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Shishugou Formation

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wallowing of large dinosaurs. Small theropods and vertebrates then became mired in these pits, dying and being forced deeper by the activities of the large dinosaurs and the struggles of later victims. The high quality of preservation suggests a rapid burying of the carcasses. Some scavenging of the bodies took place leading to a dispersal of body parts. Smaller vertebrates were prone to getting stuck in these mudholes, and the small ceratosaur,
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reaching around 6 meters (20 feet) in length and weighed 2.5 metric tons. It had three distinguishing traits: the crowns of the teeth are symmetrical and in side view wider than tall; the spine of the axis, the second neck vertebra, has a rectangular profile in side view, instead of a triangular one;
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At the Wuwaican locality, the formation is approximately 380 m thick, the lower 30 metres of the formation predominantly consist of conglomerate, with the majority of the formation consisting of red coloured mudstone with frequent channel/sheet sandstone lenses and occasional tuffaceous deposits. It
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These 'traps' or 'bonebeds' are unusual in that they consist of vertically stacked skeletons of numerous non-avian theropods in 1–2 metres (3.3–6.6 ft) deep pits. The pits are filled with a mix of alluvial and volcanic mudstone and sandstone and appear to have been created by the trampling and
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An alvarezsaur that was the largest known definite member of the superfamily, at around 2 meters long. It had an enlarged thumb claw like other alvarezsaurs, but also retained two other functional fingers, unlike more derived alvarezsaurs, where only the thumb was significantly large and clawed.
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When the site was first excavated, new species of turtles, crocodilians, pterosaurs, and early mammals were revealed. Many of these species showed the rudiments of characteristics that would later become their hallmarks. One of the first fossil skeletons exposed proved to be that of an unknown
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Jonah N. Choiniere, James M. Clark, Catherine A. Forster and Xing Xu (2010). "A basal coelurosaur (Dinosauria: Theropoda) from the Late Jurassic (Oxfordian) of the Shishugou Formation in Wucaiwan, People’s Republic of China". Journal of Vertebrate Paleontology 30 (6): 1773–1796.
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are shedding light on this period previously notorious for its paucity of fossils, but important for its massive burst of speciation among the dinosaurs and ancestors of birds. These new branches on the dinosaur family tree led to well-known dinosaur groups, such as the horned
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On analysis of the rock matrix, large amounts of volcanic ash were found, hinting at eruptions during that period, the ash raining onto the marsh creating viscous mud and becoming subsequent death traps. To date three of these death pits have been located. In one pit a
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At the time the area was covered by marshland, adjoined a small mountain range peppered with volcanoes, and was inhabited by dinosaurs, small crocodilians and amphibians. Currently it is a sparsely settled region of dry washes, arid badlands and dunes along the
520:’s western edge. The Shishugou Formation consists of mudstone, siltstone, and sandstone, and is named for the silicified wood or petrified logs found here. ('Shishugou' = 'stone tree valley') These badlands were used for some sequences in the film 1737: 1781:
Weishampel, David B; et al. (2004). "Dinosaur distribution (Middle Jurassic, Asia)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 541–542.
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Weishampel, David B; et al. (2004). "Dinosaur distribution (Late Jurassic, Asia)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 550–552.
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period. Dinosaur remains are among the fossils that have been recovered from the formation. The Shishugou Formation is considered one of the most phylogenetically and trophically diverse Middle to Late Jurassic
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with a remarkably long neck which made up half the total body length. It is one of the largest dinosaurs known, measuring 35 meters (115 feet) in length with an 18-meter-long (59 feet) neck.
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Chengkai, Jia; Forster, Catherine A; Xing, Xu; Clark, James M. (2007). "The first stegosaur (Dinosauria, Ornithischia) from the Upper Jurassic Shishugou Formation of Xinjiang, China".
2301:"Overview of Mesozoic crocodylomorphs from the Junggar Basin, Xinjiang, Northwest China, and description of isolated crocodyliform teeth from the Late Jurassic Liuhuanggou locality" 2104:
Xu X., Clark, J.M., Forster, C. A., Norell, M.A., Erickson, G.M., Eberth, D.A., Jia, C., and Zhao, Q. (2006). "A basal tyrannosauroid dinosaur from the Late Jurassic of China".
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Choiniere, J. N.; Xu, X.; Clark, J. M.; Forster, C. A.; Guo, Y.; Han, F. (2010). "A basal alvarezsauroid theropod from the Early Late Jurassic of Xinjiang, China".
1738:"A juvenile specimen of a new coelurosaur (Dinosauria: Theropoda) from the Middle–Late Jurassic Shishugou Formation of Xinjiang, People's Republic of China" 2402: 1652: 1673:"Sedimentary record of Mesozoic intracontinental deformation in the eastern Juggar Basin, northwest China: response to orogeny at the Asian margin" 1647: 472:
Excavations since 2000 have revealed a wealth of fossils dating from a period some 165 to 155 million years ago. Palaeontologists James Clark and
2397: 786:). It was bipedal and had a total length of about 1.2 meters (3.9 feet) from nose to tail, and a weight of about 15 kilograms (33 pounds). 1845: 1310:
named for the single crest on top of its skull. It is estimated to have a length at 5.5 meters (16.4 feet), the weight at 475 kilograms.
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The Wucaiwan Member, once considered a separate, underlying formation, is now considered the lowest unit of the Shishugou Formation.
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Undescribed stegosaur is present in the Wucaiwan member. Undescribed ornithopod is present in the Wucaiwan member. Undescribed
522: 381: 493:, to explore the Junggar Basin, and in particular the Shishugou Formation, where the exposed rocks have been linked to the 2437: 2363: 587:, in another three decapitated ceratosaurs. The most interesting of the traps gave up the fossil remains of an 80 kg 407: 373: 1193:, about 3 meters (9.8 feet). It has a large distinctive crest on its head, a feature seen in many primitive tyrannosaurs 2452: 2432: 221: 559: 267: 228: 2417: 2065:"A new titanosauriform dinosaur (Dinosauria: Sauropoda) from Late Jurassic of Junggar Basin, Xinjiang (In Chinese)" 2457: 2447: 465: 994:
estimated to have been approximately 98 feet long, which would then make it one of the longest known dinosaurs.
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Xu, X.; Clark, JM; Mo, J; Choiniere, J; Forster, CA; Erickson, GM; Hone, DW; Sullivan, C; et al. (2009).
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accompanied the first expedition to the Junggar Basin in 1928, and the fossils found included the medium-size
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A short-necked sauropod only known from juvenile specimens which measured about 4.8 meters (16 feet) long.
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theropod standing nearly 3 meters (9.8 feet) tall and measuring roughly 7.6 meters (25 feet) in length.
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Wings, Oliver; Schwarz-Wings, Daniela; Pfretzschner, Hans-Ulrich; Martin, Thomas (September 2010).
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Eberth, D. A.; Xing, X.; Clark, J. M. (2010). "Dinosaur Deathpits from the Jurassic of China".
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Choiniere JN; Clark JM; Forster CM; Norell MA; Eberth DA; Erickson GM; Chu H; Xu X (2013).
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that was at best 1 meter (3.3 feet) long and weighed 2 kilograms (4.4 pounds) at most.
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are pit structures found within the formation that are noted for their fossil content.
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Found on the same slab that also preserved the holotype and an associated specimen of
2391: 2340: 2300: 2015: 1832: 1498: 1433: 1002: 973: 599: 414: 331: 87: 40: 2206: 2050: 1951: 1761: 1517: 505: 2267: 2141: 1863:"A New Taxon of Basal Ceratopsian from China and the Early Evolution of Ceratopsia" 1609: 1554: 1408:. The specimen known is probably from a juvenile individual and is quite complete. 1268: 1208: 1190: 865: 830: 673: 486: 478: 17: 2286: 1889: 1753: 1279: 1158: 1151: 1019: 942: 745: 710: 617: 580: 517: 62: 1824: 1026: 2324: 2042: 1528: 1482: 1412: 1264: 1240: 892: 664: 584: 482: 107: 72: 2332: 282: 269: 243: 230: 2190: 1405: 1349: 1307: 962: 571: 205: 112: 56: 44: 2259: 2198: 2133: 2000: 1982: 1908: 854: 2377: 1679:. Colorado, US: The Geological Society of America, Inc. pp. 354–356. 1373: 2225:"A Jurassic ceratosaur from China helps clarify avian digital homologies" 1867: 1169: 912: 678:. It is estimated at around 1.3 to 1.5 meters (4.3 to 4.9 ft) long. 593: 555: 547: 419: 403: 302: 191: 102: 97: 82: 77: 67: 2251: 2125: 1384: 1260: 1062: 790: 766: 528: 117: 92: 1516: 550:, and revealed a stack of four more theropods buried below the first. 1677:
Paleozoic and Mesozoic Tectonic Evolution of Central and Eastern Asia
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and the rear neck vertebrae have large vein openings in their sides.
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approximately 3 meters (9.8 feet) long and weighs about equal to a
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identified the following species from the Shishugou Formation.
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had a small slender body measuring about 1.7 meters in length.
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with a wingspan that has been estimated at least 1.73 meters.
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Institute of Vertebrate Paleontology and Paleoanthropology
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A very basal and most primitive ceratopsian (possibly a
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Proceedings of the Royal Society B: Biological Sciences
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with an estimated wingspan of 1.47 meters (4.8 feet).
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present in both upper Shishugou and Wucaiwan members.
2094:"Table 13.1," in Weishampel, et al. (2004). Page 271. 2085:"Table 13.1," in Weishampel, et al. (2004). Page 262. 1857:
Han, Fenglu; Forster, Catherine A.; Clark, James M.;
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The Shishugou Formation dinosaur traps also known as
1965:Xu, X.; Forster, C.A.; Clark, J.M.; Mo, J. (2006). 326: 308: 298: 259: 220: 215: 197: 187: 182: 174: 164: 154: 146: 136: 126: 34: 2354:Yaoming Hu, Jin Meng & James M. Clark (2009). 575:. In all, some 600 specimens have been collected. 1107:-like species. Indeterminate tetanuran remains. 1022:measuring 15 meters and weighing 5 metric tons. 469:, is the most common in the resulting bonebeds. 1090:A mamenchisaurid known from very few remains. 8: 2154:: CS1 maint: multiple names: authors list ( 1263:that is the first definitely known Jurassic 532:('hidden dragon') being named for the film. 2014:Andres, B.; Clark, J. M.; Xing, X. (2010). 1846:China's Junggar Basin - National Geographic 438:in the southern half of the Junggar Basin. 350: 1488: 31: 2463:Fossiliferous stratigraphic units of Asia 2376: 2305:Palaeobiodiversity and Palaeoenvironments 2218: 2216: 1990: 1898: 1888: 1653:List of dinosaur-bearing rock formations 910: 1924:Acta Geologica Sinica (English Edition) 1731: 1729: 1727: 1675:. In Hendrix MS & Davis GA (eds.). 1663: 2147: 7: 1798: 1796: 1477:A single fully articulated skeleton 1367:metriacanthosaurine metriacanthosaur 1742:Journal of Systematic Palaeontology 2023:Journal of Vertebrate Paleontology 1944:10.1111/j.1755-6724.2007.tb00959.x 1671:Vincent SJ & Allen MB (2001). 1341:A little-known early alvarezsaur. 489:. Clark joined Xu, from Beijing's 442:Shishugou Formation dinosaur traps 25: 1087:"Partial postcranial skeleton." 349: 342: 313: 2403:Geologic formations of Mongolia 434:is laterally equivalent to the 358:Shishugou Formation (Dzungaria) 1648:Jurassic paleontological sites 1051:"Partial skull and skeleton." 523:Crouching Tiger, Hidden Dragon 394: 385: 377: 1: 2364:Acta Palaeontologica Polonica 1189:that was one of the earliest 2398:Geologic formations of China 2287:10.1080/02724634.2010.520779 1890:10.1371/journal.pone.0143369 1754:10.1080/14772019.2013.781067 671:), previously classified as 540:Mamenchisaurus sinocanadorum 413:Its strata date back to the 261:Approximate paleocoordinates 560:Tienshanosaurus chitaiensis 27:Geologic formation in China 2484: 1825:10.2110/palo.2009.p09-028r 915:of the Shishugou Formation 583:was found against a small 178:380 m (1,250 ft) 2325:10.1007/s12549-010-0033-1 2043:10.1080/02724630903409220 1483:Limusaurus inextricabilis 667:(or perhaps a more basal 466:Limusaurus inextricabilis 337: 39: 2468:Paleontology in Xinjiang 1256:A toothless, omnivorous 2408:Jurassic System of Asia 2191:10.1126/science.1182143 1594:Stratigraphic position 1452:Stratigraphic position 1119:Stratigraphic position 927:Stratigraphic position 815:Stratigraphic position 632:Stratigraphic position 1983:10.1098/rspb.2006.3566 1521: 1416: 1377: 1318: 1283: 1233: 1201: 1162: 1066: 1030: 966: 896: 858: 794: 758: 721: 686: 542: 512: 2378:10.4202/app.2008.0053 1520: 1428:After an overview of 1415: 1376: 1317: 1282: 1232: 1200: 1161: 1065: 1029: 987:Complete right femur 965: 895: 857: 793: 757: 720: 685: 663:An unspecified basal 607:Vertebrate paleofauna 538: 508: 2438:Sandstone formations 569:and the carnivorous 400:geological formation 131:Geological formation 2453:Lacustrine deposits 2433:Mudstone formations 2317:2010PdPe...90..283W 2252:10.1038/nature08124 2244:2009Natur.459..940X 2183:2010Sci...327..571C 2126:10.1038/nature04511 2118:2006Natur.439..715X 2035:2010JVPal..30..163A 1977:(1598): 2135–2140. 1936:2007AcGlS..81..351J 1881:2015PLoSO..1043369H 1861:(9 December 2015). 1817:2010Palai..25..112E 1271:, including China. 655:"E. wucaiwanensis" 382:traditional Chinese 370:Shishugou Formation 279: /  240: /  169:Xishanyao Formation 41:Stratigraphic range 35:Shishugou Formation 1522: 1417: 1378: 1319: 1284: 1234: 1202: 1163: 1067: 1031: 967: 897: 859: 795: 759: 722: 687: 543: 513: 374:simplified Chinese 18:Wucaiwan Formation 2418:Jurassic Mongolia 2238:(7249): 940–944. 2177:(5965): 571–574. 2112:(7077): 715–718. 1634: 1633: 1578: 1577: 1421: 1420: 1247:L. inextricabilis 1096: 1095: 901: 900: 799: 798: 600:Tyrannosaurus rex 366: 365: 16:(Redirected from 2475: 2458:Paludal deposits 2448:Fluvial deposits 2383: 2382: 2380: 2360: 2351: 2345: 2344: 2296: 2290: 2278: 2272: 2271: 2229: 2220: 2211: 2210: 2166: 2160: 2159: 2153: 2145: 2101: 2095: 2092: 2086: 2083: 2077: 2076: 2075:: 581–588. 2019. 2061: 2055: 2054: 2020: 2011: 2005: 2004: 1994: 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176: 172: 171: 166: 162: 161: 156: 152: 151: 148: 144: 143: 138: 134: 133: 128: 124: 123: 116: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 60: 37: 36: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 2480: 2469: 2466: 2464: 2461: 2459: 2456: 2454: 2451: 2449: 2446: 2444: 2441: 2439: 2436: 2434: 2431: 2429: 2426: 2424: 2421: 2419: 2416: 2414: 2411: 2409: 2406: 2404: 2401: 2399: 2396: 2395: 2393: 2379: 2374: 2370: 2366: 2365: 2357: 2350: 2347: 2342: 2338: 2334: 2330: 2326: 2322: 2318: 2314: 2310: 2306: 2302: 2295: 2292: 2288: 2284: 2277: 2274: 2269: 2265: 2261: 2257: 2253: 2249: 2245: 2241: 2237: 2233: 2226: 2219: 2217: 2213: 2208: 2204: 2200: 2196: 2192: 2188: 2184: 2180: 2176: 2172: 2165: 2162: 2157: 2151: 2143: 2139: 2135: 2131: 2127: 2123: 2119: 2115: 2111: 2107: 2100: 2097: 2091: 2088: 2082: 2079: 2074: 2070: 2066: 2060: 2057: 2052: 2048: 2044: 2040: 2036: 2032: 2028: 2024: 2017: 2010: 2007: 2002: 1998: 1993: 1988: 1984: 1980: 1976: 1972: 1968: 1961: 1958: 1953: 1949: 1945: 1941: 1937: 1933: 1929: 1925: 1918: 1915: 1910: 1906: 1901: 1896: 1891: 1886: 1882: 1878: 1874: 1870: 1869: 1864: 1860: 1853: 1850: 1847: 1842: 1839: 1834: 1830: 1826: 1822: 1818: 1814: 1810: 1806: 1799: 1797: 1793: 1789: 1788:0-520-24209-2 1785: 1778: 1776: 1774: 1772: 1768: 1763: 1759: 1755: 1751: 1747: 1743: 1739: 1732: 1730: 1728: 1724: 1720: 1719:0-520-24209-2 1716: 1709: 1707: 1705: 1703: 1701: 1699: 1697: 1693: 1688: 1682: 1678: 1674: 1667: 1664: 1658: 1654: 1651: 1649: 1646: 1644: 1643:Jurassic Asia 1641: 1640: 1636: 1630: 1628: 1627:tritylodontid 1624: 1622: 1619: 1617: 1614: 1612: 1611: 1607: 1606: 1602: 1599: 1596: 1593: 1590: 1587: 1586: 1580: 1574: 1572: 1569: 1567: 1564: 1562: 1559: 1557: 1556: 1552: 1551: 1548: 1546: 1545:Goniopholidid 1543: 1541: 1538: 1536: 1533: 1531: 1530: 1526: 1525: 1519: 1515: 1513: 1511: 1508: 1506: 1503: 1501: 1500: 1499:Junggarsuchus 1496: 1495: 1491: 1487: 1485: 1484: 1479: 1476: 1473: 1471: 1468: 1465: 1464: 1460: 1457: 1454: 1451: 1448: 1445: 1444: 1441: 1439: 1435: 1434:Junggar Basin 1431: 1423: 1414: 1410: 1407: 1403: 1399: 1397: 1394: 1392: 1389: 1387: 1386: 1382: 1381: 1375: 1371: 1368: 1364: 1362: 1359: 1357: 1354: 1352: 1351: 1347: 1346: 1343: 1340: 1338: 1335: 1333: 1332:S. inexpectus 1330: 1328: 1327: 1323: 1322: 1316: 1312: 1309: 1305: 1303: 1300: 1298: 1295: 1293: 1292: 1288: 1287: 1281: 1277: 1274: 1270: 1266: 1262: 1259: 1255: 1253: 1250: 1248: 1245: 1243: 1242: 1238: 1237: 1231: 1227: 1223: 1221: 1218: 1216: 1213: 1211: 1210: 1206: 1205: 1199: 1195: 1192: 1188: 1184: 1182: 1179: 1177: 1174: 1172: 1171: 1167: 1166: 1160: 1156: 1153: 1149: 1147: 1144: 1142: 1139: 1137: 1136: 1132: 1131: 1127: 1124: 1121: 1118: 1115: 1112: 1111: 1108: 1106: 1098: 1092: 1089: 1086: 1083: 1081: 1078: 1076: 1075: 1071: 1070: 1064: 1060: 1057: 1053: 1050: 1047: 1045: 1042: 1040: 1039: 1035: 1034: 1028: 1024: 1021: 1017: 1015: 1012: 1010: 1007: 1005: 1004: 1003:Klamelisaurus 1000: 999: 996: 993: 989: 986: 983: 981: 980:F. qitaiensis 978: 976: 975: 974:Fushanosaurus 971: 970: 964: 960: 957: 955: 952: 950: 947: 945: 944: 940: 939: 935: 932: 929: 926: 923: 920: 919: 916: 914: 909: 903: 894: 890: 887: 884: 880: 878: 875: 873: 870: 868: 867: 863: 862: 856: 852: 849: 845: 843: 840: 838: 837:K. progenitor 835: 833: 832: 828: 827: 823: 820: 817: 814: 811: 808: 807: 801: 792: 788: 785: 781: 779: 776: 774: 771: 769: 768: 764: 763: 760: 756: 751: 747: 743: 741: 738: 736: 733: 731: 730: 726: 725: 719: 715: 712: 708: 706: 703: 701: 698: 696: 695: 691: 690: 684: 680: 677: 676:wucaiwanensis 675: 670: 666: 662: 660: 657: 654: 651: 650: 645: 644: 640: 637: 634: 631: 628: 625: 624: 621: 619: 611: 606: 604: 602: 601: 596: 595: 590: 586: 582: 576: 574: 573: 568: 567: 562: 561: 557: 553: 549: 541: 537: 533: 531: 530: 525: 524: 519: 511: 507: 500: 498: 496: 492: 488: 484: 480: 475: 470: 468: 467: 457: 455: 453: 449: 441: 439: 437: 428: 426: 423: 421: 416: 415:Late Jurassic 411: 409: 405: 401: 397: 391: 383: 375: 371: 345: 336: 333: 332:Junggar Basin 329: 325: 322: 311: 307: 304: 301: 297: 292: 264: 262: 258: 253: 244:44.5°N 90.2°E 225: 223: 219: 214: 211: 207: 203: 200: 196: 193: 190: 186: 181: 177: 173: 170: 167: 163: 160: 157: 153: 149: 145: 142: 139: 135: 132: 129: 125: 119: 114: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 58: 55:163–156  50: 46: 42: 38: 33: 30: 19: 2368: 2362: 2349: 2308: 2304: 2294: 2276: 2235: 2231: 2174: 2170: 2164: 2150:cite journal 2109: 2105: 2099: 2090: 2081: 2072: 2068: 2059: 2026: 2022: 2009: 1974: 1970: 1960: 1927: 1923: 1917: 1872: 1866: 1852: 1841: 1808: 1804: 1745: 1741: 1676: 1666: 1615: 1610:Yuanotherium 1608: 1561:N. matutinus 1560: 1555:Nominosuchus 1553: 1534: 1527: 1504: 1497: 1481: 1437: 1427: 1390: 1383: 1355: 1348: 1331: 1324: 1296: 1289: 1272: 1269:Eastern Asia 1246: 1239: 1214: 1209:Haplocheirus 1207: 1191:tyrannosaurs 1175: 1168: 1140: 1133: 1103:Undescribed 1102: 1079: 1072: 1043: 1036: 1009:K. gobiensis 1008: 1001: 979: 972: 948: 941: 911: 871: 866:Sericipterus 864: 836: 831:Kryptodrakon 829: 772: 765: 752: 734: 727: 699: 692: 674:Gongbusaurus 672: 647: 615: 598: 592: 577: 570: 564: 558: 544: 539: 527: 526:, the genus 521: 514: 509: 487:tyrannosaurs 479:ceratopsians 471: 464: 461: 458:Paleontology 451: 447: 445: 432: 424: 412: 395:Shíshùgōu Zǔ 393: 369: 367: 210:conglomerate 29: 1152:alvarezsaur 943:Bellusaurus 746:stegosaurid 711:ceratopsian 618:ankylosaurs 581:crocodilian 518:Gobi desert 481:, armoured 452:death traps 288:42.5; 100.5 286: / 247: / 222:Coordinates 2392:Categories 1659:References 1597:Abundance 1529:Sunosuchus 1455:Abundance 1391:Z. salleei 1273:Limusaurus 1265:ceratosaur 1241:Limusaurus 1215:H. sollers 1122:Abundance 1020:eusauropod 818:Abundance 802:Pterosaurs 665:ornithopod 635:Abundance 585:ceratosaur 483:stegosaurs 448:death pits 249:44.5; 90.2 141:Qigu Group 2341:129583852 2333:1867-1594 1833:131520314 1505:J. sloani 1406:gray wolf 1350:Sinraptor 1308:tetanuran 1297:M. jiangi 1176:G. wucaii 1099:Theropods 930:Material 913:Sauropods 904:Sauropods 773:Y. downsi 709:An early 572:Sinraptor 429:Lithology 330:Northern 206:sandstone 183:Lithology 175:Thickness 155:Underlies 147:Sub-units 49:Oxfordian 45:Callovian 2260:19536256 2207:36904501 2199:20110503 2134:16467836 2051:53688256 2001:16901832 1952:54510491 1909:26649770 1868:PLOS ONE 1859:Xu, Xing 1762:53538348 1637:See also 1616:Y. minor 1591:Species 1449:Species 1436:, Wings 1400:A basal 1356:S. dongi 1170:Guanlong 1150:A basal 1141:A. zhaoi 1116:Species 924:Species 846:A basal 812:Species 629:Species 594:Guanlong 556:sauropod 548:theropod 420:theropod 404:Xinjiang 303:Xinjiang 274:100°30′E 216:Location 192:Mudstone 165:Overlies 2313:Bibcode 2268:4358448 2240:Bibcode 2179:Bibcode 2171:Science 2142:4424849 2114:Bibcode 2031:Bibcode 1992:1635516 1932:Bibcode 1900:4674058 1877:Bibcode 1813:Bibcode 1805:PALAIOS 1603:Images 1469:indet. 1461:Images 1432:of the 1385:Zuolong 1261:noasaur 1128:Images 936:Images 824:Images 767:Yinlong 641:Images 529:Yinlong 474:Xu Xing 422:fauna. 398:) is a 309:Country 271:42°30′N 235:90°12′E 232:44°30′N 188:Primary 137:Unit of 2339:  2331:  2266:  2258:  2232:Nature 2205:  2197:  2140:  2132:  2106:Nature 2049:  1999:  1989:  1950:  1907:  1897:  1831:  1786:  1760:  1717:  1683:  1620:Upper 1600:Notes 1588:Genus 1565:Upper 1539:Upper 1509:Upper 1474:Upper 1458:Notes 1446:Genus 1438:et al. 1395:Upper 1360:Upper 1336:Upper 1251:Upper 1219:Upper 1180:Upper 1125:Notes 1113:Genus 1084:Upper 1048:Upper 984:Upper 949:B. sui 933:Notes 921:Genus 876:Upper 841:Upper 821:Notes 809:Genus 777:Upper 739:Upper 704:Upper 658:Upper 638:Notes 626:Genus 591:named 485:, and 392:: 390:pinyin 384:: 376:: 327:Extent 318:  299:Region 2359:(PDF) 2337:S2CID 2264:S2CID 2228:(PDF) 2203:S2CID 2138:S2CID 2047:S2CID 2019:(PDF) 1948:S2CID 1829:S2CID 1758:S2CID 1267:from 1135:Aorun 408:China 321:China 198:Other 2329:ISSN 2256:PMID 2195:PMID 2156:link 2130:PMID 1997:PMID 1905:PMID 1784:ISBN 1715:ISBN 1681:ISBN 386:石樹溝組 378:石树沟组 368:The 202:Tuff 127:Type 63:PreꞒ 2373:doi 2321:doi 2283:doi 2248:doi 2236:459 2187:doi 2175:327 2122:doi 2110:439 2039:doi 1987:PMC 1979:doi 1975:273 1940:doi 1895:PMC 1885:doi 1821:doi 1750:doi 450:or 402:in 2394:: 2369:54 2367:. 2361:. 2335:. 2327:. 2319:. 2309:90 2307:. 2303:. 2262:. 2254:. 2246:. 2234:. 2230:. 2215:^ 2201:. 2193:. 2185:. 2173:. 2152:}} 2148:{{ 2136:. 2128:. 2120:. 2108:. 2073:38 2071:. 2067:. 2045:. 2037:. 2027:30 2025:. 2021:. 1995:. 1985:. 1973:. 1969:. 1946:. 1938:. 1928:81 1926:. 1903:. 1893:. 1883:. 1873:10 1871:. 1865:. 1827:. 1819:. 1809:25 1807:. 1795:^ 1770:^ 1756:. 1746:12 1744:. 1740:. 1726:^ 1695:^ 1625:A 1365:A 1306:A 1185:A 1054:A 1018:A 990:A 881:A 744:A 713:. 652:" 603:. 497:. 410:. 406:, 388:; 380:; 208:, 204:, 113:Pg 57:Ma 43:: 2381:. 2375:: 2343:. 2323:: 2315:: 2289:. 2285:: 2270:. 2250:: 2242:: 2209:. 2189:: 2181:: 2158:) 2144:. 2124:: 2116:: 2053:. 2041:: 2033:: 2003:. 1981:: 1954:. 1942:: 1934:: 1911:. 1887:: 1879:: 1835:. 1823:: 1815:: 1790:. 1764:. 1752:: 1721:. 1689:. 646:" 372:( 118:N 108:K 103:J 98:T 93:P 88:C 83:D 78:S 73:O 68:Ꞓ 52:~ 47:- 20:)

Index

Wucaiwan Formation
Stratigraphic range
Callovian
Oxfordian
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N
Geological formation
Qigu Group
Kalaza Formation
Xishanyao Formation
Mudstone
Tuff
sandstone
conglomerate
Coordinates
44°30′N 90°12′E / 44.5°N 90.2°E / 44.5; 90.2
Approximate paleocoordinates
42°30′N 100°30′E / 42.5°N 100.5°E / 42.5; 100.5
Xinjiang

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