2547:
2553:
118:
3103:
1256:
139:
1277:
3133:
487:
467:
3109:
434:
1475:
479:
453:
can be distinguished based on the following characteristics: the nasal horncore is caudally positioned, almost entirely behind external naris; the supraorbital horncores are short, robust, dorsolaterally directed, and oblate in shape with a blunt tip; the episquamosals on the mid-portion of the
1726:
Eaton, J.G., 2002. Multituberculate mammals from the
Wahweap (Campanian, Aquilan) and Kaiparowits (Campanian, Judithian) formations, within and near Grand StaircaseāEscalante National Monument, southern Utah. Miscellaneous Publication 02-4, UtahGeological Survey, 66
1320:
swamps, ponds and lakes, and was bordered by highlands. The climate was wet and humid, and supported an abundant and diverse range of organisms. This formation contains one of the best and most continuous records of Late
Cretaceous terrestrial life in the world.
1716:
Roberts EM, Deino AL, Chan MA (2005) 40Ar/39Ar age of the
Kaiparowits Formation, southern Utah, and correlation of contemporaneous Campanian strata and vertebrate faunas along the margin of the Western Interior Basin. Cretaceous Res 26:
441:
The holotype specimen UMNH VP 16784, consists of only a partial skull. This genus is known from six specimens, including two partial skulls, which when taken together preserve about 96% of the skull and 70% of the postcranial skeleton.
3285:
1736:
Titus, Alan L. and Mark A. Loewen (editors). At the Top of the Grand
Staircase: The Late Cretaceous of Southern Utah. 2013. Indiana University Press. Hardbound: 634 pp.
406:
1633:
Raia, P.; Passaro, F.; Carotenuto, F.; Maiorino, L.; Piras, P.; Teresi, L.; Meiri, S.; Itescu, Y.; Novosolov, M.; Baiano, M. A.; Martinez, R.; Fortelius, M. (2015).
3226:
1836:
1682:
Longrich, N. R. (2014). "The horned dinosaurs
Pentaceratops and Kosmoceratops from the upper Campanian of Alberta and implications for dinosaur biogeography".
3300:
3213:
3295:
3290:
2546:
1617:
1868:
1777:
Zanno, Lindsay E.; Sampson, Scott D. (2005). "A new oviraptorosaur (Theropoda; Maniraptora) from the Late
Cretaceous (Campanian) of Utah".
405:, is derived from the name of Mike Getty, who discovered the holotype and has played a pivotal role in the recovery of fossils from the
1549:
Sampson, S. D.; Loewen, M. A.; Farke, A. A.; Roberts, E. M.; Forster, C. A.; Smith, J. A.; Titus, A. L. (2010). Stepanova, Anna (ed.).
1300:
indicates that the
Kaiparowits Formation was deposited between 76.6 and 74.5 million years ago, during the Campanian stage of the Late
2552:
1820:
1747:
1526:
3126:
1488:
454:
lateral frill margin are low and extremely elongate (some >10 cm long); and the median portion of transverse bar of the
3310:
126:
1811:=1999 (1999). "Cretaceous vertebrate faunas from the Kaiparowits Plateau, south-central Utah". In Gillete, David D. (ed.).
1304:
period. During the Late
Cretaceous period, the site of the Kaiparowits Formation was located near the western shore of the
3320:
3305:
1423:, but the existing assessment of the material is not conclusive. Paleofauna present in the Kaiparowits Formation included
1316:
to the east. The plateau where dinosaurs lived was an ancient floodplain dominated by large channels and abundant wetland
3102:
138:
1512:
3315:
775:
1790:
1959:
1861:
3280:
2077:
1517:
1305:
399:
3153:
117:
446:
are estimated to have measured 4.5ā5 metres (15ā16 ft) in length and 2 tonnes (2.2 tons) in weight.
3252:
3136:
1986:
1386:
1293:
307:
3108:
1804:
Eaton, Jeffrey G.; Cifelli, Richard L.; Hutchinson, J. Howard; Kirkland, James I.; Parrish, J. Michael
3200:
1691:
1562:
1508:
418:
284:
2242:
1854:
1333:
1808:
2962:
2912:
2344:
1830:
1297:
133:
3257:
1255:
3191:
3239:
3231:
3045:
2983:
2946:
2420:
2307:
1816:
1664:
1613:
1590:
1522:
414:
357:
3244:
1748:"Presidential Proclamation: Establishment of the Grand StaircaseāEscalante National Monument"
3075:
2873:
2698:
2400:
2095:
1815:. Miscellaneous Publication 99-1. Salt Lake City: Utah Geological Survey. pp. 345ā353.
1786:
1699:
1654:
1646:
1580:
1570:
1480:
1452:
1397:
1276:
970:
410:
364:
277:
2969:
2921:
2880:
2816:
2719:
2670:
2372:
2296:
2251:
2125:
2111:
2070:
1944:
1928:
1371:
1337:
1003:
538:
361:
30:
1634:
1695:
1566:
1551:"New Horned Dinosaurs from Utah Provide Evidence for Intracontinental Dinosaur Endemism"
2976:
2935:
2866:
2853:
2824:
2809:
2779:
2740:
2726:
2643:
2604:
2483:
2393:
2358:
2351:
2231:
2181:
2104:
2084:
1585:
1550:
1424:
1353:
1348:
1030:
1016:
524:
498:
395:
257:
215:
3274:
3054:
3033:
3003:
2994:
2894:
2788:
2733:
2712:
2684:
2677:
2663:
2656:
2462:
2451:
2444:
2386:
2379:
2365:
2323:
2258:
2224:
2195:
2148:
1805:
1755:
1410:
1406:
1392:
1358:
953:
900:
831:
709:
506:
455:
68:
3068:
3019:
3012:
2928:
2901:
2887:
2795:
2768:
2754:
2705:
2691:
2583:
2570:
2501:
2490:
2475:
2437:
2429:
2411:
2316:
2282:
2265:
2217:
2210:
2166:
1971:
1919:
1419:
1415:
1382:
1363:
1344:
676:
590:
562:
466:
244:
199:
1703:
486:
1575:
398:
words "keras" (ĪŗĪĻĪ±Ļ) meaning "horn" and "ops" (į½¤Ļ) referring to the "face". The
3185:
3061:
2955:
2838:
2802:
2625:
2337:
2202:
1612:(2nd ed.). Princeton, New Jersey: Princeton University Press. p. 297.
1460:
1377:
1367:
1068:
501:
376:
43:
433:
2831:
2747:
2597:
2330:
2118:
2052:
2039:
1998:
1877:
1470:
1444:
1432:
1313:
1301:
1054:
350:
231:
88:
53:
3176:
1456:
1401:
1309:
379:
150:
93:
37:
1668:
1594:
394:, means "horned face from Utah", and is derived from the state of Utah and
478:
3170:
2632:
2618:
2611:
2174:
1910:
1901:
1340:
353:
183:
170:
83:
78:
63:
58:
48:
513:
The below cladogram follows
Longrich (2014), who named a new species of
382:, that could grow up to an estimated 4.5ā5 m (15ā16 ft) long.
3218:
3026:
2188:
2132:
1659:
1260:
343:
98:
73:
1895:
1448:
1440:
1436:
160:
3147:
1650:
1308:, a large inland sea that split North America into two landmasses,
1635:"Cope's rule and the universal scaling law of ornament complexity"
1275:
1254:
504:. It has been found to be in a clade of basal chasmosaurines with
485:
477:
465:
432:
346:
3205:
1428:
1317:
368:
3151:
2568:
2037:
1888:
1850:
1521:(Abridged ed.). United Kingdom: Oxford University Press.
1846:
1791:
10.1671/0272-4634(2005)025[0897:ANOTMF]2.0.CO;2
1280:
North
America about 75 mya, location the discovery of
1405:. Some fossil evidence suggests the presence of the
413:, Mark A. Loewen, Andrew A. Farke, Eric M. Roberts,
375:
was a large-sized, robustly-built, ground-dwelling,
3160:
3044:
3002:
2993:
2945:
2911:
2852:
2778:
2642:
2581:
2500:
2461:
2419:
2410:
2306:
2280:
2241:
2146:
2094:
2050:
1997:
1970:
1943:
517:, and included nearly all chasmosaurine species.
1862:
8:
1835:: CS1 maint: numeric names: authors list (
407:Grand StaircaseāEscalante National Monument
3286:Late Cretaceous dinosaurs of North America
3148:
2999:
2858:
2648:
2589:
2578:
2565:
2467:
2416:
2288:
2158:
2154:
2062:
2058:
2047:
2034:
1976:
1949:
1940:
1885:
1869:
1855:
1847:
116:
20:
1658:
1584:
1574:
1544:
1542:
1540:
1538:
1500:
417:, Joshua A. Smith and Alan L. Titus in
1828:
1610:The Princeton Field Guide to Dinosaurs
7:
449:According to Sampson et al. (2010),
1097:
1075:
1046:
1022:
995:
945:
938:
892:
885:
838:
826:
767:
732:
725:
701:
620:
597:
585:
578:
554:
530:
520:
356:that lived approximately 76.4~75.5
1779:Journal of Vertebrate Paleontology
14:
3132:
3131:
3107:
3101:
2551:
2545:
1608:Paul, G.S. (2016). "Genasaurs".
1489:Timeline of ceratopsian research
1473:
137:
3301:Taxa named by Catherine Forster
1813:Vertebrate Paleontology in Utah
497:has been classified as a basal
409:(GSENM). It was first named by
3296:Taxa named by Scott D. Sampson
1372:Parasaurolophus cyrtocristatus
1336:with other dinosaurs, such as
1298:Argon-argon radiometric dating
482:Selected craniofacial elements
127:Natural History Museum of Utah
1:
3291:Fossil taxa described in 2010
1704:10.1016/j.cretres.2014.06.011
474:with known elements in yellow
1576:10.1371/journal.pone.0012292
916:Almond Formation chasmosaur
1288:The only known specimen of
3337:
470:Reconstructed skeleton of
458:bone is rostrally curved.
421:, and the type species is
16:Extinct genus of dinosaurs
3121:
3099:
2861:
2651:
2592:
2577:
2564:
2542:
2470:
2291:
2161:
2157:
2065:
2061:
2046:
2033:
1979:
1952:
1939:
1884:
1393:Kosmoceratops richardsoni
1378:Gryposaurus monumentensis
1117:
1102:
1095:
1080:
1073:
1051:
1044:
1027:
1020:
1000:
993:
967:
950:
943:
936:
914:
897:
890:
883:
858:
846:Kosmoceratops richardsoni
843:
836:
824:
789:
776:Pentaceratops sternbergii
772:
765:
754:Williams Fork chasmosaur
752:
737:
730:
723:
706:
699:
673:
640:
625:
618:
602:
595:
583:
576:
559:
552:
535:
528:
313:
306:
134:Scientific classification
132:
124:
115:
23:
740:Pentaceratops aquilonius
125:Reconstructed skeleton,
2078:Micropachycephalosaurus
1639:The American Naturalist
1518:A Greek-English Lexicon
1451:were present including
1306:Western Interior Seaway
643:Chasmosaurus irvinensis
1285:
1268:
491:
483:
475:
438:
3311:Kaiparowits Formation
3253:Paleobiology Database
1509:Liddell, Henry George
1447:. A variety of early
1387:Nasutoceratops titusi
1368:duckbilled hadrosaurs
1364:armored ankylosaurids
1294:Kaiparowits Formation
1292:was recovered at the
1279:
1258:
489:
481:
469:
436:
3321:Ornithischian genera
3306:Paleontology in Utah
415:Catherine A. Forster
2243:Archaeoceratopsidae
1696:2014CrRes..51..292L
1684:Cretaceous Research
1567:2010PLoSO...512292S
1427:(sharks and rays),
1402:Hagryphus giganteus
423:Utahceratops gettyi
320:Utahceratops gettyi
2963:Crittendenceratops
2913:Albertaceratopsini
1752:September 18, 1996
1746:Clinton, William.
1420:Ornithomimus velox
1286:
1269:
628:Chasmosaurus belli
492:
484:
476:
439:
3268:
3267:
3240:Open Tree of Life
3154:Taxon identifiers
3145:
3144:
3117:
3116:
3097:
3096:
3093:
3092:
3089:
3088:
3085:
3084:
3046:Pachyrhinosaurini
2984:Yehuecauhceratops
2947:Nasutoceratopsini
2848:
2847:
2560:
2559:
2540:
2539:
2536:
2535:
2532:
2531:
2528:
2527:
2524:
2523:
2520:
2519:
2421:Protoceratopsidae
2308:Leptoceratopsidae
2276:
2275:
2142:
2141:
2029:
2028:
2025:
2024:
2021:
2020:
2017:
2016:
1758:on 28 August 2013
1619:978-0-691-16766-4
1453:multituberculates
1398:oviraptorosaurian
1247:
1246:
1238:
1237:
1229:
1228:
1220:
1219:
1211:
1210:
1202:
1201:
1193:
1192:
1184:
1183:
1175:
1174:
1166:
1165:
1157:
1156:
1148:
1147:
1139:
1138:
1130:
1129:
982:
981:
925:
924:
872:
871:
813:
812:
804:
803:
688:
687:
662:
661:
653:
652:
358:million years ago
335:
334:
287:
3328:
3316:Campanian genera
3261:
3260:
3248:
3247:
3235:
3234:
3222:
3221:
3209:
3208:
3196:
3195:
3194:
3181:
3180:
3179:
3149:
3135:
3134:
3112:
3111:
3105:
3076:Pachyrhinosaurus
3000:
2874:Machairoceratops
2859:
2699:Coahuilaceratops
2649:
2590:
2579:
2566:
2555:
2550:
2549:
2468:
2417:
2401:Zhuchengceratops
2289:
2159:
2155:
2096:Chaoyangsauridae
2063:
2059:
2048:
2035:
1977:
1950:
1941:
1934:
1933:
1886:
1871:
1864:
1857:
1848:
1841:
1840:
1834:
1826:
1809:albertosarusyear
1801:
1795:
1794:
1774:
1768:
1767:
1765:
1763:
1754:. Archived from
1743:
1737:
1734:
1728:
1724:
1718:
1714:
1708:
1707:
1679:
1673:
1672:
1662:
1630:
1624:
1623:
1605:
1599:
1598:
1588:
1578:
1546:
1533:
1532:
1505:
1483:
1481:Dinosaurs portal
1478:
1477:
1476:
1334:paleoenvironment
1312:to the west and
1098:
1076:
1047:
1023:
996:
971:Coahuilaceratops
946:
939:
893:
886:
839:
827:
768:
733:
726:
702:
621:
598:
586:
579:
555:
531:
521:
411:Scott D. Sampson
322:
318:
276:
269:
256:
243:
230:
214:
198:
142:
141:
120:
110:
40:
29:Temporal range:
21:
3336:
3335:
3331:
3330:
3329:
3327:
3326:
3325:
3271:
3270:
3269:
3264:
3256:
3251:
3243:
3238:
3230:
3225:
3217:
3212:
3204:
3199:
3190:
3189:
3184:
3175:
3174:
3169:
3156:
3146:
3141:
3113:
3106:
3081:
3040:
2989:
2970:Furcatoceratops
2941:
2922:Albertaceratops
2907:
2881:Menefeeceratops
2844:
2817:Tatankaceratops
2774:
2720:Mercuriceratops
2671:Arrhinoceratops
2638:
2586:
2573:
2556:
2544:
2516:
2496:
2457:
2406:
2373:Montanoceratops
2302:
2297:Graciliceratops
2285:
2272:
2252:Archaeoceratops
2237:
2151:
2138:
2126:Xuanhuaceratops
2112:Hualianceratops
2090:
2071:Albalophosaurus
2055:
2042:
2013:
1993:
1966:
1960:Avemetatarsalia
1945:Avemetatarsalia
1935:
1929:Marginocephalia
1891:
1890:
1880:
1875:
1845:
1844:
1827:
1823:
1803:
1802:
1798:
1776:
1775:
1771:
1761:
1759:
1745:
1744:
1740:
1735:
1731:
1725:
1721:
1715:
1711:
1681:
1680:
1676:
1632:
1631:
1627:
1620:
1607:
1606:
1602:
1548:
1547:
1536:
1529:
1507:
1506:
1502:
1497:
1479:
1474:
1472:
1469:
1425:chondrichthyans
1327:
1274:
1253:
1248:
1239:
1230:
1221:
1212:
1203:
1194:
1185:
1176:
1167:
1158:
1149:
1140:
1131:
1004:Arrhinoceratops
983:
926:
873:
814:
805:
689:
663:
654:
539:Mercuriceratops
490:Size comparison
464:
431:
390:The genus name
388:
367:in what is now
362:Late Cretaceous
331:
324:
316:
315:
302:
275:
267:
254:
241:
228:
212:
196:
136:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
36:76.4ā75.5
35:
34:
31:Late Cretaceous
27:
17:
12:
11:
5:
3334:
3332:
3324:
3323:
3318:
3313:
3308:
3303:
3298:
3293:
3288:
3283:
3281:Chasmosaurines
3273:
3272:
3266:
3265:
3263:
3262:
3249:
3236:
3223:
3210:
3197:
3182:
3166:
3164:
3158:
3157:
3152:
3143:
3142:
3140:
3139:
3129:
3122:
3119:
3118:
3115:
3114:
3100:
3098:
3095:
3094:
3091:
3090:
3087:
3086:
3083:
3082:
3080:
3079:
3072:
3065:
3058:
3050:
3048:
3042:
3041:
3039:
3038:
3030:
3023:
3016:
3008:
3006:
2997:
2991:
2990:
2988:
2987:
2980:
2977:Nasutoceratops
2973:
2966:
2959:
2951:
2949:
2943:
2942:
2940:
2939:
2936:Medusaceratops
2932:
2925:
2917:
2915:
2909:
2908:
2906:
2905:
2898:
2891:
2884:
2877:
2870:
2867:Diabloceratops
2862:
2856:
2854:Centrosaurinae
2850:
2849:
2846:
2845:
2843:
2842:
2835:
2828:
2825:Titanoceratops
2821:
2813:
2810:Regaliceratops
2806:
2799:
2792:
2784:
2782:
2780:Triceratopsini
2776:
2775:
2773:
2772:
2765:
2758:
2751:
2744:
2741:Sierraceratops
2737:
2730:
2727:Navajoceratops
2723:
2716:
2709:
2702:
2695:
2688:
2681:
2674:
2667:
2660:
2652:
2646:
2644:Chasmosaurinae
2640:
2639:
2637:
2636:
2629:
2622:
2615:
2608:
2605:Brachyceratops
2601:
2593:
2587:
2582:
2575:
2574:
2569:
2562:
2561:
2558:
2557:
2543:
2541:
2538:
2537:
2534:
2533:
2530:
2529:
2526:
2525:
2522:
2521:
2518:
2517:
2515:
2514:
2513:
2512:
2506:
2504:
2498:
2497:
2495:
2494:
2487:
2484:Turanoceratops
2480:
2471:
2465:
2459:
2458:
2456:
2455:
2448:
2441:
2434:
2425:
2423:
2414:
2408:
2407:
2405:
2404:
2397:
2394:Unescoceratops
2390:
2383:
2376:
2369:
2362:
2359:Ischioceratops
2355:
2352:Gryphoceratops
2348:
2341:
2334:
2327:
2320:
2312:
2310:
2304:
2303:
2301:
2300:
2292:
2286:
2281:
2278:
2277:
2274:
2273:
2271:
2270:
2262:
2255:
2247:
2245:
2239:
2238:
2236:
2235:
2232:Sasayamagnomus
2228:
2221:
2214:
2207:
2199:
2192:
2185:
2182:Auroraceratops
2178:
2171:
2162:
2152:
2147:
2144:
2143:
2140:
2139:
2137:
2136:
2129:
2122:
2115:
2108:
2105:Chaoyangsaurus
2100:
2098:
2092:
2091:
2089:
2088:
2085:Psittacosaurus
2081:
2074:
2066:
2056:
2051:
2044:
2043:
2038:
2031:
2030:
2027:
2026:
2023:
2022:
2019:
2018:
2015:
2014:
2012:
2011:
2010:
2009:
2003:
2001:
1995:
1994:
1992:
1991:
1990:
1989:
1980:
1974:
1968:
1967:
1965:
1964:
1963:
1962:
1953:
1947:
1937:
1936:
1932:
1931:
1922:
1913:
1904:
1898:
1889:
1882:
1881:
1876:
1874:
1873:
1866:
1859:
1851:
1843:
1842:
1821:
1796:
1785:(4): 897ā904.
1769:
1738:
1729:
1719:
1709:
1674:
1651:10.1086/682011
1645:(2): 165ā175.
1625:
1618:
1600:
1534:
1527:
1499:
1498:
1496:
1493:
1492:
1491:
1485:
1484:
1468:
1465:
1354:tyrannosaurids
1349:Talos sampsoni
1326:
1323:
1273:
1270:
1252:
1249:
1245:
1244:
1241:
1240:
1236:
1235:
1232:
1231:
1227:
1226:
1223:
1222:
1218:
1217:
1214:
1213:
1209:
1208:
1205:
1204:
1200:
1199:
1196:
1195:
1191:
1190:
1187:
1186:
1182:
1181:
1178:
1177:
1173:
1172:
1169:
1168:
1164:
1163:
1160:
1159:
1155:
1154:
1151:
1150:
1146:
1145:
1142:
1141:
1137:
1136:
1133:
1132:
1128:
1127:
1124:
1123:
1116:
1113:
1112:
1109:
1108:
1101:
1096:
1094:
1091:
1090:
1087:
1086:
1079:
1074:
1072:
1064:
1063:
1060:
1059:
1050:
1045:
1043:
1040:
1039:
1036:
1035:
1031:Titanoceratops
1026:
1021:
1019:
1017:Triceratopsini
1013:
1012:
1009:
1008:
999:
994:
992:
989:
988:
985:
984:
980:
979:
976:
975:
966:
963:
962:
959:
958:
949:
944:
942:
937:
935:
932:
931:
928:
927:
923:
922:
919:
918:
913:
910:
909:
906:
905:
896:
891:
889:
884:
882:
879:
878:
875:
874:
870:
869:
866:
865:
857:
854:
853:
850:
849:
842:
837:
835:
825:
823:
820:
819:
816:
815:
811:
810:
807:
806:
802:
801:
798:
797:
788:
785:
784:
781:
780:
771:
766:
764:
761:
760:
757:
756:
751:
748:
747:
744:
743:
736:
731:
729:
724:
722:
719:
718:
715:
714:
705:
700:
698:
695:
694:
691:
690:
686:
685:
682:
681:
672:
669:
668:
665:
664:
660:
659:
656:
655:
651:
650:
647:
646:
639:
636:
635:
632:
631:
624:
619:
617:
614:
613:
610:
609:
601:
596:
594:
584:
582:
577:
575:
572:
571:
568:
567:
558:
553:
551:
548:
547:
544:
543:
534:
529:
527:
525:Chasmosaurinae
519:
463:
460:
430:
427:
387:
384:
333:
332:
325:
311:
310:
304:
303:
299:U. gettyi
295:
293:
289:
288:
265:
261:
260:
258:Chasmosaurinae
252:
248:
247:
239:
235:
234:
226:
219:
218:
216:Neornithischia
210:
203:
202:
194:
187:
186:
181:
174:
173:
168:
164:
163:
158:
154:
153:
148:
144:
143:
130:
129:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
3333:
3322:
3319:
3317:
3314:
3312:
3309:
3307:
3304:
3302:
3299:
3297:
3294:
3292:
3289:
3287:
3284:
3282:
3279:
3278:
3276:
3259:
3254:
3250:
3246:
3241:
3237:
3233:
3228:
3224:
3220:
3215:
3211:
3207:
3202:
3198:
3193:
3187:
3183:
3178:
3172:
3168:
3167:
3165:
3163:
3159:
3155:
3150:
3138:
3130:
3128:
3124:
3123:
3120:
3110:
3104:
3078:
3077:
3073:
3071:
3070:
3066:
3064:
3063:
3059:
3057:
3056:
3055:Achelousaurus
3052:
3051:
3049:
3047:
3043:
3036:
3035:
3034:Styracosaurus
3031:
3029:
3028:
3024:
3022:
3021:
3017:
3015:
3014:
3010:
3009:
3007:
3005:
3004:Centrosaurini
3001:
2998:
2996:
2995:Eucentrosaura
2992:
2986:
2985:
2981:
2979:
2978:
2974:
2972:
2971:
2967:
2965:
2964:
2960:
2958:
2957:
2953:
2952:
2950:
2948:
2944:
2938:
2937:
2933:
2931:
2930:
2926:
2924:
2923:
2919:
2918:
2916:
2914:
2910:
2904:
2903:
2899:
2897:
2896:
2895:Wendiceratops
2892:
2890:
2889:
2885:
2883:
2882:
2878:
2876:
2875:
2871:
2869:
2868:
2864:
2863:
2860:
2857:
2855:
2851:
2841:
2840:
2836:
2834:
2833:
2829:
2827:
2826:
2822:
2819:
2818:
2814:
2812:
2811:
2807:
2805:
2804:
2800:
2798:
2797:
2793:
2791:
2790:
2789:Eotriceratops
2786:
2785:
2783:
2781:
2777:
2771:
2770:
2766:
2764:
2763:
2759:
2757:
2756:
2752:
2750:
2749:
2745:
2743:
2742:
2738:
2736:
2735:
2734:Pentaceratops
2731:
2729:
2728:
2724:
2722:
2721:
2717:
2715:
2714:
2713:Kosmoceratops
2710:
2708:
2707:
2703:
2701:
2700:
2696:
2694:
2693:
2689:
2687:
2686:
2685:Bravoceratops
2682:
2680:
2679:
2678:Bisticeratops
2675:
2673:
2672:
2668:
2666:
2665:
2664:Anchiceratops
2661:
2659:
2658:
2657:Agujaceratops
2654:
2653:
2650:
2647:
2645:
2641:
2635:
2634:
2630:
2628:
2627:
2623:
2621:
2620:
2616:
2614:
2613:
2609:
2607:
2606:
2602:
2600:
2599:
2595:
2594:
2591:
2588:
2585:
2580:
2576:
2572:
2567:
2563:
2554:
2548:
2510:
2509:
2508:
2507:
2505:
2503:
2499:
2493:
2492:
2488:
2486:
2485:
2481:
2478:
2477:
2473:
2472:
2469:
2466:
2464:
2463:Ceratopsoidea
2460:
2454:
2453:
2452:Protoceratops
2449:
2447:
2446:
2445:Breviceratops
2442:
2440:
2439:
2435:
2432:
2431:
2427:
2426:
2424:
2422:
2418:
2415:
2413:
2409:
2403:
2402:
2398:
2396:
2395:
2391:
2389:
2388:
2387:Udanoceratops
2384:
2382:
2381:
2380:Prenoceratops
2377:
2375:
2374:
2370:
2368:
2367:
2366:Leptoceratops
2363:
2361:
2360:
2356:
2354:
2353:
2349:
2347:
2346:
2342:
2340:
2339:
2335:
2333:
2332:
2328:
2326:
2325:
2324:Bainoceratops
2321:
2319:
2318:
2314:
2313:
2311:
2309:
2305:
2299:
2298:
2294:
2293:
2290:
2287:
2284:
2279:
2268:
2267:
2263:
2261:
2260:
2259:Helioceratops
2256:
2254:
2253:
2249:
2248:
2246:
2244:
2240:
2234:
2233:
2229:
2227:
2226:
2225:Mosaiceratops
2222:
2220:
2219:
2215:
2213:
2212:
2208:
2205:
2204:
2200:
2198:
2197:
2196:Koreaceratops
2193:
2191:
2190:
2186:
2184:
2183:
2179:
2177:
2176:
2172:
2169:
2168:
2164:
2163:
2160:
2156:
2153:
2150:
2149:Neoceratopsia
2145:
2135:
2134:
2130:
2128:
2127:
2123:
2121:
2120:
2116:
2114:
2113:
2109:
2107:
2106:
2102:
2101:
2099:
2097:
2093:
2087:
2086:
2082:
2080:
2079:
2075:
2073:
2072:
2068:
2067:
2064:
2060:
2057:
2054:
2049:
2045:
2041:
2036:
2032:
2007:
2006:
2005:
2004:
2002:
2000:
1996:
1988:
1984:
1983:
1982:
1981:
1978:
1975:
1973:
1969:
1961:
1957:
1956:
1955:
1954:
1951:
1948:
1946:
1942:
1938:
1930:
1926:
1923:
1921:
1917:
1914:
1912:
1908:
1905:
1903:
1899:
1897:
1893:
1892:
1887:
1883:
1879:
1872:
1867:
1865:
1860:
1858:
1853:
1852:
1849:
1838:
1832:
1824:
1822:1-55791-634-9
1818:
1814:
1810:
1807:
1806:tyrannosaurid
1800:
1797:
1792:
1788:
1784:
1780:
1773:
1770:
1757:
1753:
1749:
1742:
1739:
1733:
1730:
1723:
1720:
1713:
1710:
1705:
1701:
1697:
1693:
1689:
1685:
1678:
1675:
1670:
1666:
1661:
1656:
1652:
1648:
1644:
1640:
1636:
1629:
1626:
1621:
1615:
1611:
1604:
1601:
1596:
1592:
1587:
1582:
1577:
1572:
1568:
1564:
1561:(9): e12292.
1560:
1556:
1552:
1545:
1543:
1541:
1539:
1535:
1530:
1528:0-19-910207-4
1524:
1520:
1519:
1514:
1510:
1504:
1501:
1494:
1490:
1487:
1486:
1482:
1471:
1466:
1464:
1462:
1461:insectivorans
1458:
1454:
1450:
1446:
1442:
1438:
1434:
1430:
1426:
1422:
1421:
1417:
1413:
1412:
1411:Albertosaurus
1408:
1407:tyrannosaurid
1404:
1403:
1399:
1395:
1394:
1389:
1388:
1384:
1380:
1379:
1374:
1373:
1369:
1365:
1361:
1360:
1359:Teratophoneus
1355:
1351:
1350:
1346:
1342:
1339:
1338:dromaeosaurid
1335:
1331:
1324:
1322:
1319:
1315:
1311:
1307:
1303:
1299:
1295:
1291:
1283:
1278:
1271:
1267:
1263:
1262:
1257:
1250:
1243:
1242:
1234:
1233:
1225:
1224:
1216:
1215:
1207:
1206:
1198:
1197:
1189:
1188:
1180:
1179:
1171:
1170:
1162:
1161:
1153:
1152:
1144:
1143:
1135:
1134:
1126:
1125:
1122:
1121:
1115:
1114:
1111:
1110:
1107:
1106:
1100:
1099:
1093:
1092:
1089:
1088:
1085:
1084:
1078:
1077:
1071:
1070:
1066:
1065:
1062:
1061:
1058:
1057:
1056:
1049:
1048:
1042:
1041:
1038:
1037:
1034:
1033:
1032:
1025:
1024:
1018:
1015:
1014:
1011:
1010:
1007:
1006:
1005:
998:
997:
991:
990:
987:
986:
978:
977:
974:
973:
972:
965:
964:
961:
960:
957:
956:
955:
954:Bravoceratops
948:
947:
941:
940:
934:
933:
930:
929:
921:
920:
917:
912:
911:
908:
907:
904:
903:
902:
901:Anchiceratops
895:
894:
888:
887:
881:
880:
877:
876:
868:
867:
864:
863:sp. CMN 8301
862:
861:Kosmoceratops
856:
855:
852:
851:
848:
847:
841:
840:
834:
833:
832:Kosmoceratops
829:
828:
822:
821:
818:
817:
809:
808:
800:
799:
796:
795:
794:
787:
786:
783:
782:
779:
778:
777:
770:
769:
763:
762:
759:
758:
755:
750:
749:
746:
745:
742:
741:
735:
734:
728:
727:
721:
720:
717:
716:
713:
712:
711:
710:Agujaceratops
704:
703:
697:
696:
693:
692:
684:
683:
680:
679:
678:
671:
670:
667:
666:
658:
657:
649:
648:
645:
644:
638:
637:
634:
633:
630:
629:
623:
622:
616:
615:
612:
611:
608:
607:sp. CMN 2280
606:
600:
599:
593:
592:
588:
587:
581:
580:
574:
573:
570:
569:
566:
565:
564:
557:
556:
550:
549:
546:
545:
542:
541:
540:
533:
532:
526:
523:
522:
518:
516:
515:Pentaceratops
511:
509:
508:
507:Pentaceratops
503:
500:
499:chasmosaurine
496:
488:
480:
473:
468:
461:
459:
457:
452:
447:
445:
435:
428:
426:
424:
420:
416:
412:
408:
404:
401:
400:specific name
397:
393:
385:
383:
381:
378:
374:
370:
366:
363:
359:
355:
352:
348:
345:
341:
340:
329:
323:
321:
312:
309:
308:Binomial name
305:
301:
300:
294:
291:
290:
286:
282:
279:
274:
273:
266:
263:
262:
259:
253:
250:
249:
246:
240:
237:
236:
233:
227:
224:
221:
220:
217:
211:
208:
205:
204:
201:
195:
192:
189:
188:
185:
182:
179:
176:
175:
172:
169:
166:
165:
162:
159:
156:
155:
152:
149:
146:
145:
140:
135:
131:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
3192:Utahceratops
3162:Utahceratops
3161:
3074:
3069:Stellasaurus
3067:
3060:
3053:
3032:
3025:
3020:Coronosaurus
3018:
3013:Centrosaurus
3011:
2982:
2975:
2968:
2961:
2954:
2934:
2929:Lokiceratops
2927:
2920:
2902:Xenoceratops
2900:
2893:
2888:Sinoceratops
2886:
2879:
2872:
2865:
2837:
2830:
2823:
2815:
2808:
2801:
2796:Nedoceratops
2794:
2787:
2769:Vagaceratops
2767:
2762:Utahceratops
2761:
2760:
2755:Terminocavus
2753:
2746:
2739:
2732:
2725:
2718:
2711:
2706:Judiceratops
2704:
2697:
2692:Chasmosaurus
2690:
2683:
2676:
2669:
2662:
2655:
2631:
2624:
2617:
2610:
2603:
2596:
2584:Ceratopsidae
2571:Ceratopsidae
2502:Ceratopsidae
2491:Zuniceratops
2489:
2482:
2476:Ajkaceratops
2474:
2450:
2443:
2438:Bagaceratops
2436:
2430:Ajkaceratops
2428:
2412:Coronosauria
2399:
2392:
2385:
2378:
2371:
2364:
2357:
2350:
2343:
2336:
2329:
2322:
2317:Asiaceratops
2315:
2295:
2283:Euceratopsia
2266:Yamaceratops
2264:
2257:
2250:
2230:
2223:
2218:Microceratus
2216:
2211:Liaoceratops
2209:
2201:
2194:
2187:
2180:
2173:
2167:Ajkaceratops
2165:
2131:
2124:
2117:
2110:
2103:
2083:
2076:
2069:
1987:Ornithischia
1972:Ornithischia
1924:
1920:Ornithischia
1915:
1906:
1812:
1799:
1782:
1778:
1772:
1760:. Retrieved
1756:the original
1751:
1741:
1732:
1722:
1712:
1687:
1683:
1677:
1642:
1638:
1628:
1609:
1603:
1558:
1554:
1516:
1513:Robert Scott
1503:
1445:crocodilians
1418:
1416:ornithomimid
1409:
1400:
1391:
1385:
1383:ceratopsians
1376:
1370:
1357:
1347:
1330:Utahceratops
1329:
1328:
1290:Utahceratops
1289:
1287:
1282:Utahceratops
1281:
1266:Utahceratops
1265:
1259:
1251:Paleoecology
1119:
1118:
1104:
1103:
1083:T. utahensis
1082:
1081:
1067:
1053:
1052:
1029:
1028:
1002:
1001:
969:
968:
952:
951:
915:
899:
898:
860:
859:
845:
844:
830:
793:Utahceratops
792:
791:
790:
774:
773:
753:
739:
738:
708:
707:
677:Mojoceratops
675:
674:
642:
641:
627:
626:
605:Chasmosaurus
604:
603:
591:Chasmosaurus
589:
563:Judiceratops
561:
560:
537:
536:
514:
512:
505:
495:Utahceratops
494:
493:
472:Utahceratops
471:
451:Utahceratops
450:
448:
444:Utahceratops
443:
440:
422:
402:
392:Utahceratops
391:
389:
373:Utahceratops
372:
339:Utahceratops
338:
337:
336:
327:
319:
314:
298:
297:
280:
272:Utahceratops
271:
270:
245:Ceratopsidae
222:
206:
200:Ornithischia
190:
177:
25:Utahceratops
24:
18:
3186:Wikispecies
3062:Einiosaurus
2956:Avaceratops
2839:Triceratops
2803:Ojoceratops
2626:Monoclonius
2338:Ferrisaurus
2203:Kulceratops
1690:: 292ā308.
1660:11336/55178
1433:salamanders
1332:shared its
1296:, in Utah.
1105:T. horridus
1069:Triceratops
502:ceratopsian
437:Restoration
429:Description
377:quadrupedal
360:during the
351:ceratopsian
251:Subfamily:
3275:Categories
3125:See also:
2832:Torosaurus
2748:Spiclypeus
2598:Agathaumas
2511:see belowā
2331:Cerasinops
2119:Stenopelix
2053:Ceratopsia
2040:Ceratopsia
2008:see belowā
1999:Ceratopsia
1911:Dinosauria
1878:Ceratopsia
1762:9 November
1495:References
1457:marsupials
1345:troodontid
1325:Paleofauna
1314:Appalachia
1302:Cretaceous
1120:T. prorsus
1055:Torosaurus
232:Ceratopsia
184:Dinosauria
1894:Kingdom:
1831:cite book
1341:theropods
1310:Laramidia
462:Phylogeny
386:Discovery
380:herbivore
292:Species:
157:Kingdom:
151:Eukaryota
3206:17924513
3171:Wikidata
3137:Category
3127:Timeline
2633:Polyonax
2619:Dysganus
2612:Ceratops
2175:Aquilops
1902:Chordata
1900:Phylum:
1896:Animalia
1717:307ā318.
1669:26655146
1595:20877459
1555:PLOS ONE
1515:(1980).
1467:See also
1414:and the
1396:and the
456:parietal
354:dinosaur
326:Sampson
238:Family:
171:Chordata
167:Phylum:
161:Animalia
147:Domain:
3245:6150301
3232:1479007
3219:8527035
3177:Q134308
3027:Spinops
2345:Gremlin
2133:Yinlong
1692:Bibcode
1586:2929175
1563:Bibcode
1449:mammals
1441:lizards
1437:turtles
1272:Habitat
1261:Mirarce
344:extinct
317:†
296:†
278:Sampson
264:Genus:
105:↓
3258:243363
1819:
1667:
1616:
1593:
1583:
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1459:, and
1381:, the
1366:, the
1343:, the
403:gettyi
365:period
342:is an
330:, 2010
328:et al.
281:et al.
3227:IRMNG
1925:Clade
1916:Clade
1907:Clade
1429:frogs
1356:like
1284:shown
396:Greek
347:genus
223:Clade
207:Clade
191:Clade
178:Clade
3214:GBIF
1985:see
1958:see
1837:link
1817:ISBN
1764:2013
1665:PMID
1614:ISBN
1591:PMID
1523:ISBN
1511:and
1443:and
1390:and
1375:and
1318:peat
419:2010
369:Utah
285:2010
44:Preź
3201:EoL
2189:Beg
1787:doi
1727:pp.
1700:doi
1655:hdl
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1581:PMC
1571:doi
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