1236:
571:
5554:
19:
331:
5633:
5621:
5543:
5549:
1061:
5560:
481:
1249:
417:
281:
970:
5583:
647:
1223:
712:
1993:
873:
793:
2150:
2057:
2297:
5609:
4704:
205:
1028:
or warm-blooded animals. They found that the probable course of airflow through the snout of a living, breathing dromaeosaurid would have been more consistent with that of a cold blooded animal. However, the anatomical reconstructions the researchers used to draw these conclusions have been
3615:
I. Yu. Bolotskii; Yu. L. Bolotskii; A. P. Sorokin (2019). "The first find of an ungual phalanx of a dromaeosaurid dinosaur (Dinosauria: Dromaeosauridae) from the
Blagoveshchensk area of Late Cretaceous dinosaurs (Amur Region, Russia)".
1328:. The authors observed that many aspects of the distribution of feather homologues in the dinosaur family tree met the expectations of earlier phylogenetic hypotheses, including a gradual transition from primitive filaments in
4091:
LĂĽ, J.-C.; Xu, L.; Zhang, X.-L.; Ji, Q.; Jia, S.-H.; Hu, W.-Y.; Zhang, J.-M.; Wu, Y.-H. (2007). "New dromaeosaurid dinosaur from the Late
Cretaceous Qiupa Formation of Luanchuan area, western Henan, China".
195:
in 2000. Three years later, Xu and others would report a new species in this genus that exhibited a bizarre "four winged" body plan with long pennaceous flight feathers on both its front and hind limbs.
1024:
and others attempted to reconstruct the way air would flow through the snouts of dromaeosaurids during respiration to see whether or not it was consistent with the airflow patterns of modern
1147:
that covered the animal in life. The presence of a feathery covering is consistent with the idea that dromaeosaurids were warm-blooded. They also performed a cladistic analysis of the
1288:. Although a specific identification cannot be made, the shape of the preserved serration marks implicate a juvenile individual of one of the formation's tyrannosaurids, like
232:
to make red pigments. Since the red pigments are richest in the same layers of the formation that preserve dinosaur fossils, it is likely that Native
Americans encountered
945:. However, he proposed differing definitions for these subfamilies and argued that they had different members than were included under Barsbold's classification scheme.
3812:
Evans, DC; Larson, DW; Currie, PJ (2013). "A new dromaeosaurid (Dinosauria: Theropoda) with Asian affinities from the latest
Cretaceous of North America".
164:
led Ostrom to follow Edwin
Colbert and Dale Russel's suggestion that the Dromaeosaurinae be regarded as its own family separate from the Deinodontidae.
4736:
1358:-style true feathers would be expected due to the family's phylogenetic position. Also noted were the lack of known integumentary structures among
5658:
4042:"A large, short-armed, winged dromaeosaurid (Dinosauria: Theropoda) from the Early Cretaceous of China and its implications for feather evolution"
1007:
argued that dromaeosaurids lived in trees based on the anatomy of their hands and the backward orientation of the pubic bones in their pelvis.
120:, considering it a new type within the family Deinodontidae, a now-defunct family name that once applied to the tyrannosaurs. Not long after,
3951:
3942:
Horner, John R.; Weishampel, David B.; Forster, Catherine A. (2004). "Hadrosauridae". In
Weishampel, D. B.; Dodson, P.; Osmolska, H. (eds.).
221:
71:
452:
with its first detailed information about the anatomy of the dromaeosaurid body, as previous work focused on the skull. He proposed that
4639:
Xu, X.; Wang, X.-L. (2004). "A New
Dromaeosaur (Dinosauria: Theropoda) from the Early Cretaceous Yixian Formation of Western Liaoning".
783:
found strong support for Ostrom's hypothesis that dromaeosaurids were closely related to birds in "a pioneering and thorough analysis".
1306:. All of the marks on the jawbone seem to have been left by the same animal because the serration marks all share the same morphology.
4562:; Choinere, J.; Pittman, M.; Tan, Q.; Xiao, D.; Li, Z.; Tan, L.; Clark, J.; Norell, M.; Hone, D.W.E.; Sullivan, C. (19 March 2010).
4197:
4178:
4114:
3932:
4124:
Makovicky, Peter J.; ApesteguĂa, Sebastián; AgnolĂn, Federico L. (2005). "The earliest dromaeosaurid theropod from South
America".
1235:
5653:
4105:
Makovicky, Peter J.; Norell, Mark A. (2004). "Troodontidae". In
Weishampel, David B.; Dodson, Peter; OsmĂłlska, Halszka (eds.).
3857:
Gang Han; Luis M. Chiappe; Shu-An Ji; Michael Habib; Alan H. Turner; Anusuya
Chinsamy; Xueling Liu; Lizhuo Han (15 July 2014).
830:
5553:
4351:
Padian, K.; Ji, Qiang; Ji, Shu-An (2001). "Feathered dinosaurs and origin of flight". In Tanke, D. H.; Carpenter, K. (eds.).
2028:
131:
5542:
3961:
Jacobsen, A. R. (2001). "Tooth-marked small theropod bone: An extremely rare trace". In Tanke, D. H.; Carpenter, K. (eds.).
4408:"New Dromaeosaurids (Dinosauria: Theropoda) from the Lower Cretaceous of Utah, and the Evolution of the Dromaeosaurid Tail"
4729:
816:
Ostrom published further arguments building a case for a close evolutionary relationship between dromaeosaurids and birds.
766:
Ostrom published further arguments building a case for a close evolutionary relationship between dromaeosaurids and birds.
594:
Ostrom published further arguments building a case for a close evolutionary relationship between dromaeosaurids and birds.
566:
Ostrom published further arguments building a case for a close evolutionary relationship between dromaeosaurids and birds.
5559:
4188:
Norell, M.A. & Makovicky, P.J. (2004). "Dromaeosauridae". In Weishampel, D.B.; Dodson, P. & OsmĂłlska, H. (eds.).
171:, evidence continued to mount for a close evolutionary relationship between dromaeosaurids and birds. The dromaeosaurid
5548:
2372:
2072:
613:
4564:"A new dromaeosaurid (Dinosauria: Theropoda) from the Upper Cretaceous Wulansuhai Formation of Inner Mongolia, China"
3859:"A new raptorial dinosaur with exceptionally long feathering provides insights into dromaeosaurid flight performance"
3900:
Gong; Martin; Burnham; Falk; Hou (2012). "A new species of Microraptor from the Jehol Biota of northeastern China".
5599:
4708:
2141:
860:
as being of two animals killed and preserved while locked in mortal combat. She reinterpreted the association as a
5637:
2214:
2092:
1139:
486:
385:
358:
336:
317:
173:
116:
570:
127:
4833:
4722:
4654:"A short-armed dromaeosaurid from the Jehol Group of China with implications for early dromaeosaurid evolution"
4559:
2998:
2946:
2332:
2101:
1189:
267:
4475:
4220:
1087:
as preserving the aftermath of an ancient feeding frenzy rather than as evidence of pack hunting behavior in
2993:
1285:
1130:
633:
544:
509:
463:
444:
140:
75:
42:
189:. Discoveries of feathered dromaeosaurids continued into the 2000s. Xu, Zhou, and Wang named the new genus
2937:
2239:
2048:
843:
744:
652:
492:
380:
353:
107:
1105:
5663:
5586:
4460:
2917:
2312:
1325:
1198:
1052:
997:
671:
449:
330:
18:
748:. Unlike most dromaeosaurs, famous for the "killing claws" on their second toe, the second toe claw of
1029:
criticized for positioning the choana too far forward and for excluding the animals' secondary palate.
965:
Unwin and others followed the fighting-to-the-death interpretation of the fighting dinosaurs specimen.
850:
Osmolska challenged Kielan-Jaworowska and Barsbold's 1972 interpretation of an associated skeleton of
4887:
4603:
4521:
4419:
4362:"A new small deinonychosaur (Dinosauria: Theropoda) from the Late Cretaceous of Patagonia, Argentina"
4295:
4238:"A bizarre Cretaceous theropod dinosaur from Patagonia and the evolution of Gondwanan dromaeosaurids"
4133:
4053:
4000:
3870:
3821:
3795:
DePalma, Robert A.; Burnham, David A.; Martin, Larry D.; Larson, Peter L.; Bakker, Robert T. (2015).
3730:
3625:
2955:
2926:
2361:
2341:
2321:
2288:
2277:
2257:
2248:
2228:
2203:
2194:
2183:
2130:
2110:
2081:
2037:
2017:
1976:
1262:
1178:
1096:
1034:
983:
924:
891:
821:
807:
771:
757:
703:
680:
660:
622:
599:
585:
557:
533:
500:
430:
394:
371:
344:
299:
253:
178:
145:
104:
1271:
1074:
5625:
5304:
2426:
2390:
2352:
1321:
1214:
262:
157:
3667:, gen. et sp. nov. (Theropoda), from the upper Campanian Wapiti Formation, west-central Alberta".
5041:
4860:
4627:
4547:
4487:
4319:
4157:
3845:
3684:
3641:
2268:
2165:
2008:
1144:
229:
209:
186:
83:
4406:
Senter, P.; Kirkland, J. I.; Deblieux, D. D.; Madsen, S.; Toth, N. (2012). Dodson, Peter (ed.).
3786:
Czerkas, S. A.; Zhang, D.; Li, J; Li, Y (2002). "Flying dromaeosaurs". In Czerkas, S. J. (ed.).
3772:"A new dromaeosaurid from the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta, Canada"
1317:
1060:
1047:
1043:
5501:
5392:
5328:
5224:
5017:
4683:
4619:
4539:
4447:
4394:
4386:
4311:
4267:
4193:
4174:
4149:
4110:
4079:
4028:
3947:
3928:
3888:
3837:
3758:
2661:
2381:
1802:
1313:
1280:
1004:
576:
5613:
5425:
5371:
5231:
5209:
4673:
4665:
4611:
4529:
4479:
4471:
4437:
4427:
4376:
4303:
4257:
4249:
4224:
4216:
4141:
4069:
4061:
4018:
4008:
3911:
3878:
3829:
3748:
3738:
3705:
3676:
3633:
2576:
1998:
1958:
1350:
1156:
1121:
1066:
942:
736:
724:
694:
608:
513:
4236:
Novas, Fernando E.; Pol, Diego; Canale, Juan I.; Porfiri, Juan D.; Calvo, Jorge O. (2008).
5485:
5339:
5242:
4955:
4933:
4899:
4818:
4745:
4506:
4505:
Turner, Alan H.; Pol, Diego; Clarke, Julia A.; Erickson, Gregory M.; Norell, Mark (2007).
4208:
2976:
2972:
2766:
2630:
2412:
1895:
1248:
1209:
1152:
1148:
938:
934:
906:
hunted in packs to subdue prey larger than itself based on an association between several
732:
728:
363:
312:
153:
4207:
Norell, M.A.; Clark, J.M.; Turner, A.H.; Makovicky, P.J.; Barsbold, R.; Rowe, T. (2006).
3989:"A microraptorine (Dinosauria–Dromaeosauridae) from the Late Cretaceous of North America"
3972:"A new dromaeosaurid (Theropoda: Dromaeosauridae) from the Late Cretaceous of New Mexico"
152:
was obviously a vigorous, active animal, and exhibited characteristics linking it to the
4607:
4525:
4423:
4299:
4137:
4057:
4004:
3874:
3825:
3734:
3629:
416:
5143:
4678:
4653:
4442:
4407:
4262:
4237:
4074:
4041:
4023:
3988:
3753:
3718:
2121:
1776:
1602:
1576:
1296:
992:
480:
280:
50:
4588:
4280:
969:
646:
458:
killed its prey with the enlarged, sharply curved claws on the second toe of its feet.
5647:
5464:
5378:
5270:
5255:
5194:
5187:
5180:
5170:
5105:
5098:
5027:
3709:
3688:
3645:
2831:
2736:
2542:
2525:
2302:
2174:
1961:
1913:
1862:
1837:
1629:
915:
877:
716:
524:
286:
30:
4563:
4491:
4381:
4161:
3849:
1222:
1118:
based on the famous "fighting dinosaurs" specimen, since this was only one specimen.
5523:
5434:
5355:
5348:
5313:
5292:
5277:
5216:
5083:
5076:
5069:
5059:
5006:
4991:
4631:
4551:
4361:
4323:
4281:"New Evidence on Deinonychosaurian Dinosaurs from the Late Cretaceous of Patagonia"
3797:"The first giant raptor (Theropoda: Dromaeosauridae) from the Hell Creek Formation"
3719:"An aberrant island-dwelling theropod dinosaur from the Late Cretaceous of Romania"
2968:
2814:
2780:
2753:
2593:
2484:
2467:
2155:
2062:
1950:
1662:
1551:
1393:
1345:
1309:
1205:
1110:
1015:
852:
752:
was relatively small and not sharply curved. It is similar to those of troodontids.
518:
308:
122:
111:
66:
46:
3796:
3680:
1014:
to see whether or not they were consistent with the idea that dromaeosaurids were
4432:
3915:
156:. As such it brought support for controversial reinterpretations of dinosaurs as
96:
hundreds of years before these fossils came to the attention of formally trained
5478:
5456:
5407:
5363:
5320:
5285:
5263:
5202:
5113:
5048:
4872:
4331:
3902:
2712:
2644:
2559:
2330:
D. C. Evans, D. W. Larson, and P. J. Currie described the new genus and species
2172:
A. S. Turner, S. H. Hwang, and M. A. Norell described the new genus and species
1957:
feathers at the time of the study. Taxa marked with F were known to bear simple
1750:
1678:
1525:
1499:
1417:
1363:
1359:
1344:. However, they also noted, only filamentous protofeathers had been observed in
1290:
1254:
1241:
1079:
950:
902:
883:
467:, providing further details about the anatomy of its body. Ostrom proposed that
454:
439:
422:
234:
217:
191:
135:
92:
24:
3723:
Proceedings of the National Academy of Sciences of the United States of America
1992:
1163:
the sister group to both, and therefore the most primitive known dromaeosaurid.
958:
remains and argued that these provided further support for the hypothesis that
872:
792:
711:
5471:
5448:
5400:
5151:
5120:
4998:
4976:
4784:
4775:
3833:
3637:
2797:
2149:
2056:
1954:
1482:
1302:
1228:
1025:
1021:
911:
798:
97:
4587:
Xu, X.; Zhou, Z.; Wang, X.; Kuang, X.; Zhang, F.; Du, X. (January 23, 2003).
4390:
3781:. Life of the Past. Indianapolis: Indiana University Press. pp. 112–132.
5509:
5441:
5136:
5128:
5034:
4845:
4793:
4534:
4192:(2nd ed.). Berkeley: University of California Press. pp. 196–210.
4109:(2nd ed.). Berkeley: University of California Press. pp. 184–195.
4013:
3743:
3663:
Phil R. Bell & Philip J. Currie (2015). "A high-latitude dromaeosaurid,
2695:
2501:
2450:
1733:
975:
542:
Ostrom first discussed the apparent close evolutionary relationship between
404:
182:
79:
61:
4687:
4669:
4623:
4543:
4461:"A small derived theropod from Öösh, Early Cretaceous, Baykhangor Mongolia"
4451:
4398:
4315:
4271:
4253:
4153:
4083:
4032:
3892:
3841:
3777:. In Currie, P. J.; Koppelhus, E. B.; Shugar, M. A.; Wright, J. L. (eds.).
3762:
2296:
4703:
3946:(2 ed.). Berkeley: University of California Press. pp. 438–463.
3696:
Christiansen P. & Bonde N. (2003). "The first dinosaur from Denmark".
5493:
5414:
5385:
5159:
4969:
4769:
3771:
780:
689:
87:
58:
4615:
4307:
4145:
204:
5516:
4802:
4652:
Zheng, Xiaoting; Xu, Xing; You, Hailu; Zhao, Qi; Dong, Zhiming (2010).
4571:
3883:
3858:
2678:
1985:
Hwang and others performed a cladistic analysis of the Dromaeosauridae.
1275:
225:
4065:
2090:
Makovicky, ApesteguĂa and Agnolin described the new genus and species
181:
by Xu, Wang, and Wu, is a notable example as the fine-grained Chinese
5091:
4983:
4763:
4483:
4228:
2980:
2965:
1125:
838:
4209:"A new dromaeosaurid theropod from Ukhaa Tolgod (Ömnögov, Mongolia)"
3971:
1988:
Xu and others performed a cladistic analysis of the Dromaeosauridae.
1159:. The researchers found both groups to be monophyletic clades, with
4714:
2295:
2148:
2055:
1991:
1247:
1234:
1221:
1059:
968:
871:
834:
791:
710:
645:
569:
479:
415:
329:
279:
203:
17:
4507:"A basal dromaeosaurid and size evolution preceding avian flight"
3208:
3206:
3204:
3202:
3200:
1434:
1108:
and others argued that it was impossible to know whether or not
549:
54:
4756:
4718:
4355:. Life of the Past. Indiana University Press. pp. 117–135.
3225:
3223:
3221:
3198:
3196:
3194:
3192:
3190:
3188:
3186:
3184:
3182:
3180:
4360:
Porfiri, Juan D.; Jorge O. Calvo; Domenica dos Santos (2011).
3790:. Vol. 1. Blanding: The Dinosaur Museum. pp. 96–126.
3717:
Z., Csiki; Vremir, M.; Brusatte, S. L.; Norell, M. A. (2010).
3093:
3091:
3089:
3087:
3085:
3083:
3081:
3079:
3077:
2266:
Porfiri, Calvo and Santos described the new genus and species
1362:. The authors proposed that either science's understanding of
160:
and ancestral to birds. Its distinct nature and similarity to
4338:, an unusual theropod from the Lower Cretaceous of Montana".
3965:. Life of the Past. Indiana University Press. pp. 58–63.
3126:
407:
of dromaeosaurids were large for reptiles of their body size.
3167:
3165:
3163:
3161:
3159:
3157:
3155:
3124:
3122:
3120:
3118:
3116:
3114:
3112:
3110:
3108:
3106:
3036:
3034:
3032:
3030:
3028:
3698:
Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen
3055:; "History of Dinosaur Collecting in Montana," pages 53–54.
2026:
Christiansen and Bonde described the new genus and species
1332:
to the shared filaments and "rudimentary" true feathers in
238:
fossils long before scientifically trained paleontologists.
3314:
3312:
3310:
2046:
Currie and Varricchio described the new genus and species
4476:
10.1206/0003-0082(2007)3557[1:ASDTFS]2.0.CO;2
4221:
10.1206/0003-0082(2006)3545[1:andtfu]2.0.co;2
3976:
New Mexico Museum of Natural History and Science Bulletin
1374:
Cladogram of feathered dinosaurs from Padian et al. 2001
1077:
and others reinterpreted the association between several
90:, they may have encountered remains of the dromaeosaurid
2212:
Longrich and Currie described the new genus and species
144:. This discovery played a major role in setting off the
2006:
Czerkas and others described the new genus and species
1366:
evolution is faulty or feathers evolved multiple times.
1187:
Burnham and others described the new genus and species
3378:
2402:
Cladogram of dromaeosaurids from LĂĽ and Brusatte 2015
2310:
Senter and others described the new genus and species
2163:
Turner and others described the new genus and species
2119:
Norell and others described the new genus and species
1196:
Xu, Zhou and Wang described the new genus and species
1018:. However, the results of his study were inconclusive.
864:
that had been killed while scavenging an already dead
5597:
2246:
Zheng and others described the new genus and species
1143:. The specimen actually preserved the remains of the
1137:
Xu, Wang, and Wu described the new genus and species
315:
rocks that would later serve as the type specimen of
3015:
3013:
2915:
Bell and Currie described the new genus and species
2379:
LĂĽ and Brusatte described the new genus and species
1284:
with tooth marks. The specimen was preserved in the
1151:, finding more support for dividing the family into
1010:
Reid examined histological sections of the bones of
5424:
5338:
5303:
5241:
5169:
5058:
5016:
4953:
4898:
4871:
4844:
4817:
2944:Cau and others described the new genus and species
2388:DePalma et al. described the new genus and species
2350:Han and others described the new genus and species
2935:Xu and others described the new genus and species
2237:Xu and others described the new genus and species
2139:LĂĽ and others described the new genus and species
2099:Novas and Pol described the new genus and species
948:Maxwell and Ostrom reported on the association of
130:. Dromaeosaur research was fairly quiet until the
2192:Novas et al. described the new genus and species
4459:Turner, A.S.; Hwang, S.H.; Norell, M.A. (2007).
2070:Xu and Wang described the new genus and species
3993:Proceedings of the National Academy of Sciences
2983:) by Bolotskii, Bolotskii & Sorokin (2019).
995:and Puerta described the new genus and species
528:apparently killed and preserved while fighting.
4040:Junchang LĂĽ & Stephen L. Brusatte (2015).
3498:
1128:and Clark described the new genus and species
356:and Brown described the new genus and species
4730:
3534:
3438:
742:Barsbold described the new genus and species
8:
3788:Feathered Dinosaurs and the Origin of Flight
3342:
41:is a chronological listing of events in the
3022:; "Crow Fossil Collections," pages 272–273.
669:Jensen described the new genus and species
631:Powell described the new genus and species
5247:
4961:
4950:
4877:
4850:
4823:
4814:
4753:
4737:
4723:
4715:
4340:Peabody Museum of Natural History Bulletin
3354:
3229:
3212:
3174:; "Table 10.1: Dromaeosauridae", page 199.
3171:
3133:; "Table 10.1: Dromaeosauridae", page 198.
3130:
3097:
3040:
2398:
2286:Gong and others described the new species
1370:
403:Matthew and Brown first observed that the
185:from which it was collected preserved its
4677:
4533:
4441:
4431:
4380:
4261:
4073:
4022:
4012:
3882:
3770:Currie, P. J.; Varricchio, D. J. (2004).
3752:
3742:
3462:
4369:Anais da Academia Brasileira de CiĂŞncias
3379:Makovicky, Apesteguia and Agnolin (2005)
3232:; "Systematics and Evolution", page 207.
3100:; "Systematics and Evolution", page 206.
1949:Taxa marked with a P were known to bear
461:Ostrom further described the anatomy of
5604:
4279:Novas, Fernando E.; Pol, Diego (2005).
3009:
933:Currie followed Barsbold's division of
3582:
3330:
3318:
3301:
3289:
311:collected a partial skull and foot in
4171:Fossil Legends of the First Americans
3987:Longrich, N.R.; Currie, P.J. (2009).
3927:. Mountain Press Publishing Company.
3598:
3570:
3522:
3426:
3402:
516:reported the associated remains of a
484:The "Fighting Dinosaurs" specimen of
7:
3486:
3450:
1212:described the new genus and species
1050:described the new genus and species
841:described the new genus and species
692:described the new genus and species
611:described the new genus and species
442:described the new genus and species
383:described the new genus and species
138:described the new genus and species
3510:
3390:
2772:
2728:
2636:
2626:
2517:
2442:
2418:
2408:
2370:Jasinski described the new species
1887:
1725:
1718:
1711:
1655:
1475:
1468:
1461:
1454:
1447:
1440:
1430:
1423:
1413:
1406:
1399:
1389:
167:After Ostrom's initial research on
4658:Proceedings of the Royal Society B
4589:"Four-winged dinosaurs from China"
4242:Proceedings of the Royal Society B
3669:Journal of Vertebrate Paleontology
3558:
3546:
3277:
3265:
3253:
3241:
126:was discovered in Mongolia by the
39:timeline of dromaeosaurid research
14:
3474:
3414:
3366:
3142:
5631:
5619:
5607:
5582:
5581:
5558:
5552:
5547:
5541:
4702:
3535:Evans, Larson, and Currie (2013)
3499:Porfiri, Calvo and Santos (2011)
3439:Turner, Hwang, and Norell (2007)
3064:
3052:
910:skeletons and a specimen of the
448:. This description provided the
114:named the new genus and species
4382:10.1590/S0001-37652011000100007
3304:; "Conclusions," pages 131–132.
3019:
2975:theropod is described from the
1324:and their implications for the
471:only walked on two of its toes.
5659:Timelines of theropod research
4173:. Princeton University Press.
2029:Dromaeosauroides bornholmensis
1:
3801:Paleontological Contributions
3681:10.1080/02724634.2015.1034359
3343:Christiansen and Bonde (2003)
1274:published a description of a
4433:10.1371/journal.pone.0036790
4094:Geological Bulletin of China
3916:10.1016/j.palwor.2012.05.003
3355:Currie and Varricchio (2004)
1320:published a review of known
53:, a group of sickle-clawed,
3970:Steven E. Jasinski (2015).
3925:Dinosaurs Under the Big Sky
3230:Norell and Makovicky (2004)
3215:; "Paleoecology", page 209.
3213:Norell and Makovicky (2004)
3172:Norell and Makovicky (2004)
3131:Norell and Makovicky (2004)
3098:Norell and Makovicky (2004)
3043:; "Introduction", page 196.
3041:Norell and Makovicky (2004)
2373:Saurornitholestes sullivani
2073:Graciliraptor lujiatunensis
1239:Skeletal reconstruction of
1064:Feathered type specimen of
614:Saurornitholestes langstoni
284:Skeletal reconstruction of
228:used to use rocks from the
5680:
3710:10.1127/njgpa/227/2003/287
3463:Longrich and Currie (2009)
3321:; "Conclusions," page 132.
3256:; "Introduction," page 59.
2142:Luanchuanraptor henanensis
1226:Known skeletal remains of
881:may have been the prey of
731:into two subfamilies; the
265:described the new species
128:Central Asiatic Expedition
5571:
5539:
5250:
4964:
4949:
4930:
4880:
4853:
4826:
4813:
4752:
4468:American Museum Novitates
4213:American Museum Novitates
3834:10.1007/s00114-013-1107-5
3638:10.1134/S1028334X19010100
3319:Padian, Ji, and Ji (2001)
3302:Padian, Ji, and Ji (2001)
3290:Padian, Ji, and Ji (2001)
2828:
2811:
2794:
2777:
2770:
2750:
2733:
2726:
2709:
2692:
2675:
2658:
2641:
2634:
2624:
2607:
2590:
2573:
2556:
2539:
2522:
2515:
2498:
2481:
2464:
2447:
2440:
2423:
2416:
2215:Hesperonychus elizabethae
2093:Buitreraptor gonzalezorum
1910:
1892:
1885:
1859:
1835:
1799:
1774:
1747:
1730:
1723:
1716:
1709:
1676:
1660:
1653:
1626:
1599:
1574:
1548:
1522:
1496:
1480:
1473:
1466:
1459:
1452:
1445:
1438:
1428:
1421:
1411:
1404:
1397:
1140:Sinornithosaurus millenii
487:Velociraptor mongoliensis
386:Velociraptor mongoliensis
359:Dromaeosaurus albertensis
337:Velociraptor mongoliensis
318:Dromaeosaurus albertensis
187:life covering of feathers
174:Sinornithosaurus milennii
117:Dromaeosaurus albertensis
4353:Mesozoic Vertebrate Life
4169:Mayor, Adrienne (2005).
4052:: Article number 11775.
3963:Mesozoic Vertebrate Life
3923:Horner, John R. (2001).
3537:; "Abstract," page 1041.
3465:; "Abstract," page 5002.
3453:; "Abstract," page 1101.
3429:; "Abstract," page 1378.
3381:; "Abstract," page 1007.
3280:; "Discussion," page 60.
3268:; "Discussion," page 61.
2999:Timeline of paleontology
2947:Halszkaraptor escuilliei
2333:Acheroraptor temertyorum
2300:Artist's restoration of
2153:Artist's restoration of
2102:Neuquenraptor argentinus
1996:Artist's restoration of
1340:, to flight feathers in
1190:Bambiraptor feinbergorum
1161:Sinornithosaurus milenii
973:Artist's restoration of
650:Artistic restoration of
574:Artist's restoration of
420:Artistic restoration of
268:Ornithodesmus cluniculus
22:Artistic restoration of
4535:10.1126/science.1144066
4336:Deinonychus antirrhopus
4014:10.1073/pnas.0811664106
3744:10.1073/pnas.1006970107
3501:; "Abstract," page 109.
3489:; "Abstract," page 211.
3417:; "Abstract," page 777.
3393:; "Abstract," page 858.
3357:; "Abstract," page 112.
3345:; "Abstract," page 287.
3292:; "Abstract," page 117.
2994:History of paleontology
1286:Dinosaur Park Formation
1131:Achillobator giganticus
900:Ostrom speculated that
634:Unquillosaurus ceibalii
545:Deinonychus antirrhopus
510:Zofia Kielan-Jaworowska
464:Deinonychus antirrhopus
445:Deinonychus antirrhopus
141:Deinonychus antirrhopus
64:including animals like
5654:Paleontology timelines
4670:10.1098/rspb.2009.1178
4641:Vertebrata PalAsiatica
4334:(1969). "Osteology of
4254:10.1098/rspb.2008.1554
3618:Doklady Earth Sciences
3599:Bell and Currie (2015)
3571:LĂĽ and Brusatte (2015)
3561:; "Abstract", page 79.
3525:; "Abstract," page 58.
3369:; "Abstract," page 11.
3333:; "Abstract," page 96.
3244:; "Abstract," page 58.
2938:Zhongjianosaurus yangi
2912:
2306:
2240:Linheraptor exquisitus
2159:
2066:
2049:Atrociraptor marshalli
2002:
1973:
1258:
1245:
1232:
1070:
979:
887:
844:Utahraptor ostrommaysi
803:
745:Adasaurus mongoliensis
720:
656:
653:Adasaurus mongoliensis
581:
496:
493:Protoceratops andrewsi
426:
381:Henry Fairfield Osborn
340:
290:
212:
34:
3863:Nature Communications
3665:Boreonykus certekorum
3583:DePalma et al. (2015)
3573:; "Abstract", page 1.
3513:; "Abstract," page 1.
3477:; "Abstract," page 1.
3441:; "Abstract," page 1.
3405:; "Abstract," page 1.
3331:Czerkas et al. (2002)
2918:Boreonykus certekorum
2397:
2313:Yurgovuchia doellingi
2299:
2152:
2059:
1995:
1369:
1251:
1238:
1225:
1199:Microraptor zhaoianus
1063:
1053:Variraptor mechinorum
998:Unenlagia comahuensis
972:
875:
802:with a human to scale
795:
714:
672:Palaeopteryx thomsoni
649:
573:
483:
450:scientific literature
419:
333:
283:
207:
21:
4711:at Wikimedia Commons
3523:Senter et al. (2012)
3427:Turner et al. (2007)
3403:Norell et al. (2006)
3391:Novas and Pol (2005)
3067:; "6. Affinities of
2289:Microraptor hanqingi
2249:Tianyuraptor ostromi
2195:Austroraptor cabazai
1383:Feathered Dinosaurs
1091:as argued by Ostrom.
146:Dinosaur Renaissance
5305:Saurornitholestinae
4616:10.1038/nature01342
4608:2003Natur.421..335X
4526:2007Sci...317.1378T
4520:(5843): 1378–1381.
4424:2012PLoSO...736790S
4308:10.1038/nature03285
4300:2005Natur.433..858N
4248:(1659): 1101–1107.
4146:10.1038/nature03996
4138:2005Natur.437.1007M
4132:(7061): 1007–1011.
4058:2015NatSR...511775L
4005:2009PNAS..106.5002L
3875:2014NatCo...5.4382H
3826:2013NW....100.1041E
3814:Naturwissenschaften
3735:2010PNAS..10715357C
3729:(35): 15357–15361.
3630:2019DokES.484...18B
3487:Zheng et al. (2010)
3451:Novas et al. (2008)
2391:Dakotaraptor steini
2353:Changyuraptor yangi
1322:feathered dinosaurs
1215:Pyroraptor olympius
580:feeding on a mammal
263:Harry Govier Seeley
5638:History of science
4046:Scientific Reports
3884:10.1038/ncomms5382
3511:Gong et al. (2012)
3367:Xu and Wang (2004)
2307:
2269:Pamparaptor micros
2166:Mahakala omnogovae
2160:
2067:
2060:Skeletal mount of
2009:Cryptovolans pauli
2003:
1259:
1246:
1233:
1114:was a predator of
1071:
980:
962:was a pack hunter.
888:
804:
796:Illustration of a
721:
657:
582:
497:
427:
341:
334:The type skull of
291:
230:Cloverly Formation
224:groups inhabiting
213:
210:Cloverly Formation
208:An outcrop of the
84:Cloverly Formation
35:
5595:
5594:
5567:
5566:
5537:
5536:
5533:
5532:
5502:Saurornitholestes
5393:Saurornitholestes
5329:Saurornitholestes
5018:Halszkaraptorinae
4945:
4944:
4941:
4940:
4926:
4925:
4922:
4921:
4918:
4917:
4707:Media related to
4664:(1679): 211–217.
4602:(6921): 335–340.
4294:(7028): 858–861.
4066:10.1038/srep11775
3999:(13): 5002–5007.
3953:978-0-520-25408-4
3820:(11): 1041–1049.
3779:Feathered Dragons
3547:Han et al. (2014)
3071:," pages 147–148.
2911:
2910:
2906:
2905:
2897:
2896:
2888:
2887:
2879:
2878:
2870:
2869:
2861:
2860:
2852:
2851:
2843:
2842:
2662:Saurornitholestes
2382:Zhenyuanlong suni
1972:
1971:
1968:
1967:
1943:
1942:
1934:
1933:
1925:
1924:
1874:
1873:
1848:
1847:
1824:
1823:
1815:
1814:
1803:Protarchaeopteryx
1788:
1787:
1763:
1762:
1698:
1697:
1689:
1688:
1642:
1641:
1615:
1614:
1588:
1587:
1563:
1562:
1537:
1536:
1511:
1510:
1338:Protarchaeopteryx
1281:Saurornitholestes
1012:Saurornitholestes
577:Saurornitholestes
5671:
5636:
5635:
5634:
5624:
5623:
5622:
5612:
5611:
5610:
5603:
5585:
5584:
5562:
5556:
5551:
5545:
5426:Velociraptorinae
5372:Dromaeosauroides
5248:
5232:Zhongjianosaurus
5210:Sinornithosaurus
4962:
4951:
4878:
4851:
4824:
4815:
4808:
4807:
4754:
4739:
4732:
4725:
4716:
4706:
4691:
4681:
4648:
4635:
4593:
4583:
4581:
4580:
4568:
4555:
4537:
4511:
4501:
4499:
4498:
4465:
4455:
4445:
4435:
4402:
4384:
4366:
4356:
4347:
4327:
4285:
4275:
4265:
4232:
4203:
4184:
4165:
4120:
4101:
4087:
4077:
4036:
4026:
4016:
3983:
3966:
3957:
3938:
3919:
3896:
3886:
3853:
3808:
3791:
3782:
3776:
3766:
3756:
3746:
3713:
3692:
3650:
3649:
3612:
3606:
3596:
3590:
3580:
3574:
3568:
3562:
3556:
3550:
3544:
3538:
3532:
3526:
3520:
3514:
3508:
3502:
3496:
3490:
3484:
3478:
3475:Xu et al. (2010)
3472:
3466:
3460:
3454:
3448:
3442:
3436:
3430:
3424:
3418:
3415:LĂĽ et al. (2007)
3412:
3406:
3400:
3394:
3388:
3382:
3376:
3370:
3364:
3358:
3352:
3346:
3340:
3334:
3328:
3322:
3316:
3305:
3299:
3293:
3287:
3281:
3275:
3269:
3263:
3257:
3251:
3245:
3239:
3233:
3227:
3216:
3210:
3175:
3169:
3150:
3143:Xu et al. (2003)
3140:
3134:
3128:
3101:
3095:
3072:
3062:
3056:
3050:
3044:
3038:
3023:
3017:
2958:
2929:
2773:
2729:
2637:
2627:
2577:Sinornithosaurus
2518:
2443:
2419:
2409:
2399:
2364:
2344:
2324:
2280:
2260:
2231:
2206:
2186:
2133:
2113:
2084:
2040:
2020:
1999:Dromaeosauroides
1979:
1888:
1726:
1719:
1712:
1656:
1476:
1469:
1462:
1455:
1448:
1441:
1431:
1424:
1414:
1407:
1400:
1390:
1380:
1379:
1371:
1351:Sinornithosaurus
1326:origin of flight
1265:
1181:
1157:Velociraptorinae
1122:Altangerel Perle
1099:
1083:specimens and a
1067:Sinornithosaurus
1037:
986:
943:Velociraptorinae
927:
894:
824:
810:
774:
760:
737:Velociraptorinae
725:Rinchen Barsbold
706:
695:Hulsanpes perlei
683:
663:
625:
602:
588:
560:
536:
514:Rinchen Barsbold
503:
433:
397:
374:
347:
302:
256:
72:Native Americans
5679:
5678:
5674:
5673:
5672:
5670:
5669:
5668:
5644:
5643:
5642:
5632:
5630:
5620:
5618:
5608:
5606:
5598:
5596:
5591:
5563:
5529:
5486:Luanchuanraptor
5420:
5340:Dromaeosaurinae
5334:
5299:
5243:Eudromaeosauria
5237:
5165:
5054:
5012:
4958:
4956:Dromaeosauridae
4937:
4934:Dromaeosauridae
4914:
4900:Dromaeosauridae
4894:
4867:
4840:
4834:Avemetatarsalia
4819:Avemetatarsalia
4809:
4759:
4758:
4748:
4746:Dromaeosauridae
4743:
4709:Dromaeosauridae
4699:
4694:
4651:
4638:
4591:
4586:
4578:
4576:
4566:
4558:
4509:
4504:
4496:
4494:
4463:
4458:
4405:
4364:
4359:
4350:
4332:Ostrom, John H.
4330:
4283:
4278:
4235:
4206:
4200:
4187:
4181:
4168:
4123:
4117:
4104:
4090:
4039:
3986:
3969:
3960:
3954:
3941:
3935:
3922:
3899:
3856:
3811:
3794:
3785:
3774:
3769:
3716:
3695:
3662:
3658:
3653:
3614:
3613:
3609:
3597:
3593:
3581:
3577:
3569:
3565:
3559:Jasinski (2015)
3557:
3553:
3545:
3541:
3533:
3529:
3521:
3517:
3509:
3505:
3497:
3493:
3485:
3481:
3473:
3469:
3461:
3457:
3449:
3445:
3437:
3433:
3425:
3421:
3413:
3409:
3401:
3397:
3389:
3385:
3377:
3373:
3365:
3361:
3353:
3349:
3341:
3337:
3329:
3325:
3317:
3308:
3300:
3296:
3288:
3284:
3278:Jacobsen (2001)
3276:
3272:
3266:Jacobsen (2001)
3264:
3260:
3254:Jacobsen (2001)
3252:
3248:
3242:Jacobsen (2001)
3240:
3236:
3228:
3219:
3211:
3178:
3170:
3153:
3141:
3137:
3129:
3104:
3096:
3075:
3063:
3059:
3051:
3047:
3039:
3026:
3018:
3011:
3007:
2990:
2977:Blagoveshchensk
2956:
2927:
2907:
2898:
2889:
2880:
2871:
2862:
2853:
2844:
2767:Dromaeosaurinae
2631:Eudromaeosauria
2413:Dromaeosauridae
2362:
2342:
2322:
2278:
2258:
2229:
2225:
2204:
2184:
2131:
2111:
2082:
2038:
2018:
1977:
1944:
1935:
1926:
1896:Sinosauropteryx
1875:
1849:
1825:
1816:
1789:
1764:
1699:
1690:
1643:
1616:
1589:
1564:
1538:
1512:
1330:Sinosauropteryx
1263:
1179:
1175:
1170:
1153:Dromaeosaurinae
1149:Dromaeosauridae
1097:
1035:
984:
939:Dromaeosaurinae
935:Dromaeosauridae
925:
892:
822:
808:
790:
772:
758:
750:A. mongoliensis
733:Dromaeosaurinae
729:Dromaeosauridae
704:
681:
661:
644:
623:
600:
586:
558:
534:
501:
478:
431:
414:
395:
372:
364:Dromaeosauridae
345:
328:
313:Late Cretaceous
300:
296:
278:
254:
250:
245:
222:Native American
202:
177:, described in
154:origin of birds
49:focused on the
12:
11:
5:
5677:
5675:
5667:
5666:
5661:
5656:
5646:
5645:
5641:
5640:
5628:
5616:
5593:
5592:
5590:
5589:
5579:
5572:
5569:
5568:
5565:
5564:
5540:
5538:
5535:
5534:
5531:
5530:
5528:
5527:
5520:
5513:
5506:
5498:
5490:
5482:
5475:
5468:
5461:
5453:
5445:
5438:
5430:
5428:
5422:
5421:
5419:
5418:
5411:
5404:
5397:
5389:
5382:
5375:
5368:
5360:
5352:
5344:
5342:
5336:
5335:
5333:
5332:
5325:
5317:
5309:
5307:
5301:
5300:
5298:
5297:
5289:
5282:
5274:
5267:
5260:
5251:
5245:
5239:
5238:
5236:
5235:
5228:
5221:
5213:
5206:
5199:
5191:
5184:
5176:
5174:
5167:
5166:
5164:
5163:
5156:
5148:
5144:Unquillosaurus
5140:
5133:
5125:
5117:
5110:
5102:
5095:
5088:
5080:
5073:
5065:
5063:
5056:
5055:
5053:
5052:
5045:
5038:
5031:
5023:
5021:
5014:
5013:
5011:
5010:
5003:
4995:
4988:
4980:
4973:
4965:
4959:
4954:
4947:
4946:
4943:
4942:
4939:
4938:
4931:
4928:
4927:
4924:
4923:
4920:
4919:
4916:
4915:
4913:
4912:
4911:
4910:
4904:
4902:
4896:
4895:
4893:
4892:
4891:
4890:
4881:
4875:
4869:
4868:
4866:
4865:
4864:
4863:
4854:
4848:
4842:
4841:
4839:
4838:
4837:
4836:
4827:
4821:
4811:
4810:
4806:
4805:
4796:
4787:
4778:
4772:
4766:
4757:
4750:
4749:
4744:
4742:
4741:
4734:
4727:
4719:
4713:
4712:
4698:
4697:External links
4695:
4693:
4692:
4649:
4636:
4584:
4556:
4502:
4470:(3557): 1–27.
4456:
4403:
4375:(1): 109–116.
4357:
4348:
4328:
4276:
4233:
4215:(3545): 1–51.
4204:
4198:
4190:The Dinosauria
4185:
4179:
4166:
4121:
4115:
4107:The Dinosauria
4102:
4088:
4037:
3984:
3967:
3958:
3952:
3944:The Dinosauria
3939:
3933:
3920:
3897:
3854:
3809:
3792:
3783:
3767:
3714:
3704:(2): 287–299.
3693:
3659:
3657:
3654:
3652:
3651:
3607:
3591:
3575:
3563:
3551:
3539:
3527:
3515:
3503:
3491:
3479:
3467:
3455:
3443:
3431:
3419:
3407:
3395:
3383:
3371:
3359:
3347:
3335:
3323:
3306:
3294:
3282:
3270:
3258:
3246:
3234:
3217:
3176:
3151:
3149:pages 335–340.
3135:
3102:
3073:
3057:
3045:
3024:
3008:
3006:
3003:
3002:
3001:
2996:
2989:
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2877:
2876:
2873:
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2868:
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2864:
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2859:
2858:
2855:
2854:
2850:
2849:
2846:
2845:
2841:
2840:
2837:
2836:
2827:
2824:
2823:
2820:
2819:
2810:
2807:
2806:
2803:
2802:
2793:
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2776:
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2763:
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2759:
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2727:
2725:
2722:
2721:
2718:
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2708:
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2704:
2701:
2700:
2691:
2688:
2687:
2684:
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2674:
2671:
2670:
2667:
2666:
2657:
2654:
2653:
2650:
2649:
2640:
2635:
2633:
2625:
2623:
2620:
2619:
2616:
2615:
2606:
2603:
2602:
2599:
2598:
2589:
2586:
2585:
2582:
2581:
2572:
2569:
2568:
2565:
2564:
2555:
2552:
2551:
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2547:
2538:
2535:
2534:
2531:
2530:
2521:
2516:
2514:
2511:
2510:
2507:
2506:
2497:
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2477:
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2473:
2472:
2463:
2460:
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2422:
2417:
2415:
2407:
2404:
2403:
2396:
2395:
2386:
2377:
2358:
2357:
2338:
2337:
2318:
2317:
2294:
2293:
2274:
2273:
2254:
2253:
2244:
2224:
2221:
2220:
2219:
2200:
2199:
2180:
2179:
2170:
2147:
2146:
2127:
2126:
2122:Tsaagan mangas
2107:
2106:
2097:
2078:
2077:
2054:
2053:
2034:
2033:
2014:
2013:
1990:
1989:
1986:
1970:
1969:
1966:
1965:
1946:
1945:
1941:
1940:
1937:
1936:
1932:
1931:
1928:
1927:
1923:
1922:
1919:
1918:
1909:
1906:
1905:
1902:
1901:
1891:
1886:
1884:
1881:
1880:
1877:
1876:
1872:
1871:
1868:
1867:
1858:
1855:
1854:
1851:
1850:
1846:
1845:
1842:
1841:
1834:
1831:
1830:
1827:
1826:
1822:
1821:
1818:
1817:
1813:
1812:
1809:
1808:
1798:
1795:
1794:
1791:
1790:
1786:
1785:
1782:
1781:
1777:Therizinosaurs
1773:
1770:
1769:
1766:
1765:
1761:
1760:
1757:
1756:
1746:
1743:
1742:
1739:
1738:
1729:
1724:
1722:
1717:
1715:
1710:
1708:
1705:
1704:
1701:
1700:
1696:
1695:
1692:
1691:
1687:
1686:
1683:
1682:
1675:
1672:
1671:
1668:
1667:
1659:
1654:
1652:
1649:
1648:
1645:
1644:
1640:
1639:
1636:
1635:
1625:
1622:
1621:
1618:
1617:
1613:
1612:
1609:
1608:
1603:Confuciusornis
1598:
1595:
1594:
1591:
1590:
1586:
1585:
1582:
1581:
1577:Enantiornithes
1573:
1570:
1569:
1566:
1565:
1561:
1560:
1557:
1556:
1547:
1544:
1543:
1540:
1539:
1535:
1534:
1531:
1530:
1521:
1518:
1517:
1514:
1513:
1509:
1508:
1505:
1504:
1495:
1492:
1491:
1488:
1487:
1479:
1474:
1472:
1467:
1465:
1460:
1458:
1453:
1451:
1446:
1444:
1439:
1437:
1429:
1427:
1422:
1420:
1412:
1410:
1405:
1403:
1398:
1396:
1388:
1385:
1384:
1376:
1375:
1368:
1367:
1307:
1297:Daspletosaurus
1272:A. R. Jacobsen
1220:
1219:
1203:
1194:
1174:
1171:
1169:
1166:
1165:
1164:
1135:
1119:
1093:
1092:
1075:D. L. Brinkman
1058:
1057:
1031:
1030:
1019:
1008:
1002:
967:
966:
963:
946:
921:
920:
870:
869:
848:
818:
817:
789:
786:
785:
784:
768:
767:
754:
753:
740:
700:
699:
677:
676:
643:
640:
639:
638:
619:
618:
596:
595:
568:
567:
554:
553:
530:
529:
477:
474:
473:
472:
459:
413:
410:
409:
408:
391:
390:
368:
367:
362:and named the
327:
324:
323:
322:
295:
292:
277:
274:
273:
272:
249:
246:
244:
241:
240:
239:
201:
198:
51:dromaeosaurids
13:
10:
9:
6:
4:
3:
2:
5676:
5665:
5662:
5660:
5657:
5655:
5652:
5651:
5649:
5639:
5629:
5627:
5617:
5615:
5605:
5601:
5588:
5580:
5578:
5574:
5573:
5570:
5561:
5557:
5555:
5550:
5544:
5526:
5525:
5521:
5519:
5518:
5514:
5512:
5511:
5507:
5504:
5503:
5499:
5496:
5495:
5491:
5488:
5487:
5483:
5481:
5480:
5476:
5474:
5473:
5469:
5467:
5466:
5465:Kansaignathus
5462:
5459:
5458:
5454:
5451:
5450:
5446:
5444:
5443:
5439:
5437:
5436:
5432:
5431:
5429:
5427:
5423:
5417:
5416:
5412:
5410:
5409:
5405:
5403:
5402:
5398:
5395:
5394:
5390:
5388:
5387:
5383:
5381:
5380:
5379:Dromaeosaurus
5376:
5374:
5373:
5369:
5366:
5365:
5361:
5358:
5357:
5353:
5351:
5350:
5346:
5345:
5343:
5341:
5337:
5331:
5330:
5326:
5323:
5322:
5318:
5316:
5315:
5311:
5310:
5308:
5306:
5302:
5295:
5294:
5290:
5288:
5287:
5283:
5280:
5279:
5275:
5273:
5272:
5271:Dineobellator
5268:
5266:
5265:
5261:
5258:
5257:
5253:
5252:
5249:
5246:
5244:
5240:
5234:
5233:
5229:
5227:
5226:
5222:
5219:
5218:
5214:
5212:
5211:
5207:
5205:
5204:
5200:
5197:
5196:
5195:Hesperonychus
5192:
5190:
5189:
5188:Graciliraptor
5185:
5183:
5182:
5181:Changyuraptor
5178:
5177:
5175:
5172:
5171:Microraptoria
5168:
5162:
5161:
5157:
5154:
5153:
5149:
5146:
5145:
5141:
5139:
5138:
5134:
5131:
5130:
5126:
5123:
5122:
5118:
5116:
5115:
5111:
5108:
5107:
5106:Ornithodesmus
5103:
5101:
5100:
5099:Neuquenraptor
5096:
5094:
5093:
5089:
5086:
5085:
5081:
5079:
5078:
5074:
5072:
5071:
5067:
5066:
5064:
5061:
5057:
5051:
5050:
5046:
5044:
5043:
5039:
5037:
5036:
5032:
5030:
5029:
5028:Halszkaraptor
5025:
5024:
5022:
5019:
5015:
5009:
5008:
5004:
5001:
5000:
4996:
4994:
4993:
4989:
4986:
4985:
4981:
4979:
4978:
4974:
4972:
4971:
4967:
4966:
4963:
4960:
4957:
4952:
4948:
4936:
4935:
4929:
4908:
4907:
4906:
4905:
4903:
4901:
4897:
4889:
4885:
4884:
4883:
4882:
4879:
4876:
4874:
4870:
4862:
4858:
4857:
4856:
4855:
4852:
4849:
4847:
4843:
4835:
4831:
4830:
4829:
4828:
4825:
4822:
4820:
4816:
4812:
4804:
4800:
4797:
4795:
4791:
4788:
4786:
4782:
4779:
4777:
4773:
4771:
4767:
4765:
4761:
4760:
4755:
4751:
4747:
4740:
4735:
4733:
4728:
4726:
4721:
4720:
4717:
4710:
4705:
4701:
4700:
4696:
4689:
4685:
4680:
4675:
4671:
4667:
4663:
4659:
4655:
4650:
4646:
4642:
4637:
4633:
4629:
4625:
4621:
4617:
4613:
4609:
4605:
4601:
4597:
4590:
4585:
4574:
4573:
4565:
4561:
4557:
4553:
4549:
4545:
4541:
4536:
4531:
4527:
4523:
4519:
4515:
4508:
4503:
4493:
4489:
4485:
4481:
4477:
4473:
4469:
4462:
4457:
4453:
4449:
4444:
4439:
4434:
4429:
4425:
4421:
4418:(5): e36790.
4417:
4413:
4409:
4404:
4400:
4396:
4392:
4388:
4383:
4378:
4374:
4370:
4363:
4358:
4354:
4349:
4345:
4341:
4337:
4333:
4329:
4325:
4321:
4317:
4313:
4309:
4305:
4301:
4297:
4293:
4289:
4282:
4277:
4273:
4269:
4264:
4259:
4255:
4251:
4247:
4243:
4239:
4234:
4230:
4226:
4222:
4218:
4214:
4210:
4205:
4201:
4199:0-520-24209-2
4195:
4191:
4186:
4182:
4180:0-691-11345-9
4176:
4172:
4167:
4163:
4159:
4155:
4151:
4147:
4143:
4139:
4135:
4131:
4127:
4122:
4118:
4116:0-520-24209-2
4112:
4108:
4103:
4100:(7): 777–786.
4099:
4095:
4089:
4085:
4081:
4076:
4071:
4067:
4063:
4059:
4055:
4051:
4047:
4043:
4038:
4034:
4030:
4025:
4020:
4015:
4010:
4006:
4002:
3998:
3994:
3990:
3985:
3981:
3977:
3973:
3968:
3964:
3959:
3955:
3949:
3945:
3940:
3936:
3934:0-87842-445-8
3930:
3926:
3921:
3917:
3913:
3909:
3905:
3904:
3898:
3894:
3890:
3885:
3880:
3876:
3872:
3868:
3864:
3860:
3855:
3851:
3847:
3843:
3839:
3835:
3831:
3827:
3823:
3819:
3815:
3810:
3806:
3802:
3798:
3793:
3789:
3784:
3780:
3773:
3768:
3764:
3760:
3755:
3750:
3745:
3740:
3736:
3732:
3728:
3724:
3720:
3715:
3711:
3707:
3703:
3699:
3694:
3690:
3686:
3682:
3678:
3674:
3670:
3666:
3661:
3660:
3655:
3647:
3643:
3639:
3635:
3631:
3627:
3623:
3619:
3611:
3608:
3604:
3600:
3595:
3592:
3588:
3584:
3579:
3576:
3572:
3567:
3564:
3560:
3555:
3552:
3549:; "Abstract".
3548:
3543:
3540:
3536:
3531:
3528:
3524:
3519:
3516:
3512:
3507:
3504:
3500:
3495:
3492:
3488:
3483:
3480:
3476:
3471:
3468:
3464:
3459:
3456:
3452:
3447:
3444:
3440:
3435:
3432:
3428:
3423:
3420:
3416:
3411:
3408:
3404:
3399:
3396:
3392:
3387:
3384:
3380:
3375:
3372:
3368:
3363:
3360:
3356:
3351:
3348:
3344:
3339:
3336:
3332:
3327:
3324:
3320:
3315:
3313:
3311:
3307:
3303:
3298:
3295:
3291:
3286:
3283:
3279:
3274:
3271:
3267:
3262:
3259:
3255:
3250:
3247:
3243:
3238:
3235:
3231:
3226:
3224:
3222:
3218:
3214:
3209:
3207:
3205:
3203:
3201:
3199:
3197:
3195:
3193:
3191:
3189:
3187:
3185:
3183:
3181:
3177:
3173:
3168:
3166:
3164:
3162:
3160:
3158:
3156:
3152:
3148:
3144:
3139:
3136:
3132:
3127:
3125:
3123:
3121:
3119:
3117:
3115:
3113:
3111:
3109:
3107:
3103:
3099:
3094:
3092:
3090:
3088:
3086:
3084:
3082:
3080:
3078:
3074:
3070:
3066:
3065:Ostrom (1969)
3061:
3058:
3054:
3053:Horner (2001)
3049:
3046:
3042:
3037:
3035:
3033:
3031:
3029:
3025:
3021:
3016:
3014:
3010:
3004:
3000:
2997:
2995:
2992:
2991:
2987:
2982:
2978:
2974:
2973:dromaeosaurid
2970:
2967:
2963:
2962:
2961:
2960:
2959:
2949:
2948:
2943:
2940:
2939:
2934:
2933:
2932:
2931:
2930:
2920:
2919:
2914:
2913:
2902:
2901:
2893:
2892:
2884:
2883:
2875:
2874:
2866:
2865:
2857:
2856:
2848:
2847:
2839:
2838:
2835:
2834:
2833:
2832:Dromaeosaurus
2826:
2825:
2822:
2821:
2818:
2817:
2816:
2809:
2808:
2805:
2804:
2801:
2800:
2799:
2792:
2791:
2788:
2787:
2784:
2783:
2782:
2775:
2774:
2768:
2765:
2764:
2761:
2760:
2757:
2756:
2755:
2748:
2747:
2744:
2743:
2740:
2739:
2738:
2731:
2730:
2724:
2723:
2720:
2719:
2716:
2715:
2714:
2707:
2706:
2703:
2702:
2699:
2698:
2697:
2690:
2689:
2686:
2685:
2682:
2681:
2680:
2673:
2672:
2669:
2668:
2665:
2664:
2663:
2656:
2655:
2652:
2651:
2648:
2647:
2646:
2639:
2638:
2632:
2629:
2628:
2622:
2621:
2618:
2617:
2614:
2613:
2612:
2605:
2604:
2601:
2600:
2597:
2596:
2595:
2588:
2587:
2584:
2583:
2580:
2579:
2578:
2571:
2570:
2567:
2566:
2563:
2562:
2561:
2554:
2553:
2550:
2549:
2546:
2545:
2544:
2543:Changyuraptor
2537:
2536:
2533:
2532:
2529:
2528:
2527:
2526:Graciliraptor
2520:
2519:
2513:
2512:
2509:
2508:
2505:
2504:
2503:
2496:
2495:
2492:
2491:
2488:
2487:
2486:
2479:
2478:
2475:
2474:
2471:
2470:
2469:
2462:
2461:
2458:
2457:
2454:
2453:
2452:
2445:
2444:
2438:
2437:
2434:
2433:
2430:
2429:
2428:
2421:
2420:
2414:
2411:
2410:
2406:
2405:
2401:
2400:
2393:
2392:
2387:
2384:
2383:
2378:
2375:
2374:
2369:
2368:
2367:
2366:
2365:
2355:
2354:
2349:
2348:
2347:
2346:
2345:
2335:
2334:
2329:
2328:
2327:
2326:
2325:
2315:
2314:
2309:
2308:
2305:
2304:
2303:Changyuraptor
2298:
2291:
2290:
2285:
2284:
2283:
2282:
2281:
2271:
2270:
2265:
2264:
2263:
2262:
2261:
2251:
2250:
2245:
2242:
2241:
2236:
2235:
2234:
2233:
2232:
2222:
2217:
2216:
2211:
2210:
2209:
2208:
2207:
2197:
2196:
2191:
2190:
2189:
2188:
2187:
2177:
2176:
2175:Shanag ashile
2171:
2168:
2167:
2162:
2161:
2158:
2157:
2151:
2144:
2143:
2138:
2137:
2136:
2135:
2134:
2124:
2123:
2118:
2117:
2116:
2115:
2114:
2104:
2103:
2098:
2095:
2094:
2089:
2088:
2087:
2086:
2085:
2075:
2074:
2069:
2068:
2065:
2064:
2058:
2051:
2050:
2045:
2044:
2043:
2042:
2041:
2031:
2030:
2025:
2024:
2023:
2022:
2021:
2011:
2010:
2005:
2004:
2001:
2000:
1994:
1987:
1984:
1983:
1982:
1981:
1980:
1963:
1962:integumentary
1960:
1956:
1952:
1948:
1947:
1939:
1938:
1930:
1929:
1921:
1920:
1917:
1916:
1915:
1914:Compsognathus
1908:
1907:
1904:
1903:
1900:
1898:
1897:
1890:
1889:
1883:
1882:
1879:
1878:
1870:
1869:
1866:
1865:
1864:
1863:Tyrannosaurus
1857:
1856:
1853:
1852:
1844:
1843:
1840:
1839:
1838:Ornithomimids
1833:
1832:
1829:
1828:
1820:
1819:
1811:
1810:
1807:
1805:
1804:
1797:
1796:
1793:
1792:
1784:
1783:
1780:
1778:
1772:
1771:
1768:
1767:
1759:
1758:
1755:
1753:
1752:
1745:
1744:
1741:
1740:
1737:
1736:
1735:
1728:
1727:
1721:
1720:
1714:
1713:
1707:
1706:
1703:
1702:
1694:
1693:
1685:
1684:
1681:
1680:
1674:
1673:
1670:
1669:
1666:
1664:
1658:
1657:
1651:
1650:
1647:
1646:
1638:
1637:
1634:
1632:
1631:
1630:Archaeopteryx
1624:
1623:
1620:
1619:
1611:
1610:
1607:
1605:
1604:
1597:
1596:
1593:
1592:
1584:
1583:
1580:
1578:
1572:
1571:
1568:
1567:
1559:
1558:
1555:
1554:
1553:
1546:
1545:
1542:
1541:
1533:
1532:
1529:
1528:
1527:
1520:
1519:
1516:
1515:
1507:
1506:
1503:
1502:
1501:
1494:
1493:
1490:
1489:
1486:
1484:
1478:
1477:
1471:
1470:
1464:
1463:
1457:
1456:
1450:
1449:
1443:
1442:
1436:
1433:
1432:
1426:
1425:
1419:
1416:
1415:
1409:
1408:
1402:
1401:
1395:
1392:
1391:
1387:
1386:
1382:
1381:
1378:
1377:
1373:
1372:
1365:
1361:
1357:
1353:
1352:
1347:
1343:
1342:Archaeopteryx
1339:
1335:
1331:
1327:
1323:
1319:
1315:
1311:
1308:
1305:
1304:
1299:
1298:
1293:
1292:
1287:
1283:
1282:
1277:
1273:
1270:
1269:
1268:
1267:
1266:
1257:
1256:
1250:
1244:
1243:
1237:
1231:
1230:
1224:
1217:
1216:
1211:
1207:
1204:
1201:
1200:
1195:
1192:
1191:
1186:
1185:
1184:
1183:
1182:
1172:
1167:
1162:
1158:
1154:
1150:
1146:
1142:
1141:
1136:
1133:
1132:
1127:
1123:
1120:
1117:
1116:Protoceratops
1113:
1112:
1107:
1104:
1103:
1102:
1101:
1100:
1090:
1086:
1085:Tenontosaurus
1082:
1081:
1076:
1073:
1072:
1069:
1068:
1062:
1055:
1054:
1049:
1045:
1042:
1041:
1040:
1039:
1038:
1027:
1023:
1020:
1017:
1013:
1009:
1006:
1003:
1000:
999:
994:
991:
990:
989:
988:
987:
978:
977:
971:
964:
961:
957:
956:Tenontosaurus
953:
952:
947:
944:
940:
936:
932:
931:
930:
929:
928:
918:
917:
916:Tenontosaurus
913:
909:
905:
904:
899:
898:
897:
896:
895:
886:
885:
880:
879:
878:Tenontosaurus
874:
867:
866:Protoceratops
863:
859:
858:Protoceratops
855:
854:
849:
846:
845:
840:
836:
832:
829:
828:
827:
826:
825:
815:
814:
813:
812:
811:
801:
800:
794:
787:
782:
779:
778:
777:
776:
775:
765:
764:
763:
762:
761:
751:
747:
746:
741:
738:
734:
730:
726:
723:
722:
719:
718:
717:Archaeopteryx
713:
709:
708:
707:
697:
696:
691:
688:
687:
686:
685:
684:
674:
673:
668:
667:
666:
665:
664:
655:
654:
648:
641:
636:
635:
630:
629:
628:
627:
626:
616:
615:
610:
607:
606:
605:
604:
603:
593:
592:
591:
590:
589:
579:
578:
572:
565:
564:
563:
562:
561:
551:
547:
546:
541:
540:
539:
538:
537:
527:
526:
525:Protoceratops
521:
520:
515:
511:
508:
507:
506:
505:
504:
495:
494:
489:
488:
482:
475:
470:
466:
465:
460:
457:
456:
451:
447:
446:
441:
438:
437:
436:
435:
434:
425:
424:
418:
411:
406:
402:
401:
400:
399:
398:
388:
387:
382:
379:
378:
377:
376:
375:
365:
361:
360:
355:
352:
351:
350:
349:
348:
339:
338:
332:
325:
320:
319:
314:
310:
307:
306:
305:
304:
303:
293:
289:
288:
287:Dromaeosaurus
282:
275:
270:
269:
264:
261:
260:
259:
258:
257:
247:
242:
237:
236:
231:
227:
223:
219:
215:
214:
211:
206:
200:Prescientific
199:
197:
194:
193:
188:
184:
180:
176:
175:
170:
165:
163:
162:Dromaeosaurus
159:
155:
151:
147:
143:
142:
137:
133:
129:
125:
124:
119:
118:
113:
109:
106:
101:
99:
95:
94:
89:
85:
81:
77:
73:
69:
68:
63:
60:
56:
52:
48:
44:
40:
33:
32:
31:Zephyrosaurus
27:
26:
20:
16:
5664:Dromaeosaurs
5626:Paleontology
5576:
5546:
5524:Velociraptor
5522:
5515:
5508:
5500:
5492:
5484:
5477:
5470:
5463:
5455:
5447:
5440:
5435:Acheroraptor
5433:
5413:
5406:
5399:
5391:
5384:
5377:
5370:
5362:
5356:Dakotaraptor
5354:
5349:Achillobator
5347:
5327:
5319:
5314:Atrociraptor
5312:
5293:Zhenyuanlong
5291:
5284:
5278:Tianyuraptor
5276:
5269:
5262:
5254:
5230:
5223:
5217:Tianyuraptor
5215:
5208:
5201:
5193:
5186:
5179:
5158:
5150:
5142:
5135:
5127:
5119:
5112:
5104:
5097:
5090:
5084:Dakotaraptor
5082:
5077:Buitreraptor
5075:
5070:Austroraptor
5068:
5060:Unenlagiinae
5047:
5040:
5033:
5026:
5007:Zhenyuanlong
5005:
4997:
4992:Tianyuraptor
4990:
4982:
4975:
4968:
4932:
4798:
4789:
4780:
4661:
4657:
4647:(2): 11–119.
4644:
4640:
4599:
4595:
4577:. Retrieved
4570:
4517:
4513:
4495:. Retrieved
4467:
4415:
4411:
4372:
4368:
4352:
4343:
4339:
4335:
4291:
4287:
4245:
4241:
4212:
4189:
4170:
4129:
4125:
4106:
4097:
4093:
4049:
4045:
3996:
3992:
3979:
3975:
3962:
3943:
3924:
3910:(2): 81–91.
3907:
3901:
3866:
3862:
3817:
3813:
3804:
3800:
3787:
3778:
3726:
3722:
3701:
3697:
3675:: e1034359.
3672:
3668:
3664:
3624:(1): 18–20.
3621:
3617:
3610:
3602:
3594:
3586:
3578:
3566:
3554:
3542:
3530:
3518:
3506:
3494:
3482:
3470:
3458:
3446:
3434:
3422:
3410:
3398:
3386:
3374:
3362:
3350:
3338:
3326:
3297:
3285:
3273:
3261:
3249:
3237:
3146:
3138:
3068:
3060:
3048:
3020:Mayor (2005)
2954:
2953:
2945:
2936:
2925:
2924:
2916:
2830:
2829:
2815:Atrociraptor
2813:
2812:
2796:
2795:
2781:Achillobator
2779:
2778:
2754:Velociraptor
2752:
2751:
2735:
2734:
2711:
2710:
2694:
2693:
2677:
2676:
2660:
2659:
2643:
2642:
2611:Zhenyuanlong
2610:
2609:
2608:
2594:Tianyuraptor
2592:
2591:
2575:
2574:
2558:
2557:
2541:
2540:
2524:
2523:
2500:
2499:
2485:Buitreraptor
2483:
2482:
2468:Austroraptor
2466:
2465:
2449:
2448:
2425:
2424:
2389:
2380:
2371:
2360:
2359:
2351:
2340:
2339:
2331:
2320:
2319:
2311:
2301:
2287:
2276:
2275:
2267:
2256:
2255:
2247:
2238:
2227:
2226:
2213:
2202:
2201:
2193:
2182:
2181:
2173:
2164:
2156:Austroraptor
2154:
2140:
2129:
2128:
2120:
2109:
2108:
2100:
2091:
2080:
2079:
2071:
2063:Buitreraptor
2061:
2047:
2036:
2035:
2027:
2016:
2015:
2007:
1997:
1975:
1974:
1964:structures.
1951:plumulaceous
1912:
1911:
1894:
1893:
1861:
1860:
1836:
1801:
1800:
1775:
1749:
1748:
1732:
1731:
1677:
1663:Dromaeosaurs
1661:
1628:
1627:
1601:
1600:
1575:
1552:Patagopteryx
1550:
1549:
1524:
1523:
1498:
1497:
1481:
1394:Coelurosaurs
1355:
1349:
1346:dromaeosaurs
1341:
1337:
1333:
1329:
1310:Kevin Padian
1301:
1295:
1289:
1279:
1278:referred to
1261:
1260:
1253:
1240:
1227:
1213:
1197:
1188:
1177:
1176:
1168:21st century
1160:
1138:
1129:
1115:
1111:Velociraptor
1109:
1095:
1094:
1088:
1084:
1078:
1065:
1051:
1033:
1032:
1016:warm blooded
1011:
996:
982:
981:
974:
959:
955:
949:
923:
922:
914:
907:
901:
890:
889:
882:
876:
865:
862:Velociraptor
861:
857:
853:Velociraptor
851:
842:
820:
819:
806:
805:
797:
770:
769:
756:
755:
749:
743:
727:divided the
715:
702:
701:
693:
679:
678:
670:
659:
658:
651:
632:
621:
620:
612:
598:
597:
584:
583:
575:
556:
555:
543:
532:
531:
523:
519:Velociraptor
517:
499:
498:
491:
485:
468:
462:
453:
443:
429:
428:
421:
393:
392:
384:
370:
369:
357:
343:
342:
335:
316:
309:Barnum Brown
298:
297:
285:
276:20th century
266:
252:
251:
243:19th century
233:
190:
172:
168:
166:
161:
158:warm-blooded
149:
139:
123:Velociraptor
121:
115:
102:
91:
70:. Since the
67:Velociraptor
65:
47:paleontology
38:
36:
29:
23:
15:
5479:Linheraptor
5457:Deinonychus
5408:Yurgovuchia
5364:Deinonychus
5321:Bambiraptor
5286:Vectiraptor
5264:Bambiraptor
5203:Microraptor
5114:Pamparaptor
5049:Natovenator
4888:Maniraptora
4873:Maniraptora
4575:(2403): 1–9
3903:Palaeoworld
3069:Deinonychus
2713:Deinonychus
2645:Bambiraptor
2560:Microraptor
1959:filamentous
1751:Caudipteryx
1679:Troodontids
1526:Hesperornis
1500:Ichthyornis
1418:Maniraptora
1364:coelurosaur
1360:troodontids
1356:Caudipteryx
1334:Caudipteryx
1291:Gorgosaurus
1255:Microraptor
1242:Bambiraptor
1089:Deinonychus
1080:Deinonychus
960:Deinonychus
951:Deinonychus
908:Deinonychus
903:Deinonychus
884:Deinonychus
469:Deinonychus
455:Deinonychus
440:John Ostrom
423:Deinonychus
235:Deinonychus
218:Crow people
192:Microraptor
169:Deinonychus
150:Deinonychus
136:John Ostrom
93:Deinonychus
86:to produce
28:preying on
25:Deinonychus
5648:Categories
5575:See also:
5449:Boreonykus
5401:Utahraptor
5152:Variraptor
5121:Pyroraptor
4999:Variraptor
4977:Pyroraptor
4909:see below↓
4785:Dinosauria
4776:Sauropsida
4579:2010-03-19
4497:2007-03-29
3656:References
2798:Utahraptor
1955:pennaceous
1483:Neornithes
1303:Aublysodon
1252:Fossil of
1229:Pyroraptor
1005:Chatterjee
954:teeth and
912:ornithopod
799:Utahraptor
220:and other
98:scientists
5614:Dinosaurs
5442:Adasaurus
5137:Unenlagia
5129:Rahonavis
5035:Hulsanpes
4861:Theropoda
4846:Theropoda
4794:Theropoda
4762:Kingdom:
4484:2246/5845
4391:0001-3765
4229:2246/5823
3689:131680329
3646:134803475
3603:in passim
3587:in passim
3147:in passim
3005:Footnotes
2696:Adasaurus
2502:Rahonavis
2451:Unenlagia
1734:Oviraptor
1318:Ji Shu-an
1048:Buffetaut
1044:Le Leouff
976:Unenlagia
405:braincase
183:limestone
80:sediments
78:used the
62:dinosaurs
5587:Category
5577:Timeline
5494:Nuthetes
5415:Zapsalis
5386:Itemirus
5160:Ypupiara
5042:Mahakala
4970:Daurlong
4770:Chordata
4768:Phylum:
4764:Animalia
4688:19692406
4624:12540892
4544:17823350
4492:31096081
4452:22615813
4412:PLOS ONE
4399:21437378
4346:: 1–165.
4316:15729340
4272:19129109
4162:27078534
4154:16222297
4084:26181289
4033:19289829
3982:: 79–88.
3893:25025742
3869:: 4382.
3850:14978813
3842:24248432
3763:20805514
2988:See also
2427:Mahakala
1314:Ji Qiang
1145:feathers
831:Kirkland
781:Gauthier
690:Osmolska
148:because
88:pigments
59:theropod
5600:Portals
5517:Tsaagan
4803:Paraves
4774:Class:
4679:2842677
4632:1160118
4604:Bibcode
4572:Zootaxa
4560:Xu Xing
4552:2519726
4522:Bibcode
4514:Science
4443:3352940
4420:Bibcode
4324:4413316
4296:Bibcode
4263:2679073
4134:Bibcode
4075:4504142
4054:Bibcode
4024:2664043
4001:Bibcode
3871:Bibcode
3822:Bibcode
3807:: 1–16.
3754:2932599
3731:Bibcode
3626:Bibcode
2969:phalanx
2679:Tsaagan
1354:, when
1276:dentary
354:Matthew
226:Montana
134:, when
108:Matthew
82:of the
76:Montana
43:history
5256:Balaur
5225:Wulong
5092:Diuqin
4984:Shanag
4686:
4676:
4630:
4622:
4596:Nature
4550:
4542:
4490:
4450:
4440:
4397:
4389:
4322:
4314:
4288:Nature
4270:
4260:
4196:
4177:
4160:
4152:
4126:Nature
4113:
4082:
4072:
4031:
4021:
3950:
3931:
3891:
3848:
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