1041:; Hopfe, Charlotte; Weiß, Clemens L.; Mafessoni, Fabrizio; de la Rasilla, Marco; Lalueza-Fox, Carles; Rosas, Antonio; Soressi, Marie; Knul, Monika V.; Miller, Rebecca; Stewart, John R.; Derevianko, Anatoly P.; Jacobs, Zenobia; Li, Bo; Roberts, Richard G.; Shunkov, Michael V.; de Lumley, Henry; Perrenoud, Christian; Gušić, Ivan; Kućan, Željko; Rudan, Pavao; Aximu-Petri, Ayinuer; Essel, Elena; Nagel, Sarah; Nickel, Birgit; Schmidt, Anna; Prüfer, Kay; Kelso, Janet; Burbano, Hernán A.; Pääbo, Svante; Meyer, Matthias (2017).
79:
51:
72:
767:
A.; Butthof, A.; Hober, B.; Hoffner, B.; Siegemund, M.; Weihmann, A.; Nusbaum, C.; Lander, E. S.; Russ, C.; Novod, N.; Affourtit, J.; Egholm, M.; Verna, C.; Rudan, P.; Brajkovic, D.; Kucan, Z.; Gusic, I.; Doronichev, V. B.; Golovanova, L. V.; Lalueza-Fox, C.; de la
Rasilla, M.; Fortea, J.; Rosas, A.; Schmitz, R. W.; Johnson, P. L. F.; Eichler, E. E.; Falush, D.; Birney, E.; Mullikin, J. C.; Slatkin, M.; Nielsen, R.; Kelso, J.; Lachmann, M.; Reich, D.; Paabo, S. (2010-05-06).
5476:
327:. This led to suggestions that Neanderthals might have survived longer than previously thought and that the Neanderthals at Vindija might have lived concurrently with modern humans. However, later dating methods using more advanced techniques revealed that these earlier dating results were erroneous. The erroneous dates were due to contamination by modern carbon, as minute amounts of modern contamination may result in large errors for very old samples.
301:
44:
5710:
5723:
27:
979:
Pontus; Chintalapati, Manjusha; Dannemann, Michael; Nelson, Bradley J.; Key, Felix M.; Rudan, Pavao; Kućan, Željko; Gušić, Ivan; Golovanova, Liubov V.; Doronichev, Vladimir B.; Patterson, Nick; Reich, David; Eichler, Evan E.; Slatkin, Montgomery; Schierup, Mikkel H.; Andrés, Aida; Kelso, Janet; Meyer, Matthias; Pääbo, Svante (2017).
766:
Green, R. E.; Krause, J.; Briggs, A. W.; Maricic, T.; Stenzel, U.; Kircher, M.; Patterson, N.; Li, H.; Zhai, W.; Fritz, M. H. Y.; Hansen, N. F.; Durand, E. Y.; Malaspinas, A. S.; Jensen, J. D.; Marques-Bonet, T.; Alkan, C.; Prufer, K.; Meyer, M.; Burbano, H. A.; Good, J. M.; Schultz, R.; Aximu-Petri,
978:
Prüfer, Kay; de
Filippo, Cesare; Grote, Steffi; Mafessoni, Fabrizio; Korlević, Petra; Hajdinjak, Mateja; Vernot, Benjamin; Skov, Laurits; Hsieh, Pinghsun; Peyrégne, Stéphane; Reher, David; Hopfe, Charlotte; Nagel, Sarah; Maricic, Tomislav; Fu, Qiaomei; Theunert, Christoph; Rogers, Rebekah; Skoglund,
342:
to directly date several samples from
Vindija Cave. Their direct AMS dating results show that the Neanderthal finds at Vindija are older than 44,000 BP. Since this is earlier than the arrival of the first modern humans to the region, the Vindija Neanderthals most likely did not intermix with modern
270:, reduced facial size, and narrower front teeth. Though some have put these differences down to the small size of the Vindija individuals, a study conducted in 1995 established that the Vindija Neanderthals, though small, were of comparable size to more morphologically classic Neanderthals such as
826:
Hajdinjak, Mateja; Fu, Qiaomei; Hübner, Alexander; Petr, Martin; Mafessoni, Fabrizio; Grote, Steffi; Skoglund, Pontus; Narasimham, Vagheesh; Rougier, Hélène; Crevecoeur, Isabelle; Semal, Patrick; Soressi, Marie; Talamo, Sahra; Hublin, Jean-Jacques; Gušić, Ivan; Kućan, Željko; Rudan, Pavao;
942:
Janković, Ivor; Karavanić, Ivor; Ahern, James C. M.; Brajković, Dejana; Lenardić, Jadranka Mauch; Smith, Fred H. (2011). "Archaeological, Paleontological and
Genomic Perspectives on Late European Neandertals at Vindija Cave, Croatia". In Condemi, Silvana; Weniger, Gerd-Christian (eds.).
265:
The hominid specimens at level 3G are regarded as unquestionably
Neanderthal in overall morphology but exhibit a number of traits that sit closer to anatomically modern Europeans than to the traditional Neanderthal. These include a thinner and less projecting
827:
Golovanova, Liubov V.; Doronichev, Vladimir B.; Posth, Cosimo; Krause, Johannes; Korlević, Petra; Nagel, Sarah; Nickel, Birgit; Slatkin, Montgomery; Patterson, Nick; Reich, David; Prüfer, Kay; Meyer, Matthias; Pääbo, Svante; Kelso, Janet (2018).
1602:
5559:
2512:
713:
Devièse, Thibaut; Karavanić, Ivor; Comeskey, Daniel; Kubiak, Cara; Korlević, Petra; Hajdinjak, Mateja; Radović, Siniša; Procopio, Noemi; Buckley, Michael; Pääbo, Svante; Higham, Tom (2017).
1216:
458:, was female. The Vindija Neanderthals were also found to be genetically closer to other late European Neanderthals, to the exclusion of the Mezmaiskaya Neanderthals from
5765:
2593:
1276:
585:
5575:
1926:
409:, to sort through several unidentified fossil fragments recovered from Vindija Cave. They successfully identified a new Neanderthal fossil fragment,
5750:
1198:
4096:
2316:
5583:
262:. It is estimated that Neanderthals used the cave 40,000 years ago; approximately 8000 years before modern humans lived in that part of Europe.
1142:
5755:
5591:
5398:
4951:
4771:
4761:
4736:
4486:
4460:
960:
526:
71:
2728:
1307:
521:
4403:
4901:
1297:
331:
4312:
4308:
305:
1768:
1397:
715:"Direct dating of Neanderthal remains from the site of Vindija Cave and implications for the Middle to Upper Paleolithic transition"
5112:
3318:
1213:
124:
5459:
3308:
1269:
5671:
1242:
4931:
4911:
3389:
2636:
43:
4906:
5531:
5619:
4984:
3514:
3379:
2811:
5475:
3760:
3631:
1631:
1322:
450:, were genetically closest to each other than any other Neanderthal individuals on record. DNA analysis revealed that
402:
5451:
5419:
4481:
4171:
2656:
1798:
1550:
5760:
5567:
5283:
5077:
4851:
4811:
2753:
2040:
1262:
1234:
506:
243:
5599:
394:(directly dated to around 47,000 BP) and concluded that the fragment most likely came from the same individual as
5745:
5391:
4781:
3889:
2914:
227:
2424:
5499:
5275:
4751:
4746:
4398:
3993:
3226:
2193:
1493:
511:
292:. More likely, the Vindija Neanderthals were in transition from the classic robust form to a more gracile one.
5515:
5022:
319:
artefacts. Several of the
Neanderthal samples from Vindija also yielded surprisingly late dates when directly
5635:
5443:
5122:
4871:
3936:
3139:
2543:
1976:
1810:
1647:
1302:
5435:
1254:
1188:
5539:
5427:
5210:
5205:
4941:
4826:
4623:
4161:
3799:
3647:
3180:
3129:
2698:
2177:
2112:
1850:
1713:
1613:
5627:
5491:
2748:
5523:
5117:
4796:
4766:
4362:
4222:
4086:
3652:
2866:
2535:
2326:
2311:
5699:
5667:
5547:
5167:
4633:
2599:
2004:
1865:
1415:
5713:
5384:
5032:
5012:
4689:
4388:
4372:
4248:
3657:
3348:
2948:
2891:
2876:
2738:
2646:
2383:
2357:
2321:
1686:
1624:
1545:
1054:
992:
840:
780:
5507:
5017:
945:
Continuity and
Discontinuity in the Peopling of Europe: One Hundred Fifty Years of Neanderthal Study
5770:
5147:
5042:
4921:
4866:
4527:
4393:
4264:
4227:
4176:
4051:
3695:
3026:
2733:
2676:
2651:
2616:
2611:
2586:
2548:
2137:
1936:
1910:
1870:
1698:
886:
494:
371:
352:
351:
In 2010, the first draft sequence of the
Neanderthal genome was reconstructed primarily from three
5722:
5177:
5062:
5007:
5331:
5142:
4886:
4876:
4861:
4537:
4522:
4330:
3711:
3556:
3498:
3399:
3384:
2836:
2681:
2571:
2559:
2522:
2481:
2378:
2142:
2127:
1946:
1753:
1738:
1134:
335:
320:
5467:
4542:
2846:
2362:
1555:
516:
4532:
413:. Most of the fossil fragments that were identifiable by ZooMS were classified as belonging to
5679:
5659:
5127:
4961:
4946:
4836:
4648:
4638:
4439:
4418:
4408:
4367:
4181:
4101:
3926:
2861:
2841:
2806:
2703:
2527:
2398:
2159:
1981:
1966:
1875:
1508:
1436:
1193:
1122:
1090:
1082:
1026:
1008:
966:
956:
930:
912:
874:
856:
814:
796:
754:
736:
418:
5647:
5182:
5137:
5037:
4926:
4856:
4831:
4116:
4061:
4009:
3873:
3868:
3852:
3595:
3436:
3221:
2881:
2871:
2769:
2743:
2631:
2576:
2517:
2465:
2434:
2403:
2275:
2009:
1951:
1820:
1788:
1763:
1743:
1728:
1513:
1488:
1382:
1165:
1157:
1114:
1072:
1062:
1016:
1000:
948:
920:
902:
864:
848:
804:
788:
744:
726:
593:
459:
112:
5152:
300:
5315:
5102:
5002:
4731:
4684:
4669:
4346:
4206:
4186:
4166:
4071:
3951:
3778:
3671:
3530:
3364:
3262:
3216:
3154:
3113:
2686:
2641:
2604:
2408:
2393:
2331:
2296:
2235:
2045:
2030:
1793:
1718:
1691:
1642:
1457:
1452:
1246:
1220:
427:
390:. In 2018, researchers sequenced a low coverage genome from an undiagnosed bone fragment,
5200:
5057:
3931:
2831:
2666:
1971:
1733:
1565:
1340:
238:. Remains of three Neanderthals were selected as the primary sources for the first draft
1058:
996:
844:
784:
5726:
5611:
5226:
5157:
5047:
4791:
4786:
4628:
4613:
4592:
4434:
4201:
4191:
4146:
4056:
3988:
3847:
3732:
3369:
3190:
3097:
3066:
3056:
2984:
2886:
2856:
2826:
2723:
2708:
2691:
2671:
2661:
2581:
2455:
2347:
2075:
2065:
2050:
1956:
1931:
1885:
1837:
1758:
1657:
1360:
1021:
980:
925:
890:
869:
828:
809:
768:
749:
714:
339:
324:
5739:
5360:
5355:
5236:
5097:
5092:
5087:
5082:
4891:
4846:
4841:
4821:
4721:
4643:
4618:
4517:
4413:
4151:
4076:
3941:
3910:
3616:
3472:
3452:
3394:
3267:
3144:
3046:
2713:
2388:
2352:
2301:
2280:
2225:
2149:
2132:
2117:
2014:
1778:
1773:
1652:
1560:
1355:
1286:
1102:
586:"Neandertals and early modern humans probably didn't meet at rumored rendezvous site"
490:
417:. This was confirmed by Slon et al. (2017) who tested a soil sample from Vindija for
387:
278:
231:
102:
3737:
3685:
5265:
5244:
5027:
4956:
4936:
4896:
4576:
4285:
4269:
4136:
4121:
4111:
3946:
3626:
3585:
3546:
3457:
3415:
3333:
3303:
3288:
3211:
3206:
3175:
3149:
3092:
3051:
2979:
2974:
2851:
2816:
2790:
2091:
2035:
1900:
1860:
1855:
1803:
1723:
1703:
1345:
1239:
1038:
272:
223:
2429:
952:
5407:
5260:
5067:
5052:
4916:
4806:
4801:
4756:
4741:
4196:
4156:
4141:
4091:
3831:
3716:
3690:
3600:
3590:
3580:
3564:
3482:
3477:
3431:
3374:
3323:
3313:
3293:
3134:
3071:
3061:
3036:
3010:
2932:
2821:
2774:
2621:
2460:
2243:
2154:
2122:
2096:
2055:
1986:
1880:
1673:
1529:
1472:
1467:
1431:
1387:
1138:
489:
genome, researchers revised upwards the percentage of human DNA in non-Africans
316:
255:
211:
188:
2513:
Cave of
Altamira and Paleolithic Cave Art of Northern Spain World Heritage Site
5365:
5107:
5072:
4881:
4816:
4726:
4679:
4674:
4664:
4560:
4552:
4496:
4491:
4455:
4232:
4126:
3905:
3343:
3328:
3298:
3231:
3041:
3031:
2994:
2969:
2964:
2496:
2486:
2070:
2060:
1941:
1815:
1783:
1665:
1498:
1228:
312:
288:
267:
1126:
1086:
1012:
970:
916:
860:
800:
779:(5979). American Association for the Advancement of Science (AAAS): 710–722.
740:
552:
550:
548:
546:
139:
126:
5299:
4776:
4035:
4030:
4025:
3815:
3467:
3283:
3185:
3087:
2491:
2259:
1996:
1905:
1895:
1748:
1586:
1462:
1350:
1161:
1067:
1042:
1004:
907:
792:
731:
644:
642:
640:
638:
597:
246:
in 2010. Additional research was done on the samples and published in 2017.
4501:
1118:
1094:
1030:
934:
878:
818:
758:
382:
is the second high coverage
Neanderthal genome to be sequenced, after the
26:
5172:
5132:
4705:
4608:
4106:
4066:
3956:
3621:
3338:
2989:
2439:
2306:
2209:
1890:
1570:
1410:
1405:
1392:
1170:
463:
239:
852:
254:
The cave is located roughly 20 km (12 mi) west of the city of
4476:
4081:
3794:
3462:
3236:
1842:
1681:
235:
1077:
567:
565:
3972:
3246:
3241:
2169:
470:
259:
5162:
3159:
2718:
1143:"Upper Pleistocene Remains from Vindija Cave, Croatia, Yugoslavia"
406:
299:
1105:; Smith, Fred H. (1995). "Body size of the Vindija Neandertals".
315:
context; some of the remains occurred in a level with some mixed
5192:
981:"A high-coverage Neandertal genome from Vindija Cave in Croatia"
5380:
4982:
4306:
3758:
2912:
2594:
Rock art of the Iberian Mediterranean Basin World Heritage Site
1320:
1258:
556:
672:
648:
1927:
Caves and Ice Age Art in the Swabian Jura World Heritage Site
5474:
5376:
1043:"Neandertal and Denisovan DNA from Pleistocene sediments"
829:"Reconstructing the genetic history of late Neanderthals"
617:
571:
421:(mtDNA). Most of the classifiable mtDNA was ascribed to
493:
from Neanderthals and were able to identify additional
1191:[Secrets of evolution from the Vindija Cave].
684:
696:
5576:
Franciscan monastery and St. John the Baptist Church
5686:
5654:
5642:
5606:
5554:
5486:
5414:
5324:
5308:
5292:
5274:
5253:
5235:
5219:
5191:
4995:
4714:
4698:
4657:
4601:
4585:
4569:
4551:
4510:
4469:
4448:
4427:
4381:
4355:
4339:
4323:
4278:
4257:
4241:
4215:
4044:
4018:
4002:
3981:
3965:
3919:
3898:
3882:
3861:
3840:
3824:
3808:
3787:
3771:
3725:
3704:
3678:
3666:
3640:
3609:
3573:
3555:
3539:
3523:
3507:
3491:
3445:
3424:
3408:
3357:
3276:
3255:
3199:
3168:
3122:
3106:
3080:
3019:
3003:
2957:
2941:
2925:
2799:
2783:
2762:
2505:
2474:
2448:
2417:
2371:
2340:
2289:
2268:
2252:
2234:
2218:
2202:
2186:
2168:
2105:
2084:
2023:
1995:
1919:
1595:
1579:
1538:
1522:
1481:
1445:
1424:
1369:
1333:
434:Hajdinjak et al. (2018) found that the individuals
311:The Neanderthal remains at Vindija were found in a
207:
202:
194:
184:
179:
171:
163:
155:
118:
108:
98:
5584:Ursuline monastery and Nativity of Jesus Church
895:Proceedings of the National Academy of Sciences
719:Proceedings of the National Academy of Sciences
497:in humans that are derived from Neanderthals.
304:Neanderthal limb fragments from Vindija Cave,
5392:
1270:
629:
8:
1240:Dekodiran DNK neandertalca iz špilje Vindija
19:
5592:Capuchine monastery and Holy Trinity Church
1141:; Radovčić, Jakov; Rukavina, Darko (1981).
769:"A Draft Sequence of the Neandertal Genome"
407:Zooarchaeology by Mass Spectrometry (ZooMS)
338:ultrafiltration based on the extraction of
5460:Liberation of the Varaždin barracks (1991)
5399:
5385:
5377:
4992:
4979:
4320:
4303:
3768:
3755:
2922:
2909:
1330:
1317:
1285:Prehistoric cave sites, rock shelters and
1277:
1263:
1255:
323:, yielding dates as late as 28,000–29,000
222:is an archaeological site associated with
18:
1169:
1150:American Journal of Physical Anthropology
1076:
1066:
1020:
924:
906:
868:
808:
748:
730:
4052:Cradle of Humankind, World Heritage Site
542:
401:Devièse et al. (2017) applied collagen
16:Cave and archaeological site in Croatia
5766:Tourist attractions in Varaždin County
660:
370:Prüfer et al. (2017) sequenced a high
355:genomes from Vindija Cave, taken from
258:and 10 km (6.2 mi) north of
527:Multiregional origin of modern humans
7:
522:History of Croatia before the Croats
78:
50:
1189:"Tajne evolucije iz špilje Vindije"
947:. Dordrecht: Springer Netherlands.
584:Price, Michael (4 September 2017).
332:Oxford Radiocarbon Accelerator Unit
1201:from the original on 24 March 2012
1187:Marina Biluš (11 September 2007).
306:National Museum of Natural History
14:
2440:Mala Balanica and Velika Balanica
1603:Vézère Valley World Heritage Site
1398:Neolithic flint mines of Spiennes
230:, located in the municipality of
5721:
5709:
5708:
77:
70:
49:
42:
25:
378:. At around 30-fold coverage,
334:applied a new technique using
330:In 2017, researchers from the
1:
1197:(in Croatian). No. 617.
5756:Prehistoric sites in Croatia
5751:Landforms of Varaždin County
953:10.1007/978-94-007-0492-3_23
425:, coming predominately from
5452:Varaždin County (1850–1924)
2628:Other caves with decoration
1710:Other caves with decoration
1632:Les Eyzies-de-Tayac-Sireuil
1308:Caves containing pictograms
1113:(2). Elsevier BV: 201–208.
485:. With the addition of the
403:peptide mass fingerprinting
5787:
2612:Other World Heritage Sites
1699:Other World Heritage Sites
1298:Paleoanthropological sites
1107:Journal of Human Evolution
507:Neanderthal genome project
244:Neanderthal genome project
198:Neanderthals, Paleo-humans
5706:
5620:Varaždin Baroque Evenings
5532:Croatian National Theatre
5472:
5348:
4991:
4978:
4319:
4302:
3767:
3754:
2921:
2908:
1329:
1316:
1293:
630:Trinkaus & Smith 1995
469:Modern humans share more
36:
24:
5516:Prašinski–Sermage Palace
5479:Coat of arms of Varaždin
5420:Siege of Varaždin (1527)
5276:Northern Mariana Islands
512:Krapina Neanderthal site
5487:Memorials and landmarks
1162:10.1002/ajpa.1330540407
1068:10.1126/science.aam9695
1005:10.1126/science.aao1887
908:10.1073/pnas.1714533114
793:10.1126/science.1188021
732:10.1073/pnas.1709235114
598:10.1126/science.aap8521
454:, and thus most likely
159:50 metres (160 ft)
5636:Good Shepherd Festival
5480:
5444:Croatian Royal Council
3800:Manyana Rock Paintings
1851:La Chapelle-aux-Saints
1119:10.1006/jhev.1995.1015
891:"The last Neanderthal"
308:
175:20 metres (66 ft)
167:28 metres (92 ft)
5478:
4363:Actun Tunichil Muknal
697:Hajdinjak et al. 2018
303:
140:46.29917°N 16.07056°E
86:Vindija Cave (Europe)
3937:Ifri N'Amr Ou Moussa
2877:Red Lady of Paviland
1769:Mayrières supérieure
887:Hublin, Jean-Jacques
557:Janković et al. 2011
195:Associated with
58:Shown within Croatia
5568:St. Nicholas Church
5428:Varaždin Generalate
4752:Colorado Millennial
3390:Shiraho Saonetabaru
2194:Bukuroshja e Fjetur
1230:Video about Vindija
1135:Wolpoff, Milford H.
1059:2017Sci...356..605S
997:2017Sci...358..655P
901:(40): 10520–10522.
853:10.1038/nature26151
845:2018Natur.555..652H
785:2010Sci...328..710G
725:(40): 10606–10611.
673:Devièse et al. 2017
618:Wolpoff et al. 1981
495:phenotypic variants
136: /
61:Show map of Croatia
21:
5600:St. Florian Church
5481:
4570:Dominican Republic
4331:Cueva de las Manos
3227:Huto and Kamarband
2637:Barranc del Migdia
2523:Caves in Cantabria
1947:Hohlenstein-Stadel
1754:Grotte de Gabillou
1739:La Chaire a Calvin
1245:2010-05-22 at the
1219:2010-01-22 at the
1137:; Smith, Fred H.;
649:Prüfer et al. 2017
309:
145:46.29917; 16.07056
89:Show map of Europe
5761:Neanderthal sites
5733:
5732:
5714:Category:Varaždin
5700:Drava Park-Forest
5374:
5373:
5344:
5343:
5340:
5339:
4974:
4973:
4970:
4969:
4487:Piedras del Tunjo
4419:Toca da Tira Peia
4298:
4297:
4294:
4293:
3750:
3749:
3746:
3745:
2904:
2903:
2900:
2899:
2540:Hornos de la Peña
1053:(6338): 605–608.
991:(6363): 655–658.
962:978-94-007-0491-6
839:(7698): 652–656.
572:Green et al. 2010
483:Altai Neanderthal
419:mitochondrial DNA
384:Altai Neanderthal
217:
216:
5778:
5746:Caves of Croatia
5725:
5712:
5711:
5648:University North
5555:Church buildings
5401:
5394:
5387:
5378:
5293:Papua New Guinea
4993:
4980:
4932:Stanfield-Worley
4907:Rockhouse Cliffs
4321:
4304:
4172:Howieson's Poort
3869:Enkapune Ya Muto
3769:
3756:
3222:Eshkaft-e Siahoo
2923:
2910:
2677:Peñas de Cabrera
1977:Kleine Feldhofer
1937:Geissenklösterle
1789:Roc-aux-Sorciers
1331:
1318:
1279:
1272:
1265:
1256:
1251:
1231:
1225:
1210:
1208:
1206:
1175:
1173:
1147:
1130:
1098:
1080:
1070:
1034:
1024:
974:
938:
928:
910:
882:
872:
822:
812:
762:
752:
734:
700:
694:
688:
685:Slon et al. 2017
682:
676:
670:
664:
658:
652:
646:
633:
627:
621:
615:
609:
608:
606:
604:
581:
575:
569:
560:
554:
460:Mezmaiskaya cave
151:
150:
148:
147:
146:
141:
137:
134:
133:
132:
129:
113:Hrvatsko Zagorje
90:
81:
80:
74:
62:
53:
52:
46:
29:
22:
5786:
5785:
5781:
5780:
5779:
5777:
5776:
5775:
5736:
5735:
5734:
5729:
5702:
5682:
5650:
5638:
5602:
5550:
5482:
5470:
5468:Varaždin County
5410:
5405:
5375:
5370:
5336:
5320:
5304:
5288:
5270:
5249:
5231:
5215:
5187:
5103:Mackintosh 90/1
5023:Beginner's Luck
4987:
4966:
4782:Dutchess Quarry
4722:Arnold Research
4710:
4694:
4653:
4597:
4581:
4565:
4547:
4506:
4465:
4444:
4423:
4377:
4373:Midnight Terror
4351:
4335:
4315:
4290:
4274:
4253:
4237:
4211:
4040:
4014:
3998:
3977:
3961:
3952:Kelif el Boroud
3915:
3894:
3878:
3857:
3836:
3820:
3804:
3783:
3763:
3742:
3721:
3700:
3674:
3662:
3636:
3605:
3569:
3551:
3535:
3519:
3503:
3487:
3441:
3420:
3404:
3353:
3272:
3251:
3195:
3164:
3118:
3102:
3076:
3015:
2999:
2953:
2937:
2917:
2896:
2795:
2779:
2758:
2657:los Murciélagos
2605:Roca dels Moros
2501:
2470:
2444:
2413:
2384:Chertovy Vorota
2367:
2336:
2285:
2264:
2248:
2236:North Macedonia
2230:
2214:
2198:
2182:
2164:
2101:
2080:
2019:
1991:
1915:
1678:Abri du Poisson
1643:Les Combarelles
1591:
1575:
1534:
1518:
1477:
1441:
1420:
1416:Trou de l'Abîme
1365:
1325:
1312:
1289:
1283:
1249:
1247:Wayback Machine
1229:
1223:
1221:Wayback Machine
1204:
1202:
1186:
1183:
1178:
1145:
1133:
1101:
1037:
977:
963:
941:
885:
825:
765:
712:
708:
703:
695:
691:
683:
679:
671:
667:
659:
655:
647:
636:
628:
624:
616:
612:
602:
600:
583:
582:
578:
570:
563:
555:
544:
540:
535:
517:Peștera cu Oase
503:
428:Ursus ingressus
349:
347:Archaeogenetics
298:
252:
144:
142:
138:
135:
130:
127:
125:
123:
122:
94:
93:
92:
91:
88:
87:
84:
83:
82:
65:
64:
63:
60:
59:
56:
55:
54:
32:
17:
12:
11:
5:
5784:
5782:
5774:
5773:
5768:
5763:
5758:
5753:
5748:
5738:
5737:
5731:
5730:
5727:Portal:Croatia
5707:
5704:
5703:
5690:
5688:
5684:
5683:
5660:Varaždin Arena
5658:
5656:
5652:
5651:
5646:
5644:
5640:
5639:
5610:
5608:
5604:
5603:
5558:
5556:
5552:
5551:
5508:Patačić Palace
5490:
5488:
5484:
5483:
5473:
5471:
5418:
5416:
5412:
5411:
5406:
5404:
5403:
5396:
5389:
5381:
5372:
5371:
5369:
5368:
5363:
5358:
5353:
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5342:
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5338:
5337:
5335:
5334:
5328:
5326:
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5321:
5319:
5318:
5312:
5310:
5306:
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5302:
5296:
5294:
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5286:
5280:
5278:
5272:
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5269:
5268:
5263:
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5255:
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5239:
5233:
5232:
5230:
5229:
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5217:
5216:
5214:
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5208:
5203:
5197:
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5189:
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5180:
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5140:
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5125:
5120:
5115:
5110:
5105:
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5095:
5090:
5085:
5080:
5075:
5070:
5065:
5060:
5055:
5050:
5045:
5040:
5035:
5030:
5025:
5020:
5018:Beeton Shelter
5015:
5010:
5005:
4999:
4997:
4989:
4988:
4983:
4976:
4975:
4972:
4971:
4968:
4967:
4965:
4964:
4959:
4954:
4949:
4944:
4939:
4934:
4929:
4924:
4919:
4914:
4909:
4904:
4899:
4894:
4889:
4884:
4879:
4874:
4869:
4864:
4859:
4854:
4849:
4844:
4839:
4834:
4829:
4824:
4819:
4814:
4809:
4804:
4799:
4794:
4789:
4784:
4779:
4774:
4769:
4764:
4759:
4754:
4749:
4744:
4739:
4734:
4729:
4724:
4718:
4716:
4712:
4711:
4709:
4708:
4702:
4700:
4696:
4695:
4693:
4692:
4690:T'uqu T'uquyuq
4687:
4682:
4677:
4672:
4667:
4661:
4659:
4655:
4654:
4652:
4651:
4646:
4641:
4636:
4631:
4626:
4621:
4616:
4611:
4605:
4603:
4599:
4598:
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4595:
4589:
4587:
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4582:
4580:
4579:
4573:
4571:
4567:
4566:
4564:
4563:
4557:
4555:
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4546:
4545:
4540:
4535:
4530:
4525:
4520:
4514:
4512:
4508:
4507:
4505:
4504:
4499:
4494:
4489:
4484:
4479:
4473:
4471:
4467:
4466:
4464:
4463:
4458:
4452:
4450:
4446:
4445:
4443:
4442:
4437:
4431:
4429:
4425:
4424:
4422:
4421:
4416:
4411:
4406:
4401:
4396:
4391:
4385:
4383:
4379:
4378:
4376:
4375:
4370:
4365:
4359:
4357:
4353:
4352:
4350:
4349:
4343:
4341:
4337:
4336:
4334:
4333:
4327:
4325:
4317:
4316:
4307:
4300:
4299:
4296:
4295:
4292:
4291:
4289:
4288:
4282:
4280:
4276:
4275:
4273:
4272:
4267:
4261:
4259:
4255:
4254:
4252:
4251:
4245:
4243:
4239:
4238:
4236:
4235:
4230:
4225:
4219:
4217:
4213:
4212:
4210:
4209:
4204:
4199:
4194:
4192:Pinnacle Point
4189:
4184:
4179:
4174:
4169:
4164:
4159:
4154:
4149:
4144:
4139:
4134:
4130:
4129:
4124:
4119:
4114:
4109:
4104:
4099:
4094:
4089:
4084:
4079:
4074:
4069:
4064:
4059:
4054:
4048:
4046:
4042:
4041:
4039:
4038:
4033:
4028:
4022:
4020:
4016:
4015:
4013:
4012:
4006:
4004:
4000:
3999:
3997:
3996:
3994:The White Lady
3991:
3985:
3983:
3979:
3978:
3976:
3975:
3969:
3967:
3963:
3962:
3960:
3959:
3954:
3949:
3944:
3939:
3934:
3929:
3923:
3921:
3917:
3916:
3914:
3913:
3908:
3902:
3900:
3896:
3895:
3893:
3892:
3886:
3884:
3880:
3879:
3877:
3876:
3871:
3865:
3863:
3859:
3858:
3856:
3855:
3850:
3844:
3842:
3838:
3837:
3835:
3834:
3828:
3826:
3822:
3821:
3819:
3818:
3812:
3810:
3806:
3805:
3803:
3802:
3797:
3791:
3789:
3785:
3784:
3782:
3781:
3775:
3773:
3765:
3764:
3759:
3752:
3751:
3748:
3747:
3744:
3743:
3741:
3740:
3735:
3729:
3727:
3723:
3722:
3720:
3719:
3714:
3708:
3706:
3702:
3701:
3699:
3698:
3693:
3688:
3682:
3680:
3676:
3675:
3670:
3668:
3664:
3663:
3661:
3660:
3655:
3650:
3644:
3642:
3638:
3637:
3635:
3634:
3629:
3624:
3619:
3613:
3611:
3607:
3606:
3604:
3603:
3598:
3593:
3588:
3583:
3577:
3575:
3571:
3570:
3568:
3567:
3561:
3559:
3553:
3552:
3550:
3549:
3543:
3541:
3537:
3536:
3534:
3533:
3527:
3525:
3521:
3520:
3518:
3517:
3515:Khoit Tsenkher
3511:
3509:
3505:
3504:
3502:
3501:
3495:
3493:
3489:
3488:
3486:
3485:
3480:
3475:
3470:
3465:
3460:
3455:
3449:
3447:
3443:
3442:
3440:
3439:
3434:
3428:
3426:
3422:
3421:
3419:
3418:
3412:
3410:
3406:
3405:
3403:
3402:
3397:
3392:
3387:
3382:
3377:
3372:
3367:
3361:
3359:
3355:
3354:
3352:
3351:
3346:
3341:
3336:
3331:
3326:
3321:
3316:
3311:
3306:
3301:
3296:
3291:
3286:
3280:
3278:
3274:
3273:
3271:
3270:
3265:
3259:
3257:
3253:
3252:
3250:
3249:
3244:
3239:
3234:
3229:
3224:
3219:
3214:
3209:
3203:
3201:
3197:
3196:
3194:
3193:
3188:
3183:
3178:
3172:
3170:
3166:
3165:
3163:
3162:
3157:
3152:
3147:
3142:
3137:
3132:
3126:
3124:
3120:
3119:
3117:
3116:
3110:
3108:
3104:
3103:
3101:
3100:
3095:
3090:
3084:
3082:
3078:
3077:
3075:
3074:
3069:
3064:
3059:
3054:
3049:
3044:
3039:
3034:
3029:
3023:
3021:
3017:
3016:
3014:
3013:
3007:
3005:
3001:
3000:
2998:
2997:
2992:
2987:
2982:
2977:
2972:
2967:
2961:
2959:
2955:
2954:
2952:
2951:
2945:
2943:
2939:
2938:
2936:
2935:
2929:
2927:
2919:
2918:
2913:
2906:
2905:
2902:
2901:
2898:
2897:
2895:
2894:
2889:
2884:
2879:
2874:
2869:
2864:
2859:
2854:
2849:
2844:
2839:
2834:
2829:
2827:Creswell Crags
2824:
2819:
2814:
2809:
2807:Aveline's Hole
2803:
2801:
2800:United Kingdom
2797:
2796:
2794:
2793:
2787:
2785:
2781:
2780:
2778:
2777:
2772:
2766:
2764:
2760:
2759:
2757:
2756:
2751:
2749:Santa Catalina
2746:
2741:
2736:
2731:
2726:
2721:
2716:
2711:
2706:
2701:
2695:
2694:
2689:
2684:
2679:
2674:
2669:
2664:
2659:
2654:
2649:
2644:
2639:
2634:
2629:
2625:
2624:
2619:
2614:
2608:
2607:
2602:
2597:
2590:
2589:
2584:
2579:
2574:
2569:
2568:
2567:
2564:
2563:
2562:
2557:
2554:
2551:
2544:Monte Castillo
2541:
2538:
2533:
2530:
2520:
2515:
2509:
2507:
2503:
2502:
2500:
2499:
2494:
2489:
2484:
2478:
2476:
2472:
2471:
2469:
2468:
2463:
2458:
2452:
2450:
2446:
2445:
2443:
2442:
2437:
2432:
2427:
2425:Hadži-Prodan's
2421:
2419:
2415:
2414:
2412:
2411:
2406:
2401:
2396:
2391:
2386:
2381:
2375:
2373:
2369:
2368:
2366:
2365:
2360:
2355:
2350:
2344:
2342:
2338:
2337:
2335:
2334:
2329:
2324:
2319:
2314:
2309:
2304:
2299:
2293:
2291:
2287:
2286:
2284:
2283:
2278:
2272:
2270:
2266:
2265:
2263:
2262:
2256:
2254:
2250:
2249:
2247:
2246:
2240:
2238:
2232:
2231:
2229:
2228:
2222:
2220:
2216:
2215:
2213:
2212:
2206:
2204:
2200:
2199:
2197:
2196:
2190:
2188:
2184:
2183:
2181:
2180:
2174:
2172:
2166:
2165:
2163:
2162:
2157:
2152:
2147:
2140:
2135:
2130:
2125:
2120:
2115:
2109:
2107:
2103:
2102:
2100:
2099:
2094:
2088:
2086:
2082:
2081:
2079:
2078:
2073:
2068:
2063:
2058:
2053:
2048:
2043:
2038:
2033:
2027:
2025:
2021:
2020:
2018:
2017:
2012:
2007:
2001:
1999:
1993:
1992:
1990:
1989:
1984:
1979:
1974:
1969:
1964:
1960:
1959:
1954:
1949:
1944:
1939:
1934:
1929:
1923:
1921:
1917:
1916:
1914:
1913:
1908:
1903:
1898:
1893:
1888:
1883:
1878:
1873:
1868:
1863:
1858:
1853:
1848:
1845:
1840:
1835:
1832:
1829:
1826:
1823:
1818:
1813:
1807:
1806:
1801:
1796:
1791:
1786:
1781:
1776:
1771:
1766:
1761:
1756:
1751:
1746:
1741:
1736:
1731:
1726:
1721:
1716:
1711:
1707:
1706:
1701:
1695:
1694:
1689:
1684:
1679:
1676:
1671:
1668:
1663:
1662:Laugerie-Haute
1660:
1658:Laugerie-Basse
1655:
1650:
1645:
1640:
1639:Abri Chadourne
1637:
1634:
1629:
1628:
1627:
1622:
1616:
1611:
1608:
1605:
1599:
1597:
1593:
1592:
1590:
1589:
1583:
1581:
1577:
1576:
1574:
1573:
1568:
1563:
1558:
1553:
1548:
1542:
1540:
1539:Czech Republic
1536:
1535:
1533:
1532:
1526:
1524:
1520:
1519:
1517:
1516:
1511:
1509:Romuald's Cave
1506:
1501:
1496:
1491:
1485:
1483:
1479:
1478:
1476:
1475:
1470:
1465:
1460:
1455:
1449:
1447:
1443:
1442:
1440:
1439:
1434:
1428:
1426:
1422:
1421:
1419:
1418:
1413:
1408:
1403:
1400:
1395:
1390:
1385:
1380:
1377:
1373:
1371:
1367:
1366:
1364:
1363:
1358:
1353:
1348:
1343:
1337:
1335:
1327:
1326:
1321:
1314:
1313:
1311:
1310:
1305:
1303:Cave paintings
1300:
1294:
1291:
1290:
1287:cave paintings
1284:
1282:
1281:
1274:
1267:
1259:
1253:
1252:
1237:
1226:
1214:Špilja Vindija
1211:
1182:
1181:External links
1179:
1177:
1176:
1131:
1103:Trinkaus, Erik
1099:
1035:
975:
961:
939:
883:
823:
763:
709:
707:
704:
702:
701:
689:
677:
665:
653:
634:
622:
610:
576:
561:
541:
539:
536:
534:
531:
530:
529:
524:
519:
514:
509:
502:
499:
481:than with the
348:
345:
340:hydroxyproline
297:
294:
273:La Ferrassie 2
251:
248:
215:
214:
209:
208:Archaeologists
205:
204:
200:
199:
196:
192:
191:
186:
182:
181:
177:
176:
173:
169:
168:
165:
161:
160:
157:
153:
152:
120:
116:
115:
110:
106:
105:
100:
96:
95:
85:
76:
75:
69:
68:
67:
66:
57:
48:
47:
41:
40:
39:
38:
37:
34:
33:
30:
15:
13:
10:
9:
6:
4:
3:
2:
5783:
5772:
5769:
5767:
5764:
5762:
5759:
5757:
5754:
5752:
5749:
5747:
5744:
5743:
5741:
5728:
5724:
5719:
5715:
5705:
5701:
5697:
5693:
5689:
5685:
5681:
5677:
5673:
5669:
5665:
5661:
5657:
5653:
5649:
5645:
5641:
5637:
5633:
5629:
5625:
5621:
5617:
5613:
5609:
5605:
5601:
5597:
5593:
5589:
5585:
5581:
5577:
5573:
5569:
5565:
5561:
5557:
5553:
5549:
5545:
5541:
5537:
5533:
5529:
5525:
5521:
5517:
5513:
5509:
5505:
5501:
5500:County Palace
5497:
5493:
5489:
5485:
5477:
5469:
5465:
5461:
5457:
5453:
5449:
5445:
5441:
5437:
5433:
5429:
5425:
5421:
5417:
5413:
5409:
5402:
5397:
5395:
5390:
5388:
5383:
5382:
5379:
5367:
5364:
5362:
5361:Cave painting
5359:
5357:
5356:List of caves
5354:
5351:
5350:
5347:
5333:
5330:
5329:
5327:
5323:
5317:
5314:
5313:
5311:
5307:
5301:
5298:
5297:
5295:
5291:
5285:
5282:
5281:
5279:
5277:
5273:
5267:
5264:
5262:
5259:
5258:
5256:
5252:
5246:
5243:
5242:
5240:
5238:
5237:New Caledonia
5234:
5228:
5225:
5224:
5222:
5218:
5212:
5209:
5207:
5204:
5202:
5199:
5198:
5196:
5194:
5190:
5184:
5181:
5179:
5176:
5174:
5171:
5169:
5166:
5164:
5161:
5159:
5156:
5154:
5151:
5149:
5146:
5144:
5143:New Guinea II
5141:
5139:
5136:
5134:
5131:
5129:
5126:
5124:
5121:
5119:
5118:Mannalargenna
5116:
5114:
5111:
5109:
5106:
5104:
5101:
5099:
5096:
5094:
5091:
5089:
5086:
5084:
5081:
5079:
5076:
5074:
5071:
5069:
5066:
5064:
5061:
5059:
5056:
5054:
5051:
5049:
5046:
5044:
5041:
5039:
5036:
5034:
5031:
5029:
5026:
5024:
5021:
5019:
5016:
5014:
5011:
5009:
5006:
5004:
5001:
5000:
4998:
4994:
4990:
4986:
4981:
4977:
4963:
4960:
4958:
4955:
4953:
4950:
4948:
4945:
4943:
4940:
4938:
4935:
4933:
4930:
4928:
4925:
4923:
4920:
4918:
4915:
4913:
4910:
4908:
4905:
4903:
4900:
4898:
4895:
4893:
4890:
4888:
4887:On Your Knees
4885:
4883:
4880:
4878:
4875:
4873:
4870:
4868:
4865:
4863:
4860:
4858:
4855:
4853:
4850:
4848:
4845:
4843:
4840:
4838:
4835:
4833:
4830:
4828:
4825:
4823:
4820:
4818:
4815:
4813:
4812:Hidden Valley
4810:
4808:
4805:
4803:
4800:
4798:
4795:
4793:
4790:
4788:
4785:
4783:
4780:
4778:
4775:
4773:
4770:
4768:
4765:
4763:
4760:
4758:
4755:
4753:
4750:
4748:
4745:
4743:
4740:
4738:
4735:
4733:
4730:
4728:
4725:
4723:
4720:
4719:
4717:
4715:United States
4713:
4707:
4704:
4703:
4701:
4697:
4691:
4688:
4686:
4683:
4681:
4678:
4676:
4673:
4671:
4668:
4666:
4663:
4662:
4660:
4656:
4650:
4647:
4645:
4642:
4640:
4637:
4635:
4634:Guilá Naquitz
4632:
4630:
4627:
4625:
4622:
4620:
4619:Dzibilchaltún
4617:
4615:
4612:
4610:
4607:
4606:
4604:
4600:
4594:
4591:
4590:
4588:
4584:
4578:
4575:
4574:
4572:
4568:
4562:
4559:
4558:
4556:
4554:
4550:
4544:
4541:
4539:
4536:
4534:
4531:
4529:
4526:
4524:
4521:
4519:
4516:
4515:
4513:
4509:
4503:
4500:
4498:
4495:
4493:
4490:
4488:
4485:
4483:
4480:
4478:
4475:
4474:
4472:
4468:
4462:
4459:
4457:
4454:
4453:
4451:
4447:
4441:
4438:
4436:
4433:
4432:
4430:
4426:
4420:
4417:
4415:
4412:
4410:
4407:
4405:
4402:
4400:
4399:Pedra Pintada
4397:
4395:
4392:
4390:
4387:
4386:
4384:
4380:
4374:
4371:
4369:
4366:
4364:
4361:
4360:
4358:
4354:
4348:
4345:
4344:
4342:
4338:
4332:
4329:
4328:
4326:
4322:
4318:
4314:
4313:South America
4310:
4305:
4301:
4287:
4284:
4283:
4281:
4277:
4271:
4268:
4266:
4263:
4262:
4260:
4256:
4250:
4247:
4246:
4244:
4240:
4234:
4231:
4229:
4226:
4224:
4221:
4220:
4218:
4214:
4208:
4205:
4203:
4200:
4198:
4195:
4193:
4190:
4188:
4185:
4183:
4180:
4178:
4177:Klasies River
4175:
4173:
4170:
4168:
4165:
4163:
4160:
4158:
4155:
4153:
4152:Byneskranskop
4150:
4148:
4145:
4143:
4140:
4138:
4135:
4132:
4131:
4128:
4125:
4123:
4120:
4118:
4115:
4113:
4110:
4108:
4105:
4103:
4100:
4098:
4095:
4093:
4090:
4088:
4085:
4083:
4080:
4078:
4075:
4073:
4070:
4068:
4065:
4063:
4060:
4058:
4055:
4053:
4050:
4049:
4047:
4043:
4037:
4034:
4032:
4029:
4027:
4024:
4023:
4021:
4017:
4011:
4008:
4007:
4005:
4001:
3995:
3992:
3990:
3987:
3986:
3984:
3980:
3974:
3971:
3970:
3968:
3964:
3958:
3955:
3953:
3950:
3948:
3945:
3943:
3942:Ifri Oudadane
3940:
3938:
3935:
3933:
3930:
3928:
3925:
3924:
3922:
3918:
3912:
3911:Uan Muhuggiag
3909:
3907:
3904:
3903:
3901:
3897:
3891:
3888:
3887:
3885:
3881:
3875:
3872:
3870:
3867:
3866:
3864:
3860:
3854:
3851:
3849:
3846:
3845:
3843:
3839:
3833:
3830:
3829:
3827:
3823:
3817:
3814:
3813:
3811:
3807:
3801:
3798:
3796:
3793:
3792:
3790:
3786:
3780:
3777:
3776:
3774:
3770:
3766:
3762:
3757:
3753:
3739:
3736:
3734:
3731:
3730:
3728:
3724:
3718:
3715:
3713:
3710:
3709:
3707:
3703:
3697:
3694:
3692:
3689:
3687:
3684:
3683:
3681:
3677:
3673:
3669:
3665:
3659:
3656:
3654:
3651:
3649:
3646:
3645:
3643:
3639:
3633:
3630:
3628:
3625:
3623:
3620:
3618:
3617:Batadombalena
3615:
3614:
3612:
3608:
3602:
3599:
3597:
3594:
3592:
3589:
3587:
3584:
3582:
3579:
3578:
3576:
3572:
3566:
3563:
3562:
3560:
3558:
3554:
3548:
3545:
3544:
3542:
3538:
3532:
3529:
3528:
3526:
3522:
3516:
3513:
3512:
3510:
3506:
3500:
3497:
3496:
3494:
3490:
3484:
3481:
3479:
3476:
3474:
3473:Ras Baalbek I
3471:
3469:
3466:
3464:
3461:
3459:
3456:
3454:
3451:
3450:
3448:
3444:
3438:
3435:
3433:
3430:
3429:
3427:
3423:
3417:
3414:
3413:
3411:
3407:
3401:
3398:
3396:
3393:
3391:
3388:
3386:
3383:
3381:
3378:
3376:
3373:
3371:
3368:
3366:
3363:
3362:
3360:
3356:
3350:
3347:
3345:
3342:
3340:
3337:
3335:
3332:
3330:
3327:
3325:
3322:
3320:
3317:
3315:
3312:
3310:
3309:Nahal Me'arot
3307:
3305:
3302:
3300:
3297:
3295:
3292:
3290:
3287:
3285:
3282:
3281:
3279:
3275:
3269:
3266:
3264:
3261:
3260:
3258:
3254:
3248:
3245:
3243:
3240:
3238:
3235:
3233:
3230:
3228:
3225:
3223:
3220:
3218:
3215:
3213:
3210:
3208:
3205:
3204:
3202:
3198:
3192:
3189:
3187:
3184:
3182:
3179:
3177:
3174:
3173:
3171:
3167:
3161:
3158:
3156:
3153:
3151:
3148:
3146:
3143:
3141:
3138:
3136:
3133:
3131:
3128:
3127:
3125:
3121:
3115:
3112:
3111:
3109:
3105:
3099:
3096:
3094:
3091:
3089:
3086:
3085:
3083:
3079:
3073:
3070:
3068:
3065:
3063:
3060:
3058:
3055:
3053:
3050:
3048:
3045:
3043:
3040:
3038:
3035:
3033:
3030:
3028:
3025:
3024:
3022:
3018:
3012:
3009:
3008:
3006:
3002:
2996:
2993:
2991:
2988:
2986:
2983:
2981:
2978:
2976:
2973:
2971:
2968:
2966:
2963:
2962:
2960:
2956:
2950:
2947:
2946:
2944:
2940:
2934:
2931:
2930:
2928:
2924:
2920:
2916:
2911:
2907:
2893:
2892:Windmill Hill
2890:
2888:
2885:
2883:
2880:
2878:
2875:
2873:
2870:
2868:
2865:
2863:
2860:
2858:
2855:
2853:
2850:
2848:
2845:
2843:
2840:
2838:
2837:Heathery Burn
2835:
2833:
2830:
2828:
2825:
2823:
2820:
2818:
2815:
2813:
2810:
2808:
2805:
2804:
2802:
2798:
2792:
2789:
2788:
2786:
2782:
2776:
2773:
2771:
2768:
2767:
2765:
2761:
2755:
2752:
2750:
2747:
2745:
2742:
2740:
2737:
2735:
2732:
2730:
2727:
2725:
2722:
2720:
2717:
2715:
2712:
2710:
2707:
2705:
2702:
2700:
2697:
2696:
2693:
2690:
2688:
2685:
2683:
2680:
2678:
2675:
2673:
2670:
2668:
2665:
2663:
2660:
2658:
2655:
2653:
2650:
2648:
2645:
2643:
2640:
2638:
2635:
2633:
2630:
2627:
2626:
2623:
2620:
2618:
2615:
2613:
2610:
2609:
2606:
2603:
2601:
2598:
2595:
2592:
2591:
2588:
2585:
2583:
2580:
2578:
2575:
2573:
2572:Tito Bustillo
2570:
2565:
2561:
2558:
2555:
2553:Las Chimeneas
2552:
2550:
2547:
2546:
2545:
2542:
2539:
2537:
2534:
2531:
2529:
2526:
2525:
2524:
2521:
2519:
2516:
2514:
2511:
2510:
2508:
2504:
2498:
2495:
2493:
2490:
2488:
2485:
2483:
2480:
2479:
2477:
2473:
2467:
2464:
2462:
2459:
2457:
2454:
2453:
2451:
2447:
2441:
2438:
2436:
2433:
2431:
2428:
2426:
2423:
2422:
2420:
2416:
2410:
2407:
2405:
2402:
2400:
2397:
2395:
2392:
2390:
2387:
2385:
2382:
2380:
2379:Akhshtyrskaya
2377:
2376:
2374:
2370:
2364:
2361:
2359:
2356:
2354:
2351:
2349:
2346:
2345:
2343:
2339:
2333:
2330:
2328:
2327:Pego do Diabo
2325:
2323:
2320:
2318:
2315:
2313:
2310:
2308:
2305:
2303:
2300:
2298:
2295:
2294:
2292:
2288:
2282:
2279:
2277:
2274:
2273:
2271:
2267:
2261:
2258:
2257:
2255:
2251:
2245:
2242:
2241:
2239:
2237:
2233:
2227:
2224:
2223:
2221:
2217:
2211:
2208:
2207:
2205:
2201:
2195:
2192:
2191:
2189:
2185:
2179:
2176:
2175:
2173:
2171:
2167:
2161:
2158:
2156:
2153:
2151:
2148:
2146:
2145:
2141:
2139:
2136:
2134:
2131:
2129:
2126:
2124:
2121:
2119:
2118:Arene Candide
2116:
2114:
2111:
2110:
2108:
2104:
2098:
2095:
2093:
2090:
2089:
2087:
2083:
2077:
2074:
2072:
2069:
2067:
2064:
2062:
2059:
2057:
2054:
2052:
2049:
2047:
2044:
2042:
2039:
2037:
2034:
2032:
2029:
2028:
2026:
2022:
2016:
2013:
2011:
2008:
2006:
2005:Devil's Tower
2003:
2002:
2000:
1998:
1994:
1988:
1985:
1983:
1980:
1978:
1975:
1973:
1970:
1968:
1965:
1962:
1961:
1958:
1955:
1953:
1950:
1948:
1945:
1943:
1940:
1938:
1935:
1933:
1930:
1928:
1925:
1924:
1922:
1918:
1912:
1909:
1907:
1904:
1902:
1899:
1897:
1894:
1892:
1889:
1887:
1884:
1882:
1879:
1877:
1874:
1872:
1869:
1867:
1864:
1862:
1859:
1857:
1854:
1852:
1849:
1846:
1844:
1841:
1839:
1836:
1833:
1830:
1827:
1824:
1822:
1819:
1817:
1814:
1812:
1809:
1808:
1805:
1802:
1800:
1797:
1795:
1792:
1790:
1787:
1785:
1782:
1780:
1779:Pair-non-Pair
1777:
1775:
1772:
1770:
1767:
1765:
1764:Le Mas-d'Azil
1762:
1760:
1757:
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487:Vindija 33.19
484:
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479:Mezmaiskaya 1
476:
475:Vindija 33.19
472:
467:
465:
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456:Vindija 33.19
453:
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448:Vindija 33.19
445:
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441:
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436:Vindija 33.16
432:
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404:
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396:Vindija 33.19
393:
389:
388:Denisova Cave
385:
381:
380:Vindija 33.19
377:
376:Vindija 33.19
373:
368:
366:
365:Vindija 33.26
362:
361:Vindija 33.25
358:
357:Vindija 33.16
354:
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228:modern humans
225:
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101:
97:
73:
45:
35:
28:
23:
5717:
5695:
5692:Vindija Cave
5691:
5675:
5663:
5631:
5623:
5615:
5595:
5587:
5579:
5571:
5563:
5543:
5535:
5527:
5519:
5511:
5503:
5495:
5463:
5455:
5447:
5439:
5431:
5423:
5168:Wargata Mina
5128:Murrawijinie
5063:Eagles Reach
5058:Devil's Lair
4962:Wilson Butte
4747:Cherry Creek
4737:Bull Thistle
4482:Chiribiquete
4440:Charlie Lake
4368:Barton Creek
4182:Melkhoutboom
4122:Sterkfontein
4112:Plovers Lake
4045:South Africa
3947:Jebel Irhoud
3932:Ifri N'Ammar
3667:Turkmenistan
3632:Hunugalagala
3416:Iraq ed-Dubb
3181:Jeriji Saléh
2842:High Pasture
2399:Shulgan-Tash
2322:Pedra Furada
2143:
1982:Lichtenstein
1972:Brillenhöhle
1901:Le Regourdou
1861:La Ferrassie
1856:Combe Grenal
1799:Trois Frères
1734:Fontéchevade
1687:La Madeleine
1503:
1341:Drachenhöhle
1203:. Retrieved
1192:
1153:
1149:
1139:Malez, Mirko
1110:
1106:
1050:
1046:
988:
984:
944:
898:
894:
836:
832:
776:
772:
722:
718:
706:Bibliography
692:
680:
668:
656:
625:
613:
601:. Retrieved
589:
579:
491:introgressed
486:
482:
478:
474:
468:
455:
451:
447:
443:
439:
435:
433:
426:
422:
414:
410:
400:
395:
391:
383:
379:
375:
374:genome from
369:
364:
360:
356:
353:low coverage
350:
329:
310:
287:
283:
277:
271:
264:
253:
224:Neanderthals
220:Vindija Cave
219:
218:
31:Vindija Cave
20:Vindija Cave
5668:NK Varaždin
5612:Špancirfest
5540:Town Museum
5254:New Zealand
5078:Gwion Gwion
4947:Trail Creek
4872:Meadowcroft
4832:Last Supper
4762:Daugherty's
4624:Las Flechas
4409:Rei do Mato
4133:Other caves
4117:Rising Star
4092:Makapansgat
4057:Bolt's Farm
3874:Njoro River
3717:Teshik-Tash
3712:Obi-Rakhmat
3696:Yarımburgaz
3596:Lapuz Lapuz
3574:Philippines
3483:Nachcharini
3478:Ras El Kelb
3437:Tam Pa Ling
3432:Tham An Mah
3314:Nahal Hemar
3062:Zhoukoudian
3037:Guanyindong
3011:Laang Spean
2933:Darra-e Kur
2926:Afghanistan
2872:Portbraddon
2862:Marble Arch
2775:Wildkirchli
2763:Switzerland
2699:Other caves
2672:Ojo Guareña
2652:Maltravieso
2647:Los Casares
2622:Siega Verde
2587:Los Aviones
2582:Santimamiñe
2556:Las Monedas
2549:El Castillo
2456:Čertova pec
2404:Mezmaiskaya
2312:Lagar Velho
2244:Cocev Kamen
2123:Balzi Rossi
1963:Other caves
1952:Sirgenstein
1881:Le Moustier
1871:Fontbrégoua
1831:Bonne-Femme
1811:Other caves
1530:Aetokremnos
1473:Utroba Cave
1379:Claminforge
1376:Belle-Roche
1156:: 499–545.
661:Hublin 2017
411:Vindija *28
317:Aurignacian
250:Description
234:, northern
212:Mirko Malez
189:Paleolithic
143: /
119:Coordinates
5771:Mousterian
5740:Categories
5680:NK Varteks
5628:Town Guard
5366:Speleology
5183:Wellington
5138:Naracoorte
5123:Mudgegonga
5108:Madjedbebe
5073:Gabarnmung
5013:Ballawinne
5003:Abrakurrie
4902:Pictograph
4680:Qillqatani
4675:Pikimachay
4670:Guitarrero
4665:Chivateros
4644:Oxtotitlán
4497:Tequendama
4207:Wonderwerk
4187:Nelson Bay
4167:Elands Bay
4127:Swartkrans
4072:Gladysvale
4019:Somaliland
3966:Mozambique
3906:Haua Fteah
3705:Uzbekistan
3370:Minatogawa
3365:Kosegasawa
3319:Nahal Oren
3263:Hazar Merd
3232:Qaleh Bozi
3217:Do-Ashkaft
3114:Satsurblia
3081:East Timor
2958:Azerbaijan
2847:Kendrick's
2812:Bontnewydd
2734:Don Gaspar
2687:Praileaitz
2642:Las Caldas
2560:La Pasiega
2487:Divje Babe
2409:Okladnikov
2394:Ignatievka
2317:Pala Pinta
2226:Għar Dalam
2203:Luxembourg
2178:St Brelade
2160:Villabruna
2071:Stravomyti
2061:Megalakkos
2046:Eileithyia
2031:Alepotrypa
1942:Hohle Fels
1828:Balauzière
1784:Pech Merle
1692:Rouffignac
1666:La Micoque
1648:Cro-Magnon
1607:Bara Bahau
1551:Býčí skála
1499:Vela Spila
1458:Devetashka
1453:Bacho Kiro
1078:1887/74733
533:References
452:Vindija 87
392:Vindija 87
313:Mousterian
279:Shanidar 1
268:brow ridge
232:Donja Voća
203:Site notes
128:46°17′57″N
103:Donja Voća
5643:Education
5560:Cathedral
5524:Town Hall
5436:1776 Fire
5332:Nanumanga
5316:Falemauga
5227:Makauwahi
4996:Australia
4952:Trinchera
4827:La Grange
4797:Gatecliff
4792:Franktown
4787:Fort Rock
4767:Dry Creek
4685:Toquepala
4649:Sac Actun
4639:Ox Bel Ha
4629:Frightful
4614:Coxcatlan
4593:Long Mile
4347:Quadiriki
4324:Argentina
4162:Diepkloof
4147:Boomplaas
4102:Minnaar's
4087:Kromdraai
4036:Laas Geel
4031:Dhaymoole
4026:Dhambalin
3989:Apollo 11
3890:Liphofung
3816:Shum Laka
3779:Gueldaman
3733:Con Moong
3610:Sri Lanka
3557:Palestine
3531:Padah-Lin
3468:Ksar Akil
3400:Yamashita
3385:Pinza-Abu
3191:Pettakere
3186:Liang Bua
3169:Indonesia
3155:Pahargarh
3140:Bhimbetka
3098:Lene Hara
3088:Jerimalai
3067:Zengpiyan
3057:Yuchanyan
2857:Long Hole
2754:del Valle
2729:dels Bous
2682:la Pileta
2617:Atapuerca
2532:Covalanas
2430:Pešturina
2358:Muierilor
2348:Coliboaia
2260:Svarthola
2076:Theopetra
2066:Petralona
2051:Franchthi
2041:Asphendou
1997:Gibraltar
1967:Baumann's
1957:Vogelherd
1932:Bockstein
1911:Vallonnet
1906:Rochereil
1896:Raymonden
1886:Noisetier
1838:Bruniquel
1759:Marsoulas
1749:Lombrives
1744:La Marche
1670:La Mouthe
1636:Abri Audi
1614:Cap Blanc
1556:Koněprusy
1546:Amatérská
1514:Veternica
1463:Kozarnika
1437:Ledenjača
1361:Tischofer
1351:Lurgrotte
1127:0047-2484
1087:0036-8075
1013:0036-8075
971:1877-9077
917:0027-8424
861:0028-0836
801:0036-8075
741:0027-8424
538:Citations
131:16°4′14″E
5674:)
5548:Cemetery
5492:Old City
5408:Varaždin
5173:Warratyi
5158:Tarragal
5148:Nunamira
5133:Murujuga
5098:Kutikina
5093:Koonalda
5088:Koongine
5048:Cliefden
5043:Cave Bay
4942:Tomaquag
4927:Sisyphus
4857:Mantle's
4847:Lovelock
4842:LoDaisKa
4822:Humboldt
4706:Werehpai
4699:Suriname
4609:Chan Hol
4528:Centella
4518:Ambrosio
4470:Colombia
4435:Bluefish
4279:Zimbabwe
4216:Tanzania
4202:Stadsaal
4107:Motsetsi
4077:Gondolin
4067:Drimolen
4062:Cooper's
3957:Taforalt
3927:Hercules
3853:Swimmers
3825:DR Congo
3809:Cameroon
3788:Botswana
3658:Tham Lod
3648:Pha Taem
3641:Thailand
3622:Belilena
3540:Pakistan
3508:Mongolia
3492:Malaysia
3453:Antelias
3395:Tobayama
3349:Zuttiyeh
3268:Shanidar
3130:Ambadevi
3047:Tianyuan
3027:Baishiya
3004:Cambodia
2744:El Mirón
2739:Guanches
2714:Armintxe
2632:Bacinete
2577:Altxerri
2566:El Pendo
2536:La Garma
2518:Altamira
2475:Slovenia
2466:Jasovská
2449:Slovakia
2435:Risovača
2389:Denisova
2353:Cuciulat
2307:Furninha
2302:Escoural
2290:Portugal
2281:Paradise
2276:Obłazowa
2210:Loschbur
2150:Paglicci
2133:Corbeddu
2015:Vanguard
2010:Gorham's
1891:La Quina
1834:Bouillon
1821:Aurignac
1625:Reverdit
1610:Bernifal
1489:Grapčeva
1446:Bulgaria
1406:Scladina
1393:Naulette
1356:Salzofen
1243:Archived
1217:Archived
1199:Archived
1194:Nacional
1095:28450384
1031:28982794
935:28973864
889:(2017).
879:29562232
819:20448178
759:28874524
501:See also
464:Caucasus
372:coverage
343:humans.
256:Varaždin
240:sequence
99:Location
5607:Culture
5415:History
5284:Chugai'
5266:Ruakuri
5201:Gadao's
5178:Warreen
5113:Mammoth
5083:Jenolan
5028:Blanche
5008:Acheron
4985:Oceania
4957:Ventana
4937:Tainter
4912:Russell
4897:Pendejo
4892:Paisley
4852:Mammoth
4772:Durango
4732:Bonfire
4586:Jamaica
4553:Curaçao
4492:Sáchica
4477:El Abra
4461:Milodón
4404:Peruaçu
4394:Maquiné
4286:Bambata
4265:Kalemba
4137:Blombos
4082:Haasgat
4003:Nigeria
3982:Namibia
3920:Morocco
3883:Lesotho
3795:Tsodilo
3772:Algeria
3726:Vietnam
3686:Belbaşı
3672:Dzhebel
3627:Fa Hien
3586:Kalanay
3547:Sanghao
3524:Myanmar
3463:Kaukaba
3446:Lebanon
3334:Raqefet
3304:Misliya
3289:HaYonim
3237:Warwasi
3212:Darband
3207:Bisitun
3176:Harimau
3150:Gudiyam
3145:Edakkal
3107:Georgia
3052:Xianren
2980:Damjili
2975:Buzeyir
2949:Areni-1
2942:Armenia
2867:Oldbury
2832:Gough's
2817:Cathole
2791:Verteba
2784:Ukraine
2341:Romania
2332:Salemas
2297:Aroeira
2128:Cavallo
2113:Addaura
2092:Baradla
2085:Hungary
2036:Apidima
1920:Germany
1876:Lazaret
1804:Villars
1724:Cosquer
1704:Chauvet
1682:Lascaux
1580:Finland
1504:Vindija
1494:Krapina
1482:Croatia
1402:Ramioul
1370:Belgium
1346:Gudenus
1334:Austria
1235:YouTube
1205:15 July
1055:Bibcode
1047:Science
1022:6185897
993:Bibcode
985:Science
926:5635937
870:6485383
841:Bibcode
810:5100745
781:Bibcode
773:Science
750:5635904
590:Science
471:alleles
462:in the
289:Tabun 1
242:of the
236:Croatia
185:Periods
180:History
5718:·
5716:
5696:·
5694:
5687:Nature
5678:
5676:·
5672:former
5664:·
5662:
5632:·
5630:
5624:·
5622:
5616:·
5614:
5596:·
5594:
5588:·
5586:
5580:·
5578:
5572:·
5570:
5564:·
5562:
5544:·
5542:
5536:·
5534:
5528:·
5526:
5520:·
5518:
5512:·
5510:
5504:·
5502:
5496:·
5494:
5464:·
5462:
5456:·
5454:
5448:·
5446:
5440:·
5438:
5432:·
5430:
5424:·
5422:
5325:Tuvalu
5261:Moncks
5245:Pindai
5220:Hawaii
5211:Talagi
5206:Mahlac
5068:Fossil
5053:Cloggs
5038:Burrup
4917:Sandia
4862:Marmes
4807:Hidden
4802:Graham
4757:Danger
4742:Burnet
4602:Mexico
4577:Pomier
4538:Patana
4523:Calero
4502:Tibitó
4428:Canada
4389:Gentio
4382:Brazil
4356:Belize
4270:Mumbwa
4258:Zambia
4242:Uganda
4223:Kondoa
4197:Sibudu
4142:Border
4097:Malapa
3973:Ngalue
3848:Beasts
3832:Matupi
3761:Africa
3691:Karain
3679:Turkey
3653:Spirit
3591:Minori
3581:Callao
3565:Shuqba
3409:Jordan
3380:Ōzakai
3375:Muroya
3324:Qafzeh
3294:Kebara
3277:Israel
3247:Yafteh
3242:Wezmeh
3072:Zhiren
2985:Tağlar
2887:Thor's
2822:Coygan
2770:Bichon
2724:Bedmar
2692:Sidrón
2528:Chufín
2461:Domica
2418:Serbia
2372:Russia
2269:Poland
2253:Norway
2187:Kosovo
2170:Jersey
2155:Romito
2097:Szelim
2056:Kleidi
2024:Greece
1729:Cussac
1719:Gargas
1674:Pataud
1596:France
1566:Mladeč
1523:Cyprus
1468:Magura
1432:Badanj
1425:Bosnia
1323:Europe
1125:
1093:
1085:
1029:
1019:
1011:
969:
959:
933:
923:
915:
877:
867:
859:
833:Nature
817:
807:
799:
757:
747:
739:
603:2 June
405:, and
296:Dating
286:, and
260:Ivanec
172:Height
156:Length
109:Region
5720:
5698:
5666:
5655:Sport
5634:
5626:
5618:
5598:
5590:
5582:
5574:
5566:
5546:
5538:
5530:
5522:
5514:
5506:
5498:
5466:
5458:
5450:
5442:
5434:
5426:
5309:Samoa
5163:Ubirr
5153:ORS 7
4922:Shoup
4882:Mummy
4877:Modoc
4867:Martz
4817:Hogup
4727:Baker
4543:Pluma
4449:Chile
4414:Santo
4340:Aruba
4309:North
4249:Nyero
4233:Mumba
4157:Cango
3899:Libya
3862:Kenya
3841:Egypt
3738:Ngườm
3601:Tabon
3458:Jeita
3358:Japan
3344:Tabun
3339:Skhul
3329:Qesem
3299:Manot
3160:Tikla
3135:Belum
3123:India
3093:Laili
3042:Luobi
3032:Fuyan
3020:China
2995:Gazma
2970:Azykh
2965:Allar
2852:Kents
2719:Axlor
2709:Antón
2704:Ángel
2662:Nerja
2600:Araña
2506:Spain
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2492:Pekel
2482:Betal
2219:Malta
2144:Felci
2106:Italy
1987:Ofnet
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1843:Calès
1816:Arago
1794:Renne
1774:Niaux
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3284:Amud
3256:Iraq
3200:Iran
2915:Asia
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2667:Niño
2363:Oase
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