2484:Timeline
1990:Polyonax
1976:Dysganus
1969:Ceratops
1532:Aquilops
1259:Chordata
1257:Phylum:
1253:Animalia
1111:32547873
1031:See also
345:holotype
321:dinosaur
235:Family:
168:Chordata
164:Phylum:
158:Animalia
144:Domain:
2384:Spinops
1702:Gremlin
1490:Yinlong
1172:Portals
1102:7278894
566:and a "
417:sealeyi
400:adult.
395:of the
261:Genus:
105:↓
2548:420729
1109:
1099:
1010:, and
328:Period
1282:Clade
1273:Clade
1264:Clade
1079:PeerJ
385:NMMNH
314:genus
312:is a
220:Clade
204:Clade
188:Clade
175:Clade
1342:see
1315:see
1160:2019
1107:PMID
1021:and
590:and
537:and
498:and
476:and
361:and
276:2020
44:PreꞒ
1546:Beg
1097:PMC
1087:doi
444:or
351:of
316:of
33:, ~
2567::
2545::
2530::
1284::
1275::
1266::
1151:.
1134:45
1132:.
1128:.
1105:.
1095:.
1081:.
1077:.
1051:^
1006:,
1002:,
998:,
502:.
222::
206::
190::
177::
94:Pg
38:Ma
2394:?
2177:?
1836:?
1790:?
1626:?
1563:?
1527:?
1227:e
1220:t
1213:v
1174::
1162:.
1136:.
1113:.
1089::
1083:8
291:†
265:†
252:†
239:†
226:†
210:†
194:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
