2656:
2662:
3212:
146:
120:
3242:
3218:
1776:
661:
Chasmosaurines evolved in western North
America (Laramidia). They are currently known definitively from rocks in western Canada, the western United States, and northern Mexico. They were highly diverse and among the most species-rich groups of dinosaurs, with new species frequently described. This
1320:" lineage with a pinched shut indentation in the heart-shaped frill margin. According to the cladogram below, the Triceratopsini may have derived from this later lineage. To improve resolution, certain species based on partial or immature remains (
1760:". Triceratopsins were the largest of the chasmosaurines; suggesting that gigantism had evolved in the Ceratopsidae once. In addition there is an evolutionary trend in the solidification of the frills, the most extreme being in
1311:
The findings of a phylogenetic study done by Fowler and
Freedman in 2020 are given below. The authors proposed that two distinct and roughly contemporaneous lineages of chasmosaurines existed in the late Cretaceous: a northern
662:
high diversity of named species is likely a result of the frill. The distinctive shape of the frill with the hornlets on its edges (epoccipitals) make it possible to recognize species from incomplete or fragmentary remains.
1336:
being misclassified, and possibly referable to other species pending further study. The authors also noted that some newer species included in the previous analysis by Mallon et al. (
1815:
gen. et sp. nov., a Boldly
Audacious New Chasmosaurine Ceratopsid (Dinosauria: Ornithischia) from the Judith River Formation (Upper Cretaceous: Campanian) of Montana, USA"
1332:
lineage," was not recovered by this revised analysis as they had predicted based on frill shape, the authors speculated that this may be due to some specimens included as
692:
were removed because it was found that they decrease resolution in the analysis because of the authors' new interpretation of epiparietal configurations.
651:
2655:
133:
1977:
2661:
3235:
1789:
654:. Broadly, the most distinguishing features of chasmosaurines are prominent brow horns and long frills lacking long spines;
145:
3211:
1543:
1009:
891:
658:
generally had short brow horns and relatively shorter frills, and often had long spines projecting from their frills.
1874:"Transitional evolutionary forms in chasmosaurine ceratopsid dinosaurs: evidence from the Campanian of New Mexico"
1756:
1648:
1517:
1444:
1083:
1059:
832:
763:
2068:
1970:
1576:
3342:
2186:
1427:
780:
3262:
1733:
1141:
915:
3309:
3245:
2095:
1750:
1624:
1377:
1158:
1042:
956:
671:
3217:
3337:
1826:
1744:
1600:
1316:" lineage with a heart-shaped frill margin that flattens and curls over onto itself, and a southern "
1233:
806:
676:
299:
273:
3300:
2351:
1963:
1500:
1328:) were excluded, as in the Mallon et al. study above. While this new study did not yield a single "
1216:
1124:
858:
674:
by Mallon et al., following the traditional epiparietal homology scheme from their description of
3071:
3021:
2453:
1809:
Jordan C. Mallon; Christopher J. Ott; Peter L. Larson; Edward M. Iuliano; David C. Evans (2016).
1394:
939:
593:
292:
140:
128:
3314:
3154:
3092:
3055:
2529:
2416:
1854:
3184:
2982:
2807:
2509:
2204:
1928:
1895:
1885:
1844:
1834:
1781:
1175:
378:
119:
3078:
3030:
2989:
2925:
2828:
2779:
2481:
2405:
2360:
2234:
2220:
2179:
2053:
2037:
521:
408:
338:
39:
1830:
3085:
3044:
2975:
2962:
2933:
2918:
2888:
2849:
2835:
2713:
2592:
2502:
2467:
2460:
2340:
2290:
2213:
2193:
1900:
1873:
1849:
1810:
1738:
1664:
1103:
699:
694:
655:
561:
510:
468:
438:
418:
269:
222:
3331:
3163:
3142:
3112:
3103:
3003:
2897:
2842:
2821:
2793:
2786:
2772:
2765:
2571:
2560:
2553:
2495:
2488:
2474:
2432:
2367:
2333:
2304:
2257:
688:
682:
541:
428:
398:
358:
348:
328:
318:
77:
3177:
3128:
3121:
3037:
3010:
2996:
2904:
2877:
2870:
2863:
2814:
2800:
2692:
2679:
2610:
2599:
2584:
2546:
2538:
2520:
2425:
2391:
2374:
2326:
2319:
2275:
2080:
2028:
632:
531:
489:
479:
458:
388:
368:
281:
251:
206:
1932:
1839:
3294:
3170:
3064:
2947:
2911:
2734:
2446:
2311:
581:
551:
52:
2940:
2856:
2706:
2439:
2227:
2161:
2148:
2107:
1986:
1771:
1192:
639:
571:
448:
238:
97:
62:
3285:
17:
643:
628:
157:
102:
46:
1858:
3279:
2741:
2727:
2720:
2283:
2019:
2010:
635:
602:
190:
177:
92:
87:
72:
67:
57:
3135:
2297:
2241:
1950:
1890:
107:
82:
2004:
167:
3256:
1919:, a giant horned dinosaur from the Late Campanian of New Mexico".
647:
3260:
2677:
2146:
1997:
1959:
1955:
1344:, etc.) had yet to be coded into their revised dataset.
642:
of their time. Chasmosaurines appeared in the early
577:
567:
557:
547:
537:
527:
517:
506:
495:
485:
475:
464:
454:
444:
434:
424:
414:
404:
394:
384:
374:
364:
354:
344:
334:
324:
314:
288:
3269:
3153:
3111:
3102:
3054:
3020:
2961:
2887:
2751:
2690:
2609:
2570:
2528:
2519:
2415:
2389:
2350:
2255:
2203:
2159:
2106:
2079:
2052:
638:. They were one of the most successful groups of
646:, and became extinct, along with all other non-
1971:
1742:, which he defined as "all species closer to
8:
3257:
3108:
2967:
2757:
2698:
2687:
2674:
2576:
2525:
2397:
2267:
2263:
2171:
2167:
2156:
2143:
2085:
2058:
2049:
1994:
1978:
1964:
1956:
118:
31:
1899:
1889:
1848:
1838:
1801:
1872:Fowler DW, Freedman Fowler EA (2020).
652:Cretaceous–Paleogene extinction event
134:Royal Tyrrell Museum of Palaeontology
7:
1951:http://paleodb.org/cgi-bin/bridge.pl
698:was not resolved as a member of the
1640:
1616:
1592:
1568:
1492:
1485:
1478:
1419:
1369:
1362:
1355:
1347:
1208:
1116:
1109:
1099:
1075:
1034:
931:
907:
883:
755:
748:
741:
734:
727:
720:
713:
705:
25:
3241:
3240:
3216:
3210:
2660:
2654:
1790:Timeline of ceratopsian research
1774:
144:
1736:in 2011 for the description of
27:Extinct subfamily of dinosaurs
1:
1933:10.1016/j.cretres.2010.12.007
1840:10.1371/journal.pone.0154218
1732:Triceratopsini was named by
1544:Coahuilaceratops magnacuerna
1010:Coahuilaceratops magnacuerna
608:Marsh, 1888 sensu Abel, 1919
892:Agujaceratops mariscalensis
3359:
3230:
3208:
2970:
2760:
2701:
2686:
2673:
2651:
2579:
2400:
2270:
2266:
2174:
2170:
2155:
2142:
2088:
2061:
2048:
1993:
1757:Arrhinoceratops brachyops
1662:
1649:Arrhinoceratops brachyops
1645:
1638:
1621:
1614:
1597:
1590:
1573:
1566:
1540:
1518:Pentaceratops sternbergii
1514:
1497:
1490:
1483:
1476:
1445:Kosmoceratops richardsoni
1441:
1424:
1417:
1391:
1374:
1367:
1360:
1353:
1334:Pentaceratops sternbergii
1230:
1213:
1206:
1189:
1172:
1155:
1138:
1121:
1114:
1107:
1097:
1084:Arrhinoceratops brachyops
1080:
1073:
1060:Regaliceratops peterhewsi
1056:
1039:
1032:
1006:
953:
936:
929:
912:
905:
888:
881:
855:
833:Pentaceratops sternbergii
829:
803:
777:
764:Kosmoceratops richardsoni
760:
753:
746:
739:
732:
725:
718:
711:
670:Below is the result of a
599:
592:
311:
306:
287:
280:
141:Scientific classification
139:
126:
117:
34:
1915:Longrich, N.R. (2011). "
1577:Navajoceratops sullivani
2187:Micropachycephalosaurus
1428:Vagaceratops irvinensis
781:Vagaceratops irvinensis
1917:Titanoceratops ouranos
1193:"Torosaurus" utahensis
1142:Titanoceratops ouranos
916:Mojoceratops perifania
650:dinosaurs, during the
3310:Paleobiology Database
1751:Anchiceratops ornatus
1625:Anchiceratops ornatus
1378:Chasmosaurus russelli
1159:Nedoceratops hatcheri
1043:Anchiceratops ornatus
957:Chasmosaurus russelli
672:phylogenetic analysis
1813:Spiclypeus shipporum
1745:Triceratops horridus
1734:Nicholas R. Longrich
1601:Terminocavus sealeyi
1234:Triceratops horridus
807:Spiclypeus shipporum
677:Spiclypeus shipporum
2352:Archaeoceratopsidae
1921:Cretaceous Research
1831:2016PLoSO..1154218M
1501:Utahceratops gettyi
1217:Triceratops prorsus
1125:Ojoceratops fowleri
859:Utahceratops gettyi
3072:Crittendenceratops
3022:Albertaceratopsini
1891:10.7717/peerj.9251
1395:Chasmosaurus belli
940:Chasmosaurus belli
293:Chasmosaurus belli
129:Chasmosaurus belli
3325:
3324:
3263:Taxon identifiers
3254:
3253:
3226:
3225:
3206:
3205:
3202:
3201:
3198:
3197:
3194:
3193:
3155:Pachyrhinosaurini
3093:Yehuecauhceratops
3056:Nasutoceratopsini
2957:
2956:
2669:
2668:
2649:
2648:
2645:
2644:
2641:
2640:
2637:
2636:
2633:
2632:
2629:
2628:
2530:Protoceratopsidae
2417:Leptoceratopsidae
2385:
2384:
2251:
2250:
2138:
2137:
2134:
2133:
2130:
2129:
2126:
2125:
1729:
1728:
1720:
1719:
1711:
1710:
1702:
1701:
1693:
1692:
1684:
1683:
1675:
1674:
1555:
1554:
1529:
1528:
1465:
1464:
1456:
1455:
1406:
1405:
1308:
1307:
1299:
1298:
1290:
1289:
1281:
1280:
1272:
1271:
1263:
1262:
1254:
1253:
1245:
1244:
1021:
1020:
995:
994:
986:
985:
977:
976:
968:
967:
870:
869:
844:
843:
818:
817:
792:
791:
622:
621:
616:
609:
503:
276:
16:(Redirected from
3350:
3318:
3317:
3305:
3304:
3303:
3290:
3289:
3288:
3258:
3244:
3243:
3221:
3220:
3214:
3185:Pachyrhinosaurus
3109:
2983:Machairoceratops
2968:
2808:Coahuilaceratops
2758:
2699:
2688:
2675:
2664:
2659:
2658:
2577:
2526:
2510:Zhuchengceratops
2398:
2268:
2264:
2205:Chaoyangsauridae
2172:
2168:
2157:
2144:
2086:
2059:
2050:
2043:
2042:
1995:
1980:
1973:
1966:
1957:
1937:
1936:
1912:
1906:
1905:
1903:
1893:
1869:
1863:
1862:
1852:
1842:
1806:
1784:
1782:Dinosaurs portal
1779:
1778:
1777:
1641:
1617:
1593:
1569:
1493:
1486:
1479:
1420:
1370:
1363:
1356:
1348:
1209:
1176:Torosaurus latus
1117:
1110:
1100:
1076:
1035:
932:
908:
884:
756:
749:
742:
735:
728:
721:
714:
706:
614:
607:
579:
569:
559:
549:
539:
529:
519:
508:
501:
497:
487:
477:
466:
456:
446:
436:
426:
416:
406:
396:
386:
379:Coahuilaceratops
376:
366:
356:
346:
336:
326:
316:
290:
268:
263:
250:
237:
221:
205:
149:
148:
122:
112:
49:
38:Temporal range:
32:
21:
3358:
3357:
3353:
3352:
3351:
3349:
3348:
3347:
3328:
3327:
3326:
3321:
3313:
3308:
3299:
3298:
3293:
3284:
3283:
3278:
3265:
3255:
3250:
3222:
3215:
3190:
3149:
3098:
3079:Furcatoceratops
3050:
3031:Albertaceratops
3016:
2990:Menefeeceratops
2953:
2926:Tatankaceratops
2883:
2829:Mercuriceratops
2780:Arrhinoceratops
2747:
2695:
2682:
2665:
2653:
2625:
2605:
2566:
2515:
2482:Montanoceratops
2411:
2406:Graciliceratops
2394:
2381:
2361:Archaeoceratops
2346:
2260:
2247:
2235:Xuanhuaceratops
2221:Hualianceratops
2199:
2180:Albalophosaurus
2164:
2151:
2122:
2102:
2075:
2069:Avemetatarsalia
2054:Avemetatarsalia
2044:
2038:Marginocephalia
2000:
1999:
1989:
1984:
1946:
1941:
1940:
1914:
1913:
1909:
1871:
1870:
1866:
1825:(5): e0154218.
1808:
1807:
1803:
1798:
1780:
1775:
1773:
1770:
1730:
1721:
1712:
1703:
1694:
1685:
1676:
1556:
1530:
1466:
1457:
1407:
1351:Chasmosaurinae
1309:
1300:
1291:
1282:
1273:
1264:
1255:
1246:
1022:
996:
987:
978:
969:
871:
845:
819:
793:
709:Chasmosaurinae
668:
613:
612:Eoceratopsinae
606:
522:Tatankaceratops
409:Mercuriceratops
339:Arrhinoceratops
302:
296:
267:
261:
248:
235:
219:
203:
143:
113:
111:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
44:
43:
40:Late Cretaceous
36:
28:
23:
22:
15:
12:
11:
5:
3356:
3354:
3346:
3345:
3343:Chasmosaurines
3340:
3330:
3329:
3323:
3322:
3320:
3319:
3306:
3301:Chasmosaurinae
3291:
3275:
3273:
3271:Chasmosaurinae
3267:
3266:
3261:
3252:
3251:
3249:
3248:
3238:
3231:
3228:
3227:
3224:
3223:
3209:
3207:
3204:
3203:
3200:
3199:
3196:
3195:
3192:
3191:
3189:
3188:
3181:
3174:
3167:
3159:
3157:
3151:
3150:
3148:
3147:
3139:
3132:
3125:
3117:
3115:
3106:
3100:
3099:
3097:
3096:
3089:
3086:Nasutoceratops
3082:
3075:
3068:
3060:
3058:
3052:
3051:
3049:
3048:
3045:Medusaceratops
3041:
3034:
3026:
3024:
3018:
3017:
3015:
3014:
3007:
3000:
2993:
2986:
2979:
2976:Diabloceratops
2971:
2965:
2963:Centrosaurinae
2959:
2958:
2955:
2954:
2952:
2951:
2944:
2937:
2934:Titanoceratops
2930:
2922:
2919:Regaliceratops
2915:
2908:
2901:
2893:
2891:
2889:Triceratopsini
2885:
2884:
2882:
2881:
2874:
2867:
2860:
2853:
2850:Sierraceratops
2846:
2839:
2836:Navajoceratops
2832:
2825:
2818:
2811:
2804:
2797:
2790:
2783:
2776:
2769:
2761:
2755:
2753:Chasmosaurinae
2749:
2748:
2746:
2745:
2738:
2731:
2724:
2717:
2714:Brachyceratops
2710:
2702:
2696:
2691:
2684:
2683:
2678:
2671:
2670:
2667:
2666:
2652:
2650:
2647:
2646:
2643:
2642:
2639:
2638:
2635:
2634:
2631:
2630:
2627:
2626:
2624:
2623:
2622:
2621:
2615:
2613:
2607:
2606:
2604:
2603:
2596:
2593:Turanoceratops
2589:
2580:
2574:
2568:
2567:
2565:
2564:
2557:
2550:
2543:
2534:
2532:
2523:
2517:
2516:
2514:
2513:
2506:
2503:Unescoceratops
2499:
2492:
2485:
2478:
2471:
2468:Ischioceratops
2464:
2461:Gryphoceratops
2457:
2450:
2443:
2436:
2429:
2421:
2419:
2413:
2412:
2410:
2409:
2401:
2395:
2390:
2387:
2386:
2383:
2382:
2380:
2379:
2371:
2364:
2356:
2354:
2348:
2347:
2345:
2344:
2341:Sasayamagnomus
2337:
2330:
2323:
2316:
2308:
2301:
2294:
2291:Auroraceratops
2287:
2280:
2271:
2261:
2256:
2253:
2252:
2249:
2248:
2246:
2245:
2238:
2231:
2224:
2217:
2214:Chaoyangsaurus
2209:
2207:
2201:
2200:
2198:
2197:
2194:Psittacosaurus
2190:
2183:
2175:
2165:
2160:
2153:
2152:
2147:
2140:
2139:
2136:
2135:
2132:
2131:
2128:
2127:
2124:
2123:
2121:
2120:
2119:
2118:
2112:
2110:
2104:
2103:
2101:
2100:
2099:
2098:
2089:
2083:
2077:
2076:
2074:
2073:
2072:
2071:
2062:
2056:
2046:
2045:
2041:
2040:
2031:
2022:
2013:
2007:
1998:
1991:
1990:
1985:
1983:
1982:
1975:
1968:
1960:
1954:
1953:
1945:
1944:External links
1942:
1939:
1938:
1927:(3): 264–276.
1907:
1864:
1800:
1799:
1797:
1794:
1793:
1792:
1786:
1785:
1769:
1766:
1739:Titanoceratops
1727:
1726:
1723:
1722:
1718:
1717:
1714:
1713:
1709:
1708:
1705:
1704:
1700:
1699:
1696:
1695:
1691:
1690:
1687:
1686:
1682:
1681:
1678:
1677:
1673:
1672:
1669:
1668:
1665:Triceratopsini
1661:
1658:
1657:
1654:
1653:
1644:
1639:
1637:
1634:
1633:
1630:
1629:
1620:
1615:
1613:
1610:
1609:
1606:
1605:
1596:
1591:
1589:
1586:
1585:
1582:
1581:
1572:
1567:
1565:
1562:
1561:
1558:
1557:
1553:
1552:
1549:
1548:
1539:
1536:
1535:
1532:
1531:
1527:
1526:
1523:
1522:
1513:
1510:
1509:
1506:
1505:
1496:
1491:
1489:
1484:
1482:
1477:
1475:
1472:
1471:
1468:
1467:
1463:
1462:
1459:
1458:
1454:
1453:
1450:
1449:
1440:
1437:
1436:
1433:
1432:
1423:
1418:
1416:
1413:
1412:
1409:
1408:
1404:
1403:
1400:
1399:
1390:
1387:
1386:
1383:
1382:
1373:
1368:
1366:
1361:
1359:
1354:
1352:
1346:
1342:Regaliceratops
1306:
1305:
1302:
1301:
1297:
1296:
1293:
1292:
1288:
1287:
1284:
1283:
1279:
1278:
1275:
1274:
1270:
1269:
1266:
1265:
1261:
1260:
1257:
1256:
1252:
1251:
1248:
1247:
1243:
1242:
1239:
1238:
1229:
1226:
1225:
1222:
1221:
1212:
1207:
1205:
1202:
1201:
1198:
1197:
1188:
1185:
1184:
1181:
1180:
1171:
1168:
1167:
1164:
1163:
1154:
1151:
1150:
1147:
1146:
1137:
1134:
1133:
1130:
1129:
1120:
1115:
1113:
1108:
1106:
1104:Triceratopsini
1098:
1096:
1093:
1092:
1089:
1088:
1079:
1074:
1072:
1069:
1068:
1065:
1064:
1055:
1052:
1051:
1048:
1047:
1038:
1033:
1031:
1028:
1027:
1024:
1023:
1019:
1018:
1015:
1014:
1005:
1002:
1001:
998:
997:
993:
992:
989:
988:
984:
983:
980:
979:
975:
974:
971:
970:
966:
965:
962:
961:
952:
949:
948:
945:
944:
935:
930:
928:
925:
924:
921:
920:
911:
906:
904:
901:
900:
897:
896:
887:
882:
880:
877:
876:
873:
872:
868:
867:
864:
863:
854:
851:
850:
847:
846:
842:
841:
838:
837:
828:
825:
824:
821:
820:
816:
815:
812:
811:
802:
799:
798:
795:
794:
790:
789:
786:
785:
776:
773:
772:
769:
768:
759:
754:
752:
747:
745:
740:
738:
733:
731:
726:
724:
719:
717:
712:
710:
704:
700:Triceratopsini
695:Regaliceratops
667:
666:Classification
664:
656:centrosaurines
625:Chasmosaurinae
620:
619:
618:
617:
610:
597:
596:
590:
589:
588:
587:
586:
585:
575:
565:
562:Regaliceratops
555:
545:
535:
525:
515:
511:Titanoceratops
499:Triceratopsini
493:
483:
473:
469:Titanoceratops
462:
452:
442:
439:Sierraceratops
432:
422:
419:Navajoceratops
412:
402:
392:
382:
372:
362:
352:
342:
332:
322:
309:
308:
304:
303:
297:
285:
284:
278:
277:
265:Chasmosaurinae
259:
255:
254:
246:
242:
241:
233:
226:
225:
223:Neornithischia
217:
210:
209:
201:
194:
193:
188:
181:
180:
175:
171:
170:
165:
161:
160:
155:
151:
150:
137:
136:
124:
123:
115:
114:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
50:
37:
35:Chasmosaurines
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
3355:
3344:
3341:
3339:
3336:
3335:
3333:
3316:
3311:
3307:
3302:
3296:
3292:
3287:
3281:
3277:
3276:
3274:
3272:
3268:
3264:
3259:
3247:
3239:
3237:
3233:
3232:
3229:
3219:
3213:
3187:
3186:
3182:
3180:
3179:
3175:
3173:
3172:
3168:
3166:
3165:
3164:Achelousaurus
3161:
3160:
3158:
3156:
3152:
3145:
3144:
3143:Styracosaurus
3140:
3138:
3137:
3133:
3131:
3130:
3126:
3124:
3123:
3119:
3118:
3116:
3114:
3113:Centrosaurini
3110:
3107:
3105:
3104:Eucentrosaura
3101:
3095:
3094:
3090:
3088:
3087:
3083:
3081:
3080:
3076:
3074:
3073:
3069:
3067:
3066:
3062:
3061:
3059:
3057:
3053:
3047:
3046:
3042:
3040:
3039:
3035:
3033:
3032:
3028:
3027:
3025:
3023:
3019:
3013:
3012:
3008:
3006:
3005:
3004:Wendiceratops
3001:
2999:
2998:
2994:
2992:
2991:
2987:
2985:
2984:
2980:
2978:
2977:
2973:
2972:
2969:
2966:
2964:
2960:
2950:
2949:
2945:
2943:
2942:
2938:
2936:
2935:
2931:
2928:
2927:
2923:
2921:
2920:
2916:
2914:
2913:
2909:
2907:
2906:
2902:
2900:
2899:
2898:Eotriceratops
2895:
2894:
2892:
2890:
2886:
2880:
2879:
2875:
2873:
2872:
2868:
2866:
2865:
2861:
2859:
2858:
2854:
2852:
2851:
2847:
2845:
2844:
2843:Pentaceratops
2840:
2838:
2837:
2833:
2831:
2830:
2826:
2824:
2823:
2822:Kosmoceratops
2819:
2817:
2816:
2812:
2810:
2809:
2805:
2803:
2802:
2798:
2796:
2795:
2794:Bravoceratops
2791:
2789:
2788:
2787:Bisticeratops
2784:
2782:
2781:
2777:
2775:
2774:
2773:Anchiceratops
2770:
2768:
2767:
2766:Agujaceratops
2763:
2762:
2759:
2756:
2754:
2750:
2744:
2743:
2739:
2737:
2736:
2732:
2730:
2729:
2725:
2723:
2722:
2718:
2716:
2715:
2711:
2709:
2708:
2704:
2703:
2700:
2697:
2694:
2689:
2685:
2681:
2676:
2672:
2663:
2657:
2619:
2618:
2617:
2616:
2614:
2612:
2608:
2602:
2601:
2597:
2595:
2594:
2590:
2587:
2586:
2582:
2581:
2578:
2575:
2573:
2572:Ceratopsoidea
2569:
2563:
2562:
2561:Protoceratops
2558:
2556:
2555:
2554:Breviceratops
2551:
2549:
2548:
2544:
2541:
2540:
2536:
2535:
2533:
2531:
2527:
2524:
2522:
2518:
2512:
2511:
2507:
2505:
2504:
2500:
2498:
2497:
2496:Udanoceratops
2493:
2491:
2490:
2489:Prenoceratops
2486:
2484:
2483:
2479:
2477:
2476:
2475:Leptoceratops
2472:
2470:
2469:
2465:
2463:
2462:
2458:
2456:
2455:
2451:
2449:
2448:
2444:
2442:
2441:
2437:
2435:
2434:
2433:Bainoceratops
2430:
2428:
2427:
2423:
2422:
2420:
2418:
2414:
2408:
2407:
2403:
2402:
2399:
2396:
2393:
2388:
2377:
2376:
2372:
2370:
2369:
2368:Helioceratops
2365:
2363:
2362:
2358:
2357:
2355:
2353:
2349:
2343:
2342:
2338:
2336:
2335:
2334:Mosaiceratops
2331:
2329:
2328:
2324:
2322:
2321:
2317:
2314:
2313:
2309:
2307:
2306:
2305:Koreaceratops
2302:
2300:
2299:
2295:
2293:
2292:
2288:
2286:
2285:
2281:
2278:
2277:
2273:
2272:
2269:
2265:
2262:
2259:
2258:Neoceratopsia
2254:
2244:
2243:
2239:
2237:
2236:
2232:
2230:
2229:
2225:
2223:
2222:
2218:
2216:
2215:
2211:
2210:
2208:
2206:
2202:
2196:
2195:
2191:
2189:
2188:
2184:
2182:
2181:
2177:
2176:
2173:
2169:
2166:
2163:
2158:
2154:
2150:
2145:
2141:
2116:
2115:
2114:
2113:
2111:
2109:
2105:
2097:
2093:
2092:
2091:
2090:
2087:
2084:
2082:
2078:
2070:
2066:
2065:
2064:
2063:
2060:
2057:
2055:
2051:
2047:
2039:
2035:
2032:
2030:
2026:
2023:
2021:
2017:
2014:
2012:
2008:
2006:
2002:
2001:
1996:
1992:
1988:
1981:
1976:
1974:
1969:
1967:
1962:
1961:
1958:
1952:
1948:
1947:
1943:
1934:
1930:
1926:
1922:
1918:
1911:
1908:
1902:
1897:
1892:
1887:
1883:
1879:
1875:
1868:
1865:
1860:
1856:
1851:
1846:
1841:
1836:
1832:
1828:
1824:
1820:
1816:
1814:
1805:
1802:
1795:
1791:
1788:
1787:
1783:
1772:
1767:
1765:
1763:
1759:
1758:
1753:
1752:
1747:
1746:
1741:
1740:
1735:
1725:
1724:
1716:
1715:
1707:
1706:
1698:
1697:
1689:
1688:
1680:
1679:
1671:
1670:
1667:
1666:
1660:
1659:
1656:
1655:
1652:
1651:
1650:
1643:
1642:
1636:
1635:
1632:
1631:
1628:
1627:
1626:
1619:
1618:
1612:
1611:
1608:
1607:
1604:
1603:
1602:
1595:
1594:
1588:
1587:
1584:
1583:
1580:
1579:
1578:
1571:
1570:
1564:
1563:
1560:
1559:
1551:
1550:
1547:
1546:
1545:
1538:
1537:
1534:
1533:
1525:
1524:
1521:
1520:
1519:
1512:
1511:
1508:
1507:
1504:
1503:
1502:
1495:
1494:
1488:
1487:
1481:
1480:
1474:
1473:
1470:
1469:
1461:
1460:
1452:
1451:
1448:
1447:
1446:
1439:
1438:
1435:
1434:
1431:
1430:
1429:
1422:
1421:
1415:
1414:
1411:
1410:
1402:
1401:
1398:
1397:
1396:
1389:
1388:
1385:
1384:
1381:
1380:
1379:
1372:
1371:
1365:
1364:
1358:
1357:
1350:
1349:
1345:
1343:
1339:
1335:
1331:
1330:Pentaceratops
1327:
1326:Agujaceratops
1323:
1322:Bravoceratops
1319:
1318:Pentaceratops
1315:
1304:
1303:
1295:
1294:
1286:
1285:
1277:
1276:
1268:
1267:
1259:
1258:
1250:
1249:
1241:
1240:
1237:
1236:
1235:
1228:
1227:
1224:
1223:
1220:
1219:
1218:
1211:
1210:
1204:
1203:
1200:
1199:
1196:
1195:
1194:
1187:
1186:
1183:
1182:
1179:
1178:
1177:
1170:
1169:
1166:
1165:
1162:
1161:
1160:
1153:
1152:
1149:
1148:
1145:
1144:
1143:
1136:
1135:
1132:
1131:
1128:
1127:
1126:
1119:
1118:
1112:
1111:
1105:
1102:
1101:
1095:
1094:
1091:
1090:
1087:
1086:
1085:
1078:
1077:
1071:
1070:
1067:
1066:
1063:
1062:
1061:
1054:
1053:
1050:
1049:
1046:
1045:
1044:
1037:
1036:
1030:
1029:
1026:
1025:
1017:
1016:
1013:
1012:
1011:
1004:
1003:
1000:
999:
991:
990:
982:
981:
973:
972:
964:
963:
960:
959:
958:
951:
950:
947:
946:
943:
942:
941:
934:
933:
927:
926:
923:
922:
919:
918:
917:
910:
909:
903:
902:
899:
898:
895:
894:
893:
886:
885:
879:
878:
875:
874:
866:
865:
862:
861:
860:
853:
852:
849:
848:
840:
839:
836:
835:
834:
827:
826:
823:
822:
814:
813:
810:
809:
808:
801:
800:
797:
796:
788:
787:
784:
783:
782:
775:
774:
771:
770:
767:
766:
765:
758:
757:
751:
750:
744:
743:
737:
736:
730:
729:
723:
722:
716:
715:
708:
707:
703:
701:
697:
696:
691:
690:
689:Eotriceratops
685:
684:
683:Bravoceratops
679:
678:
673:
665:
663:
659:
657:
653:
649:
645:
641:
637:
634:
630:
626:
611:
604:
601:
600:
598:
595:
591:
584:
583:
576:
574:
573:
566:
564:
563:
556:
554:
553:
546:
544:
543:
542:Eotriceratops
536:
534:
533:
526:
524:
523:
516:
513:
512:
505:
504:
500:
494:
492:
491:
484:
482:
481:
474:
471:
470:
463:
461:
460:
453:
451:
450:
443:
441:
440:
433:
431:
430:
429:Pentaceratops
423:
421:
420:
413:
411:
410:
403:
401:
400:
399:Kosmoceratops
393:
391:
390:
383:
381:
380:
373:
371:
370:
363:
361:
360:
359:Bravoceratops
353:
351:
350:
349:Bisticeratops
343:
341:
340:
333:
331:
330:
329:Anchiceratops
323:
321:
320:
319:Agujaceratops
313:
312:
310:
305:
301:
295:
294:
286:
283:
279:
275:
271:
266:
260:
257:
256:
253:
247:
244:
243:
240:
234:
231:
228:
227:
224:
218:
215:
212:
211:
208:
202:
199:
196:
195:
192:
189:
186:
183:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
147:
142:
138:
135:
131:
130:
125:
121:
116:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
48:
41:
33:
30:
19:
18:Triceratopsin
3270:
3183:
3178:Stellasaurus
3176:
3169:
3162:
3141:
3134:
3129:Coronosaurus
3127:
3122:Centrosaurus
3120:
3091:
3084:
3077:
3070:
3063:
3043:
3038:Lokiceratops
3036:
3029:
3011:Xenoceratops
3009:
3002:
2997:Sinoceratops
2995:
2988:
2981:
2974:
2946:
2939:
2932:
2924:
2917:
2910:
2905:Nedoceratops
2903:
2896:
2878:Vagaceratops
2876:
2871:Utahceratops
2869:
2864:Terminocavus
2862:
2855:
2848:
2841:
2834:
2827:
2820:
2815:Judiceratops
2813:
2806:
2801:Chasmosaurus
2799:
2792:
2785:
2778:
2771:
2764:
2752:
2740:
2733:
2726:
2719:
2712:
2705:
2693:Ceratopsidae
2680:Ceratopsidae
2611:Ceratopsidae
2600:Zuniceratops
2598:
2591:
2585:Ajkaceratops
2583:
2559:
2552:
2547:Bagaceratops
2545:
2539:Ajkaceratops
2537:
2521:Coronosauria
2508:
2501:
2494:
2487:
2480:
2473:
2466:
2459:
2452:
2445:
2438:
2431:
2426:Asiaceratops
2424:
2404:
2392:Euceratopsia
2375:Yamaceratops
2373:
2366:
2359:
2339:
2332:
2327:Microceratus
2325:
2320:Liaoceratops
2318:
2310:
2303:
2296:
2289:
2282:
2276:Ajkaceratops
2274:
2240:
2233:
2226:
2219:
2212:
2192:
2185:
2178:
2096:Ornithischia
2081:Ornithischia
2033:
2029:Ornithischia
2024:
2015:
1924:
1920:
1916:
1910:
1881:
1877:
1867:
1822:
1818:
1812:
1804:
1761:
1755:
1749:
1743:
1737:
1731:
1663:
1647:
1646:
1623:
1622:
1599:
1598:
1575:
1574:
1542:
1541:
1516:
1515:
1499:
1498:
1443:
1442:
1426:
1425:
1393:
1392:
1376:
1375:
1341:
1337:
1333:
1329:
1325:
1321:
1317:
1314:Chasmosaurus
1313:
1310:
1232:
1231:
1215:
1214:
1191:
1190:
1174:
1173:
1157:
1156:
1140:
1139:
1123:
1122:
1082:
1081:
1058:
1057:
1041:
1040:
1008:
1007:
955:
954:
938:
937:
914:
913:
890:
889:
857:
856:
831:
830:
805:
804:
779:
778:
762:
761:
693:
687:
681:
675:
669:
660:
624:
623:
603:Ceratopsinae
580:
570:
560:
550:
540:
532:Nedoceratops
530:
520:
509:
498:
490:Vagaceratops
488:
480:Utahceratops
478:
467:
459:Terminocavus
457:
447:
437:
427:
417:
407:
397:
389:Judiceratops
387:
377:
369:Chasmosaurus
367:
357:
347:
337:
327:
317:
291:
282:Type species
264:
252:Ceratopsidae
229:
213:
207:Ornithischia
197:
184:
127:
29:
3338:Ceratopsids
3295:Wikispecies
3171:Einiosaurus
3065:Avaceratops
2948:Triceratops
2912:Ojoceratops
2735:Monoclonius
2447:Ferrisaurus
2312:Kulceratops
1762:Triceratops
615:Lambe, 1915
582:Triceratops
552:Ojoceratops
502:Marsh, 1889
258:Subfamily:
45:78–66
3332:Categories
3234:See also:
2941:Torosaurus
2857:Spiclypeus
2707:Agathaumas
2620:see below↓
2440:Cerasinops
2228:Stenopelix
2162:Ceratopsia
2149:Ceratopsia
2117:see below↓
2108:Ceratopsia
2020:Dinosauria
1987:Ceratopsia
1796:References
1338:Spiclypeus
640:herbivores
633:ceratopsid
572:Torosaurus
449:Spiclypeus
307:Subgroups
239:Ceratopsia
191:Dinosauria
132:skeleton,
2003:Kingdom:
1884:. e9251.
644:Campanian
636:dinosaurs
629:subfamily
164:Kingdom:
158:Eukaryota
3280:Wikidata
3246:Category
3236:Timeline
2742:Polyonax
2728:Dysganus
2721:Ceratops
2284:Aquilops
2011:Chordata
2009:Phylum:
2005:Animalia
1859:27191389
1819:PLOS ONE
1768:See also
1748:than to
594:Synonyms
245:Family:
178:Chordata
174:Phylum:
168:Animalia
154:Domain:
3286:Q137441
3136:Spinops
2454:Gremlin
2242:Yinlong
1901:7278894
1850:4871577
1827:Bibcode
298:Lambe,
1898:
1857:
1847:
3315:52773
2034:Clade
2025:Clade
2016:Clade
1878:PeerJ
648:avian
627:is a
270:Lambe
230:Clade
214:Clade
198:Clade
185:Clade
2094:see
2067:see
1855:PMID
1324:and
686:and
300:1902
274:1915
53:PreꞒ
2298:Beg
1929:doi
1896:PMC
1886:doi
1845:PMC
1835:doi
1754:or
631:of
3334::
3312::
3297::
3282::
2036::
2027::
2018::
1949:1.
1925:32
1923:.
1894:.
1880:.
1876:.
1853:.
1843:.
1833:.
1823:11
1821:.
1817:.
1764:.
1340:,
702:.
680:.
605:?
272:,
232::
216::
200::
187::
103:Pg
47:Ma
42:,
3146:?
2929:?
2588:?
2542:?
2378:?
2315:?
2279:?
1979:e
1972:t
1965:v
1935:.
1931::
1904:.
1888::
1882:8
1861:.
1837::
1829::
1811:"
1312:"
578:†
568:†
558:†
548:†
538:†
528:†
518:†
514:?
507:†
496:†
486:†
476:†
472:?
465:†
455:†
445:†
435:†
425:†
415:†
405:†
395:†
385:†
375:†
365:†
355:†
345:†
335:†
325:†
315:†
289:†
262:†
249:†
236:†
220:†
204:†
108:N
98:K
93:J
88:T
83:P
78:C
73:D
68:S
63:O
58:Ꞓ
20:)
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