1389:
744:
94:
756:
732:
42:
69:
582:
503:
541:, produces a much larger, darker and more robust shell that can reach 120 mm and has the characteristic winged protrusions. It is usually less elongated than that of the "southern" form and lacks its porcelain-like shine. Further research is needed to determine whether these forms represent two separate species or not.
603:
is known to cling to objects floating on the surface of the sea, including other argonauts. Chains of up to 20-30 argonauts of similar size have been reported. The first female in such chains usually clings to some inanimate object, while the other females hold on to the ventral part of the shell of
656:. The argonaut was observed using the jellyfish as cover, rotating the animal to hide itself from potential predators (in this case the photographer). The argonaut was also seen using the jellyfish as a 'hunting platform', as it "manoeuvered its host close to a smaller
768:
498:
along the keel, and less than 40 smooth ribs across the sides of the shell. The shell is usually approximately 80 mm in length, although it can exceed 120 mm in exceptional specimens; the world record size is 121.5 mm.
529:. It is a much smaller animal, with a shell that rarely exceeds 80 mm and lacks the winged protrusions for which this species is named. The "northern" form, which is found in the waters surrounding
521:
waters worldwide. It is an extremely variable species and there appear to exist at least two distinct forms; a "southern" form and "northern" form (see images). The former is most abundant in the
767:
578:
have laid their eggs. However, the size at which this takes place differs across the animal's range. Females grow to 50 mm ML, while males do not exceed 20 mm ML.
486:. The common name comes from the grey to brown coloured shell. The Chinese name for this species translates as "grey sea-horse's nest". The female of the species, like all
743:
568:
Males of this species reach sexual maturity at a mantle length (ML) of about 7 mm, presumably the maximum size attained. Females mature at about half the size of
1527:
1566:
1646:
731:
629:
in the
Philippines. This behaviour has been known for a long time, although little was understood about the relationship prior to the work of Heeger
574:. They begin to secrete an eggcase at 6.5–7 mm ML. Egg laying usually commences when females reach 14–15 mm ML; by 18–20 mm ML female
1501:
755:
1540:
1579:
1203:
773:
1656:
1269:
1228:
305:
288:
1618:
176:
1388:
490:, creates a paper-thin eggcase that coils around the octopus much like the way a nautilus lives in its shell (hence the name
1545:
1462:
1449:
1571:
93:
341:
1262:
714:
1134:
1404:
1467:
694:
345:
198:
1165:
1623:
1488:
1436:
1098:
1063:
718:
625:
594:
250:
237:
1377:
1370:
1349:
706:
58:
819:
559:. The species is preyed on by numerous predators. It has been reported in the stomach contents of
1651:
1610:
1356:
1342:
1335:
1300:
1255:
1103:
in Japan, from the middle
Pliocene Sadowara Formation, southwest Japan, and its depositional age"
668:
645:
223:
214:
88:
1192:
1532:
1592:
1475:
1307:
1279:
892:(Cephalopoda, Octopoda), in the western Pacific and the seas of the East Indian Archipelago".
678:
561:
494:). The eggcase is characterised by a wide keel that gives it a square appearance, few rounded
487:
165:
1597:
1032:
1558:
1328:
1321:
1232:
1114:
1071:
1025:
Kramp, P. L. (1956). "Pelagic Fauna". In A. Bruun; SV. Greve; H. Mielche; R. Spärck (eds.).
979:
814:
1363:
702:
605:
526:
1441:
1067:
41:
1454:
1293:
570:
491:
1640:
1553:
1026:
983:
805:
683:
623:
is often observed attached to jellyfish. It has been photographed atop the jellyfish
172:
78:
73:
796:
1480:
673:
581:
155:
325:
1605:
1514:
649:
522:
518:
1427:
17:
657:
502:
446:
135:
749:
Apparently immature male (5.0 mm ML) from the equatorial South
Atlantic
641:
613:
552:
534:
105:
1181:
1118:
636:
Underwater photographer Mark
Strickland observed and photographed a female
1421:
1239:
938:. Fauna Flora Golf. Napoli (35) (in German). Vol. 1. pp. 1–863.
691:
495:
125:
1001:
1506:
1076:
1047:
653:
548:
483:
480:
476:
272:
145:
1519:
1096:
Iwatani, H., K. Murai, T. Irizuki, H. Hayashi & Y. Tanaka (2009).
660:, quickly grasped it with another pair of tentacles and devoured it".
530:
115:
1398:
1247:
776:
of juvenile male (5.0 mm ML) from the equatorial South
Atlantic
737:
Juvenile female (6.5 mm ML) from the equatorial South
Atlantic
580:
538:
501:
1584:
1493:
1402:
1251:
1147:
Mitteilungen des naturwissenschaftlichen
Vereins fĂĽr Steiermark
761:
Oral view of the same animal; note the modified hectocotylus
1193:
Taxa
Associated with the Family Argonautidae Tryon, 1879.
608:
and Gordon
Williamson observed six freshly mated female
970:
David, P. M. (1965). "The surface fauna of the ocean".
884:
Nesis, K. N. (1977). "The biology of paper nautiluses,
697:
of southwestern Japan. In terms of eggcase morphology,
1411:
911:Rancurel, P. (1970). "Les contenus stomacaux d'
956:. Hong Kong: Government Press. p. 138 pp.
1263:
1172:. Librairie F. Savy, Paris. pp. 380–381.
1046:Heeger, T.; U. Piatkowski; H. Möller (1992).
915:dans le sud-ouest Pacifique (CĂ©phalopodes)".
820:10.2305/IUCN.UK.2014-3.RLTS.T163211A985065.en
666:appears to be closely related to the smaller
8:
1182:Current Classification of Recent Cephalopoda
1000:Mangold, K. M.; M. Vecchione; R. E. Young.
517:is cosmopolitan, occurring in tropical and
1399:
1270:
1256:
1248:
1108:Journal of the Geological Society of Japan
1048:"Predation on jellyfish by the cephalopod
1028:The Galathea Deep Sea Expedition 1950–1952
67:
40:
31:
1089:
1087:
1075:
995:
993:
818:
1135:"Der älteste bekannte und erste miozäne
851:
849:
847:
845:
843:
841:
839:
837:
585:One of the earliest known depictions of
555:have been reported from the stomachs of
965:
963:
879:
877:
875:
873:
871:
869:
867:
787:
727:
947:
945:
686:. The oldest known fossil material of
616:that were swimming along in a string.
1208:Australian Biological Resources Study
7:
1647:IUCN Red List least concern species
1164:Hoernes, R. & L. Dollo (1886).
1099:"Discovery of the oldest fossil of
952:Voss, G. L.; G. Williamson (1971).
806:IUCN Red List of Threatened Species
25:
1387:
766:
754:
742:
730:
565:from the south-western Pacific.
92:
860:. ConchBooks. pp. 189–195.
774:Funnel–mantle locking apparatus
1056:Marine Ecology Progress Series
919:. Ser. Océanogr. (in French).
1:
984:10.1016/0160-9327(65)90007-4
936:Die Cephalopoden, Systematik
1229:"CephBase: Argonauta hians"
690:originates from the middle
591:Index Testarum Conchyliorum
547:feeds primarily on pelagic
1673:
1657:Molluscs described in 1786
1240:Tree of Life web project:
858:Cephalopods: A World Guide
1385:
1288:
229:
222:
204:
197:
89:Scientific classification
87:
65:
56:
48:
39:
34:
1196:Tree of Life web project
1006:Tree of Life web project
954:Cephalopods of Hong Kong
1191:Sweeney, M. J. (2002).
1170:Manuel de Paléontologie
1119:10.5575/geosoc.115.548
894:Zoologicheskii Zhurnal
604:the preceding animal.
597:
511:
584:
505:
284:Argonauta crassicosta
813:: e.T163211A985065.
795:Allcock, L. (2014).
682:from the north-east
626:Phyllorhiza punctata
473:brown paper nautilus
463:, also known as the
1378:Argonauta tokunagai
1371:Argonauta sismondai
1350:Argonauta itoigawai
1132:Hilber, V. (1915).
1068:1992MEPS...88..293H
917:Cahiers de l'ORSTOM
886:Argonauta boettgeri
856:Norman, M. (2000).
619:In the open ocean,
510:, Taiwan (121.5 mm)
506:"Northern" form of
297:Argonauta raricosta
59:Conservation status
1357:Argonauta joanneus
1343:Argonauta absyrtus
1336:Argonauta pacifica
1301:Argonauta bottgeri
1077:10.3354/meps088293
695:Sadowara Formation
646:Mergui Archipelago
598:
512:
357:Argonauta kochiana
246:Argonauta haustrum
27:Species of mollusc
1634:
1633:
1593:Open Tree of Life
1405:Taxon identifiers
1396:
1395:
1340:Extinct species:
1308:Argonauta cornuta
934:Naef, A. (1923).
913:Alepisaurus ferox
707:A. sismondai
595:Niccolò Gualtieri
562:Alepisaurus ferox
456:
455:
450:
443:
439:Argonauta expansa
434:
433:Von Martens, 1867
420:
419:Von Martens, 1867
406:
405:Von Martens, 1867
392:
383:
372:
367:Argonauta cornuta
362:
353:
333:
309:
292:
280:
264:
254:
241:
233:Argonauta gondola
179:
82:
16:(Redirected from
1664:
1627:
1626:
1614:
1613:
1601:
1600:
1588:
1587:
1575:
1574:
1562:
1561:
1549:
1548:
1536:
1535:
1523:
1522:
1510:
1509:
1497:
1496:
1484:
1483:
1471:
1470:
1458:
1457:
1445:
1444:
1432:
1431:
1430:
1400:
1391:
1329:Argonauta nouryi
1322:Argonauta nodosa
1272:
1265:
1258:
1249:
1236:
1231:. Archived from
1217:
1215:
1214:
1184:
1179:
1173:
1163:
1159:
1153:
1145:
1143:
1131:
1127:
1121:
1106:
1095:
1091:
1082:
1081:
1079:
1043:
1037:
1036:
1022:
1016:
1015:
1013:
1012:
997:
988:
987:
967:
958:
957:
949:
940:
939:
931:
925:
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862:
861:
853:
832:
831:
829:
827:
822:
792:
770:
758:
746:
734:
669:A. bottgeri
445:
442:
432:
418:
404:
390:
387:Argonauta polita
380:
377:Argonauta dispar
370:
360:
340:
337:Argonauta owenii
324:
300:
287:
271:
268:Argonauta nitida
262:
259:Ocythoe cranchii
249:
236:
210:
171:
97:
96:
76:
71:
70:
44:
35:Winged argonaut
32:
21:
1672:
1671:
1667:
1666:
1665:
1663:
1662:
1661:
1637:
1636:
1635:
1630:
1622:
1617:
1609:
1604:
1596:
1591:
1583:
1578:
1570:
1565:
1557:
1552:
1544:
1539:
1531:
1526:
1518:
1513:
1505:
1500:
1492:
1487:
1479:
1474:
1466:
1461:
1455:Argonauta_hians
1453:
1448:
1442:Argonauta_hians
1440:
1435:
1426:
1425:
1420:
1413:Argonauta hians
1407:
1397:
1392:
1383:
1364:Argonauta oweri
1339:
1315:Argonauta hians
1284:
1276:
1227:
1224:
1212:
1210:
1201:
1188:
1187:
1180:
1176:
1161:
1160:
1156:
1141:
1133:
1129:
1128:
1124:
1113:(10): 548–551.
1101:Argonauta hians
1097:
1093:
1092:
1085:
1045:
1044:
1040:
1024:
1023:
1019:
1010:
1008:
999:
998:
991:
969:
968:
961:
951:
950:
943:
933:
932:
928:
910:
909:
905:
883:
882:
865:
855:
854:
835:
825:
823:
799:Argonauta hians
794:
793:
789:
784:
777:
771:
762:
759:
750:
747:
738:
735:
719:type repository
679:A. cornuta
606:Gilbert L. Voss
527:South China Sea
508:Argonauta hians
465:winged argonaut
460:Argonauta hians
444:
441:
431:
425:Argonauta hians
417:
411:Argonauta hians
403:
397:Argonauta hians
389:
381:
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369:
359:
339:
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299:
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270:
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248:
235:
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208:Argonauta hians
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193:
170:
91:
83:
72:
68:
61:
51:Argonauta hians
28:
23:
22:
18:Winged Argonaut
15:
12:
11:
5:
1670:
1668:
1660:
1659:
1654:
1649:
1639:
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1353:
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1332:
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1294:Argonauta argo
1289:
1286:
1285:
1277:
1275:
1274:
1267:
1260:
1252:
1246:
1245:
1237:
1235:on 2005-08-17.
1223:
1222:External links
1220:
1219:
1218:
1204:"Argonautidae"
1199:
1186:
1185:
1174:
1154:
1122:
1083:
1050:Argonauta argo
1038:
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989:
978:(92): 95–100.
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748:
741:
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701:resembles the
640:clinging to a
571:Argonauta argo
492:paper nautilus
469:muddy argonaut
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1094:(in Japanese)
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733:
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725:are unknown.
724:
720:
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715:type locality
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699:A. hians
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693:
689:
688:A. hians
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551:. Remains of
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199:Binomial name
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190:A. hians
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74:Least Concern
64:
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1313:
1306:
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1241:
1233:the original
1211:. Retrieved
1207:
1195:
1177:
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1157:
1149:
1146:
1136:
1125:
1110:
1107:
1100:
1059:
1055:
1049:
1041:
1027:
1020:
1009:. Retrieved
1005:
975:
971:
953:
935:
929:
920:
916:
912:
906:
900:: 1004–1014.
897:
893:
889:
885:
857:
824:. Retrieved
810:
804:
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790:
722:
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705:
698:
687:
677:
674:Indian Ocean
667:
663:
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507:
472:
468:
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438:
428:
424:
414:
410:
400:
396:
391:Conrad, 1854
386:
382:Conrad, 1854
376:
371:Conrad, 1854
366:
361:Dunker, 1852
356:
349:
336:
329:
320:
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312:
302:
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156:Argonautidae
50:
29:
1606:SeaLifeBase
1515:iNaturalist
1162:(in French)
1130:(in German)
1062:: 293–296.
1031:. pp.
1002:"Argonauta"
826:12 November
650:Andaman Sea
523:Philippines
519:subtropical
429:obtusangula
263:Leach, 1817
136:Cephalopoda
49:Eggcase of
1641:Categories
1213:2006-09-13
1202:Lu, C. C.
1152:: 107–110.
1011:2006-09-13
923:(4): 4–87.
782:References
658:comb jelly
593:(1742) by
553:heteropods
317:Philonexis
306:Blainville
289:Blainville
1652:Argonauta
1280:Argonauta
1242:Argonauta
1137:Argonauta
972:Endeavour
672:from the
642:jellyfish
633:in 1992.
614:Hong Kong
535:Hong Kong
496:tubercles
488:argonauts
352:1848-1850
332:1834-1847
326:d'Orbigny
279:1815-1822
215:Lightfoot
184:Species:
167:Argonauta
112:Kingdom:
106:Eukaryota
1533:10838053
1428:Q2578622
1422:Wikidata
1166:Octopoda
890:A. hians
723:A. hians
692:Pliocene
664:A. hians
638:A. hians
621:A. hians
610:A. hians
601:A. hians
587:A. hians
576:A. hians
557:A. hians
549:molluscs
545:A. hians
515:A. hians
342:A. Adams
224:Synonyms
173:Linnaeus
152:Family:
146:Octopoda
126:Mollusca
122:Phylum:
116:Animalia
102:Domain:
79:IUCN 3.1
1598:2868106
1572:2052491
1507:5189604
1283:species
1064:Bibcode
703:extinct
684:Pacific
654:Myanmar
644:in the
589:, from
484:octopus
481:pelagic
477:species
475:, is a
348:, 1848
328:, 1834
321:minimus
313:Octopus
275:, 1822
273:Lamarck
251:Dillwyn
238:Dillwyn
162:Genus:
142:Order:
132:Class:
77: (
1624:215276
1585:215276
1559:163211
1520:292208
1494:401012
1468:386815
1168:. In:
631:et al.
531:Taiwan
449:, 1872
415:mutica
401:aurita
344:&
308:, 1826
301:Leach
291:, 1826
253:, 1817
240:, 1817
217:, 1786
1619:WoRMS
1611:58225
1546:82682
1528:IRMNG
1481:67R4R
1142:(PDF)
1033:65–86
539:Japan
346:Reeve
1580:OBIS
1567:NCBI
1554:IUCN
1541:ITIS
1502:GBIF
1463:BOLD
888:and
828:2021
811:2014
717:and
713:The
676:and
612:off
537:and
525:and
447:Dall
177:1758
1489:EoL
1476:CoL
1450:AFD
1437:ADW
1115:doi
1111:115
1072:doi
980:doi
815:doi
721:of
479:of
471:or
427:f.
413:f.
399:f.
1643::
1621::
1608::
1595::
1582::
1569::
1556::
1543::
1530::
1517::
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1439::
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1206:.
1150:51
1086:^
1070:.
1060:88
1058:.
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467:,
350:in
330:in
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303:in
277:in
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1117::
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1080:.
1074::
1066::
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830:.
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81:)
20:)
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