Knowledge (XXG)

Tinirau clackae

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is elongated along the anteroposterior (forward-backward) axis of the body. The lengthening of the glenoid corresponds with a flattening of the proximal end of the humerus, a feature common in the forelimbs of more advanced stem tetrapods. Although the pectoral limb bones and girdle were not strong
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as a branch directly connected to the stem of Tetrapoda. In other words, it is a close relative of the lineage leading to land vertebrates (a prehistoric taxon can rarely be identified as an ancestor of another taxon, but is instead considered a side branch in a larger lineage).
1787:; Matt Friedman (2014). "A large, anatomically primitive tristichopterid (Sarcopterygii: Tetrapodomorpha) from the Late Devonian (Frasnian) Alves Beds, Upper Old Red Sandstone, Moray, Scotland". 4509: 4371: 530: 396:
was a fairly large, predatory fish about a meter long and with a deep, compact body. The head was large, with large terminal mout and numerous teeth. The tail was
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out of water, glenoid lengthening and other changes to the proximal forelimb were among the first steps in the transformation from pectoral fin to forelimb.
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marine environment. Six fossils are known, all preserving bones of the skull. Two fossils preserve postcranial bones in articulation. The
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is the earliest known stem tetrapod to have a significantly reduced postaxial process, and a fibula more like those of later tetrapods.
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that are transitional between fish, representing life in the water, and tetrapods, representing life on land. The specific name honors
588:, traditionally considered a direct ancestor of tetrapods, or a close relative of such an ancestor, was placed deep within the family 498: 4126: 4119: 547:
legend is the guardian of ocean life, and whose form is half human and half fish. The connection alludes to the features in
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had hindlimb configurations very similar to the first land-living tetrapods, some early forms such as
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appeared, many stem tetrapods had already developed the three major hindlimb bones of tetrapods: the
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is positioned more crownward, or closer to Tetrapoda, than other stem tetrapods like
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were situated behind the middle of the body similar to the situation seen in
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stage of the Middle Devonian. The Red Hill I beds include a series of
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It shares many advanced features with later tetrapodomorphs in the
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close to the ancestry of land-living vertebrates in the
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was named by paleontologist Brian Swartz in 2012 after
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modified from Swartz (2012) showing the placement of
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Relative to more well-known stem tetrapods, 1896: 350:is more closely related to Tetrapoda than is 8: 136:Holotype fossil and interpretive drawing of 4306: 4157: 4146: 4011: 3684: 3584: 3440: 3375: 3215: 2989: 2977: 2952: 2789: 2603: 2068: 2057: 2044: 1990: 1981: 1926: 1903: 1889: 1881: 592:, away from the tetrapod stem. Below is a 127: 31: 4510:Devonian sarcopterygians of North America 1860: 1850: 1778: 1776: 493:were first discovered by paleontologist 1772: 533:118605, is a mostly complete skeleton. 1828: 1826: 1824: 1822: 1820: 1818: 1759:(2014) considered it more likely that 356:, but farther from Tetrapoda than is 7: 435:. While derived stem tetrapods like 4500:Prehistoric lobe-finned fish genera 1547: 1524: 1499: 1475: 1453: 1431: 1407: 1383: 1359: 1335: 1285: 1278: 1254: 1227: 1203: 1099: 1075: 1051: 1027: 1003: 979: 969: 962: 884: 860: 836: 812: 802: 725: 701: 677: 667: 660: 638: 614: 603: 1783:Matthew Parfitt; Zerina Johanson; 25: 499:University of California Berkeley 497:in 1970, who then worked for the 27:Extinct species of tetrapodomorph 4290: 4289: 4132: 4125: 4118: 4111: 4104: 4097: 2938: 2932: 2927: 2920: 477:enough to support the weight of 460:Like those of the Late Devonian 419:(shoulder sockets). By the time 148: 4505:Middle Devonian sarcopterygians 1: 4254:Polydactyly in stem-tetrapods 1852:10.1371/journal.pone.0033683 1789:Scottish Journal of Geology 1755:On the other hand, Parfitt 4526: 301:Schultze & Reed (2012) 4285: 4156: 4145: 4095: 3687: 3587: 3443: 3378: 3218: 2992: 2976: 2951: 2917: 2792: 2606: 2071: 2056: 2043: 1993: 1980: 1925: 1567: 1552: 1545: 1529: 1522: 1504: 1497: 1480: 1473: 1458: 1451: 1436: 1429: 1412: 1405: 1388: 1381: 1364: 1357: 1340: 1333: 1307: 1290: 1283: 1276: 1259: 1252: 1232: 1225: 1208: 1201: 1121: 1104: 1097: 1080: 1073: 1056: 1049: 1032: 1025: 1008: 1001: 984: 977: 967: 960: 906: 889: 882: 865: 858: 841: 834: 817: 810: 800: 747: 730: 723: 706: 699: 682: 675: 665: 658: 643: 636: 619: 612: 521:, likely deposited in an 295: 288: 270: 263: 145:Scientific classification 143: 135: 126: 34: 523:outer continental shelf 4249:Evolution of tetrapods 503:Simpson Park Mountains 390: 4467:Paleobiology Database 4398:Paleobiology Database 507:Eureka County, Nevada 386:3D reconstruction of 385: 298:Bruehnopteron murphyi 366:and only species of 4269:Zachelmie trackways 3449:Pseudophlegethontia 1833:Swartz, B. (2012). 1801:10.1144/sjg2013-013 4191:Megalichthyiformes 2753:Platycephalichthys 1236:Platycephalichthys 807:Megalichthyiformes 391: 4482: 4481: 4385:Open Tree of Life 4312:Taxon identifiers 4303: 4302: 4281: 4280: 4277: 4276: 4197:Tristichopteridae 4141: 4140: 4093: 4092: 4089: 4088: 4085: 4084: 3751: 3750: 3643: 3642: 3489: 3488: 3485: 3484: 3436:Phlegethontioidea 3199: 3198: 3167:Elginerpetontidae 2947: 2946: 2915: 2914: 2911: 2910: 2907: 2906: 2903: 2902: 2832:Parapanderichthys 2630:Tristichopteridae 2599:Eotetrapodiformes 2039: 2038: 2035: 2034: 2031: 2030: 1752: 1751: 1743: 1742: 1734: 1733: 1725: 1724: 1716: 1715: 1707: 1706: 1698: 1697: 1689: 1688: 1680: 1679: 1671: 1670: 1662: 1661: 1653: 1652: 1644: 1643: 1635: 1634: 1626: 1625: 1617: 1616: 1608: 1607: 1599: 1598: 1590: 1589: 1581: 1580: 1322: 1321: 1190: 1189: 1181: 1180: 1172: 1171: 1163: 1162: 1154: 1153: 1145: 1144: 1136: 1135: 974:Tristichopteridae 957:Eotetrapodiformes 946: 945: 937: 936: 928: 927: 919: 918: 789: 788: 780: 779: 771: 770: 762: 761: 590:Tristichopteridae 573:megalichthyiforms 553:Jennifer A. Clack 495:Joseph T. Gregory 472:, the glenoid of 449:hanging over the 374:, named in 2012. 308: 307: 244: 221:Eotetrapodiformes 16:(Redirected from 4517: 4495:Tristichopterids 4475: 4474: 4462: 4461: 4449: 4448: 4447: 4434: 4433: 4432: 4406: 4405: 4393: 4392: 4380: 4379: 4367: 4366: 4354: 4353: 4352: 4339: 4338: 4337: 4307: 4293: 4292: 4203:Labyrinthodontia 4158: 4147: 4136: 4130: 4129: 4123: 4122: 4116: 4115: 4109: 4108: 4101: 4058: 4054: 4020: 4012: 4004: 3943: 3940: 3903: 3758: 3685: 3678: 3650: 3585: 3578: 3534: 3496: 3460:Phlegethontiidae 3441: 3376: 3369: 3354: 3344: 3334: 3324: 3314: 3304: 3294: 3284: 3274: 3264: 3254: 3244: 3234: 3224: 3216: 3212: 3206: 3165: 3150: 3140: 3130: 3120: 3110: 3099: 3089: 3079: 3069: 3059: 3049: 3039: 3028: 3018: 3008: 2998: 2990: 2986: 2978: 2971: 2962: 2953: 2942: 2936: 2931: 2925: 2924: 2886: 2877: 2859: 2849: 2839: 2829: 2819: 2809: 2798: 2790: 2628: 2612: 2604: 2488: 2435: 2312: 2297: 2287: 2277: 2267: 2257: 2247: 2237: 2227: 2217: 2207: 2197: 2187: 2177: 2167: 2157: 2147: 2137: 2127: 2117: 2107: 2097: 2087: 2077: 2069: 2058: 2045: 1991: 1982: 1975: 1974: 1927: 1919: 1915: 1912:Tetrapodomorpha 1905: 1898: 1891: 1882: 1875: 1874: 1864: 1854: 1830: 1813: 1812: 1780: 1548: 1525: 1500: 1476: 1454: 1432: 1408: 1384: 1360: 1336: 1286: 1279: 1255: 1228: 1204: 1100: 1076: 1052: 1028: 1004: 980: 970: 963: 885: 861: 837: 813: 803: 726: 702: 678: 668: 661: 639: 615: 604: 577:tristichopterids 302: 279: 275: 239: 232: 153: 152: 131: 121: 51: 40:Temporal range: 32: 21: 4525: 4524: 4520: 4519: 4518: 4516: 4515: 4514: 4485: 4484: 4483: 4478: 4470: 4465: 4457: 4452: 4445:Tinirau clackae 4443: 4442: 4437: 4428: 4427: 4422: 4415:Tinirau clackae 4409: 4401: 4396: 4388: 4383: 4375: 4370: 4362: 4357: 4348: 4347: 4342: 4333: 4332: 4327: 4314: 4304: 4299: 4273: 4237: 4213:Ichthyostegalia 4181:Osteolepiformes 4165: 4152: 4137: 4124: 4117: 4110: 4103: 4081: 4056: 4052: 4051:Reptiliomorpha 4045: 4018: 4005: 4002: 3997: 3941: 3933: 3905:Gephyrostegidae 3896: 3797:Carbonoherpeton 3747: 3706: 3671: 3639: 3634:Palaeomolgophis 3598: 3571: 3527: 3481: 3454: 3430: 3409:Oestocephalidae 3403: 3384:Andersonerpeton 3362: 3357:Tantallognathus 3210: 3204: 3195: 3183:Obruchevichthys 3158: 2984: 2972: 2966: 2960: 2943: 2919: 2899: 2881: 2875: 2867: 2785:Elpistostegalia 2779: 2774:Tristichopterus 2621: 2615:Palaeospondylus 2593: 2587:Sterropterygion 2521:Ectosteorhachis 2506:Cladarosymblema 2490:Megalichthyidae 2481: 2428: 2365:Propycnoctenion 2305: 2065: 2063:Tetrapodomorpha 2052: 2050:Tetrapodomorpha 2027: 2013:Tetrapodomorpha 2007: 1976: 1932: 1931: 1921: 1917: 1913: 1909: 1879: 1878: 1832: 1831: 1816: 1782: 1781: 1774: 1769: 1753: 1744: 1735: 1726: 1717: 1708: 1699: 1690: 1681: 1672: 1663: 1654: 1645: 1636: 1627: 1618: 1609: 1600: 1591: 1582: 1323: 1249:Elpistostegalia 1191: 1182: 1173: 1164: 1155: 1146: 1137: 1012:Tristichopterus 947: 938: 929: 920: 909:Megalichthyidae 790: 781: 772: 763: 608:Tetrapodomorpha 561: 537:Tinirau clackae 487: 453:bone below it. 380: 325:Middle Devonian 300: 284: 281: 277:Tinirau clackae 273: 272: 259: 256:T. clackae 238: 230: 208:Tetrapodomorpha 147: 138:Tinirau clackae 122: 120: 119: 118: 117: 112: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 46: 45: 42:Middle Devonian 38: 36:Tinirau clackae 28: 23: 22: 15: 12: 11: 5: 4523: 4521: 4513: 4512: 4507: 4502: 4497: 4487: 4486: 4480: 4479: 4477: 4476: 4463: 4450: 4435: 4419: 4417: 4411: 4410: 4408: 4407: 4394: 4381: 4368: 4355: 4340: 4324: 4322: 4316: 4315: 4310: 4301: 4300: 4298: 4297: 4286: 4283: 4282: 4279: 4278: 4275: 4274: 4272: 4271: 4266: 4261: 4256: 4251: 4245: 4243: 4239: 4238: 4236: 4235: 4233:Anthracosauria 4230: 4228:Phyllospondyli 4225: 4220: 4215: 4210: 4205: 4200: 4194: 4188: 4186:Osteolepididae 4183: 4178: 4176:Stem tetrapoda 4172: 4170: 4154: 4153: 4151:Related topics 4150: 4143: 4142: 4139: 4138: 4096: 4094: 4091: 4090: 4087: 4086: 4083: 4082: 4080: 4079: 4078: 4077: 4071: 4069:Reptiliomorpha 4062: 4060: 4047: 4046: 4044: 4043: 4042: 4041: 4035: 4026: 4024: 4022:Batrachomorpha 4009: 3999: 3998: 3996: 3995: 3988: 3981: 3974: 3967: 3959: 3951: 3949: 3935: 3934: 3932: 3931: 3924: 3917: 3914:Bruktererpeton 3909: 3907: 3898: 3897: 3895: 3894: 3891:Spondylerpeton 3887: 3880: 3873: 3870:Proterogyrinus 3866: 3859: 3852: 3849:Palaeoherpeton 3845: 3838: 3831: 3823: 3816: 3813:Diplovertebron 3809: 3800: 3793: 3790:Calligenethlon 3786: 3779: 3772: 3769:Anthracosaurus 3764: 3762: 3753: 3752: 3749: 3748: 3746: 3745: 3742:Megalocephalus 3738: 3731: 3724: 3716: 3714: 3708: 3707: 3705: 3704: 3701:Spathicephalus 3697: 3688: 3682: 3673: 3672: 3670: 3669: 3668: 3667: 3658: 3656: 3645: 3644: 3641: 3640: 3638: 3637: 3630: 3627:Dolichopareias 3623: 3620:Adelospondylus 3616: 3608: 3606: 3604:Adelogyrinidae 3600: 3599: 3597: 3596: 3588: 3582: 3573: 3572: 3570: 3569: 3562: 3555: 3548: 3540: 3538: 3529: 3528: 3526: 3525: 3518: 3511: 3502: 3500: 3498:Whatcheeriidae 3491: 3490: 3487: 3486: 3483: 3482: 3480: 3479: 3472: 3464: 3462: 3456: 3455: 3453: 3452: 3444: 3438: 3432: 3431: 3429: 3428: 3421: 3418:Coloraderpeton 3413: 3411: 3405: 3404: 3402: 3401: 3394: 3387: 3379: 3373: 3364: 3363: 3361: 3360: 3350: 3340: 3330: 3320: 3310: 3300: 3290: 3280: 3270: 3260: 3250: 3240: 3230: 3219: 3213: 3201: 3200: 3197: 3196: 3194: 3193: 3186: 3179: 3171: 3169: 3160: 3159: 3157: 3156: 3146: 3136: 3126: 3116: 3106: 3095: 3085: 3075: 3065: 3055: 3045: 3035: 3024: 3014: 3004: 2993: 2987: 2974: 2973: 2956: 2949: 2948: 2945: 2944: 2918: 2916: 2913: 2912: 2909: 2908: 2905: 2904: 2901: 2900: 2898: 2897: 2896: 2895: 2889: 2887: 2869: 2868: 2866: 2865: 2855: 2845: 2835: 2825: 2815: 2805: 2793: 2787: 2781: 2780: 2778: 2777: 2770: 2763: 2756: 2749: 2742: 2738:Marsdenichthys 2734: 2727: 2719: 2712: 2705: 2698: 2691: 2688:Eusthenopteron 2684: 2677: 2670: 2663: 2656: 2653:Callistipterus 2649: 2642: 2634: 2632: 2623: 2622: 2620: 2619: 2607: 2601: 2595: 2594: 2592: 2591: 2583: 2580:Sengoerichthys 2576: 2569: 2566:Palatinichthys 2562: 2554: 2546: 2539: 2532: 2524: 2517: 2509: 2502: 2494: 2492: 2483: 2482: 2480: 2479: 2471: 2467:Marsdenichthys 2463: 2456: 2449: 2441: 2439: 2430: 2429: 2427: 2426: 2418: 2410: 2403: 2396: 2389: 2382: 2375: 2368: 2361: 2354: 2347: 2340: 2333: 2326: 2318: 2316: 2307: 2306: 2304: 2303: 2293: 2283: 2273: 2263: 2253: 2250:Platyethmoidia 2243: 2233: 2223: 2213: 2210:Megadonichthys 2203: 2193: 2183: 2173: 2163: 2153: 2150:Harajicadectes 2143: 2133: 2123: 2113: 2103: 2093: 2083: 2072: 2066: 2061: 2054: 2053: 2048: 2041: 2040: 2037: 2036: 2033: 2032: 2029: 2028: 2026: 2025: 2024: 2023: 2017: 2015: 2009: 2008: 2006: 2005: 2004: 2003: 1994: 1988: 1978: 1977: 1973: 1972: 1963: 1954: 1945: 1939: 1930: 1923: 1922: 1910: 1908: 1907: 1900: 1893: 1885: 1877: 1876: 1814: 1771: 1770: 1768: 1765: 1750: 1749: 1746: 1745: 1741: 1740: 1737: 1736: 1732: 1731: 1728: 1727: 1723: 1722: 1719: 1718: 1714: 1713: 1710: 1709: 1705: 1704: 1701: 1700: 1696: 1695: 1692: 1691: 1687: 1686: 1683: 1682: 1678: 1677: 1674: 1673: 1669: 1668: 1665: 1664: 1660: 1659: 1656: 1655: 1651: 1650: 1647: 1646: 1642: 1641: 1638: 1637: 1633: 1632: 1629: 1628: 1624: 1623: 1620: 1619: 1615: 1614: 1611: 1610: 1606: 1605: 1602: 1601: 1597: 1596: 1593: 1592: 1588: 1587: 1584: 1583: 1579: 1578: 1575: 1574: 1566: 1563: 1562: 1559: 1558: 1551: 1546: 1544: 1541: 1540: 1537: 1536: 1528: 1523: 1521: 1515: 1514: 1511: 1510: 1503: 1498: 1496: 1493: 1492: 1489: 1488: 1479: 1474: 1472: 1469: 1468: 1465: 1464: 1457: 1452: 1450: 1447: 1446: 1443: 1442: 1439:Whatcheeriidae 1435: 1430: 1428: 1425: 1424: 1421: 1420: 1411: 1406: 1404: 1401: 1400: 1397: 1396: 1387: 1382: 1380: 1377: 1376: 1373: 1372: 1363: 1358: 1356: 1353: 1352: 1349: 1348: 1339: 1334: 1332: 1329: 1328: 1325: 1324: 1320: 1319: 1316: 1315: 1306: 1303: 1302: 1299: 1298: 1289: 1284: 1282: 1277: 1275: 1272: 1271: 1268: 1267: 1258: 1253: 1251: 1245: 1244: 1241: 1240: 1231: 1226: 1224: 1221: 1220: 1217: 1216: 1207: 1202: 1200: 1197: 1196: 1193: 1192: 1188: 1187: 1184: 1183: 1179: 1178: 1175: 1174: 1170: 1169: 1166: 1165: 1161: 1160: 1157: 1156: 1152: 1151: 1148: 1147: 1143: 1142: 1139: 1138: 1134: 1133: 1130: 1129: 1120: 1117: 1116: 1113: 1112: 1103: 1098: 1096: 1093: 1092: 1089: 1088: 1079: 1074: 1072: 1069: 1068: 1065: 1064: 1055: 1050: 1048: 1045: 1044: 1041: 1040: 1036:Eusthenopteron 1031: 1026: 1024: 1021: 1020: 1017: 1016: 1007: 1002: 1000: 997: 996: 993: 992: 983: 978: 976: 968: 966: 961: 959: 953: 952: 949: 948: 944: 943: 940: 939: 935: 934: 931: 930: 926: 925: 922: 921: 917: 916: 913: 912: 905: 902: 901: 898: 897: 888: 883: 881: 878: 877: 874: 873: 864: 859: 857: 854: 853: 850: 849: 840: 835: 833: 830: 829: 826: 825: 816: 811: 809: 801: 799: 796: 795: 792: 791: 787: 786: 783: 782: 778: 777: 774: 773: 769: 768: 765: 764: 760: 759: 756: 755: 746: 743: 742: 739: 738: 729: 724: 722: 719: 718: 715: 714: 705: 700: 698: 695: 694: 691: 690: 686:Marsdenichthys 681: 676: 674: 666: 664: 659: 657: 654: 653: 650: 649: 642: 637: 635: 632: 631: 628: 627: 618: 613: 611: 602: 586:Eusthenopteron 560: 557: 486: 483: 442:Eusthenopteron 379: 376: 353:Eusthenopteron 323:fish from the 321:sarcopterygian 315:is an extinct 306: 305: 293: 292: 286: 285: 282: 268: 267: 261: 260: 252: 250: 246: 245: 228: 224: 223: 218: 211: 210: 205: 198: 197: 192: 185: 184: 179: 175: 174: 169: 165: 164: 159: 155: 154: 141: 140: 133: 132: 124: 123: 115: 114: 113: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 52: 39: 26: 24: 18:Tinirau (fish) 14: 13: 10: 9: 6: 4: 3: 2: 4522: 4511: 4508: 4506: 4503: 4501: 4498: 4496: 4493: 4492: 4490: 4473: 4468: 4464: 4460: 4455: 4451: 4446: 4440: 4436: 4431: 4425: 4421: 4420: 4418: 4416: 4412: 4404: 4399: 4395: 4391: 4386: 4382: 4378: 4373: 4369: 4365: 4360: 4356: 4351: 4345: 4341: 4336: 4330: 4326: 4325: 4323: 4321: 4317: 4313: 4308: 4296: 4288: 4287: 4284: 4270: 4267: 4265: 4262: 4260: 4257: 4255: 4252: 4250: 4247: 4246: 4244: 4240: 4234: 4231: 4229: 4226: 4224: 4221: 4219: 4216: 4214: 4211: 4209: 4206: 4204: 4201: 4198: 4195: 4192: 4189: 4187: 4184: 4182: 4179: 4177: 4174: 4173: 4171: 4168: 4163: 4159: 4155: 4148: 4144: 4135: 4131: 4128: 4121: 4114: 4107: 4100: 4076: 4072: 4070: 4066: 4065: 4064: 4063: 4061: 4059: 4048: 4040: 4036: 4034: 4033:Temnospondyli 4030: 4029: 4028: 4027: 4025: 4023: 4017: 4016:Temnospondyli 4013: 4010: 4008: 4000: 3994: 3993: 3992:Termonerpeton 3989: 3987: 3986: 3985:Silvanerpeton 3982: 3980: 3979: 3975: 3973: 3972: 3968: 3965: 3964: 3960: 3958: 3957: 3953: 3952: 3950: 3947: 3946:anthracosaurs 3936: 3930: 3929: 3928:Gephyrostegus 3925: 3923: 3922: 3921:Eusauropleura 3918: 3916: 3915: 3911: 3910: 3908: 3906: 3899: 3893: 3892: 3888: 3886: 3885: 3881: 3879: 3878: 3874: 3872: 3871: 3867: 3865: 3864: 3863:Pholiderpeton 3860: 3858: 3857: 3853: 3851: 3850: 3846: 3844: 3843: 3839: 3837: 3836: 3835:Leptophractus 3832: 3829: 3828: 3824: 3822: 3821: 3817: 3815: 3814: 3810: 3807: 3806: 3801: 3799: 3798: 3794: 3792: 3791: 3787: 3785: 3784: 3780: 3778: 3777: 3773: 3771: 3770: 3766: 3765: 3763: 3761: 3754: 3744: 3743: 3739: 3737: 3736: 3732: 3730: 3729: 3725: 3723: 3722: 3718: 3717: 3715: 3713: 3709: 3703: 3702: 3698: 3695: 3694: 3690: 3689: 3686: 3683: 3681: 3674: 3666: 3662: 3661: 3660: 3659: 3657: 3654: 3646: 3636: 3635: 3631: 3629: 3628: 3624: 3622: 3621: 3617: 3615: 3614: 3610: 3609: 3607: 3605: 3601: 3595: 3594: 3593:Acherontiscus 3590: 3589: 3586: 3583: 3581: 3580:Adelospondyli 3574: 3568: 3567: 3566:Pholidogaster 3563: 3561: 3560: 3556: 3554: 3553: 3552:Deltaherpeton 3549: 3547: 3546: 3542: 3541: 3539: 3537: 3530: 3524: 3523: 3519: 3517: 3516: 3512: 3509: 3508: 3504: 3503: 3501: 3499: 3492: 3478: 3477: 3473: 3471: 3470: 3469:Phlegethontia 3466: 3465: 3463: 3461: 3457: 3451: 3450: 3446: 3445: 3442: 3439: 3437: 3433: 3427: 3426: 3425:Oestocephalus 3422: 3420: 3419: 3415: 3414: 3412: 3410: 3406: 3400: 3399: 3395: 3393: 3392: 3388: 3386: 3385: 3381: 3380: 3377: 3374: 3372: 3365: 3359: 3358: 3351: 3349: 3348: 3341: 3339: 3338: 3331: 3329: 3328: 3321: 3319: 3318: 3311: 3309: 3308: 3301: 3299: 3298: 3291: 3289: 3288: 3287:Ichthyerpeton 3281: 3279: 3278: 3271: 3269: 3268: 3261: 3259: 3258: 3251: 3249: 3248: 3247:Crassigyrinus 3241: 3239: 3238: 3231: 3229: 3228: 3221: 3220: 3217: 3214: 3209: 3202: 3192: 3191: 3187: 3185: 3184: 3180: 3178: 3177: 3173: 3172: 3170: 3168: 3161: 3155: 3154: 3147: 3145: 3144: 3137: 3135: 3134: 3127: 3125: 3124: 3117: 3115: 3114: 3107: 3104: 3103: 3096: 3094: 3093: 3086: 3084: 3083: 3076: 3074: 3073: 3072:Metaxygnathus 3066: 3064: 3063: 3056: 3054: 3053: 3046: 3044: 3043: 3036: 3033: 3032: 3025: 3023: 3022: 3015: 3013: 3012: 3011:Brittagnathus 3005: 3003: 3002: 2995: 2994: 2991: 2988: 2983: 2979: 2975: 2969: 2965: 2959: 2954: 2950: 2941: 2937: 2935: 2930: 2923: 2893: 2892: 2891: 2890: 2888: 2884: 2880: 2874: 2870: 2864: 2863: 2856: 2854: 2853: 2846: 2844: 2843: 2836: 2834: 2833: 2826: 2824: 2823: 2822:Panderichthys 2816: 2814: 2813: 2806: 2803: 2802: 2801:Howittichthys 2795: 2794: 2791: 2788: 2786: 2782: 2776: 2775: 2771: 2769: 2768: 2764: 2762: 2761: 2757: 2755: 2754: 2750: 2748: 2747: 2743: 2740: 2739: 2735: 2733: 2732: 2728: 2725: 2724: 2720: 2718: 2717: 2713: 2711: 2710: 2706: 2704: 2703: 2699: 2697: 2696: 2695:Heddleichthys 2692: 2690: 2689: 2685: 2683: 2682: 2678: 2676: 2675: 2671: 2669: 2668: 2664: 2662: 2661: 2657: 2655: 2654: 2650: 2648: 2647: 2646:Cabonnichthys 2643: 2641: 2640: 2639:Bruehnopteron 2636: 2635: 2633: 2631: 2624: 2617: 2616: 2609: 2608: 2605: 2602: 2600: 2596: 2589: 2588: 2584: 2582: 2581: 2577: 2575: 2574: 2570: 2568: 2567: 2563: 2560: 2559: 2555: 2552: 2551: 2547: 2545: 2544: 2540: 2538: 2537: 2533: 2530: 2529: 2525: 2523: 2522: 2518: 2515: 2514: 2510: 2508: 2507: 2503: 2501: 2500: 2496: 2495: 2493: 2491: 2484: 2477: 2476: 2472: 2469: 2468: 2464: 2462: 2461: 2457: 2455: 2454: 2450: 2448: 2447: 2443: 2442: 2440: 2438: 2437:Canowindridae 2431: 2424: 2423: 2419: 2416: 2415: 2411: 2409: 2408: 2404: 2402: 2401: 2397: 2395: 2394: 2390: 2388: 2387: 2383: 2381: 2380: 2376: 2374: 2373: 2369: 2367: 2366: 2362: 2360: 2359: 2355: 2353: 2352: 2348: 2346: 2345: 2341: 2339: 2338: 2334: 2332: 2331: 2327: 2325: 2324: 2320: 2319: 2317: 2315: 2308: 2302: 2301: 2294: 2292: 2291: 2284: 2282: 2281: 2274: 2272: 2271: 2270:Thaumatolepis 2264: 2262: 2261: 2254: 2252: 2251: 2244: 2242: 2241: 2234: 2232: 2231: 2224: 2222: 2221: 2214: 2212: 2211: 2204: 2202: 2201: 2194: 2192: 2191: 2184: 2182: 2181: 2180:Lamprotolepis 2174: 2172: 2171: 2164: 2162: 2161: 2154: 2152: 2151: 2144: 2142: 2141: 2134: 2132: 2131: 2130:Greiserolepis 2124: 2122: 2121: 2114: 2112: 2111: 2104: 2102: 2101: 2094: 2092: 2091: 2084: 2082: 2081: 2074: 2073: 2070: 2067: 2064: 2059: 2055: 2051: 2046: 2042: 2021: 2020: 2019: 2018: 2016: 2014: 2010: 2002: 2001:Gnathostomata 1998: 1997: 1996: 1995: 1992: 1989: 1987: 1986:Gnathostomata 1983: 1979: 1971: 1967: 1964: 1962: 1961:Sarcopterygii 1958: 1955: 1953: 1949: 1946: 1944: 1940: 1938: 1934: 1933: 1928: 1924: 1920: 1916:Pan-Tetrapoda 1906: 1901: 1899: 1894: 1892: 1887: 1886: 1883: 1872: 1868: 1863: 1858: 1853: 1848: 1845:(3): e33683. 1844: 1840: 1836: 1829: 1827: 1825: 1823: 1821: 1819: 1815: 1810: 1806: 1802: 1798: 1794: 1790: 1786: 1779: 1777: 1773: 1766: 1764: 1762: 1758: 1748: 1747: 1739: 1738: 1730: 1729: 1721: 1720: 1712: 1711: 1703: 1702: 1694: 1693: 1685: 1684: 1676: 1675: 1667: 1666: 1658: 1657: 1649: 1648: 1640: 1639: 1631: 1630: 1622: 1621: 1613: 1612: 1604: 1603: 1595: 1594: 1586: 1585: 1577: 1576: 1573: 1572: 1565: 1564: 1561: 1560: 1557: 1556: 1550: 1549: 1543: 1542: 1539: 1538: 1535: 1534: 1527: 1526: 1520: 1517: 1516: 1513: 1512: 1509: 1508: 1502: 1501: 1495: 1494: 1491: 1490: 1487: 1486: 1485: 1484:Crassigyrinus 1478: 1477: 1471: 1470: 1467: 1466: 1463: 1462: 1456: 1455: 1449: 1448: 1445: 1444: 1441: 1440: 1434: 1433: 1427: 1426: 1423: 1422: 1419: 1418: 1417: 1410: 1409: 1403: 1402: 1399: 1398: 1395: 1394: 1393: 1386: 1385: 1379: 1378: 1375: 1374: 1371: 1370: 1369: 1362: 1361: 1355: 1354: 1351: 1350: 1347: 1346: 1345: 1338: 1337: 1331: 1330: 1327: 1326: 1318: 1317: 1314: 1313: 1312: 1305: 1304: 1301: 1300: 1297: 1296: 1295: 1288: 1287: 1281: 1280: 1274: 1273: 1270: 1269: 1266: 1265: 1264: 1263:Panderichthys 1257: 1256: 1250: 1247: 1246: 1243: 1242: 1239: 1238: 1237: 1230: 1229: 1223: 1222: 1219: 1218: 1215: 1214: 1213: 1206: 1205: 1199: 1198: 1195: 1194: 1186: 1185: 1177: 1176: 1168: 1167: 1159: 1158: 1150: 1149: 1141: 1140: 1132: 1131: 1128: 1127: 1126: 1119: 1118: 1115: 1114: 1111: 1110: 1109: 1102: 1101: 1095: 1094: 1091: 1090: 1087: 1086: 1085: 1084:Cabonnichthys 1078: 1077: 1071: 1070: 1067: 1066: 1063: 1062: 1061: 1054: 1053: 1047: 1046: 1043: 1042: 1039: 1038: 1037: 1030: 1029: 1023: 1022: 1019: 1018: 1015: 1014: 1013: 1006: 1005: 999: 998: 995: 994: 991: 990: 989: 982: 981: 975: 972: 971: 965: 964: 958: 955: 954: 951: 950: 942: 941: 933: 932: 924: 923: 915: 914: 911: 910: 904: 903: 900: 899: 896: 895: 894: 887: 886: 880: 879: 876: 875: 872: 871: 870: 863: 862: 856: 855: 852: 851: 848: 847: 846: 839: 838: 832: 831: 828: 827: 824: 823: 822: 815: 814: 808: 805: 804: 798: 797: 794: 793: 785: 784: 776: 775: 767: 766: 758: 757: 754: 753: 752: 745: 744: 741: 740: 737: 736: 735: 728: 727: 721: 720: 717: 716: 713: 712: 711: 704: 703: 697: 696: 693: 692: 689: 688: 687: 680: 679: 673: 672:Canowindridae 670: 669: 663: 662: 656: 655: 652: 651: 648: 647: 646:Rhizodontidae 641: 640: 634: 633: 630: 629: 626: 625: 624: 617: 616: 609: 606: 605: 601: 599: 595: 591: 587: 582: 578: 574: 570: 566: 558: 556: 554: 550: 546: 542: 538: 534: 532: 528: 524: 520: 516: 512: 508: 504: 500: 496: 492: 484: 482: 480: 475: 471: 470: 465: 464: 463:Panderichthys 458: 456: 452: 448: 444: 443: 438: 437:Panderichthys 434: 430: 426: 422: 418: 414: 409: 407: 403: 402:pectoral fins 399: 395: 389: 384: 377: 375: 373: 369: 365: 361: 360: 359:Panderichthys 355: 354: 349: 345: 342: 338: 337:stem tetrapod 334: 330: 326: 322: 318: 314: 313: 304: 299: 294: 291: 287: 280: 278: 269: 266: 265:Binomial name 262: 258: 257: 251: 248: 247: 243: 237: 236: 229: 226: 225: 222: 219: 216: 213: 212: 209: 206: 203: 200: 199: 196: 195:Sarcopterygii 193: 190: 187: 186: 183: 180: 177: 176: 173: 170: 167: 166: 163: 160: 157: 156: 151: 146: 142: 139: 134: 130: 125: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 50: 43: 37: 33: 30: 19: 4414: 4319: 4242:Other topics 4223:Holospondyli 4218:Lepospondyli 4167:Polyphyletic 4162:Paraphyletic 4102: 4039:Lissamphibia 3990: 3983: 3976: 3969: 3961: 3956:Caerorhachis 3954: 3926: 3919: 3912: 3889: 3884:Seroherpeton 3882: 3875: 3868: 3861: 3854: 3847: 3842:Neopteroplax 3840: 3833: 3825: 3818: 3811: 3803: 3795: 3788: 3781: 3774: 3767: 3740: 3733: 3726: 3719: 3699: 3691: 3632: 3625: 3618: 3613:Adelogyrinus 3611: 3591: 3564: 3559:Greererpeton 3557: 3550: 3543: 3520: 3513: 3505: 3474: 3467: 3447: 3423: 3416: 3398:Ophiderpeton 3396: 3389: 3382: 3355: 3345: 3335: 3325: 3315: 3307:Mesanerpeton 3305: 3295: 3285: 3275: 3265: 3255: 3245: 3237:Aytonerpeton 3235: 3225: 3190:Webererpeton 3188: 3181: 3176:Elginerpeton 3174: 3151: 3141: 3131: 3121: 3111: 3100: 3090: 3080: 3070: 3060: 3052:Ichthyostega 3050: 3040: 3031:Elpistostege 3029: 3021:Densignathus 3019: 3009: 3001:Acanthostega 2999: 2967: 2958:Stegocephali 2926: 2882: 2873:Stegocephali 2860: 2852:Rubrognathus 2850: 2840: 2830: 2820: 2810: 2799: 2772: 2766: 2765: 2758: 2751: 2746:Notorhizodon 2744: 2736: 2729: 2721: 2714: 2707: 2700: 2693: 2686: 2679: 2672: 2665: 2658: 2651: 2644: 2637: 2613: 2585: 2578: 2571: 2564: 2558:Megistolepis 2556: 2548: 2543:Megalichthys 2541: 2534: 2526: 2519: 2511: 2504: 2499:Askerichthys 2497: 2473: 2465: 2458: 2451: 2444: 2420: 2414:Thysanolepis 2412: 2405: 2398: 2391: 2384: 2377: 2372:Pycnoctenion 2370: 2363: 2356: 2349: 2344:Gooloogongia 2342: 2335: 2328: 2321: 2314:Rhizodontida 2298: 2288: 2278: 2268: 2258: 2248: 2238: 2228: 2220:Muranjilepis 2218: 2208: 2198: 2188: 2178: 2168: 2158: 2148: 2140:Gyroptychius 2138: 2128: 2118: 2108: 2098: 2088: 2078: 1965: 1956: 1952:Osteichthyes 1947: 1842: 1838: 1795:(1): 79–85. 1792: 1788: 1760: 1756: 1754: 1568: 1553: 1533:Lissamphibia 1531:Total-group 1530: 1505: 1482: 1481: 1459: 1437: 1416:Ichthyostega 1414: 1413: 1392:Acanthostega 1390: 1389: 1366: 1365: 1344:Elginerpeton 1342: 1341: 1311:Elpistostege 1309: 1308: 1292: 1291: 1261: 1260: 1234: 1233: 1211: 1210: 1209: 1123: 1122: 1106: 1105: 1082: 1081: 1058: 1057: 1034: 1033: 1010: 1009: 986: 985: 907: 891: 890: 867: 866: 845:Gyroptychius 843: 842: 819: 818: 749: 748: 732: 731: 708: 707: 684: 683: 644: 621: 620: 597: 585: 580: 568: 565:phylogenetic 562: 548: 536: 535: 490: 488: 478: 473: 469:Ichthyostega 467: 461: 459: 454: 440: 436: 420: 410: 398:heterocercal 393: 392: 387: 371: 367: 357: 351: 347: 332: 311: 310: 309: 297: 296: 283:Swartz, 2012 276: 271: 255: 254: 234: 233: 214: 201: 188: 137: 35: 29: 4439:Wikispecies 4344:Wikispecies 4259:Romer's gap 4055:Pan-Amniota 4003:Crown group 3856:Papposaurus 3680:Baphetoidea 3536:Colosteidae 3522:Whatcheeria 3476:Sillerpeton 3267:Doragnathus 3227:Antlerpeton 2760:Spodichthys 2723:Litoptychus 2681:Eusthenodon 2674:Edenopteron 2667:Devonosteus 2573:Rhizodopsis 2536:Mahalalepis 2513:Cryptolepis 2460:Koharalepis 2446:Beelarongia 2422:Vorobjevaia 2400:Screbinodus 2393:Sauripterus 2386:Rossichthys 2358:Letognathus 2323:Archichthys 2300:Viluichthys 2170:Litoptychus 2110:Glyptopomus 2090:Chrysolepis 1970:Rhipidistia 1569:Other stem 1461:Colosteidae 1125:Eusthenodon 988:Spodichthys 751:Beelarongia 734:Koharalepis 489:Remains of 413:pelvic limb 378:Description 341:crown group 4489:Categories 3978:Eoherpeton 3827:Eoherpeton 3820:Eobaphetes 3760:Embolomeri 3712:Baphetidae 3337:Perittodus 3257:Diploradus 3143:Ventastega 3113:Tulerpeton 3082:Parmastega 3062:Jakubsonia 3042:Hynerpeton 2968:sensu lato 2894:see below↓ 2883:sensu lato 2842:Qikiqtania 2731:Mandageria 2716:Langlieria 2660:Canningius 2453:Canowindra 2407:Strepsodus 2260:Shirolepis 2230:Osteolepis 2160:Kenichthys 2100:Geptolepis 2080:Bogdanovia 2022:see below↓ 1767:References 1555:Embolomeri 1507:Baphetidae 1368:Ventastega 1108:Mandageria 869:Osteolepis 710:Canowindra 623:Kenichthys 545:Polynesian 529:specimen, 415:bones and 388:T. clackae 372:T. clackae 4430:Q20719461 4208:Rachitomi 4073:includes 4037:includes 4007:Tetrapoda 3971:Eldeceeon 3963:Casineria 3877:Pteroplax 3665:Nectridea 3653:Nectridea 3545:Colosteus 3507:Ossinodus 3391:Lethiscus 3371:Aistopoda 3347:Sigournea 3327:Ossirarus 3133:Umzantsia 3102:Tiktaalik 3092:Sinostega 2964:Tetrapoda 2879:Tetrapoda 2862:Tiktaalik 2812:Livoniana 2709:Jarvikina 2550:Megapomus 2290:Tungsenia 2240:Peregrina 2120:Gogonasus 1935:Kingdom: 1809:129313060 1785:Sam Giles 1519:Tetrapoda 1294:Tiktaalik 1060:Jarvikina 821:Gogonasus 594:cladogram 559:Phylogeny 543:, who in 515:limestone 344:Tetrapoda 249:Species: 168:Kingdom: 162:Eukaryota 47:387  4424:Wikidata 4329:Wikidata 4295:Category 4264:Timeline 3805:Cricotus 3776:Archeria 3728:Kyrinion 3721:Baphetes 3693:Eucritta 3515:Pederpes 3317:Occidens 3208:Devonian 3123:Tutusius 2982:Devonian 2528:Lohsania 2379:Rhizodus 2337:Barameda 2280:Thursius 2200:Medoevia 1943:Chordata 1941:Phylum: 1937:Animalia 1871:22448265 1839:PLOS ONE 1571:amniotes 893:Medoevia 527:holotype 519:mudstone 511:Givetian 451:fibulare 417:glenoids 290:Synonyms 240:Swartz, 182:Chordata 178:Phylum: 172:Animalia 158:Domain: 4459:8590577 4390:6375928 4377:1484753 4364:8607187 4350:Tinirau 4335:Q290682 4320:Tinirau 4075:Amniota 3783:Aversor 3735:Loxomma 3297:Koilops 3277:Gaiasia 2767:Tinirau 2702:Hyneria 2475:Owensia 2330:Aztecia 2190:Latvius 1862:3308997 1761:Tinirau 1212:Tinirau 610:  598:Tinirau 581:Tinirau 569:Tinirau 567:terms, 549:Tinirau 541:Tinirau 491:Tinirau 485:History 479:Tinirau 474:Tinirau 455:Tinirau 421:Tinirau 394:Tinirau 368:Tinirau 348:Tinirau 333:Tinirau 312:Tinirau 274:† 253:† 235:Tinirau 227:Genus: 116:↓ 4472:242676 4403:242675 4169:groups 3153:Ymeria 2351:Hongyu 1869:  1859:  1807:  1757:et al. 447:fibula 433:fibula 431:, and 362:. The 329:Nevada 4372:IRMNG 3942:Other 3205:Post- 1966:Clade 1957:Clade 1948:Clade 1805:S2CID 429:tibia 425:femur 406:pikes 335:is a 317:genus 215:Clade 202:Clade 189:Clade 4454:GBIF 4359:GBIF 4067:see 4031:see 3663:see 3211:taxa 2985:taxa 1999:see 1867:PMID 575:and 531:UCMP 517:and 466:and 364:type 242:2012 55:PreꞒ 1857:PMC 1847:doi 1797:doi 563:In 505:in 370:is 327:of 319:of 4491:: 4469:: 4456:: 4441:: 4426:: 4400:: 4387:: 4374:: 4361:: 4346:: 4331:: 1968:: 1959:: 1950:: 1865:. 1855:. 1841:. 1837:. 1817:^ 1803:. 1793:50 1791:. 1775:^ 600:: 427:, 303:? 217:: 204:: 191:: 105:Pg 49:Ma 44:, 4199:? 4193:? 4164:/ 4057:) 4053:( 4019:( 3966:? 3948:" 3944:" 3939:† 3902:† 3830:? 3808:" 3802:" 3757:† 3696:? 3677:† 3655:" 3651:" 3649:† 3577:† 3533:† 3510:? 3495:† 3368:† 3353:† 3343:† 3333:† 3323:† 3313:† 3303:† 3293:† 3283:† 3273:† 3263:† 3253:† 3243:† 3233:† 3223:† 3164:† 3149:† 3139:† 3129:† 3119:† 3109:† 3105:? 3098:† 3088:† 3078:† 3068:† 3058:† 3048:† 3038:† 3034:? 3027:† 3017:† 3007:† 2997:† 2970:) 2961:( 2885:) 2876:( 2858:† 2848:† 2838:† 2828:† 2818:† 2808:† 2804:? 2797:† 2741:? 2726:? 2627:† 2618:? 2611:† 2590:? 2561:? 2553:? 2531:? 2516:? 2487:† 2478:? 2470:? 2434:† 2425:? 2417:? 2311:† 2296:† 2286:† 2276:† 2266:† 2256:† 2246:† 2236:† 2226:† 2216:† 2206:† 2196:† 2186:† 2176:† 2166:† 2156:† 2146:† 2136:† 2126:† 2116:† 2106:† 2096:† 2086:† 2076:† 1918:) 1914:( 1904:e 1897:t 1890:v 1873:. 1849:: 1843:7 1811:. 1799:: 231:† 110:N 100:K 95:J 90:T 85:P 80:C 75:D 70:S 65:O 60:Ꞓ 20:)

Index

Tinirau (fish)
Middle Devonian
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Sarcopterygii
Tetrapodomorpha
Eotetrapodiformes
Tinirau
2012
Binomial name
Synonyms
genus
sarcopterygian

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