943:
2007:
2127:
1926:
1070:
5056:
5063:
5076:
764:
5083:
5069:
150:
1156:
566:
1278:
129:
1455:
815:, USA. He drew attention to its premaxillary teeth and deeply inset toothline which he interpreted as supporting the presence of muscular cheeks. Morris also pointed out the outwardly flaring premaxilla (which would have given it a wide beak) and large palpebrals. This skull was recognized as an unnamed hypsilophodont for many years, until Galton made it the type specimen of new genus and species
1782:
1982:, chemical elements important to life, were lacking in their samples; and cardiac cellular structures were absent. There was one possible patch with animal cellular structures. The authors found their data supported identification as a concretion of sand from the burial environment, not the heart, with the possibility that isolated areas of tissues were preserved.
4225:
673:
3960:*Hicks, J.F., Johnson, K.R., Obradovich, J. D., Miggins, D.P., and Tauxe, L. 2003. Magnetostratigraphyof Upper Cretaceous (Maastrichtian) to lower Eocene strata of the Denver Basin, Colorado. In K.R. Johnson, R.G. Raynolds and M.L. Reynolds (eds), Paleontology and Stratigraphy of Laramide Strata in the Denver Basin, Pt. II., Rocky Mountain Geology 38: 1-27.
1864:
1197:, the upper arm bone of most ornithischians articulated with the shoulder by an articular surface that consisted of the entire end of the bone, instead of a distinct ball and socket as in mammals. The orientation of the shoulder's articular surface also indicates a vertical and not horizontal upper arm in dinosaurs.
1921:
coming from it), it partially engulfs one of the ribs and has an internal structure of concentric layers in some places, and another concretion is preserved behind the right leg. The original authors defended their position; they agreed that it was a type of concretion, but one that had formed around
1850:
published in 2023, suggest its brain was small compared to other neornithischian dinosaurs, suggesting its cognitive abilities were within the range of modern reptiles. Additional analysis suggests it had poor hearing and was equipped with a highly-developed sense of smell and balance. Combined with
1378:
The dissolution of
Hypsilophodontidae has been followed by the recognition of the distinct family Thescelosauridae. This area of the dinosaur family tree has historically been complicated by a lack of research, but papers by Clint Boyd and colleagues and Caleb Brown and colleagues have specifically
1093:
Overall, the skeletal anatomy of this genus is well documented, and restorations have been published in several papers, including skeletal restorations and models. The skeleton is known well enough that a detailed reconstruction of the hip and hindlimb muscles has been made. The animal's size has
1324:
Although
Hypsilophodontidae was interpreted as a natural group in the early 1990s, this hypothesis has fallen out of favor and Hypsilophodontidae has been found to be an unnatural family composed of a variety of animals more or less closely related to Iguanodontia
512:
neornithischian is known from several partial skeletons and skulls that indicate it grew to between 2.5 and 4.0 meters (8.2 to 13.1 ft) in length on average. It had sturdy hind limbs, small wide hands, and a head with an elongate pointed snout. The
1204:. However, muscle scars or other indications of attachment have not been found for the plates, which argues against a respiratory function. Recent histological study of layered plates from a probable subadult indicates that they may have started as
1140:(tooth-bearing bone of the lower jaw). The ridges and position of the teeth, deeply internal to the outside surface of the skull, are interpreted as evidence for muscular cheeks. Aside from the long narrow beak, the skull also had teeth in the
1265:, covered in skin impressions that included elongated scales over the legs. One of the paleontologists excavating it was quoted as speculating they had a camouflage function. In a follow-up interview, Paul Barrett has noted that this means
1949:
microscopy. From these methods, the authors found the following: the object's internal structure does not include chambers but is made up of three unconnected areas of lower density material, and is not comparable to the structure of an
3594:
Brown, Caleb
Marshall; Evans, David C.; Ryan, Michael J.; Russell, Anthony P. (2013). "New data on the diversity and abundance of small-bodied ornithopods (Dinosauria, Ornithischia) from the Belly River Group (Campanian) of Alberta".
665:. The name comes from the surprise Gilmore felt at finding such a good specimen that had been unattended to for so long. He considered it to be a light, agile creature, and assigned it to the Hypsilophodontidae, a family of small
3164:
Fisher, Paul E.; Russell, Dale A.; Stoskopf, Michael K.; Barrick, Reese E.; Hammer, Michael; Kuzmitz, Aandrew A. (April 2000). "Cardiovascular evidence for an intermediate or higher metabolic rate in an ornithischian dinosaur".
1993:, the closest living relatives of non-avian dinosaurs, have four-chambered hearts (albeit modified in crocodilians), so non-avian dinosaurs probably had them as well; the structure is not necessarily tied to metabolic rate.
4026:
Pearson, Dean A.; Schaefer, Terry; Johnson, Kirk R.; Nichols, Douglas J.; and Hunter, John P. (2002). "Vertebrate biostratigraphy of the Hell Creek
Formation in southwestern North Dakota and northwestern South Dakota".
1181:
from the middle to the tip, which would have reduced the flexibility of the tail. The rib cage was broad, giving it a wide back, and the limbs were robust. The animals may have been able to move on
4165:
Weishampel, David B.; Barrett, Paul M.; Coria, Rodolfo A.; Le Loeuff, Jean; Xu Xing; Zhao Xijin; Sahni, Ashok; Gomani, Elizabeth, M.P.; and Noto, Christopher R. (2004). "Dinosaur
Distribution".
724: 8537), and drew attention to the genus' heavy build and thick bones. Due to these differences from the regular light hypsilophodont build, he suggested that the genus warranted its own
3926:; Breithaupt, Brent H. (1986). "Latest Cretaceous occurrences of nodosaurid ankylosaurs (Dinosauria, Ornithischia) in Western North America and the gradual extinction of the dinosaurs".
557:
heart. There was much discussion over whether the remains were of a heart. Many scientists now doubt the identification of the object and the implications of such an identification.
5309:
1855:
may have been possibly semi-fossorial, perhaps digging burrows underground. Alternatively, the analysis could suggest phylogenetically inherited traits from burrowing ancestors.
835:(with a question mark because he was uncertain that it belonged to the genus). LACM 33542 comprised a large partial hindlimb ("a third larger than described specimens of
5460:
661:
in 1915, describing the well-preserved skeleton. The type specimen was found largely in natural articulation and was missing only the head and neck, which were lost due to
3853:
Cleland, Timothy P.; Stoskopf, Michael K.; Schweitzer, Mary H. (2011). "Histological, chemical, and morphological reexamination of the "heart" of a small Late
Cretaceous
5540:
2983:; Manabe, Makoto (2002). "Synopsis of the Hell Creek (uppermost Cretaceous) dinosaur assemblage". In Hartman, Joseph H.; Johnson, Kirk R.; Nichols, Douglas J. (eds.).
2049:), USA. With the exception of birds, it was one of the last genera of dinosaurs, its remains being found as close as 3 meters to the boundary clay containing the
1041:. It was named by Caleb M. Brown, Clint A. Boyd and Anthony P. Russell and is known only from its holotype, a small, articulated and almost complete skeleton from the
5394:
2095:
1152:: small pointed premaxillary teeth, and leaf-shaped cheek teeth. Six small teeth were present in both premaxillae, with a toothless section at the tip of the beak.
3208:
Novas, Fernando E.; Cambiaso, Andrea V.; Ambrioso, Alfredo (2004). "A new basal iguanodontian (Dinosauria, Ornithischia) from the Upper
Cretaceous of Patagonia".
1185:, given its fairly long arms and wide hands, but this idea has not been widely discussed in the scientific literature, although it does appear in popular works.
517:
of the teeth and jaws suggest a primarily herbivorous animal. This genus of dinosaur is regarded as a specialized neornithischian, traditionally described as a
2399:"A new basal ornithopod dinosaur (Frenchman Formation, Saskatchewan, Canada), and implications for late Maastrichtian ornithischian diversity in North America"
1913:
Their conclusions have been disputed; soon after the initial description, other researchers published a paper where they asserted that the heart is really a
828:
3703:"Neuroanatomy of the late Cretaceous Thescelosaurus neglectus (Neornithischia: Thescelosauridae) reveals novel ecological specialisations within Dinosauria"
2079:. More specimens are known than have been officially described for this genus, such as the Triebold specimen, which has been the source of several skeletal
788:
4029:
The Hell Creek
Formation and the Cretaceous-Tertiary Boundary in the Northern Great Plains: An Integrated Continental Record of the End of the Cretaceous.
2985:
The Hell Creek
Formation and the Cretaceous-Tertiary Boundary in the Northern Great Plains: An Integrated Continental Record of the End of the Cretaceous
2729:; Sues, Hans-Dieter; Witmer, Larry M.; Coria, Rodolfo A. (2004). "Basal Ornithopoda". In Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (eds.).
1803:
was probably slower than other hypsilophodonts, because of its heavier build and leg structure. Compared to them, it had unusual hindlimbs, because the
502:
684:
Other remains of similar animals were found throughout the late 19th century and 20th century. Another well-preserved skeleton from the slightly older
505:
around 66 million years ago. The preservation and completeness of many of its specimens indicate that it may have preferred to live near streams.
5470:
1985:
The question of how this find reflects metabolic rate and dinosaur internal anatomy is moot, though, regardless of the object's identity. Both modern
1875:
2152:, but another suggests it preferred the opposite. The possible preference for channels is based on the relative abundance of thescelosaur fossils in
5535:
5381:
5530:
1282:
2212:
died, it may have been in or near a river, making it easier to bury and preserve for later fossilization. Russell tentatively compared it to the
1239:
he interpreted as located at least along the midline of the neck of one specimen. Scutes have not been found with other articulated specimens of
5465:
4266:
3228:
878:. Aside from the size, Morris drew attention to the way the ankle was constructed, which he considered to be unique except in comparison with
756:
is its ankle, which Galton claimed was damaged and misinterpreted, but which was regarded by
William J. Morris (1976) as truly different from
5490:
4150:
3123:
3089:
2996:
2894:
2774:
2738:
2515:
2372:
1313:. This has not been followed, with Morris arguing strongly against Galton's classification scheme. At any rate, Galton's Iguanodontidae was
1933:
A study published in 2011 applied multiple lines of inquiry to the question of the object's identity, including more advanced CT scanning,
855:, shin bones, and partial thigh bone, along with five cervical (neck) and eleven dorsal (back) vertebrae, from the Hell Creek Formation of
1301:, although recognized as being distinct among them for its robust build, unusual hindlimbs, and, more recently, its unusually long skull.
1898:, an iron mineral, by replacement of the original material. The authors interpreted the structure of the heart as indicating an elevated
5055:
2295:, and it may have been the most common dinosaur there in life, if the Lance Formation had a preservational bias against small animals.
1200:
Large thin flat mineralized plates have been found next to the ribs' sides. Their function is unknown; they may have played a role in
640:
590:
2453:
Boyd, Clint A.; Brown, Caleb M.; Scheetz, Rodney D.; Clarke, Julia A. (2009). "Taxonomic revision of the basal neornithischian taxa
1132:
was a heavily built bipedal animal, probably herbivorous, but potentially not. There was a prominent ridge along the length of both
435:
5075:
3331:
1917:. As they noted, the anatomy given for the object is incorrect (for example, the "aorta" narrows coming into the "heart" and lacks
2961:(1982). "Baby dinosaurs from the Late Cretaceous Lance and Hell Creek formations and a description of a new species of theropod".
5515:
5500:
5495:
1942:
137:
5062:
3638:
2940:
1305:
in 1974 presented one twist to the classic arrangement, suggesting that because of its hindlimb structure and heavy build (not
2692:, a conservative ornithopod dinosaur from the Upper Cretaceous of North America, with comments on ornithopod classification".
2585:
1443:, their most recent common ancestor, and all descendants) and the smaller clades Orodrominae and Thescelosaurinae. The below
1094:
been estimated in the 2.5–4.0 m range for length (8.2–13.1 ft) for various specimens, and a weight of 200–300
701:
5082:
4209:. Wyoming Geological Association Guidebook, 44th Annual Field Conference. Wyoming Geological Association. pp. 127–146.
1882:. It had been originally unearthed in 1993 in northwestern South Dakota. The authors had found the internal detail through
5510:
942:
5068:
149:
3777:
Rowe, Timothy; McBride, Earle F.; Sereno, Paul C. (February 2001). "Technical comment: dinosaur with a heart of stone".
808:
4180:
5525:
5520:
5026:
1938:
3815:; Fisher, Paul E.; Barrick, Reese E.; Stoskopf, Michael K. (February 2001). "Reply: dinosaur with a heart of stone".
1387:(2011) found North American "hypsilophodonts" of Cretaceous age to sort into two related clusters, one consisting of
1799:
in the first meter or so from the ground, feeding selectively, with food held in the mouth by cheeks while chewing.
5505:
5455:
4259:
2006:
1839:
was a more indiscriminate feeder, allowing both animals to share the same environment without competing for food.
685:
2318:
2126:
1333:
of closely related taxa. "Hypsilophodontidae" and "hypsilophodont" are better understood as informal terms for an
5480:
2646:
2611:
2548:
1186:
1014:
768:
721:
709:
1361:
and the species that had been assigned to that genus, there were at least two and possibly three species within
5450:
4354:
2287:
was also abundant in the Lance Formation. Toe bones from this genus are the most common finds after fossils of
1249:
1201:
856:
620:
616:
3759:
2013:
of the time averaged census for large-bodied dinosaurs from the entire Hell Creek Formation in the study area
5485:
2066:
1069:
636:
4205:
Derstler, Kraig (1994). "Dinosaurs of the Lance Formation in eastern Wyoming". In Nelson, Gerald E. (ed.).
2855:
1874:
In 2000, a skeleton of this genus (specimen NCSM 15728) informally known as "Willo", now on display at the
5334:
4450:
3971:
2201:
1254:
5420:
4703:
4252:
2910:
950:
736:
has, since Peter Galton's 1974 review, generally been considered a more robust individual (possibly the
4238:
744:. However, Boyd and colleagues found that they could not assign it to either of their valid species of
323:
2879:
Brett-Surman, Michael K. (1997). "Ornithopods". In Farlow, James O.; Brett-Surman, Michael K. (eds.).
1025:
per Boyd et al. (2009) is represented by at least two, and possibly three valid species: type species
5475:
4971:
4726:
4112:
4046:"Spatial niche partitioning in dinosaurs from the latest Cretaceous (Maastrichtian) of North America"
3935:
3866:
3714:
3604:
3555:
3459:
3402:
3346:
3280:
3174:
2470:
2208:
skeletons are much more complete, suggesting that this genus frequented stream channels. Thus when a
2173:
2038:
1796:
1034:
804:
791: 7210) consisting of a partial skull with heavy ridges on the lower jaw and cheek, four partial
528:
514:
343:
327:
276:
3240:
1925:
867:
5372:
5003:
4571:
3666:
3439:
3327:
2058:
2030:
1883:
1123:
1042:
1193:
oriented almost perpendicular to the body, another idea that has gone by the wayside. As noted by
5288:
5015:
3890:
3683:
3620:
3571:
3475:
3418:
2544:
2486:
2314:
2258:
2254:
2177:
2134:
2026:
1334:
1220:
931:
632:
353:
272:
144:
763:
5425:
2114:. If it is indeed a tooth from a thescelosaur-like animal, this would significantly extend the
5407:
5399:
5256:
4601:
4146:
4093:
4075:
3923:
3882:
3832:
3794:
3740:
3524:
3264:
3233:
Abstract with Programs - Geological Society of America; Southeast Section, 57th Annual Meeting
3190:
3119:
3115:
3109:
3085:
3081:
3073:
2992:
2958:
2890:
2770:
2734:
2511:
2368:
2141:
2061:
suggests an age of 69–68 Ma for the Laramie Formation. There are reports of teeth from older,
2046:
1946:
1821:
of the right foot fused at their tops, hindering swift movement. Examinations of the teeth of
1115:
1054:
852:
737:
597:
5412:
3389:
Han, Feng-Lu; Barrett, Paul M.; Butler, Richard J.; Xu, Xing (2012). "Postcranial anatomy of
2364:
1817:
and running animals in general. One specimen is known to have had a bone pathology, with the
1369:
could not yet be assigned to a species within the genus. It appears to be closely related to
1231:, with a "punctured" texture, but no regular pattern, while William J. Morris suggested that
874:
would have been about 4.5 meters (15 feet) long, greater than average specimens of
4835:
4230:
4120:
4065:
4057:
3943:
3874:
3824:
3786:
3730:
3722:
3675:
3612:
3563:
3514:
3506:
3467:
3410:
3354:
3288:
3182:
2922:
2811:
2567:
2478:
2410:
2103:
1612:
1470:
1232:
1228:
1145:
823:
being a reference to the Hell Creek Formation). Morris also named a new possible species of
729:
609:
494:
388:
252:
239:
3656:
Naylor Hudgins, Michael; Currie, Philip J.; Sullivan, Corwin (2021). "Dental assessment of
2180:
area. He described the environment of the time as a flat floodplain, with a relatively dry
2133:
and other animals and plants that coexisted in North America by the end of the Cretaceous,
1155:
5278:
4983:
4851:
4339:
4314:
4006:
3543:
3443:
2886:
2880:
2726:
2200:. Although most dinosaur skeletons from this area are incomplete, possibly due to the low
2185:
2034:
1667:
1650:
1262:
1058:
612:
565:
491:
484:
41:
2856:"Fossil of dinosaur killed in asteroid strike discovered in North Dakota, scientists say"
922:. To better accommodate this species, Galton suggested that it belonged to his new genus
4116:
3939:
3870:
3735:
3718:
3702:
3608:
3559:
3463:
3406:
3362:
3350:
3306:
3284:
3178:
2830:
2474:
1277:
527:. Several species have been suggested for this genus. Three currently are recognized as
128:
17:
5293:
5273:
5224:
5149:
4844:
4782:
4747:
4667:
4070:
4045:
3519:
3494:
2926:
2245:
2099:
1899:
1887:
1868:
1547:
1428:
1419:
1298:
1144:, or upper beak (a primitive trait among neornithischians). Long rod-like bones called
749:
518:
226:
4097:
1851:
its post-cranial skeletal features such as its strong forelimbs, it is suggested that
1345:
among the hypsilophodonts. One issue that has potentially interfered with classifying
882:, which he regarded as a separate species. Because Morris believed that the ankles of
5444:
5241:
5191:
4949:
4920:
4906:
4890:
4775:
4768:
4660:
4580:
4504:
4497:
4465:
3687:
3624:
3271:
from the Wealden (Early Cretaceous: Barremian) of the Isle of Wight: a reappraisal".
2815:
2415:
2398:
2356:
2279:
2240:
2053:
layer that closes the Cretaceous. The Laramie Formation is the oldest formation that
2022:
1813:
1581:
1514:
1401:
1395:
1293:
1171:
1148:
were present over the eyes, giving the animal heavy bony eyebrows. Its teeth were of
1110:
estimated at 4–4.5 meters (13.1–14.8 feet) long. As discussed more fully under "
1103:
847:
796:
628:
605:
523:
448:
79:
4241:– The official site for "Willo", from the North Carolina Museum of Natural Sciences.
3908:
Chinsamy, Anusuya; Hillenius, Willem J. (2004). "Physiology of nonavian dinosaurs".
3575:
3479:
3422:
3332:"Systematics of Hypsilophodontidae and Basal Iguanodontia (Dinosauria: Ornithopoda)"
2571:
2490:
1454:
5283:
5234:
5177:
5163:
5156:
5111:
4927:
4882:
4810:
4789:
4692:
4559:
4551:
4527:
4481:
4416:
4403:
4366:
4275:
4138:
3947:
3894:
3812:
3139:
3046:
3017:
2980:
2795:
2765:, Dryosauridae". In Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (eds.).
2685:
2267:
2249:
2224:
2165:
2115:
1907:
1684:
1626:
1371:
1342:
1302:
1194:
1175:
1099:
1079:
1046:
998:
981:
was essentially the same as a skull found with a postcranial skeleton that matched
864:
841:
812:
653:
594:
574:
550:
532:
368:
284:
210:
3828:
3790:
3679:
3292:
1781:
549:
The genus attracted media attention in 2000, when a specimen unearthed in 1993 in
3616:
3450:(n.g.), an unusual euornithopod dinosaur from the latest Cretaceous of Romania".
3414:
3393:(Dinosauria, Ornithischia) from the Lower Cretaceous Yixian Formation of China".
3186:
3053:
Gilmore (Ornithischia; Ornithopoda) from the Upper Cretaceous of North America".
2592:, a new species of orthopodous dinosaur from the Edmonton Formation of Alberta".
5366:
5212:
5184:
5142:
5135:
5128:
4960:
4859:
4824:
4761:
4715:
4640:
4519:
4489:
4442:
4434:
2802:
Gilmore (Ornithischia: Ornithopoda) from the Late Cretaceous of North America".
2228:
1955:
1480:
1009:, and agreed with Morris that the ankle structure was distinct, returning it to
935:
54:
3726:
1353:
necessarily belong to it. Clint Boyd and colleagues found that while the clade
5205:
5198:
4992:
4941:
4913:
4898:
4874:
4817:
4796:
4674:
4653:
4629:
4622:
4608:
4594:
4543:
4535:
4511:
4220:
3878:
3567:
3471:
3358:
3229:"The morphology and histology of thoracic plates on neornithischian dinosaurs"
3105:
2234:
2181:
2149:
2075:
1986:
1979:
1914:
1827:
1818:
1564:
1389:
1310:
1244:
1216:
1149:
1141:
1085:
720:, based on another articulated skeleton, this time including a partial skull (
666:
509:
99:
64:
5357:
3307:"Tanis: Fossil found of dinosaur killed in asteroid strike, scientists claim"
2831:"Tanis: Fossil found of dinosaur killed in asteroid strike, scientists claim"
1208:
and became bone as the animal aged. Such plates are known from several other
1005:
sp. They found that LACM 33542, although fragmentary, was a specimen of
5170:
4803:
4587:
4473:
4184:
2769:(1st ed.). Berkeley: University of California Press. pp. 498–509.
2733:(2nd ed.). Berkeley: University of California Press. pp. 393–412.
2505:
2263:
2153:
2062:
2010:
1934:
1891:
1444:
1326:
1318:
1314:
1306:
1205:
1182:
725:
658:
161:
104:
48:
4079:
4061:
3886:
3836:
3798:
3744:
3528:
3510:
3267:; Galton, Peter M. (2008). "The 'dermal armour' of the ornithopod dinosaur
3194:
2991:, 361. Boulder, Colorado: Geological Society of America. pp. 169–176.
2357:"Hypsilophodont dinosaurs: a new species and comments on their systematics"
1958:
minerals not known to be produced in biological systems, such as goethite,
1922:
and partially preserved the more muscular portions of the heart and aorta.
1878:, was described as including the remnants of a four-chambered heart and an
1061:, making it the first non-avian dinosaur fossil recovered from that date.
672:
5351:
5121:
4615:
4458:
4323:
4299:
3024:(Ornithischia: Ornithopoda) from the Upper Cretaceous of North America".
2482:
2213:
2189:
2161:
2157:
2042:
1975:
1959:
1895:
1309:, or built for running, by his definition), it should be included in the
1209:
1166:
Thescelosaurs had short, broad, five-fingered hands, four-toed feet with
1095:
969:
in 2009, using new cranial material as a starting point. They found that
792:
631:. The skeleton, however, remained in its shipping crates for years until
582:
487:
194:
181:
94:
89:
74:
69:
59:
930:, although he also noted that it could be belong to the similarly sized
5386:
5249:
4681:
4305:
3645:. St. Paul, Minnesota: The Science Museum of Minnesota. pp. 31–32.
3495:"First trace and body fossil evidence of a burrowing, denning dinosaur"
2273:
2107:
2080:
2050:
1951:
1427:
clade and several long-snouted Asian forms (previously described under
1269:
was not as feathered as hypothesized for other small neornithischians.
1190:
1137:
1133:
860:
689:
662:
624:
601:
572:
holotype at the Smithsonian Museum, with skull and neck restored after
109:
84:
4293:
3493:
Varricchio, David J.; Martin, Anthony J.; Katsura, Yoshihiro (2007).
2197:
1971:
1967:
1963:
1918:
1417:(2013) recovered similar results, with the addition of the new genus
1317:
and not a natural group, and so would not be recognized under modern
1224:
1178:
906:, Galton demonstrated that the features Morris had thought connected
693:
643:
had it prepared and described it in a short paper in 1913, naming it
554:
171:
5328:
894:. However, the scientific literature has favored Galton's view that
2947:. St. Paul, Minnesota: The Science Museum of Minnesota. p. 31.
2885:. Bloomington and Indianapolis: Indiana University Press. pp.
1021:
in some anatomical details, and may represent a new species. Thus,
3986:
2217:
2145:
2125:
2005:
1924:
1879:
1862:
1808:
1804:
1780:
1453:
1330:
1276:
1236:
1154:
1068:
941:
762:
671:
564:
498:
480:
2761:
Sues, Hans-Dieter; Norman, David B. (1990). "Hypsilophodontidae,
2164:
or accumulations of multiple individuals have yet been reported.
1126:
remains are known from several locations, mostly based on teeth.
4867:
3049:(1997). "Cranial anatomy of the basal hypsilophodontid dinosaur
2193:
1990:
1863:
1167:
5332:
5098:
4401:
4286:
4248:
4244:
4145:. Minocqua, Wisconsin: NorthWord Press, Inc. pp. 175–176.
4124:
3664:(Ornithischia: Thescelosauridae): paleoecological inferences".
27:
Ornithischian dinosaur genus from Late Cretaceous US and Canada
1119:
3972:"The vertebrate fauna of the Judith River Formation, Montana"
2913:(1927). "The pelvic musculature of ornithischian dinosaurs".
2553:, an orthopodus dinosaur from the Lance Formation of Wyoming"
2397:
Brown, Caleb M.; Boyd, Clint A.; Russell, Anthony P. (2011).
473:
466:
458:
451:
418:
2798:(1995). "The species of the basal hypsilophodontid dinosaur
394:
4181:"Cretaceous "Hell Creek Faunal Facies"; Late Maastrichtian"
4098:"Late Cretaceous dinosaurs from the Denver Basin, Colorado"
2804:
Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen
2223:
Other dinosaurs that shared its time and place include the
424:
409:
397:
752:
within the genus. The other point of contention regarding
651:: "neglected"). At the time, he thought it was related to
497:
in North America. It was a member of the last dinosaurian
2653:, n. sp., and classification of the Hypsilophodontidae".
403:
3546:(2008). "The phylogeny of the ornithischian dinosaurs".
3020:(1999). "Cranial anatomy of the hypsilophodont dinosaur
2184:
climate that supported a variety of plants ranging from
1846:, specifically the "Willo" specimen by Button and Zanno
819:("large-cheeked lizard belonging to the lower regions",
3701:
Button, David J.; Zanno, Lindsay E. (6 November 2023).
3499:
Proceedings of the Royal Society B: Biological Sciences
2140:
Conflicting reports have been made as to its preferred
1894:) under airless burial conditions, and then changed to
1825:
and comparisons with the contemporary pachycephalosaur
1136:(the tooth-bearing "cheek" bones), and a ridge on both
716:, in 1937. Sternberg also named an additional species,
2156:, representing channel environments, in comparison to
1867:"Willo" specimen, with the possible heart left of the
957:
Clint Boyd and colleagues published a reassessment of
5310:
List of stratigraphic units with ornithischian tracks
1886:(CT) imagery. They suggested that the heart had been
1037:, RSM P.1225.1 was assigned to its own species,
465:) meaning "godlike", "marvellous", or "wondrous" and
436:
427:
415:
400:
391:
2319:"A new dinosaur from the Lance Formation of Wyoming"
1435:
clade. They also formally defined Thescelosauridae (
1219:, be it scales or something else, remained unknown.
1111:
1049:. In April 2022, it was reported that a specimen of
914:
were the result of damage to the latter's ankle, so
704:
in 1926. This skeleton had notable differences from
421:
406:
333:
314:
291:
5341:
5302:
5266:
5222:
5109:
5014:
4991:
4982:
4959:
4834:
4745:
4714:
4691:
4639:
4570:
4414:
4365:
4338:
1215:For most of its history, the nature of this genus'
412:
3979:Bulletin of the American Museum of Natural History
2620:Geological Society of America Proceedings for 1936
3760:"Dig This: 'Neglected' Dinosaur Had Super Senses"
3142:(1970). "The posture of hadrosaurian dinosaurs".
2594:University of Toronto Studies (Geological Series)
902:(see above). In the same paper that he described
553:, United States, was interpreted as including a
521:, but more recently recognized as distinct from
4044:Lyson, Tyler R.; Longrich, Nicholas R. (2011).
1365:, and several specimens previously assigned to
863:, USA. The specimen was discovered by amateur
771: 8537, described as the type specimen for
4143:An Odyssey in Time: Dinosaurs of North America
1341:has been regarded as both very basal and very
1033:, and a possible unnamed species. In December
1013:. Finally, they noted that another specimen,
4260:
2392:
2390:
2388:
2386:
2384:
2069:of Alberta, but these specimens are not from
676:Charles Gilmore's 1915 reconstruction of the
8:
3643:Dinosaurs of the Science Museum of Minnesota
3589:
3587:
3585:
3384:
3382:
3227:Boyd, Clint A.; Cleland, Timothy P. (2008).
2945:Dinosaurs of the Science Museum of Minnesota
2448:
2446:
2021:remains are known definitely only from late
1261:specimen allegedly killed on the day of the
1057:, supposedly dating to the exact day of the
3848:
3846:
3159:
3157:
2989:Geological Society of America Special Paper
2444:
2442:
2440:
2438:
2436:
2434:
2432:
2430:
2428:
2426:
2172:was the most common small herbivore in the
2160:, representing floodplain environments. No
1462:NCSM 15728 ("Willo") in left and right view
1349:is that not all of the remains assigned to
939:, or be part of a third, unknown dinosaur.
787:In his paper, Morris described a specimen (
5461:Late Cretaceous dinosaurs of North America
5329:
5106:
5095:
4988:
4753:
4645:
4426:
4422:
4411:
4398:
4344:
4335:
4283:
4267:
4253:
4245:
3111:The Dinosaur Society Dinosaur Encyclopedia
3072:Lambert, David; the Diagram Group (1990).
2363:. Toronto: Royal Ontario Museum. pp.
1077:species (right) compared to its relatives
993:, they regarded the genus as a synonym of
127:
31:
4069:
4039:
4037:
3734:
3660:(Ornithischia: Pachycephalosauridae) and
3518:
3434:
3432:
2721:
2719:
2717:
2715:
2713:
2711:
2709:
2707:
2414:
2403:Zoological Journal of the Linnean Society
1876:North Carolina Museum of Natural Sciences
3041:
3039:
3012:
3010:
3008:
2790:
2788:
2786:
2680:
2678:
2676:
2674:
2672:
2670:
2668:
2618:, with a description of a new species".
2094:in 1999, he described the dentary tooth
1227:material near the shoulders as possible
5541:Multispecific non-avian dinosaur genera
3322:
3320:
3259:
3257:
2756:
2754:
2752:
2750:
2641:
2639:
2637:
2635:
2633:
2631:
2629:
2560:Proceedings of the U.S. National Museum
2539:
2537:
2535:
2533:
2531:
2529:
2527:
2510:. Oxford University Press, Oxford, UK.
2507:Greek-English Lexicon, Abridged Edition
2350:
2348:
2346:
2344:
2342:
2340:
2338:
2336:
2309:
2307:
2303:
1283:Rocky Mountain Dinosaur Resource Center
1243:, though, and Morris's scutes could be
918:could also be considered distinct from
1954:'s heart; the "walls" are composed of
1297:and other small neornithischians as a
1001:, and SDSM 7210 as an example of
593: 7757) was discovered in 1891 by
2323:Smithsonian Miscellaneous Collections
503:Cretaceous–Paleogene extinction event
490:that appeared at the very end of the
7:
3542:Butler, Richard J.; Upchurch, Paul;
3446:(2003). "Osteology and phylogeny of
1247:in origin. In his 2022 documentary,
3548:Journal of Systematic Palaeontology
3452:Journal of Systematic Palaeontology
3442:; Jianu, Coralia-Maria; Csiki, Z.;
1970:, as well as some plant fragments;
1700:
1642:
1618:
1539:
1493:
1486:
1476:
1466:
4050:Proceedings of the Royal Society B
3928:Journal of Vertebrate Paleontology
3597:Journal of Vertebrate Paleontology
3395:Journal of Vertebrate Paleontology
2927:10.1111/j.1463-6395.1927.tb00653.x
2463:Journal of Vertebrate Paleontology
2144:; two papers suggest it preferred
2073:and are much more likely those of
2065:-age rocks, particularly from the
1235:was present, in the form of small
803:, from the late Maastrichtian-age
641:National Museum of Natural History
25:
3912:(2nd ed.). pp. 643–659.
3758:Peake, Tracey (7 November 2023).
1281:Front of reconstructed skeleton,
989:could not be differentiated from
5471:Taxa named by Charles W. Gilmore
5081:
5074:
5067:
5061:
5054:
4239:Willo, the Dinosaur with a Heart
4223:
3080:. New York: Avon Books. p.
2416:10.1111/j.1096-3642.2011.00735.x
2168:, in a popular work, noted that
1943:X-ray photoelectron spectroscopy
1379:addressed these dinosaurs. Boyd
890:, he tentatively assigned it to
748:and regarded the specimen as of
387:
148:
138:Burpee Museum of Natural History
5536:Ornithischians of North America
3987:http://hdl.handle.net/2246/1099
2572:10.5479/si.00963801.49-2127.591
1112:Discovery, history, and species
886:compared favorably to those of
600:and William H. Utterback, from
561:Discovery, history, and species
5531:Late Cretaceous ornithischians
3948:10.1080/02724634.1986.10011619
3330:; Heinrich, Ronald E. (1992).
1835:was a selective feeder, while
1405:, and the other consisting of
1045:(late Maastrichtian stage) of
1017: P.1225.1, differed from
799:as an unidentified species of
1:
5466:Fossil taxa described in 1913
3829:10.1126/science.291.5505.783a
3791:10.1126/science.291.5505.783a
3680:10.1016/j.cretres.2021.105058
3293:10.1016/j.cretres.2008.02.002
3114:. Random House, Inc. p.
2829:Amos, Jonathan (2022-04-06).
2041:), Montana (Hell Creek), and
2002:Temporal and geographic range
1842:Examinations of the skull of
1709:Thescelosaurus assiniboiensis
1291:has generally been allied to
1162:protofeathered reconstruction
1039:Thescelosaurus assiniboiensis
5491:Paleontology in South Dakota
4009:. Triebold Paleontology, Inc
3617:10.1080/02724634.2013.746229
3415:10.1080/02724634.2012.694385
3187:10.1126/science.288.5465.503
2651:Thescelosaurus edmontonensis
2504:Liddell & Scott (1980).
1174:, and a long tail braced by
880:Thescelosaurus edmontonensis
845:or nearly twice as large as
474:
459:
342:Brown, Boyd, & Russell,
3391:Jeholosaurus shangyuanensis
3055:Revue Paléobiologie, Genève
3026:Revue Paléobiologie, Genève
2614:(1937). "Classification of
2359:. In Churcher, C.S. (ed.).
2355:Morris, William J. (1976).
740:of the type individual) of
5557:
4096:; Young; D. Bruce (2002).
3727:10.1038/s41598-023-45658-3
3108:; Glut, Donald F. (1993).
2816:10.1127/njgpa/198/1995/297
2025:-age rocks, from Alberta (
1929:"Willo"'s supposed "heart"
1189:reconstructed it with the
1089:(left), as well as a human
712:placed it in a new genus,
686:Horseshoe Canyon Formation
467:
452:
5105:
5094:
5051:
4756:
4648:
4429:
4425:
4410:
4397:
4347:
4334:
4282:
3879:10.1007/s00114-010-0760-1
3568:10.1017/S1477201907002271
3472:10.1017/S1477201903001032
3359:10.1080/10292389209380426
1859:Supposed fossilized heart
1722:
1705:
1698:
1681:
1664:
1647:
1640:
1623:
1616:
1578:
1561:
1544:
1537:
1511:
1498:
1491:
1484:
1474:
973:was indeed distinct from
657:. He provided a detailed
359:
352:
311:
306:
290:
283:
145:Scientific classification
143:
135:
126:
34:
4207:The Dinosaurs of Wyoming
3975:(free PDF, may not load)
3662:Thescelosaurus neglectus
3051:Thescelosaurus neglectus
2963:Contributions to Geology
2033:), Canada, and Wyoming (
1726:Thescelosaurus neglectus
1437:Thescelosaurus neglectus
1250:Dinosaurs: The Final Day
977:, and that the skull of
296:Thescelosaurus neglectus
136:Reconstructed skeleton,
18:Thescelosaurus neglectus
5516:Cretaceous South Dakota
5501:Paleontology in Alberta
5496:Paleontology in Wyoming
3144:Journal of Paleontology
2694:Journal of Paleontology
2655:Journal of Paleontology
2067:Dinosaur Park Formation
1122:larger than the other.
949:with skin impressions,
898:was not different from
637:Smithsonian Institution
4105:Rocky Mountain Geology
4062:10.1098/rspb.2010.1444
3511:10.1098/rspb.2006.0443
3239:(2): 2. Archived from
3078:The Dinosaur Data Book
3022:Bugenasaura infernalis
2590:Thescelosaurus warreni
2202:preservation potential
2137:
2086:When Galton revisited
2014:
1930:
1871:
1789:
1463:
1329:), with various small
1285:
1255:Sir David Attenborough
1163:
1090:
979:Bugenasaura infernalis
954:
817:Bugenasaura infernalis
784:
681:
578:
5421:Paleobiology Database
3970:Sahni, Ashok (1972).
2882:The Complete Dinosaur
2647:Sternberg, Charles M.
2612:Sternberg, Charles M.
2129:
2098: 46911 from the
2009:
1928:
1866:
1784:
1457:
1280:
1223:described patches of
1158:
1072:
951:Museum of the Rockies
945:
851:") including a foot,
766:
675:
619:(at the time part of
568:
5511:Ornithischian genera
3440:Weishampel, David B.
3328:Weishampel, David B.
2483:10.1671/039.029.0328
2174:Hell Creek Formation
2118:range of the group.
2039:Hell Creek Formation
2029:) and Saskatchewan (
1807:was longer than the
1337:, not a true clade.
1187:Charles M. Sternberg
1114:", it may have been
805:Hell Creek Formation
710:Charles M. Sternberg
4572:Heterodontosauridae
4117:2002RMGeo..37..237C
4056:(1709): 1158–1164.
3940:1986JVPal...6..251C
3871:2011NW.....98..203C
3859:Naturwissenschaften
3719:2023NatSR..1319224B
3667:Cretaceous Research
3609:2013JVPal..33..495B
3560:2008JSPal...6....1B
3505:(1616): 1361–1368.
3464:2003JSPal...1...65W
3407:2012JVPal..32.1370H
3351:1992HBio....6..159W
3285:2008CrRes..29..636B
3273:Cretaceous Research
3179:2000Sci...288..503F
2545:Gilmore, Charles W.
2475:2009JVPal..29..758B
2315:Gilmore, Charles W.
2255:pachycephalosaurian
2059:magnetostratigraphy
2057:is known from, and
2031:Frenchman Formation
1884:computed tomography
1357:included the genus
1043:Frenchman Formation
831: 33542: ?
750:uncertain placement
5526:Cretaceous Alberta
5521:Cretaceous Wyoming
5289:Hypsilophodontidae
5027:Pachycephalosauria
5016:Pachycephalosauria
4179:Bigelow, Phillip.
4094:Carpenter, Kenneth
3924:Carpenter, Kenneth
3707:Scientific Reports
3658:Stegoceras validum
3639:Erickson, Bruce R.
3339:Historical Biology
3265:Butler, Richard J.
2959:Carpenter, Kenneth
2941:Erickson, Bruce R.
2688:(1974). "Notes on
2259:Pachycephalosaurus
2138:
2135:Smithsonian Museum
2027:Scollard Formation
2015:
1931:
1872:
1811:, the opposite of
1790:
1500:TMP 2008.045.0002
1464:
1441:Orodromeus makelai
1335:evolutionary grade
1286:
1164:
1116:sexually dimorphic
1102:), with the large
1091:
955:
932:pachycephalosaurid
870:, hence the name.
785:
682:
633:Charles W. Gilmore
606:late Maastrichtian
579:
479:) "lizard") was a
5506:Laramie Formation
5456:Fossils of Canada
5436:
5435:
5408:Open Tree of Life
5335:Taxon identifiers
5326:
5325:
5322:
5321:
5318:
5317:
5257:Serendipaceratops
5090:
5089:
5049:
5048:
5045:
5044:
5041:
5040:
5037:
5036:
4741:
4740:
4737:
4736:
4602:Heterodontosaurus
4393:
4392:
4389:
4388:
4385:
4384:
4152:978-1-55971-038-1
3173:(5465): 503–505.
3125:978-0-679-41770-5
3091:978-0-380-75896-8
3074:"Thescelosaurids"
2998:978-0-8137-2361-7
2896:978-0-253-33349-0
2776:978-0-520-06727-1
2740:978-0-520-24209-8
2566:(2127): 591–616.
2517:978-0-19-910207-5
2374:978-0-88854-157-4
2047:Laramie Formation
2037:), South Dakota (
1947:scanning electron
1939:X-ray diffraction
1788:scaly restoration
1773:
1772:
1764:
1763:
1755:
1754:
1746:
1745:
1737:
1736:
1602:
1601:
1593:
1592:
1526:
1525:
1447:is that of Brown
1383:(2009) and Brown
1055:Tanis fossil site
1053:was found at the
997:, the species as
598:John Bell Hatcher
544:T. assiniboiensis
378:
377:
372:
346:
338:T. assiniboiensis
330:
279:
16:(Redirected from
5548:
5481:Hell Creek fauna
5429:
5428:
5416:
5415:
5403:
5402:
5390:
5389:
5377:
5376:
5375:
5362:
5361:
5360:
5330:
5107:
5096:
5085:
5079:
5078:
5072:
5071:
5065:
5059:
5058:
4989:
4836:Thescelosauridae
4754:
4646:
4427:
4423:
4412:
4399:
4345:
4336:
4329:
4328:
4284:
4269:
4262:
4255:
4246:
4233:
4231:Dinosaurs portal
4228:
4227:
4226:
4211:
4210:
4202:
4196:
4195:
4193:
4192:
4183:. Archived from
4176:
4170:
4163:
4157:
4156:
4139:Russell, Dale A.
4135:
4129:
4128:
4102:
4090:
4084:
4083:
4073:
4041:
4032:
4024:
4018:
4017:
4015:
4014:
4007:"Thescelosaurus"
4003:
3997:
3996:
3994:
3993:
3976:
3967:
3961:
3958:
3952:
3951:
3920:
3914:
3913:
3905:
3899:
3898:
3850:
3841:
3840:
3813:Russell, Dale A.
3809:
3803:
3802:
3774:
3768:
3767:
3755:
3749:
3748:
3738:
3698:
3692:
3691:
3653:
3647:
3646:
3635:
3629:
3628:
3591:
3580:
3579:
3544:Norman, David B.
3539:
3533:
3532:
3522:
3490:
3484:
3483:
3444:Norman, David B.
3436:
3427:
3426:
3401:(6): 1370–1395.
3386:
3377:
3376:
3374:
3373:
3367:
3361:. Archived from
3336:
3324:
3315:
3314:
3303:
3297:
3296:
3261:
3252:
3251:
3249:
3248:
3224:
3218:
3217:
3205:
3199:
3198:
3161:
3152:
3151:
3140:Galton, Peter M.
3136:
3130:
3129:
3102:
3096:
3095:
3069:
3063:
3062:
3047:Galton, Peter M.
3043:
3034:
3033:
3018:Galton, Peter M.
3014:
3003:
3002:
2981:Russell, Dale A.
2977:
2971:
2970:
2955:
2949:
2948:
2937:
2931:
2930:
2921:(2–3): 225–275.
2911:Romer, Alfred S.
2907:
2901:
2900:
2876:
2870:
2869:
2867:
2866:
2854:Martin, Saleen.
2851:
2845:
2844:
2842:
2841:
2826:
2820:
2819:
2796:Galton, Peter M.
2792:
2781:
2780:
2758:
2745:
2744:
2727:Norman, David B.
2723:
2702:
2701:
2686:Galton, Peter M.
2682:
2663:
2662:
2643:
2624:
2623:
2608:
2602:
2601:
2586:Parks, William A
2582:
2576:
2575:
2557:
2541:
2522:
2521:
2501:
2495:
2494:
2450:
2421:
2420:
2418:
2409:(4): 1157–1198.
2394:
2379:
2378:
2352:
2331:
2330:
2311:
1908:cold-bloodedness
1906:, not reptilian
1701:
1643:
1619:
1613:Thescelosaurinae
1540:
1494:
1487:
1477:
1471:Thescelosauridae
1467:
1299:hypsilophodontid
912:T. edmontonensis
896:T. edmontonensis
888:T. edmontonensis
773:T. edmontonensis
754:T. edmontonensis
734:T. edmontonensis
730:Thescelosaurinae
718:T. edmontonensis
610:Upper Cretaceous
477:
470:
469:
462:
455:
454:
443:
439:
434:
433:
430:
429:
426:
423:
420:
417:
414:
411:
408:
405:
402:
399:
396:
393:
367:
341:
335:
322:
316:
293:
271:
264:
253:Thescelosaurinae
251:
240:Thescelosauridae
238:
225:
209:
153:
152:
131:
121:
51:
40:Temporal range:
32:
21:
5556:
5555:
5551:
5550:
5549:
5547:
5546:
5545:
5451:Thescelosaurids
5441:
5440:
5437:
5432:
5424:
5419:
5411:
5406:
5398:
5393:
5385:
5380:
5371:
5370:
5365:
5356:
5355:
5350:
5337:
5327:
5314:
5298:
5279:Phytodinosauria
5262:
5218:
5101:
5086:
5073:
5066:
5053:
5033:
5010:
4984:Marginocephalia
4978:
4955:
4852:Changchunsaurus
4830:
4750:
4733:
4710:
4687:
4635:
4566:
4419:
4406:
4381:
4361:
4355:Avemetatarsalia
4340:Avemetatarsalia
4330:
4315:Avemetatarsalia
4289:
4288:
4278:
4273:
4229:
4224:
4222:
4219:
4214:
4204:
4203:
4199:
4190:
4188:
4178:
4177:
4173:
4169:(2nd). 517–606.
4164:
4160:
4153:
4137:
4136:
4132:
4100:
4092:
4091:
4087:
4043:
4042:
4035:
4025:
4021:
4012:
4010:
4005:
4004:
4000:
3991:
3989:
3974:
3969:
3968:
3964:
3959:
3955:
3922:
3921:
3917:
3907:
3906:
3902:
3852:
3851:
3844:
3811:
3810:
3806:
3776:
3775:
3771:
3757:
3756:
3752:
3700:
3699:
3695:
3655:
3654:
3650:
3637:
3636:
3632:
3593:
3592:
3583:
3541:
3540:
3536:
3492:
3491:
3487:
3438:
3437:
3430:
3388:
3387:
3380:
3371:
3369:
3365:
3334:
3326:
3325:
3318:
3313:. 6 April 2022.
3305:
3304:
3300:
3263:
3262:
3255:
3246:
3244:
3226:
3225:
3221:
3207:
3206:
3202:
3163:
3162:
3155:
3138:
3137:
3133:
3126:
3104:
3103:
3099:
3092:
3071:
3070:
3066:
3045:
3044:
3037:
3016:
3015:
3006:
2999:
2979:
2978:
2974:
2957:
2956:
2952:
2939:
2938:
2934:
2909:
2908:
2904:
2897:
2878:
2877:
2873:
2864:
2862:
2853:
2852:
2848:
2839:
2837:
2828:
2827:
2823:
2794:
2793:
2784:
2777:
2760:
2759:
2748:
2741:
2725:
2724:
2705:
2700:(5): 1048–1067.
2684:
2683:
2666:
2645:
2644:
2627:
2610:
2609:
2605:
2584:
2583:
2579:
2555:
2543:
2542:
2525:
2518:
2503:
2502:
2498:
2452:
2451:
2424:
2396:
2395:
2382:
2375:
2354:
2353:
2334:
2313:
2312:
2305:
2301:
2124:
2035:Lance Formation
2004:
1999:
1861:
1779:
1774:
1765:
1756:
1747:
1738:
1651:Changchunsaurus
1603:
1594:
1527:
1275:
1263:K-Pg extinction
1221:Charles Gilmore
1067:
1059:K-Pg extinction
857:Garfield County
621:Converse County
617:Niobrara County
613:Lance Formation
595:paleontologists
563:
492:Late Cretaceous
485:neornithischian
441:
437:
390:
386:
302:
299:
270:
262:
249:
236:
223:
207:
147:
122:
120:
119:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
46:
45:
42:Late Cretaceous
38:
28:
23:
22:
15:
12:
11:
5:
5554:
5552:
5544:
5543:
5538:
5533:
5528:
5523:
5518:
5513:
5508:
5503:
5498:
5493:
5488:
5486:Scollard fauna
5483:
5478:
5473:
5468:
5463:
5458:
5453:
5443:
5442:
5434:
5433:
5431:
5430:
5417:
5404:
5391:
5378:
5373:Thescelosaurus
5363:
5347:
5345:
5343:Thescelosaurus
5339:
5338:
5333:
5324:
5323:
5320:
5319:
5316:
5315:
5313:
5312:
5306:
5304:
5303:Other articles
5300:
5299:
5297:
5296:
5294:Jeholosauridae
5291:
5286:
5281:
5276:
5274:Ornithoscelida
5270:
5268:
5264:
5263:
5261:
5260:
5253:
5246:
5238:
5230:
5228:
5225:Incertae sedis
5220:
5219:
5217:
5216:
5209:
5202:
5195:
5188:
5181:
5174:
5167:
5160:
5153:
5150:Ferganocephale
5146:
5139:
5132:
5125:
5117:
5115:
5103:
5102:
5099:
5092:
5091:
5088:
5087:
5052:
5050:
5047:
5046:
5043:
5042:
5039:
5038:
5035:
5034:
5032:
5031:
5030:
5029:
5020:
5018:
5012:
5011:
5009:
5008:
5007:
5006:
4997:
4995:
4986:
4980:
4979:
4977:
4976:
4975:
4974:
4965:
4963:
4957:
4956:
4954:
4953:
4946:
4938:
4935:Thescelosaurus
4931:
4924:
4917:
4910:
4903:
4895:
4887:
4879:
4871:
4864:
4856:
4848:
4845:Albertadromeus
4840:
4838:
4832:
4831:
4829:
4828:
4821:
4814:
4807:
4800:
4793:
4786:
4783:Kulindadromeus
4779:
4772:
4765:
4757:
4751:
4748:Neornithischia
4746:
4743:
4742:
4739:
4738:
4735:
4734:
4732:
4731:
4730:
4729:
4720:
4718:
4712:
4711:
4709:
4708:
4707:
4706:
4697:
4695:
4689:
4688:
4686:
4685:
4678:
4671:
4668:Scutellosaurus
4664:
4657:
4649:
4643:
4637:
4636:
4634:
4633:
4626:
4619:
4612:
4605:
4598:
4591:
4584:
4576:
4574:
4568:
4567:
4565:
4564:
4556:
4548:
4540:
4532:
4524:
4516:
4508:
4501:
4494:
4486:
4478:
4470:
4462:
4455:
4447:
4439:
4430:
4420:
4415:
4408:
4407:
4402:
4395:
4394:
4391:
4390:
4387:
4386:
4383:
4382:
4380:
4379:
4378:
4377:
4371:
4369:
4363:
4362:
4360:
4359:
4358:
4357:
4348:
4342:
4332:
4331:
4327:
4326:
4317:
4308:
4302:
4296:
4287:
4280:
4279:
4274:
4272:
4271:
4264:
4257:
4249:
4243:
4242:
4235:
4234:
4218:
4217:External links
4215:
4213:
4212:
4197:
4171:
4167:The Dinosauria
4158:
4151:
4130:
4111:(2): 237–254.
4085:
4033:
4019:
3998:
3962:
3953:
3934:(3): 251–257.
3915:
3910:The Dinosauria
3900:
3865:(3): 203–211.
3855:Thescelosaurus
3842:
3823:(5505): 783a.
3804:
3785:(5505): 783a.
3769:
3750:
3693:
3648:
3630:
3603:(3): 495–520.
3581:
3534:
3485:
3428:
3378:
3345:(3): 159–184.
3316:
3298:
3279:(4): 636–642.
3253:
3219:
3200:
3153:
3131:
3124:
3106:Lessem, Donald
3097:
3090:
3064:
3035:
3004:
2997:
2972:
2950:
2932:
2915:Acta Zoologica
2902:
2895:
2871:
2846:
2821:
2810:(3): 297–311.
2800:Thescelosaurus
2782:
2775:
2767:The Dinosauria
2746:
2739:
2731:The Dinosauria
2703:
2690:Thescelosaurus
2664:
2625:
2616:Thescelosaurus
2603:
2577:
2551:Thescelosaurus
2549:"Osteology of
2523:
2516:
2496:
2469:(3): 758–770.
2455:Thescelosaurus
2422:
2380:
2373:
2332:
2302:
2300:
2297:
2285:Thescelosaurus
2238:, hadrosaurid
2210:Thescelosaurus
2206:Thescelosaurus
2170:Thescelosaurus
2131:Thescelosaurus
2123:
2120:
2100:Upper Jurassic
2088:Thescelosaurus
2071:Thescelosaurus
2055:Thescelosaurus
2019:Thescelosaurus
2003:
2000:
1998:
1995:
1904:Thescelosaurus
1900:metabolic rate
1869:shoulder blade
1860:
1857:
1853:Thescelosaurus
1844:Thescelosaurus
1833:Thescelosaurus
1823:Thescelosaurus
1801:Thescelosaurus
1793:Thescelosaurus
1778:
1775:
1771:
1770:
1767:
1766:
1762:
1761:
1758:
1757:
1753:
1752:
1749:
1748:
1744:
1743:
1740:
1739:
1735:
1734:
1731:
1730:
1721:
1718:
1717:
1714:
1713:
1704:
1699:
1697:
1694:
1693:
1690:
1689:
1680:
1677:
1676:
1673:
1672:
1663:
1660:
1659:
1656:
1655:
1646:
1641:
1639:
1636:
1635:
1632:
1631:
1622:
1617:
1615:
1609:
1608:
1605:
1604:
1600:
1599:
1596:
1595:
1591:
1590:
1587:
1586:
1577:
1574:
1573:
1570:
1569:
1560:
1557:
1556:
1553:
1552:
1548:Albertadromeus
1543:
1538:
1536:
1533:
1532:
1529:
1528:
1524:
1523:
1520:
1519:
1510:
1507:
1506:
1503:
1502:
1497:
1492:
1490:
1485:
1483:
1475:
1473:
1465:
1433:Thescelosaurus
1429:Jeholosauridae
1420:Albertadromeus
1411:Thescelosaurus
1363:Thescelosaurus
1355:Thescelosaurus
1347:Thescelosaurus
1339:Thescelosaurus
1311:Iguanodontidae
1289:Thescelosaurus
1274:
1273:Classification
1271:
1267:Thescelosaurus
1259:Thescelosaurus
1241:Thescelosaurus
1130:Thescelosaurus
1098:(450–660
1075:Thescelosaurus
1066:
1063:
1051:Thescelosaurus
1023:Thescelosaurus
1007:Thescelosaurus
995:Thescelosaurus
991:Thescelosaurus
983:Thescelosaurus
975:Thescelosaurus
959:Thescelosaurus
947:Thescelosaurus
920:Thescelosaurus
892:Thescelosaurus
868:Harley Garbani
865:paleontologist
825:Thescelosaurus
809:Harding County
801:Thescelosaurus
746:Thescelosaurus
587:Thescelosaurus
562:
559:
519:hypsilophodont
382:Thescelosaurus
376:
375:
374:
373:
357:
356:
350:
349:
348:
347:
331:
309:
308:
307:Other species
304:
303:
300:
288:
287:
281:
280:
267:Thescelosaurus
260:
256:
255:
247:
243:
242:
234:
230:
229:
227:Neornithischia
221:
214:
213:
205:
198:
197:
192:
185:
184:
179:
175:
174:
169:
165:
164:
159:
155:
154:
141:
140:
133:
132:
124:
123:
115:
114:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
52:
39:
36:Thescelosaurus
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
5553:
5542:
5539:
5537:
5534:
5532:
5529:
5527:
5524:
5522:
5519:
5517:
5514:
5512:
5509:
5507:
5504:
5502:
5499:
5497:
5494:
5492:
5489:
5487:
5484:
5482:
5479:
5477:
5474:
5472:
5469:
5467:
5464:
5462:
5459:
5457:
5454:
5452:
5449:
5448:
5446:
5439:
5427:
5422:
5418:
5414:
5409:
5405:
5401:
5396:
5392:
5388:
5383:
5379:
5374:
5368:
5364:
5359:
5353:
5349:
5348:
5346:
5344:
5340:
5336:
5331:
5311:
5308:
5307:
5305:
5301:
5295:
5292:
5290:
5287:
5285:
5282:
5280:
5277:
5275:
5272:
5271:
5269:
5265:
5259:
5258:
5254:
5252:
5251:
5247:
5244:
5243:
5242:Dravidosaurus
5239:
5237:
5236:
5232:
5231:
5229:
5227:
5226:
5221:
5215:
5214:
5210:
5208:
5207:
5203:
5201:
5200:
5196:
5194:
5193:
5192:Syngonosaurus
5189:
5187:
5186:
5182:
5180:
5179:
5175:
5173:
5172:
5168:
5166:
5165:
5161:
5159:
5158:
5154:
5152:
5151:
5147:
5145:
5144:
5140:
5138:
5137:
5133:
5131:
5130:
5126:
5124:
5123:
5119:
5118:
5116:
5114:
5113:
5108:
5104:
5097:
5093:
5084:
5080:
5077:
5070:
5064:
5057:
5028:
5024:
5023:
5022:
5021:
5019:
5017:
5013:
5005:
5001:
5000:
4999:
4998:
4996:
4994:
4990:
4987:
4985:
4981:
4973:
4969:
4968:
4967:
4966:
4964:
4962:
4958:
4952:
4951:
4950:Zephyrosaurus
4947:
4944:
4943:
4939:
4937:
4936:
4932:
4930:
4929:
4925:
4923:
4922:
4921:Oryctodromeus
4918:
4916:
4915:
4911:
4909:
4908:
4907:Nevadadromeus
4904:
4901:
4900:
4896:
4893:
4892:
4891:Koreanosaurus
4888:
4885:
4884:
4880:
4877:
4876:
4872:
4870:
4869:
4865:
4862:
4861:
4857:
4854:
4853:
4849:
4847:
4846:
4842:
4841:
4839:
4837:
4833:
4827:
4826:
4822:
4820:
4819:
4815:
4813:
4812:
4808:
4806:
4805:
4801:
4799:
4798:
4794:
4792:
4791:
4787:
4785:
4784:
4780:
4778:
4777:
4776:Hypsilophodon
4773:
4771:
4770:
4769:Hexinlusaurus
4766:
4764:
4763:
4759:
4758:
4755:
4752:
4749:
4744:
4728:
4724:
4723:
4722:
4721:
4719:
4717:
4713:
4705:
4701:
4700:
4699:
4698:
4696:
4694:
4690:
4684:
4683:
4679:
4677:
4676:
4672:
4670:
4669:
4665:
4663:
4662:
4661:Scelidosaurus
4658:
4656:
4655:
4651:
4650:
4647:
4644:
4642:
4638:
4632:
4631:
4627:
4625:
4624:
4620:
4618:
4617:
4613:
4611:
4610:
4606:
4604:
4603:
4599:
4597:
4596:
4592:
4590:
4589:
4585:
4583:
4582:
4581:Abrictosaurus
4578:
4577:
4575:
4573:
4569:
4562:
4561:
4557:
4554:
4553:
4549:
4546:
4545:
4541:
4538:
4537:
4533:
4530:
4529:
4525:
4522:
4521:
4517:
4514:
4513:
4509:
4507:
4506:
4505:Lesothosaurus
4502:
4500:
4499:
4498:Laquintasaura
4495:
4492:
4491:
4487:
4484:
4483:
4479:
4476:
4475:
4471:
4468:
4467:
4466:Eucoelophysis
4463:
4461:
4460:
4456:
4453:
4452:
4448:
4445:
4444:
4440:
4437:
4436:
4432:
4431:
4428:
4424:
4421:
4418:
4413:
4409:
4405:
4400:
4396:
4375:
4374:
4373:
4372:
4370:
4368:
4364:
4356:
4352:
4351:
4350:
4349:
4346:
4343:
4341:
4337:
4333:
4325:
4321:
4318:
4316:
4312:
4309:
4307:
4303:
4301:
4297:
4295:
4291:
4290:
4285:
4281:
4277:
4270:
4265:
4263:
4258:
4256:
4251:
4250:
4247:
4240:
4237:
4236:
4232:
4221:
4216:
4208:
4201:
4198:
4187:on 2007-01-24
4186:
4182:
4175:
4172:
4168:
4162:
4159:
4154:
4148:
4144:
4140:
4134:
4131:
4126:
4122:
4118:
4114:
4110:
4106:
4099:
4095:
4089:
4086:
4081:
4077:
4072:
4067:
4063:
4059:
4055:
4051:
4047:
4040:
4038:
4034:
4030:
4023:
4020:
4008:
4002:
3999:
3988:
3984:
3980:
3973:
3966:
3963:
3957:
3954:
3949:
3945:
3941:
3937:
3933:
3929:
3925:
3919:
3916:
3911:
3904:
3901:
3896:
3892:
3888:
3884:
3880:
3876:
3872:
3868:
3864:
3860:
3856:
3849:
3847:
3843:
3838:
3834:
3830:
3826:
3822:
3818:
3814:
3808:
3805:
3800:
3796:
3792:
3788:
3784:
3780:
3773:
3770:
3765:
3764:NC State News
3761:
3754:
3751:
3746:
3742:
3737:
3732:
3728:
3724:
3720:
3716:
3712:
3708:
3704:
3697:
3694:
3689:
3685:
3681:
3677:
3673:
3669:
3668:
3663:
3659:
3652:
3649:
3644:
3640:
3634:
3631:
3626:
3622:
3618:
3614:
3610:
3606:
3602:
3598:
3590:
3588:
3586:
3582:
3577:
3573:
3569:
3565:
3561:
3557:
3553:
3549:
3545:
3538:
3535:
3530:
3526:
3521:
3516:
3512:
3508:
3504:
3500:
3496:
3489:
3486:
3481:
3477:
3473:
3469:
3465:
3461:
3457:
3453:
3449:
3445:
3441:
3435:
3433:
3429:
3424:
3420:
3416:
3412:
3408:
3404:
3400:
3396:
3392:
3385:
3383:
3379:
3368:on 2007-09-30
3364:
3360:
3356:
3352:
3348:
3344:
3340:
3333:
3329:
3323:
3321:
3317:
3312:
3308:
3302:
3299:
3294:
3290:
3286:
3282:
3278:
3274:
3270:
3269:Hypsilophodon
3266:
3260:
3258:
3254:
3243:on 2011-06-08
3242:
3238:
3234:
3230:
3223:
3220:
3215:
3211:
3204:
3201:
3196:
3192:
3188:
3184:
3180:
3176:
3172:
3168:
3160:
3158:
3154:
3150:(3): 464–473.
3149:
3145:
3141:
3135:
3132:
3127:
3121:
3117:
3113:
3112:
3107:
3101:
3098:
3093:
3087:
3083:
3079:
3075:
3068:
3065:
3061:(1): 231–258.
3060:
3056:
3052:
3048:
3042:
3040:
3036:
3032:(2): 517–534.
3031:
3027:
3023:
3019:
3013:
3011:
3009:
3005:
3000:
2994:
2990:
2986:
2982:
2976:
2973:
2969:(2): 123–134.
2968:
2964:
2960:
2954:
2951:
2946:
2942:
2936:
2933:
2928:
2924:
2920:
2916:
2912:
2906:
2903:
2898:
2892:
2888:
2884:
2883:
2875:
2872:
2861:
2857:
2850:
2847:
2836:
2832:
2825:
2822:
2817:
2813:
2809:
2805:
2801:
2797:
2791:
2789:
2787:
2783:
2778:
2772:
2768:
2764:
2763:Tenontosaurus
2757:
2755:
2753:
2751:
2747:
2742:
2736:
2732:
2728:
2722:
2720:
2718:
2716:
2714:
2712:
2710:
2708:
2704:
2699:
2695:
2691:
2687:
2681:
2679:
2677:
2675:
2673:
2671:
2669:
2665:
2661:(5): 481–494.
2660:
2656:
2652:
2648:
2642:
2640:
2638:
2636:
2634:
2632:
2630:
2626:
2621:
2617:
2613:
2607:
2604:
2599:
2595:
2591:
2587:
2581:
2578:
2573:
2569:
2565:
2561:
2554:
2552:
2546:
2540:
2538:
2536:
2534:
2532:
2530:
2528:
2524:
2519:
2513:
2509:
2508:
2500:
2497:
2492:
2488:
2484:
2480:
2476:
2472:
2468:
2464:
2460:
2456:
2449:
2447:
2445:
2443:
2441:
2439:
2437:
2435:
2433:
2431:
2429:
2427:
2423:
2417:
2412:
2408:
2404:
2400:
2393:
2391:
2389:
2387:
2385:
2381:
2376:
2370:
2366:
2362:
2358:
2351:
2349:
2347:
2345:
2343:
2341:
2339:
2337:
2333:
2328:
2324:
2320:
2316:
2310:
2308:
2304:
2298:
2296:
2294:
2293:Edmontosaurus
2290:
2286:
2282:
2281:
2280:Tyrannosaurus
2276:
2275:
2270:
2269:
2265:
2261:
2260:
2256:
2252:
2251:
2247:
2243:
2242:
2241:Edmontosaurus
2237:
2236:
2231:
2230:
2226:
2221:
2219:
2215:
2211:
2207:
2203:
2199:
2195:
2191:
2187:
2183:
2179:
2175:
2171:
2167:
2163:
2159:
2155:
2151:
2147:
2143:
2136:
2132:
2128:
2121:
2119:
2117:
2116:stratigraphic
2113:
2109:
2105:
2101:
2097:
2093:
2089:
2084:
2083:for museums.
2082:
2078:
2077:
2072:
2068:
2064:
2060:
2056:
2052:
2048:
2044:
2040:
2036:
2032:
2028:
2024:
2023:Maastrichtian
2020:
2012:
2008:
2001:
1996:
1994:
1992:
1988:
1983:
1981:
1977:
1973:
1969:
1965:
1961:
1957:
1953:
1948:
1944:
1940:
1936:
1927:
1923:
1920:
1916:
1911:
1909:
1905:
1901:
1897:
1893:
1889:
1885:
1881:
1877:
1870:
1865:
1858:
1856:
1854:
1849:
1845:
1840:
1838:
1834:
1831:suggest that
1830:
1829:
1824:
1820:
1816:
1815:
1814:Hypsilophodon
1810:
1806:
1802:
1798:
1794:
1787:
1783:
1776:
1769:
1768:
1760:
1759:
1751:
1750:
1742:
1741:
1733:
1732:
1729:
1728:
1727:
1720:
1719:
1716:
1715:
1712:
1711:
1710:
1703:
1702:
1696:
1695:
1692:
1691:
1688:
1687:
1686:
1679:
1678:
1675:
1674:
1671:
1670:
1669:
1662:
1661:
1658:
1657:
1654:
1653:
1652:
1645:
1644:
1638:
1637:
1634:
1633:
1630:
1629:
1628:
1621:
1620:
1614:
1611:
1610:
1607:
1606:
1598:
1597:
1589:
1588:
1585:
1584:
1583:
1582:Zephyrosaurus
1576:
1575:
1572:
1571:
1568:
1567:
1566:
1559:
1558:
1555:
1554:
1551:
1550:
1549:
1542:
1541:
1535:
1534:
1531:
1530:
1522:
1521:
1518:
1517:
1516:
1515:Oryctodromeus
1509:
1508:
1505:
1504:
1501:
1496:
1495:
1489:
1488:
1482:
1479:
1478:
1472:
1469:
1468:
1461:
1456:
1452:
1450:
1446:
1442:
1438:
1434:
1430:
1426:
1422:
1421:
1416:
1412:
1408:
1404:
1403:
1402:Zephyrosaurus
1398:
1397:
1396:Oryctodromeus
1392:
1391:
1386:
1382:
1376:
1374:
1373:
1368:
1364:
1360:
1356:
1352:
1348:
1344:
1340:
1336:
1332:
1328:
1322:
1320:
1316:
1312:
1308:
1304:
1300:
1296:
1295:
1294:Hypsilophodon
1290:
1284:
1279:
1272:
1270:
1268:
1264:
1260:
1256:
1252:
1251:
1246:
1242:
1238:
1234:
1230:
1226:
1222:
1218:
1213:
1211:
1207:
1203:
1198:
1196:
1192:
1188:
1184:
1180:
1177:
1173:
1169:
1161:
1157:
1153:
1151:
1147:
1143:
1139:
1135:
1131:
1127:
1125:
1121:
1117:
1113:
1109:
1105:
1104:type specimen
1101:
1097:
1088:
1087:
1083:(center) and
1082:
1081:
1076:
1071:
1064:
1062:
1060:
1056:
1052:
1048:
1044:
1040:
1036:
1032:
1028:
1024:
1020:
1016:
1012:
1008:
1004:
1000:
996:
992:
988:
987:B. infernalis
984:
980:
976:
972:
968:
964:
960:
952:
948:
944:
940:
938:
937:
933:
929:
925:
921:
917:
913:
909:
905:
901:
897:
893:
889:
885:
881:
877:
873:
869:
866:
862:
858:
854:
850:
849:
848:Hypsilophodon
844:
843:
838:
834:
830:
827:for specimen
826:
822:
818:
814:
810:
806:
802:
798:
794:
790:
782:
779:sp. per Boyd
778:
774:
770:
765:
761:
759:
755:
751:
747:
743:
739:
735:
731:
727:
723:
719:
715:
711:
707:
703:
702:William Parks
699:
695:
691:
687:
680:type specimen
679:
674:
670:
668:
664:
660:
656:
655:
650:
646:
642:
638:
634:
630:
626:
622:
618:
614:
611:
607:
603:
599:
596:
592:
588:
584:
583:type specimen
577:
576:
571:
567:
560:
558:
556:
552:
547:
545:
541:
537:
534:
530:
526:
525:
524:Hypsilophodon
520:
516:
511:
506:
504:
500:
496:
493:
489:
486:
482:
478:
476:
464:
461:
450:
449:ancient Greek
446:
445:
432:
384:
383:
370:
366:
365:
361:
360:
358:
355:
351:
345:
340:
339:
332:
329:
325:
321:
320:
313:
312:
310:
305:
301:Gilmore, 1913
298:
297:
289:
286:
282:
278:
274:
269:
268:
261:
258:
257:
254:
248:
245:
244:
241:
235:
232:
231:
228:
222:
219:
216:
215:
212:
206:
203:
200:
199:
196:
193:
190:
187:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
151:
146:
142:
139:
134:
130:
125:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
50:
43:
37:
33:
30:
19:
5438:
5342:
5284:Silesauridae
5255:
5248:
5240:
5235:Ajkaceratops
5233:
5223:
5211:
5204:
5197:
5190:
5183:
5178:Notoceratops
5176:
5169:
5164:Gongbusaurus
5162:
5157:Geranosaurus
5155:
5148:
5141:
5134:
5127:
5120:
5112:Nomina dubia
5110:
5060:
4948:
4940:
4934:
4933:
4928:Parksosaurus
4926:
4919:
4912:
4905:
4897:
4889:
4883:Jeholosaurus
4881:
4873:
4866:
4858:
4850:
4843:
4823:
4816:
4811:Sanxiasaurus
4809:
4802:
4795:
4790:Minimocursor
4788:
4781:
4774:
4767:
4760:
4704:Ankylosauria
4693:Ankylosauria
4680:
4673:
4666:
4659:
4652:
4628:
4621:
4614:
4607:
4600:
4593:
4586:
4579:
4560:Technosaurus
4558:
4552:Soumyasaurus
4550:
4542:
4534:
4528:Pisanosaurus
4526:
4518:
4510:
4503:
4496:
4488:
4482:Ignotosaurus
4480:
4472:
4464:
4457:
4449:
4441:
4433:
4417:Ornithischia
4404:Ornithischia
4367:Ornithischia
4319:
4310:
4276:Ornithischia
4206:
4200:
4189:. Retrieved
4185:the original
4174:
4166:
4161:
4142:
4133:
4108:
4104:
4088:
4053:
4049:
4028:
4022:
4011:. Retrieved
4001:
3990:. Retrieved
3982:
3978:
3965:
3956:
3931:
3927:
3918:
3909:
3903:
3862:
3858:
3854:
3820:
3816:
3807:
3782:
3778:
3772:
3763:
3753:
3713:(1): 19224.
3710:
3706:
3696:
3671:
3665:
3661:
3657:
3651:
3642:
3633:
3600:
3596:
3551:
3547:
3537:
3502:
3498:
3488:
3455:
3451:
3447:
3398:
3394:
3390:
3370:. Retrieved
3363:the original
3342:
3338:
3310:
3301:
3276:
3272:
3268:
3245:. Retrieved
3241:the original
3236:
3232:
3222:
3213:
3209:
3203:
3170:
3166:
3147:
3143:
3134:
3110:
3100:
3077:
3067:
3058:
3054:
3050:
3029:
3025:
3021:
2988:
2984:
2975:
2966:
2962:
2953:
2944:
2935:
2918:
2914:
2905:
2881:
2874:
2863:. Retrieved
2859:
2849:
2838:. Retrieved
2834:
2824:
2807:
2803:
2799:
2766:
2762:
2730:
2697:
2693:
2689:
2658:
2654:
2650:
2619:
2615:
2606:
2597:
2593:
2589:
2580:
2563:
2559:
2550:
2506:
2499:
2466:
2462:
2458:
2454:
2406:
2402:
2360:
2326:
2322:
2317:(May 1913).
2292:
2288:
2284:
2278:
2272:
2268:Ornithomimus
2266:
2257:
2250:Ankylosaurus
2248:
2246:ankylosaurid
2239:
2233:
2227:
2222:
2209:
2205:
2204:of forests,
2190:bald cypress
2169:
2166:Dale Russell
2139:
2130:
2111:
2091:
2087:
2085:
2074:
2070:
2054:
2018:
2016:
1997:Paleoecology
1987:crocodilians
1984:
1932:
1912:
1903:
1873:
1852:
1847:
1843:
1841:
1836:
1832:
1826:
1822:
1812:
1800:
1792:
1791:
1786:T. neglectus
1785:
1777:Paleobiology
1725:
1724:
1723:
1708:
1707:
1706:
1685:Jeholosaurus
1683:
1682:
1666:
1665:
1649:
1648:
1627:Parksosaurus
1625:
1624:
1580:
1579:
1563:
1562:
1546:
1545:
1513:
1512:
1499:
1460:T. neglectus
1459:
1448:
1440:
1436:
1432:
1424:
1418:
1414:
1410:
1407:Parksosaurus
1406:
1400:
1394:
1388:
1384:
1380:
1377:
1372:Parksosaurus
1370:
1367:T. neglectus
1366:
1362:
1358:
1354:
1351:T. neglectus
1350:
1346:
1338:
1323:
1315:polyphyletic
1303:Peter Galton
1292:
1288:
1287:
1266:
1258:
1248:
1240:
1214:
1199:
1195:Peter Galton
1165:
1160:T. neglectus
1159:
1129:
1128:
1107:
1092:
1084:
1080:Parksosaurus
1078:
1074:
1073:Size of two
1050:
1047:Saskatchewan
1038:
1030:
1027:T. neglectus
1026:
1022:
1019:T. neglectus
1018:
1010:
1006:
1002:
994:
990:
986:
982:
978:
974:
971:Parksosaurus
970:
967:Parksosaurus
966:
962:
958:
956:
953:specimen 979
946:
934:
927:
923:
919:
915:
911:
907:
903:
900:T. neglectus
899:
895:
891:
887:
883:
879:
876:T. neglectus
875:
871:
846:
842:Parksosaurus
840:
837:T. neglectus
836:
832:
824:
820:
816:
813:South Dakota
800:
797:finger bones
786:
780:
776:
772:
758:T. neglectus
757:
753:
745:
742:T. neglectus
741:
738:opposite sex
733:
717:
714:Parksosaurus
713:
706:T. neglectus
705:
697:
696:, was named
683:
678:T. neglectus
677:
654:Camptosaurus
652:
648:
645:T. neglectus
644:
586:
580:
575:Camptosaurus
573:
570:T. neglectus
569:
551:South Dakota
548:
543:
539:
536:T. neglectus
535:
533:type species
522:
507:
472:
457:
381:
380:
379:
363:
362:
337:
336:
318:
317:
295:
294:
285:Type species
266:
265:
217:
211:Ornithischia
201:
188:
35:
29:
5476:Lance fauna
5367:Wikispecies
5213:Trimucrodon
5185:Siluosaurus
5143:Fabrosaurus
5136:Craspedodon
5129:Bienosaurus
4972:Ornithopoda
4961:Ornithopoda
4860:Changmiania
4825:Yandusaurus
4762:Agilisaurus
4727:Stegosauria
4716:Stegosauria
4641:Thyreophora
4520:Lutungutali
4490:Kwanasaurus
4443:Asilisaurus
4435:Amanasaurus
3985:: 321–412.
3554:(1): 1–40.
3458:(2): 1–56.
3216:(1): 75–82.
3210:Ameghiniana
2459:Bugenasaura
2289:Triceratops
2229:Triceratops
2225:ceratopsids
2182:subtropical
2150:floodplains
2112:Bugenasaura
2092:Bugenasaura
1956:sedimentary
1890:(turned to
1795:would have
1481:Orodrominae
1359:Bugenasaura
1257:reported a
1245:crocodilian
1202:respiration
1118:, with one
1108:T. garbanii
1065:Description
1031:T. garbanii
1011:T. garbanii
963:Bugenasaura
936:Stygimoloch
928:B. garbanii
924:Bugenasaura
916:T. garbanii
908:T. garbanii
904:Bugenasaura
884:T. garbanii
872:T. garbanii
833:T. garbanii
669:dinosaurs.
540:T. garbanii
501:before the
364:Bugenasaura
319:T. garbanii
246:Subfamily:
47:68–66
5445:Categories
5267:Other taxa
5206:Tichosteus
5199:Tatisaurus
5004:Ceratopsia
4993:Ceratopsia
4942:Yueosaurus
4914:Orodromeus
4899:Nanosaurus
4818:Xiaosaurus
4797:Nanosaurus
4675:Yuxisaurus
4654:Emausaurus
4630:Tianyulong
4623:Pegomastax
4609:Lycorhinus
4595:Fruitadens
4544:Silesaurus
4536:Sacisaurus
4512:Lewisuchus
4376:see below↓
4324:Dinosauria
4306:Sauropsida
4191:2007-01-26
4125:10.2113/11
4013:2016-06-01
3992:2007-03-10
3674:: 105058.
3372:2007-03-10
3247:2008-06-13
2865:2022-10-25
2840:2022-10-25
2299:References
2262:, and the
2235:Torosaurus
2188:trees, to
2186:angiosperm
2154:sandstones
2076:Orodromeus
1980:phosphorus
1962:minerals,
1915:concretion
1888:saponified
1837:Stegoceras
1828:Stegoceras
1819:long bones
1565:Orodromeus
1425:Orodromeus
1390:Orodromeus
1225:carbonized
1217:integument
1146:palpebrals
1142:premaxilla
1086:Orodromeus
985:. Because
821:infernalis
795:, and two
698:T. warreni
555:fossilized
195:Dinosauria
5171:Laosaurus
4804:Phyllodon
4588:Echinodon
4474:Gamatavus
4292:Kingdom:
3688:239253658
3625:129160518
2649:(1940). "
2588:(1926). "
2329:(5): 1–5.
2264:theropods
2214:capybaras
2178:Fort Peck
2158:mudstones
2063:Campanian
2011:Pie chart
1935:histology
1892:grave wax
1805:upper leg
1458:Skull of
1445:cladogram
1431:) to the
1327:paraphyly
1319:cladistic
1307:cursorial
1229:epidermis
1210:cerapodas
1206:cartilage
1191:upper arm
1183:all fours
1150:two types
1138:dentaries
1096:kilograms
793:vertebrae
726:subfamily
708:, and so
659:monograph
649:neglectus
460:theskelos
168:Kingdom:
162:Eukaryota
5352:Wikidata
5122:Alocodon
5100:See also
4616:Manidens
4459:Eocursor
4451:Diodorus
4300:Chordata
4298:Phylum:
4294:Animalia
4141:(1989).
4080:20943689
4031:145–167.
3887:21279321
3837:11157158
3799:11157158
3745:37932280
3736:10628235
3641:(2003).
3576:86728076
3529:17374596
3480:86339025
3448:Zalmoxes
3423:86754247
3311:BBC News
3195:10775107
2943:(2003).
2860:phys.org
2835:BBC News
2547:(1915).
2491:84273584
2162:bonebeds
2146:channels
2104:Weymouth
2043:Colorado
1976:nitrogen
1960:feldspar
1919:arteries
1896:goethite
1413:. Brown
1176:ossified
1172:toe tips
1134:maxillae
1124:Juvenile
767:Cast of
488:dinosaur
453:θέσκελος
354:Synonyms
233:Family:
182:Chordata
178:Phylum:
172:Animalia
158:Domain:
5413:4947345
5400:1024605
5387:4572457
5358:Q131150
5250:Jakapil
4682:Jakapil
4304:Class:
4113:Bibcode
4071:3049066
3936:Bibcode
3895:2408562
3867:Bibcode
3817:Science
3779:Science
3715:Bibcode
3605:Bibcode
3556:Bibcode
3520:2176205
3460:Bibcode
3403:Bibcode
3347:Bibcode
3281:Bibcode
3175:Bibcode
3167:Science
2887:330–346
2600:: 1–42.
2471:Bibcode
2274:Troodon
2198:ginkgos
2176:of the
2142:habitat
2122:Habitat
2110:as cf.
2108:England
2051:iridium
1952:ostrich
1848:et al.,
1797:browsed
1423:to the
1343:derived
1321:usage.
1179:tendons
999:dubious
861:Montana
690:Alberta
667:bipedal
663:erosion
635:of the
625:Wyoming
604:of the
510:bipedal
273:Gilmore
259:Genus:
116:↓
4149:
4078:
4068:
3893:
3885:
3835:
3797:
3743:
3733:
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3623:
3574:
3527:
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3193:
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3088:
2995:
2893:
2773:
2737:
2622:: 365.
2514:
2489:
2371:
2365:93–113
2361:Athlon
2277:, and
2218:tapirs
1978:, and
1972:carbon
1968:gypsum
1966:, and
1964:quartz
1945:, and
1449:et al.
1415:et al.
1399:, and
1385:et al.
1381:et al.
1331:clades
1237:scutes
1170:-like
1100:pounds
965:, and
853:tarsus
694:Canada
531:: the
495:period
475:sauros
468:σαυρος
440:-il-ə-
371:, 1995
369:Galton
324:Morris
5426:38742
5395:IRMNG
4320:Clade
4311:Clade
4101:(PDF)
3891:S2CID
3684:S2CID
3621:S2CID
3572:S2CID
3476:S2CID
3419:S2CID
3366:(PDF)
3335:(PDF)
2556:(PDF)
2487:S2CID
2194:ferns
2192:, to
2081:casts
2017:True
1991:birds
1880:aorta
1233:armor
781:et al
688:, in
608:-age
529:valid
508:This
499:fauna
481:genus
438:THESS
218:Clade
202:Clade
189:Clade
5382:GBIF
5025:see
5002:see
4970:see
4875:Haya
4868:Fona
4725:see
4702:see
4353:see
4147:ISBN
4076:PMID
3883:PMID
3833:PMID
3795:PMID
3741:PMID
3525:PMID
3191:PMID
3120:ISBN
3086:ISBN
2993:ISBN
2891:ISBN
2771:ISBN
2735:ISBN
2512:ISBN
2457:and
2369:ISBN
2291:and
2232:and
2216:and
2196:and
2096:UCMP
2090:and
1989:and
1902:for
1809:shin
1668:Haya
1409:and
1168:hoof
1035:2011
910:and
839:and
829:LACM
789:SDSM
783:., )
602:beds
591:USNM
581:The
542:and
515:form
344:2011
328:1976
277:1913
55:PreꞒ
4121:doi
4066:PMC
4058:doi
4054:278
3983:147
3944:doi
3875:doi
3857:".
3825:doi
3821:291
3787:doi
3783:291
3731:PMC
3723:doi
3676:doi
3672:130
3613:doi
3564:doi
3515:PMC
3507:doi
3503:274
3468:doi
3411:doi
3355:doi
3289:doi
3183:doi
3171:288
3116:475
3082:153
2923:doi
2812:doi
2808:198
2568:doi
2479:doi
2461:".
2411:doi
2407:163
2220:.
2148:to
2102:of
1120:sex
1106:of
1015:RSM
926:as
807:of
769:CMN
722:NMC
700:by
639:'s
629:USA
623:),
615:of
585:of
483:of
456:- (
444:-əs
442:SOR
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