2465:
1904:
1828:
120:
1918:
1334:
258:
1294:
928:
467:
1344:
1314:
1324:
482:
1304:
1932:
2031:
De Blieux, Donald D. "Analysis of Jim's hadrosaur site; a dinosaur site in the middle
Campanian (Cretaceous) Wahweap Formation of Grand Staircase–Escalante National Monument (GSENM), southern Utah." Abstracts with Programs – Geological Society of America, vol. 39, no. 5, pp.6, May
1887:
In 2010 a unique trace fossil from the
Wahweap was discovered that indicates a predator–prey relationship between dinosaurs and primitive mammals. The trace fossil includes at least two fossilized mammalian den complexes as well as associated digging grooves presumably caused by a
2041:
Kirkland, James Ian. "An inventory of paleontological resources in the lower
Wahweap Formation (lower Campanian), southern Kaiparowits Plateau, Grand Staircase–Escalante National Monument, Utah." Abstracts with Programs – Geological Society of America, vol.37, no.7, pp.114, Oct
2183:
Gardner, James D.; Eaton, Jeffrey G.; Cifelli, Richard L. (2013). "Preliminary Report on
Salamanders (Lissamphibia; Caudata) from the Late Cretaceous (Late Cenomanian–Late Campanian) of Southern Utah, U.S.A.". In Titus, Alan L.; Loewen, Mark A. (eds.).
1964:
Weishampel, David B; et al. (2004). "Dinosaur distribution (Late
Cretaceous, North America)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 574–588.
2630:
Evans, David C.; Williamson, Thomas; Loewen, Mark A.; Kirkland, James I. (2013). "Review of
Pachycephalosaurian Dinosaurs from Grand Staircase–Escalante National Monument, Southern Utah". In Titus, Alan L.; Loewen, Mark A. (eds.).
2738:
2693:
Eaton, Jeffrey G; Cifelli, Richard L. "Review of
Cretaceous mammalian paleontology; Grand Staircase–Escalante National Monument, Utah." Abstracts with Programs – Geological Society of America, vol.37, no.7, pp.115, Oct
2812:
2731:
2659:
Simpson, Edward L.; Hilbert-Wolf, Hannah L.; Wizevich, Michael C.; Tindall, Sarah E.; Fasinski, Ben R.; Storm, Lauren P.; Needle, Mattathias D. (2010). "Predatory digging behavior by dinosaurs".
2924:
2724:
1985:"Refined geochronology and revised stratigraphic nomenclature of the Upper Cretaceous Wahweap Formation, Utah, U.S.A. and the age of early Campanian vertebrates from southern Laramidia"
1876:
are also relatively abundant in the
Wahweap, and include vertebrate tracks as well as burrow activity. Tracks preserved in the capping sandstone indicate the presence of
2929:
2747:
299:
125:
2246:
Gates, T.A.; Horner, J.R.; Hanna, R.R.; Nelson, C.R. (2011). "New unadorned hadrosaurine hadrosaurid (Dinosauria, Ornithopoda) from the
Campanian of North America".
2401:
Gates, Terry A.; Lund, Eric K.; Boyd, C.A.; DeBlieux, Donald D.; Titus, Alan L.; Evans, David C.; Getty, Michael A.; Kirkland, James I.; Eaton, Jeffrey J. (2013).
2565:"A New Centrosaurine Ceratopsid, Machairoceratops cronusi gen et sp. nov., from the Upper Sand Member of the Wahweap Formation (Middle Campanian), Southern Utah"
2403:"Ornithopod Dinosaurs from the Grand Staircase–Escalante National Monument Region, Utah, and Their Role in Paleobiogeographic and Macroevolutionary Studies"
2108:
2209:
2708:
Isolated vertebrate tracks from the Upper
Cretaceous capping sandstone member of the Wahweap Formation; Grand Staircase–Escalante National Monument, Utah
1983:
Beveridge, Tegan L.; Roberts, Eric M.; Ramezani, Jahandar; Titus, Alan L.; Eaton, Jeffrey G.; Irmis, Randall B.; Sertich, Joseph J. W. (April 1, 2022).
2466:"New basal centrosaurine ceratopsian skulls from the Wahweap Formation (Middle Campanian), Grand Staircase–Escalante National Monument, southern Utah"
1945:
2899:
2934:
2863:
2109:"A new hybodontid shark (Chondrichthyes, Hybodontiformes) from the Lower Jurassic Posidonienschiefer Formation of Dotternhausen, SW Germany"
2904:
2827:
756:
A large hybodont shark reaching over 2 m (6.6 ft) in length. Normally a marine species but would enter freshwater environments
2914:
1750:
The skeleton was under excavation as of 2013. The cranium was only tentatively identified as a nodosaurid, but now can't be confirmed.
2640:
2480:
2448:
2414:
2385:
2305:
2193:
2162:
2091:
1970:
1892:
dinosaur. The proximity indicates a case of probable active predation of the burrow inhabitants by the owners of the claw marks.
357:
that characterize the shell beds. Large fossilized crabs are common at most shell bed sites in the Wahweap, and over 1,900
1846:
A fair number of mammals spanning the lower Campanian are known from the Wahweap as well, including at least 15 genera of
2294:"New Hadrosaurid (Dinosauria, Ornithopoda) Specimens from the Lower—Middle Campanian Wahweap Formation of Southern Utah"
1384:
205:
2909:
2435:
Loewen, Mark A.; Farke, Andrew A.; Sampson, Scott D.; Getty, Michael A.; Lund, Eric K.; O'Connor, Patrick M. (2013).
2496:
Loewen, Mark A.; Irmis, Randall B.; Sertich, Joseph J. W.; Currie, Philip J.; Sampson, Scott D. (November 6, 2013).
2151:"Freshwater Osteichthyes from the Cenomanian to Late Campanian of Grand Staircase–Escalante National Monument, Utah"
349:
The Wahweap Formation shows a substantial amount of invertebrate activity ranging from fossilized insect burrows in
212:
2842:
178:
1827:
2919:
2802:
1267:; "a partial articulated skull roof including lacrimals and entire braincase, and a single cervical vertebra"
1782:
The dome doesn't preserve diagnostic features, so its affinity with other pachycephalosaurids is uncertain.
2868:
2372:
Loewen, Mark A.; Burns, Michael E.; Getty, Michael E.; Kirkland, James I.; Matthew K., Vickaryous (2013).
1285:
2220:
2875:
2832:
1909:
550:
168:
1175:
322:
The Wahweap formation is divided into four members, which are listed below with their respective ages:
2053:
Preliminary study of freshwater gastropods in the Wahweap Formation, Bryce Canyon National Park, Utah.
2668:
2576:
2509:
2255:
1996:
140:
306:
in southern Utah and northern Arizona, around the Lake Powell region, whose strata date back to the
2837:
1923:
1140:
303:
150:
2374:"Review of Late Cretaceous Ankylosaurian Dinosaurs from the Grand Staircase Region, Southern Utah"
119:
2271:
1755:
1584:
614:
794:
560:
2716:
2636:
2612:
2594:
2545:
2527:
2476:
2444:
2410:
2381:
2351:
2301:
2189:
2158:
2128:
2087:
2014:
1966:
1847:
1497:
1443:
1273:
474:
2322:
2797:
2676:
2602:
2584:
2535:
2517:
2341:
2263:
2120:
2004:
1937:
1648:
1402:
970:
599:
314:
stage). Dinosaur remains are among the fossils that have been recovered from the formation.
160:
Last Chance Creek Member, Reynolds Point Member, Coyote Point Member, Pardner Canyon Member
2817:
2766:
1884:
dinosaurs. At least one possible theropod track has been identified in this area as well.
1877:
1527:
920:
307:
2710:
Abstracts with Programs – Geological Society of America, vol. 39, no. 5, pp.42, May 2007
2672:
2580:
2563:
Lund, Eric K.; O’Connor, Patrick M.; Loewen, Mark A.; Jinnah, Zubair A. (May 18, 2016).
2513:
2259:
2055:
Abstracts with Programs – Geological Society of America, vol. 39, no. 5, pp.43, May 2007
2000:
1596:; "a skull preserving the entire left side of the skull and portions of the right side"
1333:
2822:
2607:
2564:
2540:
2497:
2149:
Brinkman, Donald B.; Newbrey, Michael G.; Neuman, Andrew G.; Eaton, Jeffrey G. (2013).
1638:
1569:
1536:
1505:
1472:
1411:
1293:
927:
410:
378:
350:
466:
2893:
2402:
2293:
2150:
1881:
1787:
1558:
1395:
1106:
1007:
946:
495:
286:
263:
71:
29:
1880:, which had been previously known in this area only from teeth elements, as well as
1343:
1313:
2807:
2781:
2761:
2473:
New Perspectives on Horned Dinosaurs: The Royal Tyrell Museum Ceratopsian Symposium
2275:
1873:
1862:
1838:
digging a primitive mammal out of its burrow, as per the 2010 discovery by Simpson
1685:
1151:
1046:
871:
545:
459:
1842:
of trace fossils indicating a predator–prey relationship in the Wahweap Formation.
1323:
481:
2589:
2522:
2267:
2009:
1984:
2776:
2771:
2124:
1889:
1851:
1398:
1358:
1196:
604:
46:
2498:"Tyrant Dinosaur Evolution Tracks the Rise and Fall of Late Cretaceous Oceans"
2296:. In Eberth, David A.; Evans, David Christopher; Ralrick, Patricia E. (eds.).
1899:
1855:
1832:
1244:
1027:
999:
762:
91:
56:
2598:
2531:
2355:
2132:
2018:
227:
214:
2327:
type locality in the Upper Cretaceous (Campanian) Wahweap Formation of Utah"
2292:
Gates, Terry A.; Jinnah, Zubair; Levitt, Carolyn; Getty, Michael A. (2014).
1859:
1610:
1385:
1303:
1279:
1192:
1095:
979:
903:
658:
362:
358:
311:
193:
96:
40:
33:
2616:
2549:
1835:
1258:"near the junction of Smokey Mountain Road and Right Hand Collet Canyon"
1210:
1200:
935:
361:
specimens (of four likely genera) have been unearthed in the formation's
86:
81:
66:
61:
51:
2346:
2080:"Elasmobranchs from Upper Cretaceous Freshwater Facies in Southern Utah"
1556:
Also known as "Wahweap Centrosaurine C." Possibly has spikes similar to
2633:
At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah
2441:
At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah
2407:
At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah
2186:
At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah
2155:
At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah
2084:
At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah
852:
732:
354:
101:
76:
2378:
At the Top of the Grand Staircase-The Late Cretaceous of Southern Utah
1205:
2680:
1522:; "A partial braincase and a nearly complete parietosquamosal frill"
2436:
2373:
2079:
2078:
Kirkland, James I.; Eaton, Jeffrey G.; Brinkman, Donald B. (2013).
2236:
John Wesley Powell Memorial Museum display, visited April 30, 2009
1342:
1332:
1322:
1312:
1302:
1292:
926:
480:
465:
2437:"Ceratopsid Dinosaurs from the Grand Staircase of Southern Utah"
129:
2720:
2471:. In Ryan, M.J.; Chinnery-Allgeier, B.J.; Eberth, D.A. (eds.).
1191:
Dinosaurs known from the Wahweap include at least 2 species of
2113:
Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen
2635:. Bloomington: Indiana University Press. pp. 482–487.
2475:. Bloomington: Indiana University Press. pp. 117–140.
2443:. Bloomington: Indiana University Press. pp. 488–503.
2409:. Bloomington: Indiana University Press. pp. 463–481.
2380:. Bloomington: Indiana University Press. pp. 445–462.
2321:
Holroyd, Patricia A.; Hutchison, J. Howard (June 9, 2016).
2188:. Bloomington: Indiana University Press. pp. 237–272.
2157:. Bloomington: Indiana University Press. pp. 195–236.
2086:. Bloomington: Indiana University Press. pp. 153–194.
726:
A small guitarfish, suited for handling hard-shelled prey.
2107:
Maisch, Michael W.; Matzke, Andreas T. (June 1, 2016).
1600:
The older specimen from Nipple Butte may not belong to
548:
which inhabited freshwater. One specimen is similar to
1094:
Higher level relationships are uncertain. Similar to
1525:
Also known as "Wahweap Centrosaurine A." Similar to
2856:
2790:
2754:
282:
277:
269:
251:
243:
204:
199:
189:
184:
174:
164:
156:
146:
136:
23:
2210:"New Horned Dinosaurs from the Wahweap Formation"
1989:Palaeogeography, Palaeoclimatology, Palaeoecology
2464:Kirkland, James I.; Deblieux, Donald D. (2010).
1553:; "Partial frill and some postcranial elements"
2051:Williams, Jessica A J; Lohrengel, C Frederick.
1680:Previously known as "Wahweap Centrosaurine B."
2300:. Indiana University Press. pp. 156–173.
1002:which is adapted to eating hard-shelled prey.
2732:
2439:. In Titus, Alan L.; Loewen, Mark A. (eds.).
2405:. In Titus, Alan L.; Loewen, Mark A. (eds.).
2376:. In Titus, Alan L.; Loewen, Mark A. (eds.).
2153:. In Titus, Alan L.; Loewen, Mark A. (eds.).
2082:. In Titus, Alan L.; Loewen, Mark A. (eds.).
1461:; "Partial limb bones and a partial maxilla"
8:
2925:Grand Staircase–Escalante National Monument
2748:Grand Staircase–Escalante National Monument
1143:, a family of extinct aquatic salamanders.
300:Grand Staircase–Escalante National Monument
126:Grand Staircase–Escalante National Monument
2739:
2725:
2717:
2654:
2652:
1816:Probably represents a new distinct taxon.
372:
118:
20:
2606:
2588:
2539:
2521:
2345:
2008:
2930:Upper Cretaceous Series of North America
1946:List of dinosaur-bearing rock formations
1826:
1208:
1040:
865:
326:Last Chance Creek Member: 82.17-81.55 Ma
1957:
1516:Upper part of Last Chance Creek Member
1483:Upper part of Last Chance Creek Member
16:Geologic formation in the United States
1030:, very rare in the Wahweap Formation.
923:fish, no extant relatives live today.
2430:
2428:
2426:
2367:
2365:
2287:
2285:
1589:Middle part of Reynolds Point Member
329:Reynolds Point Member: 81.55-80.61 Ma
7:
2178:
2176:
2174:
2144:
2142:
2073:
2071:
2069:
2067:
2065:
2063:
2061:
1766:Lower part of Reynolds Point Member
1627:Lower part of Reynolds Point Member
1261:Upper part of Reynolds Point Member
1098:, but lacks sirenid synapomorphies.
1676:A centrosaurine closely related to
2248:Journal of Vertebrate Paleontology
1547:Upper part of Coyote Point Member
1455:Upper part of Coyote Point Member
1375:Lower part of Coyote Point Member
335:Pardner Canyon Member: 79-77.29 Ma
14:
1271:A hadrosaurid closely related to
973:fish, adapted to crush its prey.
1930:
1916:
1902:
1774:; incomplete frontoparietal dome
1496:but is more derived, possibly a
332:Coyote Point Member: 80.61-79 Ma
256:
273:Southern Utah, Northern Arizona
2900:Geologic formations of Arizona
2864:DomĂnguez–Escalante expedition
1:
2935:Ichnofossiliferous formations
2590:10.1371/journal.pone.0154403
2523:10.1371/journal.pone.0079420
2268:10.1080/02724634.2011.577854
2010:10.1016/j.palaeo.2022.110876
462:which inhabited freshwater.
2905:Geologic formations of Utah
1805:; partial juvenile skeleton
2951:
2915:Cretaceous geology of Utah
2323:"Fauna and setting of the
1291:
2843:Straight Cliffs Formation
1697:Last Chance Creek Member?
1049:of the Wahweap Formation
1045:
925:
874:of the Wahweap Formation
870:
464:
381:of the Wahweap Formation
377:
179:Straight Cliffs Formation
117:
28:
2803:Canyons of the Escalante
1811:; partial adult skeleton
1583:42Ka800V, south side of
1213:of the Wahweap Formation
602:ray, closely related to
2125:10.1127/njgpa/2016/0577
1492:Originally referred to
2869:Hole in the Rock Trail
2325:Adelolophus hutchisoni
2219:. 2007. Archived from
2217:Utah Geological Survey
1843:
1797:Reynolds Point Member
1624:UMNH VP Locality 1501
1350:
1340:
1330:
1320:
1310:
1300:
1286:Two Medicine Formation
939:
488:
478:
2876:Xylorhiza cronquistii
2833:Kaiparowits Formation
1910:Earth sciences portal
1830:
1700:Reynolds Point Member
1346:
1336:
1326:
1316:
1306:
1296:
1118:UMNH VP Locality 130
930:
484:
471:Chiloscyllium griseum
469:
169:Kaiparowits Formation
1756:Pachycephalosauridae
1665:Coyote Point Member
1421:Coyote Point Member
1284:Also known from the
1163:UMNH VP Locality 77
1083:UMNH VP Locality 77
774:UMNH VP Locality 82
571:UMNH VP Locality 77
141:Geological formation
2838:Kaiparowits Plateau
2673:2010Geo....38..699S
2581:2016PLoSO..1154403L
2514:2013PLoSO...879420L
2347:10.5070/P9331031196
2260:2011JVPal..31..798G
2226:on August 31, 2021.
2001:2022PPP...59110876B
1924:Paleontology portal
1777:some isolated teeth
1703:Coyote Point Member
1633:; partial skeleton
1132:UMNH VP 19199-19200
1126:UMNH VP 19194-19198
1042:
1015:MNA Locality 456-2
966:; pharyngeal tooth
958:MNA Locality 706-2
867:
810:UMNH VP Locality 82
807:UMNH VP Locality 77
787:Lonchidon selachos.
745:MNA Locality 456-2
633:UMNH VP Locality 82
630:UMNH VP Locality 77
508:UMNH VP Locality 77
429:UMNH VP Locality 82
374:
224: /
151:Kaiparowits Plateau
124:Wahweap Formation,
30:Stratigraphic range
2910:Cretaceous Arizona
1844:
1745:a partial skeleton
1585:Last Chance Canyon
1351:
1341:
1331:
1321:
1311:
1301:
1041:
940:
866:
627:MNA Locality 456-1
551:Restesia americana
511:MNA Locality 455-1
489:
486:Hybodus hauffianus
479:
426:MNA Locality 455-1
373:
2887:
2886:
2848:Wahweap Formation
1848:multituberculates
1820:
1819:
1444:Brachylophosaurus
1318:Brachylophosaurus
1274:Brachylophosaurus
1199:and at least one
1184:
1183:
1141:batrachosauroidid
1134:; trunk vertebrae
1091:; trunk vertebra
1034:
1033:
859:
858:
820:UMNH VP 18882-83;
475:grey bamboo shark
296:Wahweap Formation
292:
291:
24:Wahweap Formation
2942:
2880:
2798:Aquarius Plateau
2741:
2734:
2727:
2718:
2711:
2701:
2695:
2691:
2685:
2684:
2681:10.1130/G31019.1
2656:
2647:
2646:
2627:
2621:
2620:
2610:
2592:
2560:
2554:
2553:
2543:
2525:
2493:
2487:
2486:
2470:
2461:
2455:
2454:
2432:
2421:
2420:
2398:
2392:
2391:
2369:
2360:
2359:
2349:
2331:
2318:
2312:
2311:
2289:
2280:
2279:
2243:
2237:
2234:
2228:
2227:
2225:
2214:
2206:
2200:
2199:
2180:
2169:
2168:
2146:
2137:
2136:
2104:
2098:
2097:
2075:
2056:
2049:
2043:
2039:
2033:
2029:
2023:
2022:
2012:
1980:
1974:
1962:
1940:
1938:Dinosaurs portal
1935:
1934:
1933:
1926:
1921:
1920:
1919:
1912:
1907:
1906:
1905:
1742:a cranium (lost)
1672:; partial skull
1649:Machairoceratops
1498:nasutoceratopsin
1489:; Partial skull
1387:
1348:Machairoceratops
1206:
1043:
868:
834:UMNH VP 18889-94
449:UMNH VP 18915-16
423:MNA Locality 705
375:
302:is a geological
262:
260:
259:
239:
238:
236:
235:
234:
229:
225:
222:
221:
220:
217:
122:
113:
43:
21:
2950:
2949:
2945:
2944:
2943:
2941:
2940:
2939:
2920:Campanian Stage
2890:
2889:
2888:
2883:
2879:(plant species)
2873:
2852:
2818:Grand Staircase
2786:
2767:Escalante River
2755:Bodies of water
2750:
2745:
2715:
2714:
2703:Tester, Edward
2702:
2698:
2692:
2688:
2658:
2657:
2650:
2643:
2629:
2628:
2624:
2575:(5): e0154403.
2562:
2561:
2557:
2495:
2494:
2490:
2483:
2468:
2463:
2462:
2458:
2451:
2434:
2433:
2424:
2417:
2400:
2399:
2395:
2388:
2371:
2370:
2363:
2329:
2320:
2319:
2315:
2308:
2291:
2290:
2283:
2245:
2244:
2240:
2235:
2231:
2223:
2212:
2208:
2207:
2203:
2196:
2182:
2181:
2172:
2165:
2148:
2147:
2140:
2106:
2105:
2101:
2094:
2077:
2076:
2059:
2050:
2046:
2040:
2036:
2030:
2026:
1982:
1981:
1977:
1963:
1959:
1954:
1936:
1931:
1929:
1922:
1917:
1915:
1908:
1903:
1901:
1898:
1878:crocodylomorphs
1871:
1825:
1678:Diabloceratops.
1602:Diabloceratops.
1528:Albertaceratops
1494:Diabloceratops,
1464:Different from
1403:parasaurolophin
1389:
1195:, at least two
1189:
1176:scapherpetontid
1039:
864:
502:C. missouriense
371:
347:
342:
320:
308:Late Cretaceous
257:
255:
232:
230:
226:
223:
218:
215:
213:
211:
210:
132:
112:
111:
110:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
39:82.2–77.3
38:
37:
17:
12:
11:
5:
2948:
2946:
2938:
2937:
2932:
2927:
2922:
2917:
2912:
2907:
2902:
2892:
2891:
2885:
2884:
2882:
2881:
2871:
2866:
2860:
2858:
2854:
2853:
2851:
2850:
2845:
2840:
2835:
2830:
2825:
2823:Grosvenor Arch
2820:
2815:
2810:
2805:
2800:
2794:
2792:
2788:
2787:
2785:
2784:
2779:
2774:
2769:
2764:
2758:
2756:
2752:
2751:
2746:
2744:
2743:
2736:
2729:
2721:
2713:
2712:
2696:
2686:
2667:(8): 699–702.
2648:
2641:
2622:
2555:
2508:(11): e79420.
2488:
2481:
2456:
2449:
2422:
2415:
2393:
2386:
2361:
2313:
2306:
2281:
2254:(4): 798–811.
2238:
2229:
2201:
2194:
2170:
2163:
2138:
2119:(3): 241–257.
2099:
2092:
2057:
2044:
2034:
2024:
1975:
1956:
1955:
1953:
1950:
1949:
1948:
1942:
1941:
1927:
1913:
1897:
1894:
1870:
1867:
1824:
1821:
1818:
1817:
1814:
1813:
1812:
1806:
1798:
1795:
1793:
1792:Indeterminate
1790:
1784:
1783:
1780:
1779:
1778:
1775:
1767:
1764:
1761:
1760:Indeterminate
1758:
1752:
1751:
1748:
1747:
1746:
1743:
1740:
1722:
1706:
1705:
1704:
1701:
1698:
1693:
1691:
1690:Indeterminate
1688:
1682:
1681:
1674:
1666:
1663:
1660:
1653:
1643:
1642:
1639:tyrannosaurine
1634:
1628:
1625:
1622:
1615:
1605:
1604:
1598:
1594:UMNH VP 166999
1590:
1587:
1581:
1574:
1570:Diabloceratops
1564:
1563:
1554:
1548:
1545:
1542:
1541:Indeterminate
1539:
1537:Centrosaurinae
1533:
1532:
1523:
1517:
1514:
1511:
1510:Indeterminate
1508:
1506:Centrosaurinae
1502:
1501:
1490:
1484:
1481:
1478:
1477:Indeterminate
1475:
1473:Centrosaurinae
1469:
1468:
1462:
1456:
1453:
1450:
1449:Indeterminate
1447:
1439:
1438:
1436:
1422:
1419:
1417:
1416:Indeterminate
1414:
1412:Ankylosauridae
1408:
1407:
1391:
1383:
1376:
1373:
1370:
1363:
1353:
1352:
1328:Diabloceratops
1290:
1268:
1262:
1259:
1256:
1253:A. gagslarsoni
1249:
1239:
1238:
1235:
1232:
1229:
1226:
1223:
1220:
1216:
1215:
1188:
1185:
1182:
1181:
1179:
1172:
1166:
1164:
1161:
1155:
1147:
1146:
1144:
1137:
1136:
1135:
1129:
1121:
1119:
1116:
1110:
1102:
1101:
1099:
1092:
1086:
1084:
1081:
1078:
1074:
1073:
1070:
1067:
1064:
1061:
1058:
1055:
1051:
1050:
1047:Lissamphibians
1038:
1035:
1032:
1031:
1024:
1018:
1016:
1013:
1012:Indeterminate
1010:
1004:
1003:
996:
993:
991:
989:
983:
975:
974:
971:pycnodontiform
967:
961:
959:
956:
950:
942:
941:
924:
917:
914:
912:
910:
909:Indeterminate
907:
899:
898:
895:
892:
889:
886:
883:
880:
876:
875:
863:
860:
857:
856:
849:
848:
847:
841:
823:
815:
813:
812:
811:
808:
803:
798:
790:
789:
783:
777:
775:
772:
766:
758:
757:
754:
748:
746:
743:
737:
728:
727:
724:
723:
722:
704:
662:
638:
636:
635:
634:
631:
628:
623:
618:
610:
609:
600:sclerorhynchid
596:
574:
572:
569:
564:
556:
555:
542:
531:UMMNH VP 18886
516:
514:
513:
512:
509:
504:
499:
491:
490:
463:
456:
434:
432:
431:
430:
427:
424:
419:
414:
411:Cantioscyllium
406:
405:
402:
399:
396:
393:
390:
387:
383:
382:
370:
367:
351:petrified logs
346:
343:
341:
338:
337:
336:
333:
330:
327:
319:
316:
290:
289:
284:
280:
279:
275:
274:
271:
267:
266:
253:
249:
248:
245:
241:
240:
228:37.5°N 111.7°W
208:
202:
201:
197:
196:
191:
187:
186:
182:
181:
176:
172:
171:
166:
162:
161:
158:
154:
153:
148:
144:
143:
138:
134:
133:
123:
115:
114:
107:
106:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
44:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
2947:
2936:
2933:
2931:
2928:
2926:
2923:
2921:
2918:
2916:
2913:
2911:
2908:
2906:
2903:
2901:
2898:
2897:
2895:
2878:
2877:
2872:
2870:
2867:
2865:
2862:
2861:
2859:
2855:
2849:
2846:
2844:
2841:
2839:
2836:
2834:
2831:
2829:
2826:
2824:
2821:
2819:
2816:
2814:
2813:Devils Garden
2811:
2809:
2806:
2804:
2801:
2799:
2796:
2795:
2793:
2789:
2783:
2780:
2778:
2775:
2773:
2770:
2768:
2765:
2763:
2760:
2759:
2757:
2753:
2749:
2742:
2737:
2735:
2730:
2728:
2723:
2722:
2719:
2709:
2706:
2700:
2697:
2690:
2687:
2682:
2678:
2674:
2670:
2666:
2662:
2655:
2653:
2649:
2644:
2642:9780253008961
2638:
2634:
2626:
2623:
2618:
2614:
2609:
2604:
2600:
2596:
2591:
2586:
2582:
2578:
2574:
2570:
2566:
2559:
2556:
2551:
2547:
2542:
2537:
2533:
2529:
2524:
2519:
2515:
2511:
2507:
2503:
2499:
2492:
2489:
2484:
2482:9780253007797
2478:
2474:
2467:
2460:
2457:
2452:
2450:9780253008961
2446:
2442:
2438:
2431:
2429:
2427:
2423:
2418:
2416:9780253008961
2412:
2408:
2404:
2397:
2394:
2389:
2387:9780253008961
2383:
2379:
2375:
2368:
2366:
2362:
2357:
2353:
2348:
2343:
2339:
2335:
2328:
2326:
2317:
2314:
2309:
2307:9780253013859
2303:
2299:
2295:
2288:
2286:
2282:
2277:
2273:
2269:
2265:
2261:
2257:
2253:
2249:
2242:
2239:
2233:
2230:
2222:
2218:
2211:
2205:
2202:
2197:
2195:9780253008961
2191:
2187:
2179:
2177:
2175:
2171:
2166:
2164:9780253008961
2160:
2156:
2152:
2145:
2143:
2139:
2134:
2130:
2126:
2122:
2118:
2114:
2110:
2103:
2100:
2095:
2093:9780253008961
2089:
2085:
2081:
2074:
2072:
2070:
2068:
2066:
2064:
2062:
2058:
2054:
2048:
2045:
2038:
2035:
2028:
2025:
2020:
2016:
2011:
2006:
2002:
1998:
1994:
1990:
1986:
1979:
1976:
1972:
1971:0-520-24209-2
1968:
1961:
1958:
1951:
1947:
1944:
1943:
1939:
1928:
1925:
1914:
1911:
1900:
1895:
1893:
1891:
1885:
1883:
1882:ornithischian
1879:
1875:
1874:Trace fossils
1869:Trace fossils
1868:
1866:
1864:
1861:
1857:
1853:
1852:cladotherians
1849:
1841:
1837:
1834:
1829:
1822:
1815:
1810:
1809:UMNH VP 13881
1807:
1804:
1803:UMNH VP 21087
1801:
1800:
1799:
1796:
1794:
1791:
1789:
1788:Saurolophinae
1786:
1785:
1781:
1776:
1773:
1772:UMNH VP 11939
1770:
1769:
1768:
1765:
1762:
1759:
1757:
1754:
1753:
1749:
1744:
1741:
1738:
1737:UMNH VP 21207
1734:
1733:UMNH VP 16408
1730:
1729:UMNH VP 15664
1726:
1725:UMNH VP 13981
1723:
1720:
1716:
1712:
1709:
1708:
1707:
1702:
1699:
1696:
1695:
1694:
1692:
1689:
1687:
1684:
1683:
1679:
1675:
1673:
1671:
1670:UMNH VP 20550
1667:
1664:
1661:
1659:
1658:
1654:
1652:
1651:
1650:
1645:
1644:
1641:
1640:
1635:
1632:
1631:UMNH VP 20200
1629:
1626:
1623:
1621:
1620:
1616:
1614:
1613:
1612:
1607:
1606:
1603:
1599:
1597:
1595:
1591:
1588:
1586:
1582:
1580:
1579:
1575:
1573:
1572:
1571:
1566:
1565:
1562:
1560:
1559:Styracosaurus
1555:
1552:
1549:
1546:
1543:
1540:
1538:
1535:
1534:
1531:
1529:
1524:
1521:
1520:UMNH VP 20600
1518:
1515:
1512:
1509:
1507:
1504:
1503:
1499:
1495:
1491:
1488:
1487:UMNH VP 16704
1485:
1482:
1480:Nipple Butte
1479:
1476:
1474:
1471:
1470:
1467:
1463:
1460:
1457:
1454:
1451:
1448:
1446:
1445:
1441:
1440:
1437:
1434:
1430:
1426:
1423:
1420:
1418:
1415:
1413:
1410:
1409:
1406:
1404:
1401:, possibly a
1400:
1397:
1396:lambeosaurine
1392:
1390:
1388:
1381:
1377:
1374:
1371:
1369:
1368:
1367:A. hutchisoni
1364:
1362:
1361:
1360:
1355:
1354:
1349:
1345:
1339:
1335:
1329:
1325:
1319:
1315:
1309:
1305:
1299:
1295:
1289:
1287:
1283:
1281:
1276:
1275:
1269:
1266:
1265:UMNH VP 16607
1263:
1260:
1257:
1255:
1254:
1250:
1248:
1247:
1246:
1241:
1240:
1236:
1233:
1230:
1227:
1224:
1221:
1218:
1217:
1214:
1212:
1207:
1204:
1202:
1198:
1194:
1186:
1180:
1177:
1173:
1170:
1169:UMNH VP 19186
1167:
1165:
1162:
1159:
1156:
1154:
1153:
1149:
1148:
1145:
1142:
1138:
1133:
1130:
1127:
1124:
1123:
1122:
1120:
1117:
1114:
1111:
1109:
1108:
1107:Opisthotriton
1104:
1103:
1100:
1097:
1093:
1090:
1089:UMNH VP 19209
1087:
1085:
1082:
1079:
1076:
1075:
1071:
1068:
1065:
1062:
1059:
1056:
1053:
1052:
1048:
1044:
1036:
1029:
1025:
1022:
1019:
1017:
1014:
1011:
1009:
1008:Polyodontidae
1006:
1005:
1001:
997:
994:
992:
990:
987:
984:
982:
981:
977:
976:
972:
968:
965:
962:
960:
957:
954:
951:
949:
948:
947:Micropycnodon
944:
943:
937:
933:
932:Albula vulpes
929:
922:
918:
915:
913:
911:
908:
906:
905:
901:
900:
896:
893:
890:
887:
884:
881:
878:
877:
873:
869:
861:
854:
850:
845:
844:UMNH VP 18918
842:
839:
838:UMNH VP 18920
835:
831:
830:UMNH VP 18885
827:
826:UMNH VP 18836
824:
822:Rostral teeth
821:
818:
817:
816:
814:
809:
806:
805:
804:
802:
799:
797:
796:
792:
791:
788:
785:Smaller than
784:
781:
780:UMNH VP 18917
778:
776:
773:
770:
767:
765:
764:
760:
759:
755:
752:
749:
747:
744:
741:
738:
735:
734:
730:
729:
725:
721:; large teeth
720:
719:UMNH VP 17401
716:
712:
708:
705:
703:; small teeth
702:
701:UMNH VP 17404
698:
697:UMNH VP 17399
694:
693:UMNH VP 17395
690:
689:UMNH VP 17393
686:
682:
678:
674:
670:
666:
663:
660:
656:
655:UMNH VP 17393
652:
648:
644:
641:
640:
639:
637:
632:
629:
626:
625:
624:
622:
619:
617:
616:
612:
611:
608:
606:
601:
597:
594:
593:UMNH VP 18878
590:
589:UMNH VP 18876
586:
585:UMNH VP 18836
582:
581:UMNH VP 18879
578:
577:UMNH VP 18877
575:
573:
570:
568:
565:
563:
562:
558:
557:
554:
552:
547:
543:
540:
539:UMNH VP 18888
536:
535:UMNH VP 18887
532:
528:
527:UMNH VP 18882
524:
523:UMNH VP 18880
520:
517:
515:
510:
507:
506:
505:
503:
500:
498:
497:
496:Chiloscyllium
493:
492:
487:
483:
476:
472:
468:
461:
457:
454:
453:UMNH VP 18919
450:
446:
442:
441:MNA V10384-85
438:
437:MNA V10230-32
435:
433:
428:
425:
422:
421:
420:
418:
415:
413:
412:
408:
407:
403:
400:
397:
394:
391:
388:
385:
384:
380:
379:Elasmobranchs
376:
369:Elasmobranchs
368:
366:
364:
360:
356:
352:
345:Invertebrates
344:
339:
334:
331:
328:
325:
324:
323:
317:
315:
313:
309:
305:
301:
297:
288:
287:Wahweap Creek
285:
281:
276:
272:
268:
265:
254:
250:
247:North America
246:
242:
237:
209:
207:
203:
198:
195:
192:
188:
183:
180:
177:
173:
170:
167:
163:
159:
155:
152:
149:
145:
142:
139:
135:
131:
127:
121:
116:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
42:
35:
31:
27:
22:
19:
2874:
2847:
2828:Horse Canyon
2808:Death Hollow
2782:Willis Creek
2762:Coyote Gulch
2707:
2704:
2699:
2689:
2664:
2660:
2632:
2625:
2572:
2568:
2558:
2505:
2501:
2491:
2472:
2459:
2440:
2406:
2396:
2377:
2337:
2333:
2324:
2316:
2297:
2251:
2247:
2241:
2232:
2221:the original
2216:
2204:
2185:
2154:
2116:
2112:
2102:
2083:
2052:
2047:
2037:
2027:
1992:
1988:
1978:
1960:
1890:maniraptoran
1886:
1872:
1863:insectivores
1845:
1839:
1833:maniraptoran
1808:
1802:
1771:
1763:Clints Cove
1739:; osteoderms
1736:
1732:
1728:
1724:
1718:
1714:
1710:
1686:Nodosauridae
1677:
1669:
1668:
1656:
1655:
1647:
1646:
1636:
1630:
1618:
1617:
1609:
1608:
1601:
1593:
1592:
1577:
1576:
1568:
1567:
1557:
1551:UMNH VP 9549
1550:
1544:Death Ridge
1526:
1519:
1513:Pilot Knoll
1493:
1486:
1465:
1459:UMNH VP 9548
1458:
1452:Death Ridge
1442:
1432:
1428:
1424:
1393:
1379:
1378:
1372:UCMP V98173
1366:
1365:
1357:
1356:
1347:
1337:
1327:
1317:
1307:
1297:
1278:
1272:
1270:
1264:
1252:
1251:
1243:
1242:
1209:
1197:ceratopsians
1190:
1168:
1157:
1152:Scapherpeton
1150:
1131:
1125:
1112:
1105:
1088:
1020:
985:
978:
963:
952:
945:
931:
921:seminotiform
902:
872:Osteichthyes
862:Osteichthyes
843:
840:; Oral teeth
837:
833:
829:
825:
819:
801:T. brycensis
800:
793:
786:
779:
768:
761:
750:
739:
731:
718:
714:
710:
706:
700:
696:
692:
688:
684:
680:
676:
672:
668:
664:
654:
650:
646:
642:
620:
613:
603:
592:
588:
584:
580:
576:
567:C. deblieuxi
566:
559:
549:
546:bamboo shark
538:
534:
530:
526:
522:
518:
501:
494:
485:
470:
452:
448:
444:
440:
436:
416:
409:
348:
321:
295:
293:
278:Type section
233:37.5; -111.7
18:
2777:Paria River
2772:Harris Wash
1713:(in part),
1619:L. argestes
1466:Acristavus.
1427:(in part),
1399:hadrosaurid
1380:UCMP 152028
1359:Adelolophus
1308:Adelolophus
1037:Salamanders
1023:; denticle
621:C. ciffelli
615:Cristomylus
605:Squatirhina
460:nurse shark
353:to various
231: /
206:Coordinates
2894:Categories
2298:Hadrosaurs
1995:: 110876.
1952:References
1856:marsupials
1719:OMNH 24278
1715:OMNH 21992
1711:OMNH 21280
1662:Star Seep
1657:M. cronusi
1433:OMNH 24276
1429:OMNH 21858
1425:OMNH 21280
1382:; partial
1298:Acristavus
1245:Acristavus
1128:; atlantes
1077:Gen. nov.
1028:paddlefish
964:MNA V10336
846:; denticle
795:Texatrygon
763:Lonchidion
751:MNA V10387
659:symphysial
561:Columbusia
519:MNA V10386
340:Paleobiota
2599:1932-6203
2532:1932-6203
2356:0031-0298
2334:PaleoBios
2133:0077-7749
2019:0031-0182
1860:placental
1611:Lythronax
1578:D. eatoni
1338:Lythronax
1280:Maiasaura
1231:Material
1225:Locality
1211:Dinosaurs
1193:hadrosaur
1187:Dinosaurs
1080:sp. nov.
1066:Material
1060:Locality
1021:MNA 10356
980:Paralbula
904:Lepidotes
891:Material
885:Locality
715:MNA V9686
711:MNA V9683
707:MNA V9593
685:MNA V9652
681:MNA V9635
677:MNA V9633
673:MNA V9569
669:MNA V9525
665:MNA V9502
651:MNA V9600
647:MNA V9568
643:MNA V9531
445:MNA 10390
417:C. estesi
398:Material
392:Locality
363:siltstone
359:gastropod
312:Campanian
304:formation
283:Named for
194:sandstone
185:Lithology
165:Underlies
157:Sub-units
34:Campanian
2617:27192148
2569:PLOS ONE
2550:24223179
2502:PLOS ONE
1896:See also
1836:dinosaur
1435:; Teeth
1222:Species
1201:theropod
1171:; atlas
1096:sirenids
1057:Species
1000:bonefish
936:Bonefish
782:; Tooth
753:; Tooth
595:; teeth
541:; teeth
455:; teeth
389:Species
355:mollusks
219:111°42′W
200:Location
175:Overlies
2791:Geology
2669:Bibcode
2661:Geology
2608:4871575
2577:Bibcode
2541:3819173
2510:Bibcode
2276:8878474
2256:Bibcode
1997:Bibcode
1823:Mammals
1721:; Teeth
1386:maxilla
1237:Images
1228:Member
1072:Images
1063:Member
897:Images
888:Member
853:sawfish
733:Hybodus
404:Images
395:Member
298:of the
252:Country
216:37°30′N
190:Primary
147:Unit of
108:↓
2705:et al.
2639:
2615:
2605:
2597:
2548:
2538:
2530:
2479:
2447:
2413:
2384:
2354:
2304:
2274:
2192:
2161:
2131:
2090:
2017:
1969:
1858:, and
1840:et al.
1234:Notes
1219:Taxon
1069:Notes
1054:Taxon
995:Teeth
916:Teeth
894:Notes
882:Genus
879:Taxon
401:Notes
386:Taxon
270:Extent
261:
244:Region
2857:Other
2469:(PDF)
2330:(PDF)
2272:S2CID
2224:(PDF)
2213:(PDF)
2042:2005.
661:teeth
2694:2005
2637:ISBN
2613:PMID
2595:ISSN
2546:PMID
2528:ISSN
2477:ISBN
2445:ISBN
2411:ISBN
2382:ISBN
2352:ISSN
2302:ISBN
2190:ISBN
2159:ISBN
2129:ISSN
2088:ISBN
2032:2007
2015:ISSN
1967:ISBN
1277:and
1160:sp.
1115:sp.
988:sp.
955:sp.
771:sp.
294:The
137:Type
130:Utah
47:Preęž’
2677:doi
2603:PMC
2585:doi
2536:PMC
2518:doi
2342:doi
2264:doi
2121:doi
2117:280
2005:doi
1993:591
742:sp
691:,
318:Age
264:USA
2896::
2675:.
2665:38
2663:.
2651:^
2611:.
2601:.
2593:.
2583:.
2573:11
2571:.
2567:.
2544:.
2534:.
2526:.
2516:.
2504:.
2500:.
2425:^
2364:^
2350:.
2340:.
2338:33
2336:.
2332:.
2284:^
2270:.
2262:.
2252:31
2250:.
2215:.
2173:^
2141:^
2127:.
2115:.
2111:.
2060:^
2013:.
2003:.
1991:.
1987:.
1865:.
1854:,
1850:,
1831:A
1735:,
1731:,
1727:,
1717:,
1637:A
1500:.
1431:,
1405:.
1394:A
1288:.
1203:.
1178:.
1174:A
1158:S.
1139:A
1113:O.
1026:A
998:A
986:P.
969:A
953:M.
919:A
855:.
851:A
836:,
832:,
828:,
769:L.
740:H.
736:?
717:,
713:,
709:,
699:,
695:,
687:,
683:,
679:,
675:,
671:,
667:,
657:;
653:,
649:,
645:,
598:A
591:,
587:,
583:,
579:,
544:A
537:,
533:,
529:,
525:,
521:,
458:A
451:,
447:,
443:,
439:,
365:.
128:,
97:Pg
41:Ma
36:,
32::
2740:e
2733:t
2726:v
2683:.
2679::
2671::
2645:.
2619:.
2587::
2579::
2552:.
2520::
2512::
2506:8
2485:.
2453:.
2419:.
2390:.
2358:.
2344::
2310:.
2278:.
2266::
2258::
2198:.
2167:.
2135:.
2123::
2096:.
2021:.
2007::
1999::
1973:.
1561:.
1530:.
1282:.
938:)
934:(
607:.
553:.
477:)
473:(
310:(
102:N
92:K
87:J
82:T
77:P
72:C
67:D
62:S
57:O
52:ęž’
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
