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Trapalcotherium

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separated by larger valleys than the labial ones. At the front of the tooth is a triangular structure consisting of the first lingual and the first two labial cusps. A low crest connects the first lingual to the first labial cusp and a stronger crest, separated from the first by a relatively shallow valley, connects the second lingual to the first labial cusp. Behind this structure, a second triangle is formed by two crests passing from the second lingual cusp to two cusps at the labial side (the back of the two is broken away, but its existence is presumed from the crown pattern). The front of these two crests is interrupted by a groove. The third lingual cusp is also connected to two crests, which encircle a small depression and presumably connected to one or more labial cusps, which are missing from the fossil.
49: 2290: 2313: 483: 1500: 331:; as a first molar because it is longer than wide; and as a left tooth because the left side (interpreted as labial, in the direction of the lips) bears more cusps than the right side (lingual, the direction of the tongue). The tooth is 2.48 mm long and 2.07 mm wide. Part of the back labial corner is missing. 334:
The lingual row contains three cusps and the labial probably five (the broken corner renders the number uncertain). Transverse ridges, separated by deep valleys, connect the lingual and labial cusps; therefore, the cusps are not strongly separate, but rather fused. The lingual cusps are larger and
219:
is therefore recognized as a member of the same family Ferugliotheriidae. Ferugliotheriidae is one of two families of gondwanatheres, an enigmatic group without close relationships to any living mammals.
275:—a group of primitive mammals that dominates the Late Cretaceous mammalian faunas of Argentina. The fauna was described in a 2009 paper by Guillermo Rougier and colleagues, who named 205:, one at the inner (lingual) side and the other at the outer (labial) side, which are connected by transverse ridges separated by deep valleys. This pattern is reminiscent of 2454: 380:; and the tips of the lingual cusps are more labially placed. The evolutionary affinities of gondwanatheres, which include the Ferugliotheriidae and the higher-crowned 2393: 2494: 372:; the valley behind the front triangle is less curved; the ridges attached to the second lingual cusp form another triangle; the tooth is relatively shorter; 2479: 2499: 2484: 2380: 2474: 2489: 517: 48: 2469: 2464: 271:(84–71 mya). The mammals from the Allen Formation are known from seven teeth, six of which represent four species of 2316: 280: 2289: 2459: 510: 245: 2449: 300: 2333: 256: 2419: 1746: 144: 1067: 396:
does not provide additional information that has a bearing on the relationships of the gondwanatheres.
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from the Los Alamitos Formation, the only previously known uncontroversial member of the family
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Rougier, G.W., Chornogubsky, L., Casadio, S., Paéz Arango, N. and Giallombardo, A. 2009.
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used by the paleontologists who collected the fossils from the Allen Formation.
272: 2356: 482: 267:(latest Cretaceous, about 71–66 million years ago, mya) and perhaps partly the 2108: 2060: 2010: 2003: 1773: 1725: 1418: 1404: 1382: 1368: 1361: 1299: 1189: 1182: 1175: 1038: 887: 740: 722: 709: 664: 526: 176: 2266: 1976: 1969: 1934: 1927: 1906: 1899: 1872: 1865: 1838: 1705: 1684: 1677: 1643: 1636: 1629: 1622: 1593: 1245: 1238: 1161: 1127: 864: 582: 351: 268: 60: 368:, but differs in some characters: the triangle at the front is narrower in 2350: 2081: 2052: 2024: 1913: 1879: 1780: 1375: 1333: 1120: 1052: 1011: 872: 610: 550: 389: 355: 80: 2385: 1948: 1920: 1732: 1558: 1285: 950: 894: 637: 556: 544: 201:
in age, and was first described in 2009. The tooth bears two rows of
90: 70: 2327: 299:"beast", commonly used to mean "mammal" in scientific names. The 308: 2331: 1515: 707: 537: 499: 495: 469:
Mammals from the Allen Formation, Late Cretaceous, Argentina
388:(a large group mainly known from the northern continents of 350:—a small and enigmatic group of mammals from Cretaceous and 376:
does not have the Y-shaped valleys between cusps seen in
471:(subscription required). Cretaceous Research 30:223–238. 392:) has repeatedly been proposed; the identification of 263:). All three are probably about equally old, from the 215:
mammal from similarly aged deposits in Argentina, and
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Rougier 14: 2500:Taxa named by Andres Giallombardo 2495:Taxa named by Natalia Paéz Arango 2485:Taxa named by Laura Chornogubsky 2312: 2311: 2288: 1498: 481: 291:where Cerro Tortuga is located, 47: 248:(rock units) that have yielded 1: 2475:Fossil taxa described in 2009 346:is identified as a member of 2490:Taxa named by Silvio Casadio 455:Rougier et al., 2009, p. 233 441:Rougier et al., 2009, p. 232 414:Rougier et al., 2009, p. 225 354:of the southern continents ( 175:is a fossil mammal from the 157:Trapalcotherium matuastensis 287:, combines the name of the 179:of Argentina in the family 2516: 2301: 2286: 2165: 2047: 1833: 1539: 1524: 1511: 1496: 999: 923: 761: 735: 731: 716: 703: 645: 618: 591: 576: 533: 323:is identified as a lower 228:The only known fossil of 150: 143: 44:Scientific classification 42: 23: 16:Extinct family of mammals 183:. The single species, 2420:Paleobiology Database 261:La Colonia Formations 224:Discovery and context 2470:Fossils of Argentina 2465:Cretaceous Argentina 319:The single tooth of 197:, which is probably 163:Rougier et al., 2009 136:T. matuastensis 123:Rougier et al., 2009 2158:Djadochtatherioidea 2230:Djadochtatheriidae 1792:Microcosmodontidae 238:Río Negro Province 2435: 2434: 2407:Open Tree of Life 2334:Taxon identifiers 2325: 2324: 2297: 2296: 2284: 2283: 2280: 2279: 2276: 2275: 2152: 2151: 2034: 2033: 1891:Neoplagiaulacidae 1507: 1506: 1494: 1493: 1490: 1489: 1486: 1485: 1462: 1461: 1262: 1261: 1068:Paulchoffatioidea 1062: 1061: 908: 907: 904: 903: 786:Ferugliotheriidae 699: 698: 695: 694: 691: 690: 687: 686: 683: 682: 386:multituberculates 366:Ferugliotheriidae 295:, with the Greek 193:. It is from the 181:Ferugliotheriidae 168: 167: 124: 104:Ferugliotheriidae 2507: 2428: 2427: 2415: 2414: 2402: 2401: 2389: 2388: 2376: 2375: 2374: 2361: 2360: 2359: 2329: 2315: 2314: 2292: 2246:Djadochtatherium 2163: 2045: 2040:Taeniolabidoidea 1831: 1537: 1526: 1513: 1502: 1430:Albionbaataridae 1273: 1146:Paulchoffatiidae 1077:Pinheirodontidae 1073: 997: 921: 915:Multituberculata 759: 733: 729: 718: 705: 643: 616: 589: 578: 571: 570: 535: 520: 513: 506: 497: 486:Data related to 485: 456: 453: 442: 439: 424: 421: 415: 412: 159: 155: 122: 115: 102: 52: 51: 29:Temporal range: 21: 2515: 2514: 2510: 2509: 2508: 2506: 2505: 2504: 2460:Allen Formation 2440: 2439: 2436: 2431: 2423: 2418: 2410: 2405: 2397: 2392: 2384: 2379: 2372:Trapalcotherium 2370: 2369: 2364: 2355: 2354: 2349: 2342:Trapalcotherium 2336: 2326: 2321: 2293: 2272: 2224: 2196:Sloanbaataridae 2190: 2148: 2127:Taeniolabididae 2121: 2087: 2030: 1982: 1885: 1851: 1820: 1786: 1758:Eucosmodontidae 1752: 1711: 1656: 1607: 1533: 1520: 1503: 1482: 1458: 1453:Proalbionbaatar 1424: 1305: 1268:Plagiaulacoidea 1258: 1218:Plesiochoffatia 1204:Meketichoffatia 1155:Bathmochoffatia 1140: 1058: 1024: 986: 917: 900: 858:Gondwanatherium 829: 824:Trapalcotherium 804:Ferugliotherium 780: 747: 725: 712: 679: 659: 632: 605: 585: 572: 540: 539: 529: 524: 489:Trapalcotherium 478: 464: 459: 454: 445: 440: 427: 422: 418: 413: 406: 402: 394:Trapalcotherium 378:Ferugliotherium 374:Trapalcotherium 370:Trapalcotherium 361:Ferugliotherium 344:Trapalcotherium 341: 321:Trapalcotherium 317: 285:Trapalcotherium 277:Trapalcotherium 250:Late Cretaceous 244:, one of three 242:Allen Formation 230:Trapalcotherium 226: 217:Trapalcotherium 208:Ferugliotherium 195:Allen Formation 186:T. matuastensis 172:Trapalcotherium 164: 161: 153: 152: 139: 121: 118:Trapalcotherium 113: 100: 46: 38: 31:Late Cretaceous 27: 25:Trapalcotherium 17: 12: 11: 5: 2513: 2511: 2503: 2502: 2497: 2492: 2487: 2482: 2477: 2472: 2467: 2462: 2457: 2452: 2450:Gondwanatheres 2442: 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976: 972:Gondwanatheria 968: 961: 954: 947: 940: 932: 924: 918: 913: 910: 909: 906: 905: 902: 901: 899: 898: 891: 884: 876: 869: 861: 854: 847: 839: 837: 831: 830: 828: 827: 820: 813: 807: 800: 791: 789: 782: 781: 779: 778: 770: 762: 756: 753:Gondwanatheria 749: 748: 746: 745: 736: 726: 721: 714: 713: 708: 701: 700: 697: 696: 693: 692: 689: 688: 685: 684: 681: 680: 678: 677: 676: 675: 669: 667: 661: 660: 658: 657: 656: 655: 646: 640: 634: 633: 631: 630: 629: 628: 619: 613: 607: 606: 604: 603: 602: 601: 592: 586: 581: 574: 573: 569: 568: 559: 553: 547: 538: 531: 530: 525: 523: 522: 515: 508: 500: 494: 493: 492:at Wikispecies 477: 476:External links 474: 473: 472: 463: 460: 458: 457: 443: 425: 416: 403: 401: 398: 348:Gondwanatheria 340: 337: 316: 313: 225: 222: 166: 165: 162: 148: 147: 141: 140: 132: 130: 126: 125: 111: 107: 106: 98: 94: 93: 88: 84: 83: 78: 74: 73: 68: 64: 63: 58: 54: 53: 40: 39: 28: 15: 13: 10: 9: 6: 4: 3: 2: 2512: 2501: 2498: 2496: 2493: 2491: 2488: 2486: 2483: 2481: 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1988:Ptilodontidae 1985: 1979: 1978: 1974: 1972: 1971: 1967: 1965: 1964: 1960: 1958: 1957: 1953: 1951: 1950: 1946: 1944: 1943: 1942:Neoplagiaulax 1939: 1937: 1936: 1932: 1930: 1929: 1925: 1923: 1922: 1918: 1916: 1915: 1911: 1909: 1908: 1904: 1902: 1901: 1897: 1896: 1894: 1892: 1888: 1882: 1881: 1877: 1875: 1874: 1870: 1868: 1867: 1863: 1862: 1860: 1858: 1854: 1848: 1847: 1843: 1841: 1840: 1836: 1835: 1832: 1829: 1827: 1823: 1817: 1816: 1815:Pentacosmodon 1812: 1810: 1809: 1808:Microcosmodon 1805: 1803: 1802: 1798: 1797: 1795: 1793: 1789: 1783: 1782: 1778: 1776: 1775: 1771: 1769: 1768: 1764: 1763: 1761: 1759: 1755: 1749: 1748: 1744: 1742: 1741: 1737: 1735: 1734: 1730: 1728: 1727: 1723: 1722: 1720: 1718: 1714: 1708: 1707: 1703: 1701: 1700: 1696: 1694: 1693: 1689: 1687: 1686: 1682: 1680: 1679: 1675: 1673: 1672: 1668: 1667: 1665: 1663: 1659: 1653: 1652: 1651:Paracimexomys 1648: 1646: 1645: 1641: 1639: 1638: 1634: 1632: 1631: 1627: 1625: 1624: 1620: 1619: 1617: 1614: 1613:Paracimexomys 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131: 128: 127: 120: 119: 112: 109: 108: 105: 99: 96: 95: 92: 89: 86: 85: 82: 79: 76: 75: 72: 69: 66: 65: 62: 59: 56: 55: 50: 45: 41: 36: 35:Maastrichtian 32: 26: 22: 19: 2437: 2341: 2265: 2260:Mangasbaatar 2258: 2253:Kryptobaatar 2251: 2244: 2239:Catopsbaatar 2237: 2217: 2212:Nessovbaatar 2210: 2205:Kamptobaatar 2203: 2185:Nemegtbaatar 2183: 2176: 2171:Bulganbaatar 2169: 2141: 2134: 2116:Sphenopsalis 2114: 2107: 2102:Lambdopsalis 2100: 2080: 2075:Valenopsalis 2073: 2066: 2059: 2051: 2023: 2016: 2009: 2002: 1995: 1975: 1968: 1963:Paressonodon 1961: 1956:Parectypodus 1954: 1947: 1940: 1933: 1926: 1919: 1912: 1905: 1898: 1878: 1871: 1864: 1844: 1837: 1813: 1806: 1799: 1779: 1772: 1765: 1745: 1738: 1731: 1724: 1704: 1697: 1692:Meniscoessus 1690: 1683: 1676: 1669: 1662:Cimolomyidae 1649: 1642: 1635: 1628: 1621: 1612: 1599: 1592: 1585: 1578: 1571: 1566:Corriebaatar 1564: 1557: 1550: 1543: 1451: 1444: 1439:Albionbaatar 1437: 1417: 1412:Tedoribaatar 1410: 1403: 1396: 1389: 1381: 1374: 1367: 1360: 1353: 1346: 1339: 1332: 1327:Dolichoprion 1325: 1318: 1311:Eobaataridae 1298: 1293:Morrisonodon 1291: 1284: 1251: 1244: 1237: 1230: 1223: 1216: 1209: 1202: 1195: 1188: 1181: 1174: 1169:Guimarotodon 1167: 1160: 1153: 1133: 1126: 1119: 1112: 1105: 1098: 1091: 1084: 1051: 1044: 1037: 1030:Allodontidae 1017: 1010: 1003: 978: 970: 963: 956: 949: 944:Hahnotherium 942: 934: 927: 893: 886: 878: 871: 863: 856: 849: 842: 823: 822: 817:Magallanodon 815: 802: 796:Argentodites 794: 774:Groeberiidae 772: 767:Adalatherium 765: 739: 566:Theriiformes 561: 488: 462:Bibliography 419: 393: 377: 373: 369: 359: 343: 342: 333: 320: 318: 305:matuastensis 304: 296: 284: 281:generic name 276: 273:dryolestoids 257:Los Alamitos 229: 227: 216: 206: 185: 184: 171: 170: 169: 156: 151: 135: 134: 117: 116: 24: 18: 2366:Wikispecies 2219:Sloanbaatar 2143:Taeniolabis 2018:Prochetodon 1846:Neoliotomus 1801:Acheronodon 1767:Clemensodon 1717:Kogaionidae 1671:Buginbaatar 1587:Uzbekbaatar 1552:Ameribaatar 1531:Cimolodonta 1518:Cimolodonta 1468:Cimolodonta 1391:Nokerbaatar 1320:Cheruscodon 1135:Pinheirodon 1114:Gerhardodon 1107:Ecprepaulax 1093:Bructerodon 1086:Bernardodon 1019:Zofiabaatar 1005:Arginbaatar 810:LACM 149371 315:Description 2444:Categories 2305:See also: 2109:Prionessus 2061:Catopsalis 2011:Kimbetohia 2004:Baiotomeus 1997:Aenigmamys 1774:Eucosmodon 1726:Barbatodon 1699:Paressodon 1477:see below↓ 1419:Teutonodon 1405:Sinobaatar 1383:Monobaatar 1369:Liaobaatar 1362:Indobaatar 1300:Plagiaulax 1190:Kuehneodon 1183:Kielanodon 1176:Henkelodon 1039:Ctenacodon 936:Baidabatyr 929:Argillomys 888:Sudamerica 741:Haramiyida 723:Allotheria 710:Allotheria 674:see below↓ 665:Allotheria 626:Cynodontia 611:Cynodontia 527:Allotheria 400:References 246:formations 177:Cretaceous 2267:Tombaatar 1977:Xyronomys 1970:Xanclomys 1935:Mimetodon 1928:Mesodmops 1907:Ectypodus 1900:Cernaysia 1873:Cimolodon 1866:Anconodon 1839:Filikomys 1740:Kogaionon 1706:Parikimys 1685:Essonodon 1678:Cimolomys 1644:Dakotamys 1637:Cimexomys 1630:Cedaromys 1623:Bryceomys 1594:Viridomys 1573:Fractinus 1545:Allocodon 1246:Sunnyodon 1239:Rugosodon 1232:Renatodon 1162:Galveodon 1128:Lavocatia 1100:Cantalera 965:Tashtykia 880:Patagonia 865:Greniodon 599:Synapsida 583:Synapsida 543:Kingdom: 352:Paleogene 269:Campanian 129:Species: 67:Kingdom: 61:Eukaryota 2357:Q7835549 2351:Wikidata 2317:Category 2082:Yubaatar 2053:Bubodens 2025:Ptilodus 1914:Krauseia 1880:Liotomus 1781:Stygimys 1376:Loxaulax 1334:Eobaatar 1121:Iberodon 1053:Psalodon 1046:Passumys 1012:Glirodon 873:Lavanify 653:Mammalia 638:Mammalia 557:Mammalia 551:Chordata 549:Phylum: 545:Animalia 390:Laurasia 356:Gondwana 97:Family: 91:Mammalia 81:Chordata 77:Phylum: 71:Animalia 57:Domain: 2412:4945628 2399:1478744 2386:4968806 1949:Nidimys 1921:Mesodma 1747:Litovoi 1733:Hainina 1580:Halodon 1559:Boffius 1355:Iberica 1286:Bolodon 958:Tagaria 951:Janumys 895:Vintana 555:Class: 297:therion 154:† 133:† 110:Genus: 87:Class: 2425:140936 2394:IRMNG 1615:group 562:Clade 329:cusps 325:molar 289:basin 203:cusps 191:molar 2381:GBIF 651:see 624:see 597:see 309:shed 259:and 211:, a 236:in 2446:: 2422:: 2409:: 2396:: 2383:: 2368:: 2353:: 564:: 446:^ 428:^ 407:^ 303:, 283:, 33:(? 2056:? 1604:? 1386:? 983:? 975:? 939:? 883:? 868:? 812:? 799:? 788:? 777:? 755:? 744:? 519:e 512:t 505:v 114:† 101:† 37:)

Index

Late Cretaceous
Maastrichtian
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Mammalia
Ferugliotheriidae
Trapalcotherium
Binomial name
Cretaceous
Ferugliotheriidae
molar
Allen Formation
Maastrichtian
cusps
Ferugliotherium
gondwanathere
Cerro Tortuga
Río Negro Province
Allen Formation
formations
Late Cretaceous
gondwanathere
Los Alamitos
La Colonia Formations
Maastrichtian
Campanian
dryolestoids

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