1107:"Waeringopteridae" and the superfamily that includes it, "Waeringopteroidea" (named after eurypterid researcher Erik N. Kjellesvig-Waering), are not formally published clades. The names derive from a 2004 thesis by O. Erik Tetlie and are as such not actually technically valid names. Nevertheless, as phylogenetic analyses confirms the grouping as monophyletic and the names supply an easy way to refer to the group, they remain routinely used within studies centered around eurypterid taxonomy. In these studies, the names are usually within quotation marks and/or are noted to not be properly valid names that derive from a thesis. The group is also sometimes referred to as the "waeringopteroid clade" rather than as the "Waeringopteroidea" within studies to differentiate it from the properly described superfamilies, pending the formal publication of Waeringopteroidea.
1165:
135:
2020:
451:
2323:
113:
1133:, is known from outside of Laurentia and it is thus assumed that the group originated in Laurentia before they spread to modern-day Germany in the Devonian. Further fossil finds are required to properly study the distribution and evolutionary history of the group. The temporal range of the group extends from the
437:
Although confirmed by phylogenetic analyses as monophyletic, and routinely used within eurypterid taxonomy, "Waeringopteroidea" and "Waeringopteridae" are as of yet not formally published clades, the names instead deriving from a 2004 thesis by O. Erik Tetlie. Some researchers refer to the group more
468:
Although waeringopterid fossils remain rare and in most cases fragmentary and very little concrete information on the group can confidently be established due to the poor fossil record, the waeringopterid eurypterids are joined by a handful of known features shared by all genera within the group.
434:. It is thus considered likely that the group originated in Laurentia before spreading to Germany during the Devonian. As the fossil record of the group is extremely poor, little is known of the morphology of the taxa within the group.
573:
and the waeringopteroids), as inferred by O. Erik Tetlie and Markus
Poschmann in 2008, based on the results of a 2008 analysis specifically pertaining to the Adelophthalmoidea and a preceding 2004 analysis.
1213:
1259:
Dunlop, J. A., Penney, D. & Jekel, D. 2015. A summary list of fossil spiders and their relatives. In World Spider
Catalog. Natural History Museum Bern, online at
508:(the posteriormost segment of the body) was xiphous (long and pointed). The eyes of waeringopterid eurypterids were located close to the marginal rim of the
1205:
527:
reached 60 cm (24 in) in length. Though this is relatively long in comparison to modern arthropods, other members of
Diploperculata, such as the
2360:
561:. The cladogram below presents the inferred phylogenetic positions of most of the genera included in the three most derived superfamilies of the
2099:
2089:
1611:
2350:
1280:
Tetlie, O. Erik; Cuggy, Michael B. (2007-01-01). "Phylogeny of the basal swimming eurypterids (Chelicerata; Eurypterida; Eurypterina)".
2051:
1336:"A new Ordovician eurypterid (Arthropoda: Chelicerata) from southeast Turkey: Evidence for a cryptic Ordovician record of Eurypterida"
2066:
1115:
Out of the five known waeringopterid species, four are from the fossil deposits in areas that were part of the ancient continent of
2310:
2094:
2036:
2084:
2074:
2028:
134:
414:
Waeringopterid fossils are very rare and the family only contains five known species classified in three different genera (
2355:
2046:
2041:
1376:
2079:
2056:
493:
1712:
1604:
1178:
1527:
1238:
1402:"Cope's rule and Romer's theory: patterns of diversity and gigantism in eurypterids and Palaeozoic vertebrates"
2019:
2345:
477:
543:
The
Waeringopteridae is the only family within the superfamily Waeringopteroidea and contains the genera
2326:
1597:
1476:
1449:
1347:
1289:
1222:
1170:
1531:
1492:
1313:
328:
129:
1915:
1848:
1840:
1576:
1441:
1423:
1305:
566:
1465:"Phylogeny and palaeoecology of the Adelophthalmoidea (Arthropoda; Chelicerata; Eurypterida)"
2255:
1969:
1961:
1923:
1894:
1810:
1759:
1566:
1523:
1484:
1431:
1415:
1355:
1297:
1230:
803:
779:
746:
684:
1264:
2284:
2130:
1987:
1928:
1884:
1787:
1774:
1406:
1134:
907:
450:
388:
28:
430:, with the only species known from outside of it being a Devonian species recovered from
1480:
1351:
1293:
1226:
2200:
2158:
2116:
1979:
1906:
1866:
1858:
1782:
1436:
1144:
869:
820:
629:
581:
570:
557:
532:
460:
407:
402:
370:
318:
279:
215:
32:
504:(abdomen). In the waeringopterids, the fifth pair of appendages were spiniferous. The
2339:
2207:
2179:
2172:
2144:
2123:
1876:
1815:
1805:
1797:
1751:
1741:
1733:
1689:
1653:
1464:
1335:
1020:
698:
70:
1535:
1496:
1401:
1317:
442:" rather than as the superfamily "Waeringopteroidea", pending a formal publication.
2165:
1997:
1933:
1889:
1764:
1149:
1139:
1121:
955:
883:
646:
605:
551:
545:
393:
379:
310:
302:
268:
121:
1512:"A New Eurypterid From the Upper Ordovician of Manitoulin Island, Ontario, Canada"
1234:
2262:
2239:
2137:
1831:
1724:
1662:
1644:
729:
562:
501:
473:
374:
365:. The Waeringopteridae is the only family classified as part of the superfamily
202:
176:
45:
411:, which is named in honor of eurypterid researcher Erik N. Kjellesvig-Waering.
112:
2228:
2214:
2193:
2186:
2151:
1992:
1680:
1671:
1620:
1488:
1301:
1160:
1003:
979:
528:
358:
189:
90:
55:
1580:
1571:
1550:
1427:
1359:
1309:
2305:
2289:
2279:
1638:
1116:
427:
398:
362:
166:
146:
95:
39:
1445:
1419:
2248:
2221:
1126:
931:
509:
455:
85:
80:
65:
60:
50:
476:, and other arthropods in general, waeringopterid eurypterids possessed
1130:
497:
485:
481:
431:
100:
75:
1632:
505:
489:
384:
156:
1511:
449:
291:
1206:"Distribution and dispersal history of Eurypterida (Chelicerata)"
405:) age. The name Waeringopteridae is derived from the type genus
1593:
1589:
1377:"Lille1tv - Eurypterid phylogeny and the origin of Arachnida"
1528:
10.1666/0022-3360(2005)079[1166:ANECFT]2.0.CO;2
1334:
Lamsdell, James; HoΕgΓΆr, Izzet; Selden, Paul (2013-01-31).
426:). Four of these species are from the ancient continent of
1549:
Dunlop, Jason A.; Braddy, Simon J.; Tetlie, Erik (2008).
1265:
http://www.wsc.nmbe.ch/resources/fossils/Fossils16.0.pdf
515:
The waeringopterids were almost all small eurypterids.
1260:
535:, reached much larger sizes, surpassing two meters.
369:, which in turn is classified within the infraorder
275:
2298:
2272:
2238:
2108:
2065:
2027:
1978:
1960:
1942:
1914:
1903:
1875:
1857:
1839:
1828:
1796:
1773:
1750:
1732:
1721:
1705:
1698:
480:bodies and jointed appendages (limbs) covered in a
519:reached 15 cm (5.9 in) in length, while
1214:Palaeogeography, Palaeoclimatology, Palaeoecology
1463:Erik Tetlie, O; Poschmann, Markus (2008-06-01).
1605:
1400:Lamsdell, James C.; Braddy, Simon J. (2009).
1199:
1197:
1195:
1193:
391:) age and the latest known surviving member,
8:
492:. The chelicerate body is divided into two
1911:
1836:
1729:
1702:
1612:
1598:
1590:
377:. The earliest known member of the group,
111:
20:
1570:
1435:
1119:. Only the last known surviving species,
1189:
397:, has been recovered from deposits of
383:, has been recovered from deposits of
1255:
1253:
1251:
7:
1555:(Gross, 1933) from Overath, Germany"
1371:
1369:
1329:
1327:
1275:
1273:
1510:Stott, Tetlie; et al. (2005).
1469:Journal of Systematic Palaeontology
1282:Journal of Systematic Palaeontology
995:
971:
947:
923:
899:
875:
795:
771:
721:
714:
690:
680:
621:
597:
587:
577:
1111:Distribution and paleobiogeography
14:
523:reached 30 cm (12 in).
2322:
2321:
2018:
1163:
133:
2311:Timeline of eurypterid research
1551:"The Early Devonian eurypterid
2361:Prehistoric arthropod families
361:, an extinct group of aquatic
1:
517:Waeringopterus cumberlandicus
280:Waeringopterus cumberlandicus
1235:10.1016/j.palaeo.2007.05.011
525:Orcanopterus manitoulinensis
16:Extinct family of arthropods
2377:
2351:Silurian first appearances
2319:
2016:
1713:List of eurypterid genera
1627:
1489:10.1017/S1477201907002416
1302:10.1017/S1477201907002131
1179:List of eurypterid genera
1017:
1000:
993:
976:
969:
952:
945:
928:
921:
904:
897:
880:
873:
817:
800:
793:
776:
769:
743:
726:
719:
712:
695:
688:
678:
643:
626:
619:
602:
595:
585:
565:suborder of eurypterids (
334:
327:
297:
290:
274:
267:
130:Scientific classification
128:
119:
110:
23:
1572:10.1002/mmng.20020050107
1360:10.1016/j.gr.2012.04.006
496:(sections); the frontal
1516:Journal of Paleontology
1381:lille1tv.univ-lille1.fr
1204:O. Erik Tetlie (2007).
1420:10.1098/rsbl.2009.0700
465:
344:Not formally published
260:Not formally published
237:Not formally published
1553:Grossopterus overathi
1450:Supplemental material
1122:Grossopterus overathi
521:Grossopterus overathi
500:(head) and posterior
453:
440:waeringopteroid clade
2356:Devonian extinctions
512:(the "head" plate).
1481:2008JSPal...6..237T
1352:2013GondR..23..354L
1294:2007JSPal...5..345T
1227:2007PPP...252..557T
1171:Paleontology portal
1841:Onychopterelloidea
1261:http://wsc.nmbe.ch
466:
2333:
2332:
2014:
2013:
2010:
2009:
2006:
2005:
1962:Adelophthalmoidea
1944:Waeringopteroidea
1916:Carcinosomatoidea
1849:Onychopterellidae
1824:
1823:
1340:Gondwana Research
1104:
1103:
1095:
1094:
1086:
1085:
1077:
1076:
1068:
1067:
1059:
1058:
1050:
1049:
1041:
1040:
1032:
1031:
859:
858:
850:
849:
841:
840:
832:
831:
758:
757:
685:Adelophthalmoidea
667:
666:
658:
657:
592:Waeringopteroidea
567:Adelophthalmoidea
438:formally as the "
367:Waeringopteroidea
352:
351:
346:
340:
263:
262:
240:
239:
229:Waeringopteroidea
2368:
2325:
2324:
2299:Related articles
2256:Merostomichnites
2022:
1970:Adelophthalmidae
1952:Waeringopteridae
1924:Carcinosomatidae
1912:
1895:Strobilopteridae
1837:
1811:Hibbertopteridae
1760:Parastylonuridae
1730:
1703:
1614:
1607:
1600:
1591:
1585:
1584:
1574:
1546:
1540:
1539:
1522:(6): 1166β1174.
1507:
1501:
1500:
1460:
1454:
1453:
1439:
1397:
1391:
1390:
1388:
1387:
1373:
1364:
1363:
1331:
1322:
1321:
1277:
1268:
1257:
1246:
1245:
1243:
1237:. Archived from
1221:(3β4): 557β574.
1210:
1201:
1173:
1168:
1167:
1166:
996:
972:
948:
924:
900:
876:
804:Parahughmilleria
796:
780:Nanahughmilleria
772:
747:Pittsfordipterus
722:
715:
691:
681:
622:
598:
588:
578:
373:in the suborder
355:Waeringopteridae
342:
338:
277:
258:
255:
252:Waeringopteridae
250:
235:
232:
227:
214:
201:
188:
138:
137:
115:
105:
42:
27:Temporal range:
24:Waeringopteridae
21:
2376:
2375:
2371:
2370:
2369:
2367:
2366:
2365:
2336:
2335:
2334:
2329:
2315:
2294:
2285:Chasmataspidida
2268:
2234:
2131:Campylocephalus
2104:
2061:
2023:
2002:
1988:Hughmilleriidae
1974:
1956:
1938:
1929:Megalograptidae
1905:
1899:
1885:Dolichopteridae
1871:
1859:Moselopteroidea
1853:
1830:
1820:
1806:Drepanopteridae
1792:
1788:Hardieopteridae
1775:Kokomopteroidea
1769:
1746:
1723:
1717:
1694:
1623:
1618:
1588:
1548:
1547:
1543:
1509:
1508:
1504:
1462:
1461:
1457:
1407:Biology Letters
1399:
1398:
1394:
1385:
1383:
1375:
1374:
1367:
1333:
1332:
1325:
1279:
1278:
1271:
1263:, version 16.0
1258:
1249:
1241:
1208:
1203:
1202:
1191:
1187:
1169:
1164:
1162:
1159:
1135:Late Ordovician
1113:
1105:
1096:
1087:
1078:
1069:
1060:
1051:
1042:
1033:
908:Herefordopterus
860:
851:
842:
833:
759:
668:
659:
541:
533:carcinosomatids
472:Like all other
448:
389:Late Ordovician
357:is a family of
341:
337:Orcanopteridae
286:
283:
257:
254:
248:
234:
231:
225:
212:
199:
186:
132:
106:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
37:
36:
29:Late Ordovician
25:
17:
12:
11:
5:
2374:
2372:
2364:
2363:
2358:
2353:
2348:
2346:Diploperculata
2338:
2337:
2331:
2330:
2320:
2317:
2316:
2314:
2313:
2308:
2302:
2300:
2296:
2295:
2293:
2292:
2287:
2282:
2276:
2274:
2273:Related groups
2270:
2269:
2267:
2266:
2259:
2252:
2244:
2242:
2236:
2235:
2233:
2232:
2225:
2218:
2211:
2204:
2201:Onychopterella
2197:
2190:
2183:
2176:
2169:
2162:
2159:Hibbertopterus
2155:
2148:
2141:
2134:
2127:
2120:
2117:Adelophthalmus
2112:
2110:
2109:Notable genera
2106:
2105:
2103:
2102:
2097:
2092:
2087:
2082:
2077:
2071:
2069:
2063:
2062:
2060:
2059:
2054:
2049:
2044:
2039:
2033:
2031:
2025:
2024:
2017:
2015:
2012:
2011:
2008:
2007:
2004:
2003:
2001:
2000:
1995:
1990:
1984:
1982:
1980:Pterygotioidea
1976:
1975:
1973:
1972:
1966:
1964:
1958:
1957:
1955:
1954:
1948:
1946:
1940:
1939:
1937:
1936:
1931:
1926:
1920:
1918:
1909:
1907:Diploperculata
1901:
1900:
1898:
1897:
1892:
1887:
1881:
1879:
1873:
1872:
1870:
1869:
1867:Moselopteridae
1863:
1861:
1855:
1854:
1852:
1851:
1845:
1843:
1834:
1826:
1825:
1822:
1821:
1819:
1818:
1813:
1808:
1802:
1800:
1794:
1793:
1791:
1790:
1785:
1783:Kokomopteridae
1779:
1777:
1771:
1770:
1768:
1767:
1762:
1756:
1754:
1748:
1747:
1745:
1744:
1738:
1736:
1734:Rhenopteroidea
1727:
1719:
1718:
1716:
1715:
1709:
1707:
1700:
1696:
1695:
1693:
1692:
1683:
1674:
1665:
1656:
1647:
1641:
1635:
1628:
1625:
1624:
1619:
1617:
1616:
1609:
1602:
1594:
1587:
1586:
1541:
1502:
1475:(2): 237β249.
1455:
1414:(2): 265β269.
1392:
1365:
1346:(1): 354β366.
1323:
1288:(3): 345β356.
1269:
1247:
1244:on 2011-07-18.
1188:
1186:
1183:
1182:
1181:
1175:
1174:
1158:
1155:
1145:Early Devonian
1112:
1109:
1102:
1101:
1098:
1097:
1093:
1092:
1089:
1088:
1084:
1083:
1080:
1079:
1075:
1074:
1071:
1070:
1066:
1065:
1062:
1061:
1057:
1056:
1053:
1052:
1048:
1047:
1044:
1043:
1039:
1038:
1035:
1034:
1030:
1029:
1026:
1025:
1016:
1013:
1012:
1009:
1008:
999:
994:
992:
989:
988:
985:
984:
975:
970:
968:
965:
964:
961:
960:
951:
946:
944:
941:
940:
937:
936:
927:
922:
920:
917:
916:
913:
912:
903:
898:
896:
893:
892:
889:
888:
879:
874:
872:
870:Pterygotioidea
866:
865:
862:
861:
857:
856:
853:
852:
848:
847:
844:
843:
839:
838:
835:
834:
830:
829:
826:
825:
821:Adelophthalmus
816:
813:
812:
809:
808:
799:
794:
792:
789:
788:
785:
784:
775:
770:
768:
765:
764:
761:
760:
756:
755:
752:
751:
742:
739:
738:
735:
734:
725:
720:
718:
713:
711:
708:
707:
704:
703:
694:
689:
687:
679:
677:
674:
673:
670:
669:
665:
664:
661:
660:
656:
655:
652:
651:
642:
639:
638:
635:
634:
630:Waeringopterus
625:
620:
618:
615:
614:
611:
610:
601:
596:
594:
586:
584:
582:Diploperculata
576:
571:Pterygotioidea
558:Waeringopterus
540:
539:Classification
537:
461:Waeringopterus
447:
444:
416:Waeringopterus
408:Waeringopterus
403:Early Devonian
371:Diploperculata
350:
349:
348:
347:
332:
331:
325:
324:
323:
322:
319:Waeringopterus
314:
306:
295:
294:
288:
287:
284:
272:
271:
265:
264:
246:
242:
241:
223:
219:
218:
216:Diploperculata
210:
206:
205:
197:
193:
192:
184:
180:
179:
174:
170:
169:
164:
160:
159:
154:
150:
149:
144:
140:
139:
126:
125:
117:
116:
108:
107:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
43:
33:Early Devonian
26:
15:
13:
10:
9:
6:
4:
3:
2:
2373:
2362:
2359:
2357:
2354:
2352:
2349:
2347:
2344:
2343:
2341:
2328:
2318:
2312:
2309:
2307:
2304:
2303:
2301:
2297:
2291:
2288:
2286:
2283:
2281:
2278:
2277:
2275:
2271:
2265:
2264:
2260:
2258:
2257:
2253:
2251:
2250:
2246:
2245:
2243:
2241:
2237:
2231:
2230:
2226:
2224:
2223:
2219:
2217:
2216:
2212:
2210:
2209:
2208:Pentecopterus
2205:
2203:
2202:
2198:
2196:
2195:
2191:
2189:
2188:
2184:
2182:
2181:
2180:Megalograptus
2177:
2175:
2174:
2173:Jaekelopterus
2170:
2168:
2167:
2163:
2161:
2160:
2156:
2154:
2153:
2149:
2147:
2146:
2145:Drepanopterus
2142:
2140:
2139:
2135:
2133:
2132:
2128:
2126:
2125:
2124:Brachyopterus
2121:
2119:
2118:
2114:
2113:
2111:
2107:
2101:
2100:South America
2098:
2096:
2093:
2091:
2090:North America
2088:
2086:
2083:
2081:
2078:
2076:
2073:
2072:
2070:
2068:
2064:
2058:
2055:
2053:
2052:Carboniferous
2050:
2048:
2045:
2043:
2040:
2038:
2035:
2034:
2032:
2030:
2029:Geochronology
2026:
2021:
1999:
1996:
1994:
1991:
1989:
1986:
1985:
1983:
1981:
1977:
1971:
1968:
1967:
1965:
1963:
1959:
1953:
1950:
1949:
1947:
1945:
1941:
1935:
1932:
1930:
1927:
1925:
1922:
1921:
1919:
1917:
1913:
1910:
1908:
1902:
1896:
1893:
1891:
1888:
1886:
1883:
1882:
1880:
1878:
1877:Eurypteroidea
1874:
1868:
1865:
1864:
1862:
1860:
1856:
1850:
1847:
1846:
1844:
1842:
1838:
1835:
1833:
1827:
1817:
1816:Mycteroptidae
1814:
1812:
1809:
1807:
1804:
1803:
1801:
1799:
1798:Mycteropoidea
1795:
1789:
1786:
1784:
1781:
1780:
1778:
1776:
1772:
1766:
1763:
1761:
1758:
1757:
1755:
1753:
1752:Stylonuroidea
1749:
1743:
1742:Rhenopteridae
1740:
1739:
1737:
1735:
1731:
1728:
1726:
1720:
1714:
1711:
1710:
1708:
1704:
1701:
1697:
1691:
1690:Sclerophorata
1687:
1684:
1682:
1678:
1675:
1673:
1669:
1666:
1664:
1660:
1657:
1655:
1654:Euchelicerata
1651:
1648:
1646:
1642:
1640:
1636:
1634:
1630:
1629:
1626:
1622:
1615:
1610:
1608:
1603:
1601:
1596:
1595:
1592:
1582:
1578:
1573:
1568:
1565:(1): 93β104.
1564:
1560:
1559:Fossil Record
1556:
1554:
1545:
1542:
1537:
1533:
1529:
1525:
1521:
1517:
1513:
1506:
1503:
1498:
1494:
1490:
1486:
1482:
1478:
1474:
1470:
1466:
1459:
1456:
1451:
1447:
1443:
1438:
1433:
1429:
1425:
1421:
1417:
1413:
1409:
1408:
1403:
1396:
1393:
1382:
1378:
1372:
1370:
1366:
1361:
1357:
1353:
1349:
1345:
1341:
1337:
1330:
1328:
1324:
1319:
1315:
1311:
1307:
1303:
1299:
1295:
1291:
1287:
1283:
1276:
1274:
1270:
1266:
1262:
1256:
1254:
1252:
1248:
1240:
1236:
1232:
1228:
1224:
1220:
1216:
1215:
1207:
1200:
1198:
1196:
1194:
1190:
1184:
1180:
1177:
1176:
1172:
1161:
1156:
1154:
1152:
1151:
1146:
1142:
1141:
1136:
1132:
1128:
1124:
1123:
1118:
1110:
1108:
1100:
1099:
1091:
1090:
1082:
1081:
1073:
1072:
1064:
1063:
1055:
1054:
1046:
1045:
1037:
1036:
1028:
1027:
1024:
1023:
1022:
1021:Jaekelopterus
1015:
1014:
1011:
1010:
1007:
1006:
1005:
998:
997:
991:
990:
987:
986:
983:
982:
981:
974:
973:
967:
966:
963:
962:
959:
958:
957:
950:
949:
943:
942:
939:
938:
935:
934:
933:
926:
925:
919:
918:
915:
914:
911:
910:
909:
902:
901:
895:
894:
891:
890:
887:
886:
885:
878:
877:
871:
868:
867:
864:
863:
855:
854:
846:
845:
837:
836:
828:
827:
824:
823:
822:
815:
814:
811:
810:
807:
806:
805:
798:
797:
791:
790:
787:
786:
783:
782:
781:
774:
773:
767:
766:
763:
762:
754:
753:
750:
749:
748:
741:
740:
737:
736:
733:
732:
731:
724:
723:
717:
716:
710:
709:
706:
705:
702:
701:
700:
699:Eysyslopterus
693:
692:
686:
683:
682:
676:
675:
672:
671:
663:
662:
654:
653:
650:
649:
648:
641:
640:
637:
636:
633:
632:
631:
624:
623:
617:
616:
613:
612:
609:
608:
607:
600:
599:
593:
590:
589:
583:
580:
579:
575:
572:
568:
564:
560:
559:
554:
553:
548:
547:
538:
536:
534:
530:
526:
522:
518:
513:
511:
507:
503:
499:
495:
491:
487:
483:
479:
475:
470:
463:
462:
457:
452:
445:
443:
441:
435:
433:
429:
425:
421:
417:
412:
410:
409:
404:
400:
396:
395:
390:
386:
382:
381:
376:
372:
368:
364:
360:
356:
345:
336:
335:
333:
330:
326:
321:
320:
315:
313:
312:
307:
305:
304:
299:
298:
296:
293:
289:
282:
281:
273:
270:
266:
261:
253:
247:
244:
243:
238:
230:
224:
222:Superfamily:
221:
220:
217:
211:
208:
207:
204:
198:
195:
194:
191:
185:
182:
181:
178:
175:
172:
171:
168:
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
136:
131:
127:
124:
123:
118:
114:
109:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
41:
38:449β407
34:
30:
22:
19:
2261:
2254:
2247:
2227:
2220:
2213:
2206:
2199:
2192:
2185:
2178:
2171:
2166:Hughmilleria
2164:
2157:
2150:
2143:
2136:
2129:
2122:
2115:
1998:Pterygotidae
1951:
1943:
1934:Mixopteridae
1890:Eurypteridae
1765:Stylonuridae
1685:
1676:
1667:
1658:
1649:
1562:
1558:
1552:
1544:
1519:
1515:
1505:
1472:
1468:
1458:
1411:
1405:
1395:
1384:. Retrieved
1380:
1343:
1339:
1285:
1281:
1239:the original
1218:
1212:
1150:Grossopterus
1148:
1140:Orcanopterus
1138:
1120:
1114:
1106:
1019:
1018:
1002:
1001:
978:
977:
956:Erettopterus
954:
953:
930:
929:
906:
905:
884:Hughmilleria
882:
881:
819:
818:
802:
801:
778:
777:
745:
744:
728:
727:
697:
696:
647:Grossopterus
645:
644:
628:
627:
606:Orcanopterus
604:
603:
591:
556:
552:Orcanopterus
550:
546:Grossopterus
544:
542:
524:
520:
516:
514:
484:composed of
474:chelicerates
471:
467:
459:
439:
436:
424:Grossopterus
423:
420:Orcanopterus
419:
415:
413:
406:
394:Grossopterus
392:
380:Orcanopterus
378:
366:
354:
353:
343:
339:Tetlie, 2004
317:
311:Orcanopterus
309:
303:Grossopterus
301:
285:Swartz, 1923
278:
269:Type species
259:
256:Tetlie, 2004
251:
236:
233:Tetlie, 2004
228:
209:Infraorder:
122:Orcanopterus
120:
18:
2263:Palmichnium
2240:Ichnogenera
2138:Carcinosoma
1832:Eurypterina
1725:Stylonurina
1663:Prosomapoda
1645:Chelicerata
1643:Subphylum:
1621:Eurypterids
1125:, from the
730:Bassipterus
563:Eurypterina
529:pterygotids
502:opisthosoma
446:Description
375:Eurypterina
359:eurypterids
203:Eurypterina
190:Eurypterida
177:Chelicerata
173:Subphylum:
2340:Categories
2229:Stylonurus
2215:Pterygotus
2194:Mixopterus
2187:Megarachne
2152:Eurypterus
2037:Ordovician
1993:Slimonidae
1904:Infraorder
1681:Dekatriata
1672:Planaterga
1639:Arthropoda
1386:2018-04-02
1185:References
1004:Acutiramus
980:Pterygotus
363:arthropods
196:Suborder:
167:Arthropoda
2306:Metastoma
2290:Xiphosura
2280:Arachnida
2067:Geography
1631:Kingdom:
1581:1860-1014
1428:1744-9561
1310:1477-2019
1143:) to the
1117:Laurentia
478:segmented
456:carapaces
428:Laurentia
399:Siegenian
153:Kingdom:
147:Eukaryota
2327:Category
2249:Arcuites
2222:Slimonia
2047:Devonian
2042:Silurian
1829:Suborder
1722:Suborder
1699:Taxonomy
1637:Phylum:
1633:Animalia
1536:55014027
1497:59488956
1446:19828493
1318:88413267
1157:See also
1127:Devonian
932:Slimonia
510:carapace
486:proteins
329:Synonyms
245:Family:
163:Phylum:
157:Animalia
143:Domain:
2095:Oceania
2057:Permian
1477:Bibcode
1437:2865068
1348:Bibcode
1290:Bibcode
1223:Bibcode
1131:Germany
531:or the
498:prosoma
494:tagmata
482:cuticle
454:Fossil
432:Germany
183:Order:
2085:Europe
2075:Africa
1706:Genera
1579:
1534:
1495:
1444:
1434:
1426:
1316:
1308:
1267:(PDF).
506:telson
490:chitin
385:Katian
292:Genera
1686:Clade
1677:Clade
1668:Clade
1659:Clade
1650:Clade
1532:S2CID
1493:S2CID
1314:S2CID
1242:(PDF)
1209:(PDF)
2080:Asia
1577:ISSN
1442:PMID
1424:ISSN
1306:ISSN
555:and
488:and
422:and
46:Preκ
1567:doi
1524:doi
1485:doi
1432:PMC
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1356:doi
1298:doi
1231:doi
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1129:of
458:of
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1422:.
1410:.
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1379:.
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1272:^
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96:Pg
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316:β
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276:β
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66:D
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51:κ
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