Yeast in winemaking - Knowledge (XXG)

1672: 2121: 31: 225: 1486: 993: 2968: 4939: 611: 308: 1338: 1688:. Ideally winemakers want to add enough inoculum to have a viable cell population density of 5 million cells per milliliter. The exact amount of freeze-dried culture varies by manufacturer and strain of yeast but it is often around 1 gram per gallon (or 25 grams per 100 liters). Wines that could have potentially problematic fermentation (such as high sugar level late harvest or botryized wines) may have more yeast added. 1863: 1710: 1554: 1110: 1977:

predictability of fermentation due to the high level of survival factors that cultured yeast are assured of having without necessarily needing to expose the wine to additional levels of oxygen. Winemakers using "ambient" yeasts that are resident in their winery may not have this same assurance of survival factors and may need to compensate with other winemaking techniques.

494: 360: 772: 2229:, for metabolism. Additionally, Brett can produce a wide range of by-products that could influence the wine beyond just the 4-EP and 4-EG compounds previously discussed. Many of these compounds, such as the "footprints" of the 4-EP and 4-EG, will still remain in the wine even after yeast cells die and are removed by racking and sterile filtration. 236:, winemakers did not know the mechanism that somehow converted sugary grape juice into alcoholic wine. They could observe the fermentation process which was often described as "boiling", "seething" or the wine being "troubled" due to release of carbon dioxide that gave the wine a frothy, bubbling appearance. This history is preserved in the 1994: 251:", would uncover the connection between microscopic yeast cells and the process of the fermentation. It was Pasteur who discovered that yeast converted sugars in the must into alcohol and carbon dioxide, though the exact mechanisms of how the yeast would accomplish this task was not discovered till the 20th century with the 299:

strains that accentuate desirable features in wine, such as aromatic compounds, mouthfeel, and fermentation kinetics. This commercial availability of yeast strains has revolutionized the art of winemaking by allowing for more precise control over the fermentation process and the resultant wine's character.

330:) in the grape must into alcohol. The yeast accomplishes this by utilizing glucose through a series of metabolic pathways that, in the presence of oxygen, produces not only large amounts of energy for the cell but also many different intermediates that the cell needs to function. In the absence of oxygen ( 1691:

Similarly, re-hydration procedures will also vary depending on the manufacturer and winery. Yeast is often inoculated in a volume of water or grape must that is 5–10 times the weight of the dry yeast. This liquid is often brought to temperature of 40 °C (104 °F) prior to the introduction of

1700:

The temperature of the starter culture is then slowly reduced, often by the graduated addition of must to get within 5–10 °C (41–50 °F) of the must that the culture will be added to. This is done to avoid the sudden cold shock that the yeast cells may experience if the starter culture was

1172:

Some distinct difference among various strains include the production of certain "off-flavor" and aromas that may be temporary (but producing a "stinky fermentation") or could stay with the wine and either have to be dealt with through other winemaking means (such as the presence of volatile sulfur

1164:

identified. Not all of the strains are suitable for winemaking and even among the strains that are, there is debate among winemakers and scientists about the actual magnitude of differences between the various strains and their potential impact on the wine. Even among strains that have demonstrated

346:

needed to keep metabolism going. It is through this process of fermentation that ethanol is released by the yeast cells as a waste product. Eventually, if the yeast cells are healthy and fermentation is allowed to run to the completion, all fermentable sugars will be used up by the yeast with only

1696:

components escape the cell. Re-hydration at lower temperatures can greatly reduce the viability of the yeast with up to 60% cell death if the yeast is re-hydrated at 15 °C (59 °F). The culture is then stirred and aerated to incorporate oxygen into the culture which the yeast uses in the

1468:

wild yeast, these genera of wild yeasts have very low tolerance to both alcohol and sulfur dioxide. They are capable of starting a fermentation and often begin this process as early as the harvest bin when clusters of grapes get slightly crushed under their own weight. Some winemakers will try to

1561:

When winemakers select a cultured yeast strain, it is largely done because the winemaker wants a predictable fermentation taken to completion by a strain that has a track record of dependability. Among the particular considerations that are often important to winemakers is a yeast's tendency to:

1345:

In winemaking, the term "wild yeast" has multiple meanings. In its most basic context, it refers to yeast that has not been introduced to the must by intentional inoculation of a cultured strain. Instead, these "wild yeasts" often come into contact with the must through their presence on harvest

298:

In modern winemaking, winemakers have the option to select from a diverse range of yeast strains, each offering distinct characteristics that influence the wine's sensory profile. These strains are readily available for purchase from specialized suppliers. Winemakers can now easily access yeast

1692:

the yeast (though some yeast strains may need temperatures below 38 °C (100 °F)) to allow the cells to disperse easily rather than clump and sink to the bottom of the container. The heat activation also allows the cells to quickly reestablish their membrane barrier before soluble

1976:

Cultured yeasts that are freeze-dried and available for inoculation of wine must are deliberately grown in commercial labs in high oxygen/low sugar conditions that favor the development of these survival factors. One of the reasons that some winemakers prefer using inoculated yeast is the

1972:

and levels of alcohol in the wine. As a waste product of its own metabolism, alcohol is actually very toxic to yeast cells. Yeast with weak survival factors and lacking sterols may succumb to these conditions before fermenting a wine to complete dryness, leaving a stuck fermentation.

1717:

In order to successfully complete a fermentation with minimum to no negative attributes being added to the wine, yeast needs to have the full assortment of its nutritional needs met. These include not only an available energy source (carbon in the form of sugars such as glucose) and

389:

and a slightly sweet taste without increasing the alcohol level of the wine, some winemakers try to intentionally favor conditions that would promote glycerol production in wine. This includes selecting yeast strains that favor glycerol production (or allowing some wild yeast like

1531:

wild yeast will have a role in beginning the fermentation of virtually every wine but for the wineries that choose to allow these yeasts to continue fermenting versus minimizing their influence do so with the intent of enhancing complexity through bio-diversity. While these

1658:

but a negative attribute for many other wines), enhancement of a wine's color or certain varietal characteristics by enzymes in the yeast cells and other metabolic products produced by the yeast, foaming and flocculation tendencies, yeasticidal properties (a trait known as

543:

available in the must or by the decomposition of dead yeast cells by other yeast that releases sulfur-containing amino acids that are further broken down by the yeast. The latter often happens with wines that sit in contact with their lees for long periods of time between

1519:

in order to encourage the sustained presence of favorable strains. But compared to inoculated yeast, these ambient yeasts hold the risk of having a more unpredictable fermentation. Not only could this unpredictability include the presence of off-flavors/aromas and higher

1177:) or leave a faulty wine. Another difference includes the "vigor" or speed of fermentation (which can also be influenced by other factors beyond yeast selection) with some yeast strains having the tendency to do "fast ferments" while others may take longer to get going. 1926:

by breaking down the cell, including the amino acids. This autolysis of the cell provides an available nitrogen source for the still-fermenting and viable yeast cells. However, this autolysis can also release sulfur-link compounds (such as the breakdown of amino acid

451:

and then to the more neutral-smelling 2,3-Butanediol. Many beer and winemakers who have a wine with too much "butteriness" will often "pitch" fresh yeast cultures into the no longer fermenting tank so that the yeast will consume the diacetyl and reduce the

1650:). These strains are tested in laboratories to determine a strain's vigor, sulfur dioxide and alcohol tolerance, production levels of acetic acid and sulfur compounds, ability to re-ferment (positive for sparkling wine but a negative attribute for sweet 1683:

and packaged for commercial use. Prior to their addition into must, these yeasts need to be re-hydrated in "starter cultures" that must be carefully monitored (particularly in regards to temperature) to ensure that the yeast cells are not killed off by

484:

acids can also be formed depending on the yeast strain. Most countries have wine laws setting the legal limit of volatile acidity, usually expressed as acetic acid, to 1200–2000 mg/L. Acetic acid can also lead to the development of the wine fault

614:

The lees left over from the secondary fermentation of sparkling wine can be seen on the bottom side of this bottle being inspected. Eventually this wine will go through riddling to collect the lees in the neck, where it will be removed prior to

1477:

with skin contact. Other winemakers may allow the wild yeasts to continue fermenting until they succumb to the toxicity of the alcohol they produce which is often between 3–5% alcohol by volume and then letting either inoculated or "ambient"

1951:

process done in the absence of oxygen, early exposure of the yeast to oxygen can be a vital component in the successful completion of that fermentation. This is because oxygen is important in the synthesis of cell "survival factors" such as

311:

In the absence of oxygen, yeast cells will take the pyruvate produced by glycolysis and reduce it into acetaldehyde which is further reduced into ethanol "recharging" the NAD+ co-enzymes that is needed for various metabolic processes of the

367:

While the production of alcohol is the most noteworthy by-product of yeast metabolism from a winemaking perspective, there are a number of other products that yeast produce that can be also influence the resulting wine. This includes

380:

needed to continue other metabolic activities. This is usually produced early in the fermentation process before the mechanisms to reduce acetaldehyde into ethanol to recharge NADH becomes the cell's primary means of maintaining

2009:

can cover the surface of a wine with a film layer that not only consumed most of the free sulfur dioxide available to protect the wine but also produces high levels of acetic acid that will contribute to volatile acidity in a

278:

were identified. The differences between the vast majority of these strains are mostly minor, though individual winemakers will develop a preference for particular strains when making certain wines or working with particular

1984:

yeasts often need a much greater exposure to oxygen in order to build up survival factors which is why many of these yeasts are often found living oxidatively as "film yeast" on the surface of wines in tanks or barrels.

1536:

ferment glucose and fructose into alcohol, they also have the potential to create other intermediates that could influence the aroma and flavor profile of the wine. Some of these intermediates could be positive, such as

109:

or neutral spirits to kill off the yeast cells. If fermentation is unintentionally stopped, such as when the yeasts become exhausted of available nutrients and the wine has not yet reached dryness, this is considered a

2220:

Once Brett is in a winery, it is very difficult to control even with strict hygiene and the discarding of barrels and equipment that has previously come into contact with "Bretty" wine. This is because many species of

1890:) and even the remnant of dead or extracted yeast cells such that the fermenting yeast can break down to mine for available nitrogen and nutrients. One historical winemaking tradition that is still practiced in some 524:

will produce higher amounts that contributes to the characterized "aldehydic" aromas of Sherries. In the presence of oxygen, yeast can convert some of the ethanol presence in the wine back into acetaldehyde creating

398:

to ferment), increased oxygen exposure and aeration as well as fermenting at higher temperatures. Glycerol production is also encouraged if most available acetaldehyde is made unavailable by binding with

166:. While additions of sulfur dioxide (often added at the crusher) may limit some of the wild yeast activities, these yeasts will usually die out once the alcohol level reaches about 15% due to the 162:. These yeasts often begin the fermentation process almost as soon as the grapes are picked when the weight of the clusters in the harvest bins begin to crush the grapes, releasing the sugar-rich 2692: 1545:-like aroma. However, as with ambient yeasts, the products of these yeasts can be very unpredictable – especially in terms of the types of flavors and aromas that these yeasts can produce. 489:

which is characterized by a "nail polish remover" smell. However, small amounts of acetic acid are actually beneficial for the yeast as they use them to synthesis lipids in the cell membrane.

1354:

that have taken residence in these places over the years, sometimes being previously introduced by inoculation of prior vintages. In this context, these wild yeasts are often referred to as

363:

If a Chardonnay has too much "buttery" diacetyl notes, winemakers may add fresh yeast to the wine to consume the diacetyl and reduce it to the more neutral-smelling fusel oil 2,3-Butanediol.

2069:

on top of the wine in the tank of barrel. Allowed to go unchecked, these yeasts can rapidly deplete the available free sulfur compounds that keeps a wine protected from oxidation and other

2100:

Growth of many unfavorable wild yeasts is generally slowed at lower cellar temperatures, so many winemakers who wish to inhibit the activities of these yeasts before the more favorable

1713:

Diammonium phosphate (or DAP) is a common additive that provides two necessary nutrients for yeast to have a healthy and sustained fermentation – nitrogen and phosphate.

1701:

added directly to the must itself which can kill up to 60% of the culture. Additionally, surviving cells exposed to cold shock tend to see an increase in hydrogen sulfide production.

1421:

of wild yeasts per berry could exist in a typical vineyard. These yeasts can be carried by air currents, birds and insects through the vineyard and even into the winery (such as by

1557:

Some winemakers favor the use of freeze-dried cultured yeast (left) and yeast nutrients (right) because of their relative predictability in beginning and completing a fermentation.

697:

Most of the benefits associated with lees contact deals with the influence on the wine of the mannoproteins released during the autolysis of the yeast cells. Composed primarily of

2093:

and shifting the pH of wine upwards to levels that make the wine prone to attack by other spoilage microbes. Commonly called "film yeast", these yeasts are distinguished from the

513: – While most of the acetaldehyde produce gets reduced to ethanol or is bound by sulfur dioxide, concentrations between 50 and 100 mg/L can remain in the wine. The 2166:), a limited amount of these compounds could be considered a positive attribute that adds to the complexity of wine. To other winemakers and with other wine styles (such as 1011:

in addition to being used in brewing and breadmaking) because of the generally reliable and positive attributes it can bring to the wine. These yeasts will usually readily

1453:, itself, is actually quite rarely found in the vineyard or on the surface freshly harvested wine grapes unless the winery frequently reintroduced winery waste (such as 2585: 1497:

wild yeasts carries both potential benefits and risk. Some winemakers feel that the use of resident/indigenous yeast helps contribute to the unique expression of

2046:. Also any yeast can have a low tolerance to nutritional deficiencies, temperature fluctuation or extremes and excessive or low sugar levels that may lead to a 690:

and other compounds that were present in the yeast cell walls and membranes. This stirring also helps avoid the development of reductive sulfur compounds like

2158:(4-EG), that can have a wine being described as smelling like a "barnyard", "wet saddle" or "band-aid". To some winemakers and with some wine styles (such as 682:

produced in many wine regions across the globe. Typically when wines are left in contact with their lees, they are regularly stirred in order to release the

247:

was tasked by the French government to study what made some wines spoil. His work, which would later lead to Pasteur being considered one of the "Fathers of

458: – Like glycerol, this is often formed early in fermentation. Usually found in concentrations of 500–1200 mg/L, it is a minor acid in the overall 1489:

Wineries that wish to cultivate an "in-house" ambient yeast strain will often recycle the leftover pomace of previous vintages as compost in the vineyard.

961:, very alcohol-tolerant and can grow in wines up to 18% v/v. Additionally this yeast can survive in extremely high sugar levels (as much as 60% w/w or 60 3004: 1878:

Many of these nutrients are available in the must and skins of the grapes themselves but sometimes are supplemented by winemakers with additions such as

2686: 759:

necessary for the style. A small amount of sugared liquid is added to individual bottles, and the yeast is allowed to convert this to more alcohol and

1180:

Another less measurable difference that are subject to more debate and questions of winemakers preference is the influence of strain selection on the

1511:

and highly regarded estates will often tout the quality of their resident "chateau" strains. To this extent, wineries will often take the leftover

2902: 252: 1671: 1113:

While some strains of yeast may influence the sensory characteristics and aromas of young wine, these differences seem to fade as the wine ages.

425:

in the must by enzymes of the yeast. More commonly found in red wines than white but only in very small amounts between 20 and 200 mg/L.

2724: 1524:

but also the potential for a stuck fermentation if the indigenous yeast strains are not vigorous enough to fully convert all the sugars.

694:

and hydrogen sulfide that can appear if the lees layer is more than 10 cm (3.9 in) thick and undisturbed for more than a week.

472:

that can make a wine taste unbalanced and overly acidic. While acetic acid is the main volatile acid produced by yeast, trace amounts of

127:

which has been favored due to its predictable and vigorous fermentation capabilities, tolerance of relatively high levels of alcohol and

709:

as the cell wall breaks down. Not only does the release of mannoproteins impart sensory changes in the wine but they can contribute to

1634:

that have been identified and plated from wineries across the world (including notable producers from well-known wine regions such as

535: – Often produced by yeast during fermentation because of a nitrogen deficiency in the must. This can be done by a reduction of 2120: 1806: 560:

are known to produce higher levels of hydrogen sulfides than other strains, particularly if the must has some nutrient deficiencies.

377: 1378:. Wineries that often solely rely on these "in-house" strains will sometimes market their wines as being the product of wild or 639:

and age. During the time that the wine spends in contact with the lees, a number of changes can impact the wine due to both the

1228:. Similarly, though potentially to a much smaller extent, other varieties could be influenced by hydrolytic enzymes working on 3042: 2997: 2668: 2622: 2568: 2545: 2500: 2459: 2412: 2351: 2290: 2263: 1947:

meaning that they can exist in both the presence and absence of oxygen. While fermentation is traditionally thought of as an

627:. During fermentation, the first significant racking which removes the bulk of dead yeast cells is often referred to as the 283:. Some of these differences include the "vigor" or speed of fermentation, temperature tolerance, the production of volatile 2928: 2582: 1799:. A deficiency of this vitamin could lead into increase hydrogen sulfide production with off-aromas in the resulting wine. 1165:

distinctive difference when compared among young wines, these differences seem to fade and become less distinctive as the

4929: 815:

text under its asexual classification though some scientific and winemaking texts may describe specific species (such as

658:

The process of leaving the wine to spend some contact with the lees has a long history in winemaking, being known to the

2933: 2022:" and aromas that can be the by-product of some "wild yeast" fermentation such as those by species within the genera of 190:

is a species of yeast that can tolerate alcohol levels of 17–20% and is often used in fortified wine production such as

30: 2104:

yeast kick in, will often chill their must, such as the practice of "cold soaking" the must during a pre-fermentation

1012: 317: 2635:

Science and Civilisation in China, Volume 6 Biology and Biological Technology, Part 5: Fermentations and Food Science

1675:

Preparing a yeast starter culture and gradually cooling the culture down to the must temperature by adding some wine

4959: 2853: 2820: 1719: 228:

French scientist Louis Pasteur discovered the connection between microscopic yeast and the process of fermentation.

763:. The lees are then ricked into the neck of the bottle, frozen, and expelled via pressure of the carbonated wine. 2990: 2943: 2833: 1252: 1090: 742: 97:. This can be achieved by dropping fermentation temperatures to the point where the yeast are inactive, sterile 4964: 4916: 3057: 2938: 2717: 927: 123: 1319: 1076: 846:. Other genera of yeast that can be involved in winemaking (either beneficially or as the cause of potential 2828: 2614: 2492: 2343: 1083: 975: 224: 2034:

can be behind some wine faults with some strains of the yeast known to produce higher than ideal levels of

1485: 619:

When yeast cells die, they sink to the bottom of the fermentation vessel where they combine with insoluble

85:, the higher the potential alcohol level of the wine if the yeast are allowed to carry out fermentation to 4393: 3276: 3142: 1965: 1766: 3072: 2179: 1944: 1750:) that serve as important growth and survival factors. Among the other nutritional needs of wine yeast: 1647: 1575:

Have a high sulfur dioxide tolerance but low production of sulfur compounds such as hydrogen sulfide or

1325: 1313: 1260: 1069: 1062: 659: 186: 1469:"knock out" these yeasts with doses of sulfur dioxide, most often at the crusher before the grapes are 502: 992: 4802: 3577: 3147: 3062: 1879: 1814: 1619: 1418: 1389: 1384: 1097: 817: 403:

molecules in the wine, but it would take a substantial amount of sulfur dioxide addition (far beyond

102: 1595:

Produce minimum foaming during fermentation which may create difficulties for cap management during

4969: 2810: 2710: 1923: 1824: 1508: 1445: 941: 683: 647:

conditions that can develop if the lees are not aerated or stirred (a process that the French call

321: 78: 270:(as opposed to circular) shape of the cells. Throughout the 20th century, more than 700 different 3597: 3067: 3013: 2848: 2097:

sherry yeast that are usually welcomed by winemakers in producing the delicate fino-style wines.

2090: 2047: 1908:

of other wines into a newly fermenting batch of wine as an additional food source for the yeast.

1651: 1643: 1241: 790: 726: 714: 710: 111: 4872: 4867: 4722: 4483: 1341:

Fruit flies are a common vector that transports ambient or "wild" yeast strains within wineries.

1307:, though the commercial flor yeast available for inoculation is often from different species of 1663:") and tolerance for nutritional deficiencies in a must that may lead to a stuck fermentation. 4712: 4627: 4610: 4160: 3947: 3887: 3077: 2802: 2749: 2664: 2618: 2564: 2541: 2496: 2455: 2408: 2347: 2286: 2259: 2105: 1935:

that can contribute to a "stinky fermentation" or later development into various wine faults.

1596: 1538: 1300: 1209: 1141: 957: 946: 831:. Unless otherwise noted, this article will commonly refer to the asexual form of wine yeast. 701:

and proteins, with some glucose, mannoproteins are often bound in the cell wall of yeast with

575: 571: 1968:

of the yeast cell membrane which becomes critical as the yeast becomes exposed to increasing

1569:

Completely utilize all fermentable sugars with a predictable sugar-to-alcohol conversion rate

4531: 4505: 4260: 4140: 3095: 2967: 2074: 1969: 1792: 1576: 1521: 1433: 1174: 867: 722: 675: 640: 532: 469: 373: 288: 271: 155: 98: 82: 2130:

on the wine, in limited amounts, as added complexity, many winemakers view the presence of

666:

in the 2nd century BC. Today the practice is widely associated with any red wines that are

610: 4270: 4244: 4175: 4150: 4115: 4064: 4004: 3587: 3194: 3052: 3047: 2948: 2769: 2696: 2589: 2163: 2155: 2086: 1905: 1639: 1635: 1504: 1474: 1337: 1185: 934: 913:), usually the most common "wild yeast" found in the vineyard. Some species are known as " 794: 687: 549: 459: 331: 280: 233: 90: 407:) to prolong glycerol production beyond just these very nascent stages of fermentation. 307: 38:

it releases alcohol and carbon dioxide (seen here as the foaming bubbles) as byproducts.

4585: 4418: 4383: 4195: 4054: 2843: 2838: 2787: 2741: 2193:

can usually ferment a wine up to 10–11% alcohol levels before they die out. Sometimes

2112:, will not be inhibited and may even thrive during an extended period of cold soaking. 1802: 1566:

Quickly begin fermentation, out-competing other "wild yeasts" for nutrients in the must

1470: 1346:

equipment, transport bins, the surface winemaking equipment and as part of the natural

1233: 1217: 760: 752: 663: 594: 481: 432: 144: 128: 86: 74: 1862: 556:

that contribute to off aromas and wine faults. Some commercial yeast strains, such as

4953: 4807: 4787: 4667: 4373: 4220: 4120: 4110: 4014: 3922: 3902: 3810: 3592: 2868: 2777: 2210: 2206: 2151: 2126: 2082: 1828: 1762: 1758: 1600: 1264: 969: 918: 897: 879: 855: 807: 730: 636: 486: 455: 418: 244: 208: 176: 66: 34:

The process of fermentation at work on Pinot noir. As yeast consume the sugar in the

2256:"From Vines to Wines: The Complete Guide to Growing Grapes and Making Your Own Wine" 2175: 2014:

Either directly or indirectly, wine yeast can be a culprit behind a wide variety of

1709: 1572:

Have an alcohol-tolerance up to 15% or even higher depending on the winemaking style

1422: 143:

is rarely the only yeast species involved in a fermentation. Grapes brought in from

4943: 4519: 4155: 4074: 4059: 3942: 3376: 3266: 3261: 3152: 3137: 2171: 2108:

at temperatures between 4–15 °C (39–59 °F). Though some species, such as

2039: 2019: 1931:) which can combine with other molecules and react with alcohol to create volatile 1891: 1680: 1660: 1607: 1590: 1553: 1225: 1221: 914: 885: 839: 567: 563: 518: 510: 473: 339: 292: 248: 94: 4742: 4717: 4565: 4433: 4275: 3291: 2473: 4672: 4408: 4353: 4255: 4225: 3992: 3755: 3735: 3705: 3642: 3512: 3477: 3452: 3416: 3366: 3336: 3326: 3251: 3241: 3231: 3226: 3100: 2972: 2923: 2886: 2205:

strain for nutrients and even inhibit it due to the high levels of acetic acid,

2089:

that may also be produced) with the side-effect of substantially decreasing the

2035: 2015: 1786: 1727: 1693: 1611: 1586: 1454: 1208:

that are more prevalent in particular strains. Other aromatic varieties such as

1189: 1109: 1008: 922: 891: 847: 822: 812: 756: 718: 706: 702: 624: 498: 477: 465: 386: 191: 136: 2073:

attack. The presence of these yeasts is often identified by elevated levels of

431: – Formed by the decomposition of amino acids by the yeast. This includes 4877: 4812: 4757: 4697: 4642: 4590: 4358: 4079: 4049: 3316: 3306: 3164: 2876: 2792: 2782: 2733: 2159: 1957: 1953: 1948: 1747: 1685: 1414: 1284: 1276: 1229: 1197: 1157: 1133: 980: 802: 755:

requires a second fermentation to occur in the bottle in order to produce the

691: 679: 667: 548:. In the presence of alcohol, hydrogen sulfide can react with ethanol to form 440: 428: 343: 335: 213: 171: 135:

between 2.8 and 4. Despite its widespread use which often includes deliberate

118: 58: 2407:

Second Edition pp. 3–28 Springer Science and Business Media, New York (2010)

4862: 4827: 4797: 4657: 4368: 4130: 4024: 3987: 3765: 3695: 3572: 3552: 3487: 3406: 3281: 3120: 2754: 2070: 1851: 1772: 1754: 1735: 1427: 1398: 1166: 1137: 1129: 1122: 1043: 1023: 905: 782: 777: 748: 620: 553: 493: 404: 400: 359: 237: 195: 149: 17: 4652: 4428: 3627: 2225:

can use a wide variety of carbon sources in wine and grape must, including

212:

whose presence in a wine may be viewed by different winemakers as either a

89:. Sometimes winemakers will stop fermentation early in order to leave some 771: 4887: 4882: 4847: 4837: 4747: 4682: 4575: 4514: 4493: 4478: 4328: 4029: 3892: 3835: 3815: 3715: 3690: 3657: 3542: 3401: 3396: 3386: 3301: 3286: 3271: 2699:. International Journal of Food Microbiology 86 (2003). pp. 169–180. 2167: 1928: 1919: 1915: 1839: 1820: 1782: 1582: 1425:). The most common wild yeasts found in the vineyard are from the genera 1280: 1272: 1213: 1201: 1181: 1007:(sugar mold) is favored for winemaking (for both grapes as well as other 798: 671: 590: 526: 436: 414: 369: 167: 2142:) in their wineries as a negative influence that needs to be controlled. 1046:) which is usually present in small amount in wines as residual sugars. 4857: 4777: 4762: 4707: 4620: 4600: 4555: 4550: 4540: 4524: 4498: 4463: 4423: 4413: 4363: 4308: 4185: 4170: 4089: 4069: 4034: 3967: 3567: 3527: 3507: 3472: 3431: 3356: 3115: 3110: 3105: 2953: 2759: 2226: 2066: 1961: 1896: 1835: 1743: 1739: 1731: 1723: 1615: 1516: 1515:

and lees from winemaking and return them to the vineyard to be used as

1499: 1237: 1205: 1039: 1031: 1027: 1019: 1015: 950: 843: 827: 698: 598: 586: 545: 540: 536: 448: 422: 348: 327: 267: 70: 2982: 4897: 4822: 4752: 4732: 4727: 4687: 4398: 4323: 4318: 4303: 4280: 4265: 4240: 4230: 4200: 4190: 4180: 4044: 4009: 3999: 3957: 3952: 3932: 3912: 3865: 3840: 3830: 3680: 3547: 3537: 3517: 3492: 3421: 3296: 2918: 2881: 2054: 2043: 1932: 1901: 1843: 1778: 1512: 1458: 1439: 1413:

genera of yeasts that are present in the vineyard, on the surface of

1288: 1193: 1126: 873: 801:(or "imperfect" form) while others may be classified by their sexual 582: 521: 444: 284: 159: 106: 1396:草麴) wild yeast with rice and various herbs, including the poisonous 797:. Therefore, some winemaking yeasts are classified by their asexual 1993: 999:

as seen under a Differential Interference Contrast (DIC) microscope

326:

The primary role of yeast is to convert the sugars present (namely

4632: 4438: 4388: 4343: 4333: 4298: 4210: 4084: 4019: 3962: 3825: 3800: 3795: 3780: 3760: 3745: 3725: 3710: 3617: 3607: 3602: 3562: 3557: 3532: 3502: 3467: 3462: 3426: 3411: 3346: 3216: 3130: 2648:

Nan-fang ts'ao-mu chuang: a fourth century flora of Southeast Asia

2512: 2119: 1992: 1861: 1810: 1796: 1708: 1670: 1552: 1484: 1347: 1336: 1220:

may also be influenced by yeast strains containing high levels of

1153: 1108: 991: 770: 644: 609: 492: 382: 358: 306: 223: 206:

sugar levels. Another common yeast involved in wine production is

199: 62: 54: 51: 29: 1874:

from a previous fermentation is added to a newly fermenting wine.

871:(Teleomorphs for different species from several genera including 725:

of the wine as well as decrease the perception of bitterness and

4545: 4205: 3790: 3770: 3700: 3652: 3647: 3582: 3522: 3125: 2561:

Winery Technology & Operations A Handbook for Small Wineries

2094: 1866:

One traditional way of providing nutrients for the yeast is the

1855: 1847: 1655: 1542: 1296: 1292: 1245: 1149: 962: 514: 372:

which is produced when an intermediate of the glycolysis cycle (

203: 163: 47: 35: 4461: 3863: 3192: 3024: 2986: 2706: 1132:. This sequencing helped confirm the nearly century of work by 4772: 4378: 3977: 3637: 834:

The most common yeast generally associated with winemaking is

566: – Along with acetaldehyde, this compound can react with 240:

of the word "yeast" itself which essentially means "to boil".

2702: 1775: – important for the uptake and utilization of phosphate 651:). The length of time that a wine spends on its lees (called 1769:

which the cell uses for transferring energy for metabolism).

1279:

well, allowing the dead yeast cells to be removed easily by

643:(or self-metabolize) of the dead yeast cells as well as the 334:), the cell will continue some metabolic functions (such as 147:

are usually teeming with a variety of "wild yeast" from the

1417:

and of the grapes themselves. Anywhere from 160 to 100,000

132: 262:

was first identified in late 19th century enology text as

2174:), the presence of any Brett will be considered a fault. 1914:

can assimilate nitrogen from both inorganic (ammonia and

1809:(NAD+), a co-enzyme that is important in maintaining the 1654:), development of surface film on the wine (positive for 216:

or in limited quantities as an added note of complexity.

2446: 2444: 2442: 2440: 2197:

already present in a wine that has been inoculated with

2150:(or "Brett") produces very distinctive aroma compounds, 655:) will depend on the winemaking style and type of wine. 2683:

P. Romano, C. Fiore, M. Paraggio, M. Caruso, A. Capece

2540:

pp. 97–114 Kluwer Academic Publishers, New York (1999)

2454:

pp. 281–90 Kluwer Academic Publishers, New York (1999)

2438: 2436: 2434: 2432: 2430: 2428: 2426: 2424: 2422: 2420: 1765:(an important component of the cell membrane) and ATP ( 1409:

Another use of the term "wild yeast" refers to the non-

1192:. It is believed that these wines can be influenced by 1160:. Today there are several hundred different strains of 338:) but will rely on other pathways such as reduction of 2399: 2397: 2395: 2393: 2391: 2389: 2387: 2385: 2383: 2381: 2379: 1922:). As yeast cells die, enzymes within the cells begin 1679:

Pure culture yeasts that are grown in a lab are often

793:

includes classification of yeast species depending on

351:

leaving behind a negligible amount of residual sugar.

4927: 2687:

Function of yeast species and strains in wine flavour

2663:

p. 80 The Wine Appreciation Guild Third Edition 2009

2377: 2375: 2373: 2371: 2369: 2367: 2365: 2363: 2361: 2359: 1255:

some winemakers select strains (such as one known as

2605: 2603: 2478:

Online Etymology Dictionary Accessed: May 31st, 2012

2334: 2332: 2330: 2328: 2326: 2324: 2322: 2320: 2318: 811:(or "Brett") that is usually referenced in wine and 4291: 4100: 3877: 3668: 3442: 3206: 3088: 3035: 2911: 2895: 2867: 2819: 2801: 2768: 2740: 2316: 2314: 2312: 2310: 2308: 2306: 2304: 2302: 2300: 2298: 623:, grape seeds, skin and pulp fragments to form the 1388:has the earliest description of winemaking using " 1870:method where the leftover grape skins and pomace 1813:balance of the cell as well as in the process of 1795: – involved in the metabolism of sugars and 805:(or "perfect" form). A common example of this is 775:Film yeast on the surface of wine in a barrel of 46:is the most important element that distinguishes 131:as well as its ability to thrive in normal wine 1918:) and organic forms (amino acids, particularly 1291:production, the surface film of yeast known as 1105:Influences of different strains on fermentation 578:) that can enhance the color of some red wines. 2124:While some wine regions view the influence of 342:into ethanol (fermentation) to "recharge" the 243:In the mid-19th century, the French scientist 27:Yeasts used for alcoholic fermentation of wine 2998: 2718: 2633:Joseph Needham and Huang Hsing-Tsung (2000), 2563:pp. 67–74 The Wine Appreciation Guild (1996) 2536:B. Zoecklein, K. Fugelsang, B. Gump, F. Nury 2450:B. Zoecklein, K. Fugelsang, B. Gump, F. Nury 1882:(DAP), freeze-dried micro-nutrients (such as 1350:of a winery. Very often these are strains of 1184:flavors of certainly grape varieties such as 291:) and other compounds that may influence the 8: 2077:, particularly acetic acid. Some strains of 1026:and metabolize glucose, sucrose, raffinose, 979:found in moist cellars that can contaminate 973:, particularly the "black yeast" species of 435:which is formed by yeast that are consuming 332:and sometimes even in the presence of oxygen 4466:and infused distilled drinks by ingredients 2532: 2530: 1057:that are involved with winemaking include: 385:balance. As glycerol contributes increased 4458: 3874: 3860: 3203: 3189: 3032: 3021: 3005: 2991: 2983: 2725: 2711: 2703: 965:) and is very resistant to sulfur dioxide. 501:are caused by special yeast native to the 1303:sherries comes from different strains of 945:, the only wine yeast that reproduced by 795:the presence or absence of a sexual phase 355:Other compounds in wine produced by yeast 2186:between tanks and even nearby wineries. 2081:will metabolize acetic acid (as well as 1667:Re-hydrating freeze dried yeast cultures 1449:being the most dominant species by far. 4934: 2903:Clarification and stabilization of wine 2555: 2553: 2238: 925:that can kill off sensitive strains of 574:to create a more stable color pigment ( 468: – Considered a main component of 93:and sweetness in the wine such as with 2650:, The Chinese University Press, p. 59. 2277: 2275: 2273: 2271: 2042:and volatile sulfur compounds such as 1697:synthesis of needed survival factors. 1295:used to make the distinctive style of 117:The most common yeast associated with 2637:, Cambridge University Press, p. 183. 2250: 2248: 2246: 2244: 2242: 2018:. These can include the presence of " 1805: – involved in the synthesis of 1781: – involved in the synthesis of 1581:Produce a minimum amount of residual 1503:in the wine. In wine regions such as 949:whereas most wine yeast reproduce by 497:The distinctive "aldehydic" notes of 7: 1527:It is virtually inevitable that non- 1333:Wild yeasts and natural fermentation 917:" that produce inhibitory levels of 410:Other by-products of yeast include: 258:The yeast species commonly known as 1757: – used for the production of 635:that come as the wine continues to 2258:pp. 133–36 Storey Publishing 1999 1493:The use of both "ambient" and non- 767:Types of yeasts used in winemaking 180:species take over. In addition to 25: 1964:are important in maintaining the 1807:Nicotinamide adenine dinucleotide 1599:or cause bungs to pop out during 1482:strains finish the fermentation. 1053:, other species within the genus 4937: 2966: 2491:Third Edition pgs 267 & 508 174:while the more alcohol tolerant 101:the wine to remove the yeast or 2285:pp. 67–73 DBQA Publishing 2005 2283:"Understanding Wine Technology" 1705:Nutritional needs of wine yeast 517:yeast strains that produce the 376:) is reduced to "recharge" the 3043:History of alcoholic beverages 2611:"The Oxford Companion to Wine" 2489:"The Oxford Companion to Wine" 2340:"The Oxford Companion to Wine" 1900:method of adding the leftover 1275:strain 505) that are known to 631:as opposed to the less coarse 447:" aroma, reducing it first to 253:Embden–Meyerhof–Parnas pathway 170:of alcohol on the yeast cells 1: 2929:Glossary of viticulture terms 2030:. Even the common wine yeast 1121:was the first single-celled, 2934:Glossary of winemaking terms 2538:Wine Analysis and Production 2452:Wine Analysis and Production 1989:Wine faults related to yeast 1259:named after the town in the 1125:organism to have its entire 705:aroma compounds that become 318:Fermentation (biochemistry) 4986: 2854:Yeast assimilable nitrogen 2661:The University Wine Course 2583:Diacetyl: Homebrew Science 2136:Brettanomyces bruxellensis 1858:for healthy cell function. 1827:molecules that facilitate 1823: – involved with the 1720:yeast assimilable nitrogen 1038:cannot ferment or utilize 740: 439:, the compound that gives 315: 264:Saccharomyces ellipsoideus 4912: 4471: 4457: 3873: 3859: 3202: 3188: 3031: 3020: 2962: 2944:History of the wine press 2834:Sparkling wine production 2613:Third Edition pp. 398–99 2403:K. Fugelsang, C. Edwards 2342:Third Edition pp. 778–80 2189:As a fermentation yeast, 1253:sparkling wine production 1140:in identifying different 1091:Saccharomyces pastorianus 825:sexual classification of 743:Sparkling wine production 4315:Coconut and other palms 3687:Coconut and other palms 3058:History of Bordeaux wine 2939:Wine tasting descriptors 2199:Saccharomyces cerevisiae 2032:Saccharomyces cerevisiae 1912:Saccharomyces cerevisiae 1632:Saccharomyces cerevisiae 1630:) yeasts are strains of 1451:Saccharomyces cerevisiae 1352:Saccharomyces cerevisiae 1320:Saccharomyces fermentati 1305:Saccharomyces cerevisiae 1224:enzymes that can modify 1146:Saccharomyces cerevisiae 1119:Saccharomyces cerevisiae 1077:Saccharomyces fermentati 1051:Saccharomyces cerevisiae 997:Saccharomyces cerevisiae 928:Saccharomyces cerevisiae 836:Saccharomyces cerevisiae 572:contact with grape skins 276:Saccharomyces cerevisiae 260:Saccharomyces cerevisiae 124:Saccharomyces cerevisiae 3143:Drinking establishments 2829:Malolactic fermentation 2615:Oxford University Press 2493:Oxford University Press 2344:Oxford University Press 1084:Saccharomyces paradoxus 976:Aureobasidium pullulans 77:through the process of 4350:Sugarcane or molasses 3787:Sugarcane or molasses 3026:History and production 2143: 2011: 1966:selective permeability 1875: 1767:Adenosine triphosphate 1714: 1676: 1614:compaction that makes 1558: 1490: 1342: 1114: 1000: 838:which is also used in 786: 737:Secondary fermentation 616: 506: 417: – Caused by the 364: 313: 229: 194:and varieties such as 39: 3073:History of Rioja wine 2213:that many strains of 2201:will out compete the 2123: 1996: 1945:facultative anaerobes 1865: 1712: 1674: 1556: 1541:, which can impart a 1488: 1464:Unlike the "ambient" 1461:) into the vineyard. 1380:natural fermentations 1340: 1326:Saccharomyces bayanus 1314:Saccharomyces beticus 1261:Champagne wine region 1204:-linked compounds by 1112: 1070:Saccharomyces beticus 1063:Saccharomyces bayanus 995: 774: 613: 496: 362: 310: 227: 187:Saccharomyces bayanus 139:from cultured stock, 33: 4784:Mammee apple flower 3063:History of Champagne 1880:diammonium phosphate 1815:ethanol fermentation 1620:fining and filtering 1606:Have high levels of 1419:colony forming units 1385:Nanfang Caomu Zhuang 1366:yeast as opposed to 1269:California Champagne 1240:derivatives such as 1098:Saccharomyces uvarum 818:Dekkera bruxellensis 83:sugars in the grapes 4834:Sugarcane/molasses 3277:Moscatel de Setúbal 3170:Yeast in winemaking 3158:Yeast in winemaking 3014:Alcoholic beverages 2859:Yeast in winemaking 2811:Carbonic maceration 2646:Li Hui-Lin (1979), 2559:Dr. Yair Margalit, 2182:in the transfer of 2057:several species of 2053:In the presence of 1825:secondary messenger 1622:of the wine easier. 1446:Kloeckera apiculata 942:Schizosaccharomyces 717:, help enhance the 443:and other wines a " 322:Fermentation (wine) 287:compounds (such as 44:yeast in winemaking 3068:History of Chianti 3036:History of alcohol 2849:Traditional method 2695:2015-06-06 at the 2588:2010-02-02 at the 2144: 2091:titratable acidity 2048:stuck fermentation 2012: 1939:The role of oxygen 1876: 1789:and nucleic acids. 1715: 1677: 1656:some Sherry styles 1652:late-harvest wines 1559: 1491: 1343: 1115: 1001: 787: 747:The production of 617: 507: 365: 347:the unfermentable 314: 303:Role in winemaking 230: 202:harvested at high 112:stuck fermentation 69:of the fruit into 40: 4960:Food microbiology 4925: 4924: 4908: 4907: 4611:Chocolate liqueur 4572:Blackthorn shrub 4453: 4452: 4449: 4448: 4405:Various starches 4161:Tennessee whiskey 3855: 3854: 3851: 3850: 3822:Various starches 3184: 3183: 3180: 3179: 3078:History of Sherry 2980: 2979: 2750:Late harvest wine 2609:J. Robinson (ed) 2487:J. Robinson (ed) 2405:Wine Microbiology 2338:J. Robinson (ed) 1834:Trace amounts of 1730:or YAN) but also 1443:with the species 1399:Gelsemium elegans 1148:that are used in 958:Zygosaccharomyces 715:protein stability 674:, sparkling wine 662:and described by 576:pyranoanthocyanin 503:Jerez wine region 59:absence of oxygen 16:(Redirected from 4977: 4942: 4941: 4940: 4933: 4532:Herbs de Majorca 4506:Anisado Mallorca 4459: 4261:American whiskey 4252:Multiple grains 4141:Buckwheat whisky 3875: 3861: 3634:Multiple grains 3204: 3195:Fermented drinks 3190: 3096:Alcohol industry 3033: 3022: 3007: 3000: 2993: 2984: 2971: 2970: 2727: 2720: 2713: 2704: 2671: 2657: 2651: 2644: 2638: 2631: 2625: 2607: 2598: 2579:Brewing Science 2577: 2571: 2557: 2548: 2534: 2525: 2524: 2522: 2520: 2509: 2503: 2485: 2479: 2468: 2462: 2448: 2415: 2401: 2354: 2336: 2293: 2279: 2266: 2252: 2134:species such as 2075:volatile acidity 1997:Film yeast like 1970:osmotic pressure 1793:Pantothenic acid 1577:dimethyl sulfide 1549:Inoculated yeast 1522:volatile acidity 1271:, also known as 1196:produced by the 1175:hydrogen sulfide 1003:The yeast genus 785:region of France 668:barrel fermented 550:ethyl mercaptans 533:Hydrogen sulfide 470:volatile acidity 374:dihydroxyacetone 289:hydrogen sulfide 232:For most of the 21: 4985: 4984: 4980: 4979: 4978: 4976: 4975: 4974: 4965:Saccharomycetes 4950: 4949: 4948: 4938: 4936: 4928: 4926: 4921: 4904: 4873:Crème de cassis 4868:Crème de banane 4854:Various fruits 4723:Crème de menthe 4484:Crème de Noyaux 4467: 4445: 4287: 4271:Canadian whisky 4245:Kaoliang liquor 4151:Bourbon whiskey 4116:Japanese whisky 4096: 4065:Marillenschnaps 4041:Various fruits 3869: 3847: 3676:Agave americana 3664: 3438: 3393:Various fruits 3198: 3176: 3084: 3053:History of wine 3048:History of beer 3027: 3016: 3011: 2981: 2976: 2973:Wine portal 2965: 2958: 2949:History of wine 2907: 2891: 2863: 2815: 2797: 2778:Deacidification 2764: 2736: 2731: 2697:Wayback Machine 2680: 2675: 2674: 2658: 2654: 2645: 2641: 2632: 2628: 2608: 2601: 2590:Wayback Machine 2578: 2574: 2558: 2551: 2535: 2528: 2518: 2516: 2511: 2510: 2506: 2486: 2482: 2470:Douglas Harper 2469: 2465: 2449: 2418: 2402: 2357: 2337: 2296: 2280: 2269: 2253: 2240: 2235: 2156:4-Ethylguaiacol 2146:The wine yeast 2118: 2087:isoamyl acetate 1991: 1941: 1894:regions is the 1707: 1669: 1626:Inoculated (or 1551: 1382:. The (c. 304) 1335: 1186:Sauvignon blanc 1173:compounds like 1107: 1049:In addition to 990: 935:Saccharomycodes 769: 753:sparkling wines 745: 739: 688:polysaccharides 608: 570:extracted from 460:acidity of wine 357: 324: 305: 281:grape varieties 234:history of wine 222: 91:residual sugars 28: 23: 22: 15: 12: 11: 5: 4983: 4981: 4973: 4972: 4967: 4962: 4952: 4951: 4947: 4946: 4923: 4922: 4920: 4919: 4913: 4910: 4909: 4906: 4905: 4903: 4902: 4901: 4900: 4892: 4891: 4890: 4885: 4880: 4875: 4870: 4865: 4860: 4852: 4851: 4850: 4842: 4841: 4840: 4832: 4831: 4830: 4825: 4817: 4816: 4815: 4810: 4805: 4800: 4792: 4791: 4790: 4782: 4781: 4780: 4775: 4767: 4766: 4765: 4760: 4755: 4750: 4745: 4737: 4736: 4735: 4730: 4725: 4720: 4715: 4710: 4702: 4701: 4700: 4692: 4691: 4690: 4685: 4677: 4676: 4675: 4670: 4662: 4661: 4660: 4655: 4647: 4646: 4645: 4637: 4636: 4635: 4625: 4624: 4623: 4615: 4614: 4613: 4605: 4604: 4603: 4597:Chili peppers 4595: 4594: 4593: 4588: 4586:Cherry Heering 4580: 4579: 4578: 4570: 4569: 4568: 4560: 4559: 4558: 4553: 4548: 4543: 4538: 4537: 4536: 4535: 4534: 4522: 4517: 4512: 4511: 4510: 4509: 4508: 4496: 4488: 4487: 4486: 4481: 4472: 4469: 4468: 4462: 4455: 4454: 4451: 4450: 4447: 4446: 4444: 4443: 4442: 4441: 4436: 4431: 4426: 4421: 4416: 4411: 4403: 4402: 4401: 4396: 4391: 4386: 4384:Seco Herrerano 4381: 4376: 4371: 4366: 4361: 4356: 4348: 4347: 4346: 4338: 4337: 4336: 4331: 4326: 4321: 4313: 4312: 4311: 4306: 4295: 4293: 4289: 4288: 4286: 4285: 4284: 4283: 4278: 4273: 4268: 4263: 4258: 4250: 4249: 4248: 4235: 4234: 4233: 4228: 4223: 4215: 4214: 4213: 4208: 4203: 4198: 4196:Mixiang Baijiu 4193: 4188: 4183: 4178: 4173: 4165: 4164: 4163: 4158: 4153: 4145: 4144: 4143: 4135: 4134: 4133: 4125: 4124: 4123: 4118: 4113: 4104: 4102: 4098: 4097: 4095: 4094: 4093: 4092: 4087: 4082: 4077: 4072: 4067: 4062: 4057: 4052: 4047: 4039: 4038: 4037: 4032: 4027: 4022: 4017: 4012: 4007: 3997: 3996: 3995: 3990: 3982: 3981: 3980: 3972: 3971: 3970: 3965: 3960: 3955: 3950: 3945: 3937: 3936: 3935: 3927: 3926: 3925: 3917: 3916: 3915: 3907: 3906: 3905: 3897: 3896: 3895: 3890: 3881: 3879: 3871: 3870: 3868:by ingredients 3864: 3857: 3856: 3853: 3852: 3849: 3848: 3846: 3845: 3844: 3843: 3838: 3833: 3828: 3820: 3819: 3818: 3813: 3805: 3804: 3803: 3798: 3793: 3785: 3784: 3783: 3775: 3774: 3773: 3768: 3763: 3758: 3750: 3749: 3748: 3740: 3739: 3738: 3730: 3729: 3728: 3720: 3719: 3718: 3713: 3708: 3703: 3698: 3693: 3685: 3684: 3683: 3672: 3670: 3666: 3665: 3663: 3662: 3661: 3660: 3655: 3650: 3645: 3640: 3632: 3631: 3630: 3622: 3621: 3620: 3612: 3611: 3610: 3605: 3600: 3595: 3590: 3585: 3580: 3575: 3570: 3565: 3560: 3555: 3550: 3545: 3540: 3535: 3530: 3525: 3520: 3515: 3510: 3505: 3497: 3496: 3495: 3490: 3482: 3481: 3480: 3475: 3470: 3465: 3457: 3456: 3455: 3446: 3444: 3440: 3439: 3437: 3436: 3435: 3434: 3429: 3424: 3419: 3414: 3409: 3404: 3399: 3391: 3390: 3389: 3381: 3380: 3379: 3371: 3370: 3369: 3361: 3360: 3359: 3351: 3350: 3349: 3341: 3340: 3339: 3331: 3330: 3329: 3321: 3320: 3319: 3311: 3310: 3309: 3304: 3299: 3294: 3289: 3284: 3279: 3274: 3269: 3264: 3256: 3255: 3254: 3246: 3245: 3244: 3236: 3235: 3234: 3229: 3221: 3220: 3219: 3210: 3208: 3200: 3199: 3197:by ingredients 3193: 3186: 3185: 3182: 3181: 3178: 3177: 3175: 3174: 3173: 3172: 3162: 3161: 3160: 3155: 3145: 3140: 3135: 3134: 3133: 3128: 3123: 3118: 3113: 3103: 3098: 3092: 3090: 3086: 3085: 3083: 3082: 3081: 3080: 3075: 3070: 3065: 3060: 3050: 3045: 3039: 3037: 3029: 3028: 3025: 3018: 3017: 3012: 3010: 3009: 3002: 2995: 2987: 2978: 2977: 2963: 2960: 2959: 2957: 2956: 2951: 2946: 2941: 2936: 2931: 2926: 2921: 2915: 2913: 2909: 2908: 2906: 2905: 2899: 2897: 2893: 2892: 2890: 2889: 2884: 2879: 2873: 2871: 2865: 2864: 2862: 2861: 2856: 2851: 2846: 2841: 2839:Sugars in wine 2836: 2831: 2825: 2823: 2817: 2816: 2814: 2813: 2807: 2805: 2799: 2798: 2796: 2795: 2790: 2788:Chaptalization 2785: 2780: 2774: 2772: 2766: 2765: 2763: 2762: 2757: 2752: 2746: 2744: 2738: 2737: 2732: 2730: 2729: 2722: 2715: 2707: 2701: 2700: 2679: 2678:External links 2676: 2673: 2672: 2652: 2639: 2626: 2599: 2597:November, 2002 2572: 2549: 2526: 2504: 2480: 2463: 2416: 2355: 2294: 2267: 2237: 2236: 2234: 2231: 2117: 2114: 1990: 1987: 1940: 1937: 1860: 1859: 1832: 1818: 1803:Nicotinic acid 1800: 1790: 1776: 1770: 1706: 1703: 1668: 1665: 1648:Barossa Valley 1624: 1623: 1604: 1593: 1579: 1573: 1570: 1567: 1550: 1547: 1473:or allowed to 1376:cultured yeast 1334: 1331: 1234:norisoprenoids 1210:Gewürztraminer 1106: 1103: 1102: 1101: 1094: 1087: 1080: 1073: 1066: 989: 986: 985: 984: 966: 954: 938: 931: 902: 864: 768: 765: 761:carbon dioxide 741:Main article: 738: 735: 664:Cato the Elder 660:Ancient Romans 607: 604: 603: 602: 579: 561: 558:Montrachet 522 530: 491: 490: 463: 453: 433:2,3-Butanediol 426: 356: 353: 304: 301: 221: 218: 129:sulfur dioxide 75:carbon dioxide 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 4982: 4971: 4968: 4966: 4963: 4961: 4958: 4957: 4955: 4945: 4935: 4931: 4918: 4915: 4914: 4911: 4899: 4896: 4895: 4893: 4889: 4886: 4884: 4881: 4879: 4876: 4874: 4871: 4869: 4866: 4864: 4861: 4859: 4856: 4855: 4853: 4849: 4846: 4845: 4843: 4839: 4836: 4835: 4833: 4829: 4826: 4824: 4821: 4820: 4818: 4814: 4811: 4809: 4808:Grand Marnier 4806: 4804: 4801: 4799: 4796: 4795: 4793: 4789: 4786: 4785: 4783: 4779: 4776: 4774: 4771: 4770: 4768: 4764: 4761: 4759: 4756: 4754: 4751: 4749: 4746: 4744: 4741: 4740: 4738: 4734: 4731: 4729: 4726: 4724: 4721: 4719: 4716: 4714: 4711: 4709: 4706: 4705: 4703: 4699: 4696: 4695: 4693: 4689: 4686: 4684: 4681: 4680: 4678: 4674: 4671: 4669: 4668:Cream liqueur 4666: 4665: 4663: 4659: 4656: 4654: 4651: 4650: 4648: 4644: 4641: 4640: 4638: 4634: 4631: 4630: 4629: 4626: 4622: 4619: 4618: 4616: 4612: 4609: 4608: 4606: 4602: 4599: 4598: 4596: 4592: 4589: 4587: 4584: 4583: 4581: 4577: 4574: 4573: 4571: 4567: 4564: 4563: 4561: 4557: 4554: 4552: 4549: 4547: 4544: 4542: 4539: 4533: 4530: 4529: 4528: 4527: 4526: 4523: 4521: 4518: 4516: 4513: 4507: 4504: 4503: 4502: 4501: 4500: 4497: 4495: 4492: 4491: 4489: 4485: 4482: 4480: 4477: 4476: 4474: 4473: 4470: 4465: 4460: 4456: 4440: 4437: 4435: 4432: 4430: 4427: 4425: 4422: 4420: 4417: 4415: 4412: 4410: 4407: 4406: 4404: 4400: 4397: 4395: 4392: 4390: 4387: 4385: 4382: 4380: 4377: 4375: 4372: 4370: 4367: 4365: 4362: 4360: 4357: 4355: 4352: 4351: 4349: 4345: 4342: 4341: 4339: 4335: 4332: 4330: 4327: 4325: 4322: 4320: 4317: 4316: 4314: 4310: 4307: 4305: 4302: 4301: 4300: 4297: 4296: 4294: 4290: 4282: 4279: 4277: 4274: 4272: 4269: 4267: 4264: 4262: 4259: 4257: 4254: 4253: 4251: 4246: 4242: 4239: 4238: 4236: 4232: 4229: 4227: 4224: 4222: 4219: 4218: 4216: 4212: 4209: 4207: 4204: 4202: 4199: 4197: 4194: 4192: 4189: 4187: 4184: 4182: 4179: 4177: 4174: 4172: 4169: 4168: 4166: 4162: 4159: 4157: 4154: 4152: 4149: 4148: 4146: 4142: 4139: 4138: 4136: 4132: 4129: 4128: 4126: 4122: 4121:Scotch whisky 4119: 4117: 4114: 4112: 4111:Irish whiskey 4109: 4108: 4106: 4105: 4103: 4099: 4091: 4088: 4086: 4083: 4081: 4078: 4076: 4073: 4071: 4068: 4066: 4063: 4061: 4058: 4056: 4053: 4051: 4048: 4046: 4043: 4042: 4040: 4036: 4033: 4031: 4028: 4026: 4023: 4021: 4018: 4016: 4013: 4011: 4008: 4006: 4003: 4002: 4001: 3998: 3994: 3991: 3989: 3986: 3985: 3983: 3979: 3976: 3975: 3973: 3969: 3966: 3964: 3961: 3959: 3956: 3954: 3951: 3949: 3946: 3944: 3941: 3940: 3938: 3934: 3931: 3930: 3928: 3924: 3921: 3920: 3918: 3914: 3911: 3910: 3908: 3904: 3901: 3900: 3899:Cashew apple 3898: 3894: 3891: 3889: 3886: 3885: 3883: 3882: 3880: 3876: 3872: 3867: 3862: 3858: 3842: 3839: 3837: 3834: 3832: 3829: 3827: 3824: 3823: 3821: 3817: 3814: 3812: 3811:Fermented tea 3809: 3808: 3806: 3802: 3799: 3797: 3794: 3792: 3789: 3788: 3786: 3782: 3779: 3778: 3776: 3772: 3769: 3767: 3764: 3762: 3759: 3757: 3754: 3753: 3751: 3747: 3744: 3743: 3741: 3737: 3734: 3733: 3731: 3727: 3724: 3723: 3721: 3717: 3714: 3712: 3709: 3707: 3704: 3702: 3699: 3697: 3694: 3692: 3689: 3688: 3686: 3682: 3679: 3678: 3677: 3674: 3673: 3671: 3667: 3659: 3656: 3654: 3651: 3649: 3646: 3644: 3641: 3639: 3636: 3635: 3633: 3629: 3626: 3625: 3623: 3619: 3616: 3615: 3613: 3609: 3606: 3604: 3601: 3599: 3596: 3594: 3593:Shaoxing wine 3591: 3589: 3586: 3584: 3581: 3579: 3576: 3574: 3571: 3569: 3566: 3564: 3561: 3559: 3556: 3554: 3551: 3549: 3546: 3544: 3541: 3539: 3536: 3534: 3531: 3529: 3526: 3524: 3521: 3519: 3516: 3514: 3511: 3509: 3506: 3504: 3501: 3500: 3498: 3494: 3491: 3489: 3486: 3485: 3483: 3479: 3476: 3474: 3471: 3469: 3466: 3464: 3461: 3460: 3458: 3454: 3451: 3450: 3448: 3447: 3445: 3441: 3433: 3430: 3428: 3425: 3423: 3420: 3418: 3415: 3413: 3410: 3408: 3405: 3403: 3400: 3398: 3395: 3394: 3392: 3388: 3385: 3384: 3383:Prickly pear 3382: 3378: 3375: 3374: 3372: 3368: 3365: 3364: 3362: 3358: 3355: 3354: 3352: 3348: 3345: 3344: 3342: 3338: 3335: 3334: 3332: 3328: 3325: 3324: 3322: 3318: 3315: 3314: 3312: 3308: 3305: 3303: 3300: 3298: 3295: 3293: 3290: 3288: 3285: 3283: 3280: 3278: 3275: 3273: 3270: 3268: 3265: 3263: 3260: 3259: 3257: 3253: 3250: 3249: 3247: 3243: 3240: 3239: 3237: 3233: 3230: 3228: 3225: 3224: 3222: 3218: 3215: 3214: 3212: 3211: 3209: 3205: 3201: 3196: 3191: 3187: 3171: 3168: 3167: 3166: 3163: 3159: 3156: 3154: 3151: 3150: 3149: 3146: 3144: 3141: 3139: 3136: 3132: 3129: 3127: 3124: 3122: 3119: 3117: 3114: 3112: 3109: 3108: 3107: 3104: 3102: 3099: 3097: 3094: 3093: 3091: 3087: 3079: 3076: 3074: 3071: 3069: 3066: 3064: 3061: 3059: 3056: 3055: 3054: 3051: 3049: 3046: 3044: 3041: 3040: 3038: 3034: 3030: 3023: 3019: 3015: 3008: 3003: 3001: 2996: 2994: 2989: 2988: 2985: 2975: 2974: 2969: 2961: 2955: 2952: 2950: 2947: 2945: 2942: 2940: 2937: 2935: 2932: 2930: 2927: 2925: 2922: 2920: 2917: 2916: 2914: 2910: 2904: 2901: 2900: 2898: 2894: 2888: 2885: 2883: 2880: 2878: 2875: 2874: 2872: 2870: 2866: 2860: 2857: 2855: 2852: 2850: 2847: 2845: 2842: 2840: 2837: 2835: 2832: 2830: 2827: 2826: 2824: 2822: 2818: 2812: 2809: 2808: 2806: 2804: 2800: 2794: 2791: 2789: 2786: 2784: 2781: 2779: 2776: 2775: 2773: 2771: 2767: 2761: 2758: 2756: 2753: 2751: 2748: 2747: 2745: 2743: 2739: 2735: 2728: 2723: 2721: 2716: 2714: 2709: 2708: 2705: 2698: 2694: 2690: 2688: 2682: 2681: 2677: 2670: 2666: 2662: 2656: 2653: 2649: 2643: 2640: 2636: 2630: 2627: 2624: 2620: 2616: 2612: 2606: 2604: 2600: 2596: 2595:Brew Magazine 2593: 2591: 2587: 2584: 2576: 2573: 2570: 2566: 2562: 2556: 2554: 2550: 2547: 2543: 2539: 2533: 2531: 2527: 2514: 2508: 2505: 2502: 2498: 2494: 2490: 2484: 2481: 2477: 2475: 2467: 2464: 2461: 2457: 2453: 2447: 2445: 2443: 2441: 2439: 2437: 2435: 2433: 2431: 2429: 2427: 2425: 2423: 2421: 2417: 2414: 2410: 2406: 2400: 2398: 2396: 2394: 2392: 2390: 2388: 2386: 2384: 2382: 2380: 2378: 2376: 2374: 2372: 2370: 2368: 2366: 2364: 2362: 2360: 2356: 2353: 2349: 2345: 2341: 2335: 2333: 2331: 2329: 2327: 2325: 2323: 2321: 2319: 2317: 2315: 2313: 2311: 2309: 2307: 2305: 2303: 2301: 2299: 2295: 2292: 2288: 2284: 2278: 2276: 2274: 2272: 2268: 2265: 2261: 2257: 2251: 2249: 2247: 2245: 2243: 2239: 2232: 2230: 2228: 2224: 2223:Brettanomyces 2218: 2217:can produce. 2216: 2215:Brettanomyces 2212: 2211:octanoic acid 2208: 2207:decanoic acid 2204: 2203:Saccharomyces 2200: 2196: 2195:Brettanomyces 2192: 2191:Brettanomyces 2187: 2185: 2184:Brettanomyces 2181: 2177: 2173: 2169: 2165: 2161: 2157: 2153: 2152:4-Ethylphenol 2149: 2148:Brettanomyces 2141: 2137: 2133: 2132:Brettanomyces 2129: 2128: 2127:Brettanomyces 2122: 2116:Brettanomyces 2115: 2113: 2111: 2110:Brettanomyces 2107: 2103: 2102:Saccharomyces 2098: 2096: 2092: 2088: 2084: 2083:ethyl acetate 2080: 2076: 2072: 2068: 2065:can create a 2064: 2060: 2056: 2051: 2049: 2045: 2041: 2037: 2033: 2029: 2025: 2021: 2017: 2008: 2004: 2000: 1995: 1988: 1986: 1983: 1982:Saccharomyces 1978: 1974: 1971: 1967: 1963: 1959: 1955: 1950: 1946: 1938: 1936: 1934: 1930: 1925: 1921: 1917: 1913: 1909: 1907: 1903: 1899: 1898: 1893: 1889: 1885: 1881: 1873: 1869: 1864: 1857: 1853: 1849: 1845: 1841: 1837: 1833: 1830: 1829:cell division 1826: 1822: 1819: 1816: 1812: 1808: 1804: 1801: 1798: 1794: 1791: 1788: 1784: 1780: 1777: 1774: 1771: 1768: 1764: 1763:phospholipids 1760: 1759:nucleic acids 1756: 1753: 1752: 1751: 1749: 1745: 1741: 1737: 1733: 1729: 1725: 1721: 1711: 1704: 1702: 1698: 1695: 1689: 1687: 1682: 1673: 1666: 1664: 1662: 1657: 1653: 1649: 1645: 1641: 1637: 1633: 1629: 1628:pure cultured 1621: 1617: 1613: 1609: 1605: 1603:fermentation. 1602: 1598: 1594: 1592: 1588: 1584: 1580: 1578: 1574: 1571: 1568: 1565: 1564: 1563: 1555: 1548: 1546: 1544: 1540: 1539:phenylethanol 1535: 1534:Saccharomyces 1530: 1529:Saccharomyces 1525: 1523: 1518: 1514: 1510: 1506: 1502: 1501: 1496: 1495:Saccharomyces 1487: 1483: 1481: 1480:Saccharomyces 1476: 1472: 1467: 1466:Saccharomyces 1462: 1460: 1456: 1452: 1448: 1447: 1442: 1441: 1436: 1435: 1430: 1429: 1424: 1420: 1416: 1412: 1411:Saccharomyces 1407: 1405: 1401: 1400: 1395: 1391: 1387: 1386: 1381: 1377: 1373: 1369: 1365: 1361: 1357: 1353: 1349: 1339: 1332: 1330: 1328: 1327: 1322: 1321: 1316: 1315: 1310: 1309:Saccharomyces 1306: 1302: 1298: 1294: 1290: 1286: 1282: 1278: 1274: 1270: 1266: 1262: 1258: 1254: 1249: 1247: 1243: 1239: 1235: 1231: 1227: 1223: 1219: 1215: 1211: 1207: 1203: 1199: 1195: 1191: 1187: 1183: 1178: 1176: 1170: 1168: 1163: 1162:S. cerevisiae 1159: 1155: 1151: 1147: 1143: 1139: 1135: 1131: 1128: 1124: 1120: 1111: 1104: 1100: 1099: 1095: 1093: 1092: 1088: 1086: 1085: 1081: 1079: 1078: 1074: 1072: 1071: 1067: 1065: 1064: 1060: 1059: 1058: 1056: 1055:Saccharomyces 1052: 1047: 1045: 1041: 1037: 1036:Saccharomyces 1033: 1029: 1025: 1021: 1017: 1014: 1010: 1006: 1005:Saccharomyces 998: 994: 988:Saccharomyces 987: 982: 978: 977: 972: 971: 970:Aureobasidium 967: 964: 960: 959: 955: 952: 948: 944: 943: 939: 937: 936: 932: 930: 929: 924: 920: 919:ethyl acetate 916: 912: 911:Hanseniaspora 908: 907: 903: 900: 899: 898:Kluyveromyces 894: 893: 888: 887: 882: 881: 880:Metschnikowia 876: 875: 870: 869: 865: 862: 858: 857: 856:Brettanomyces 853: 852: 851: 849: 845: 841: 837: 832: 830: 829: 824: 820: 819: 814: 810: 809: 808:Brettanomyces 804: 800: 796: 792: 784: 780: 779: 773: 766: 764: 762: 758: 754: 750: 744: 736: 734: 732: 728: 724: 720: 716: 712: 708: 704: 700: 695: 693: 689: 685: 684:mannoproteins 681: 677: 673: 669: 665: 661: 656: 654: 650: 646: 642: 638: 634: 630: 626: 622: 612: 605: 600: 596: 592: 588: 584: 580: 577: 573: 569: 565: 562: 559: 555: 551: 547: 542: 538: 534: 531: 528: 523: 520: 516: 512: 509: 508: 504: 500: 495: 488: 487:ethyl acetate 483: 479: 475: 471: 467: 464: 461: 457: 456:Succinic acid 454: 450: 446: 442: 438: 434: 430: 427: 424: 420: 419:demethylation 416: 413: 412: 411: 408: 406: 402: 397: 396:Metschnikowia 393: 388: 384: 379: 375: 371: 361: 354: 352: 350: 345: 341: 337: 333: 329: 323: 319: 309: 302: 300: 296: 295:of the wine. 294: 290: 286: 282: 277: 273: 269: 265: 261: 256: 254: 250: 246: 245:Louis Pasteur 241: 239: 235: 226: 219: 217: 215: 211: 210: 209:Brettanomyces 205: 201: 197: 193: 189: 188: 183: 182:S. cerevisiae 179: 178: 177:Saccharomyces 173: 169: 165: 161: 158: 157: 152: 151: 146: 142: 141:S. cerevisiae 138: 134: 130: 126: 125: 120: 115: 113: 108: 104: 103:fortification 100: 96: 95:dessert wines 92: 88: 84: 80: 76: 72: 68: 65:converts the 64: 60: 56: 53: 49: 45: 37: 32: 19: 4156:Corn whiskey 4060:Himbeergeist 3913:Kirschwasser 3675: 3373:Pomegranate 3267:Marsala wine 3262:Madeira wine 3169: 3157: 3153:Simple syrup 3148:Fermentation 3138:Distillation 2964: 2858: 2821:Fermentation 2684: 2660: 2655: 2647: 2642: 2634: 2629: 2610: 2594: 2580: 2575: 2560: 2537: 2517:. Retrieved 2515:. Scott Labs 2513:"Wine Yeast" 2507: 2488: 2483: 2471: 2466: 2451: 2404: 2339: 2282: 2255: 2222: 2219: 2214: 2202: 2198: 2194: 2190: 2188: 2183: 2147: 2145: 2139: 2135: 2131: 2125: 2109: 2101: 2099: 2078: 2067:film surface 2062: 2058: 2052: 2040:acetaldehyde 2031: 2027: 2023: 2013: 2006: 2002: 1998: 1981: 1979: 1975: 1942: 1911: 1910: 1895: 1892:Italian wine 1887: 1883: 1877: 1871: 1867: 1716: 1699: 1690: 1681:freeze dried 1678: 1661:Killer yeast 1631: 1627: 1625: 1608:flocculation 1591:acetaldehyde 1560: 1533: 1528: 1526: 1498: 1494: 1492: 1479: 1465: 1463: 1450: 1444: 1438: 1432: 1426: 1410: 1408: 1403: 1397: 1393: 1390:herb ferment 1383: 1379: 1375: 1371: 1367: 1363: 1359: 1355: 1351: 1344: 1324: 1318: 1312: 1308: 1304: 1285:disgorgement 1268: 1256: 1250: 1226:monoterpenes 1222:glycosidases 1179: 1171: 1161: 1145: 1118: 1116: 1096: 1089: 1082: 1075: 1068: 1061: 1054: 1050: 1048: 1035: 1004: 1002: 996: 974: 968: 956: 940: 933: 926: 915:killer yeast 910: 909:(Teleomorph 904: 896: 890: 886:Issatchenkia 884: 878: 872: 866: 860: 859:(Teleomorph 854: 840:bread making 835: 833: 826: 821:) under its 816: 806: 788: 776: 746: 696: 657: 652: 648: 632: 628: 618: 568:anthocyanins 564:Pyruvic acid 557: 519:Spanish wine 511:Acetaldehyde 499:Sherry wines 409: 405:legal limits 395: 391: 366: 340:acetaldehyde 325: 297: 275: 263: 259: 257: 249:Microbiology 242: 231: 207: 185: 181: 175: 154: 148: 140: 122: 116: 79:fermentation 43: 42:The role of 41: 18:Yeast (wine) 4819:Star anise 4713:Bénédictine 4673:Irish cream 4409:Aguardiente 4354:Aguardiente 4226:Rye whiskey 3736:Ginger wine 3453:Barley wine 3417:Mulled wine 3367:Plum jerkum 3337:Lychee wine 3327:Longan wine 3252:Bokbunja-ju 3242:Bignay wine 3232:Banana wine 3227:Banana beer 3101:Bathtub gin 2924:Wine bottle 2896:Other steps 2887:Wine cellar 2844:Süssreserve 2178:are common 2176:Fruit flies 2154:(4-EP) and 2036:acetic acid 2020:off flavors 2016:wine faults 1943:Yeasts are 1787:fatty acids 1728:amino acids 1694:cytoplasmic 1644:Napa Valley 1587:acetic acid 1423:fruit flies 1242:polyphenols 1200:of certain 1134:mycologists 1034:. However, 1009:fruit wines 983:in barrels. 923:acetic acid 892:Torulaspora 850:) include: 848:wine faults 823:sporulating 813:viticulture 757:carbonation 727:astringency 707:volatilized 703:hydrophobic 678:as well as 466:Acetic acid 378:NADH enzyme 266:due to the 137:inoculation 81:. The more 4970:Winemaking 4954:Categories 4878:Limoncello 4813:Triple sec 4788:Eau créole 4758:Irish Mist 4743:Bärenjäger 4698:Frangelico 4628:Cloudberry 4607:Chocolate 4591:Maraschino 4137:Buckwheat 4050:Eau de vie 3353:Pineapple 3317:Duhat wine 3313:Java plum 3307:White wine 3165:Winemaking 3089:Production 2803:Maceration 2793:Wine press 2783:Destemming 2734:Winemaking 2669:0932664695 2623:0198609906 2569:0932664660 2546:0834217015 2501:0198609906 2460:0834217015 2413:0387333495 2352:0198609906 2291:1891267914 2264:1580171052 2233:References 2160:Pinot noir 2106:maceration 1958:lanosterol 1954:ergosterol 1924:autolyzing 1872:(pictured) 1748:riboflavin 1686:cold shock 1597:maceration 1509:classified 1415:grapevines 1368:inoculated 1360:indigenous 1301:manzanilla 1277:flocculate 1230:aliphatics 1198:hydrolysis 1158:winemaking 1138:enologists 1123:eukaryotic 981:aging wine 803:teleomorph 692:mercaptans 680:Chardonnay 629:gross lees 554:disulfides 441:Chardonnay 429:Fusel oils 344:co-enzymes 336:glycolysis 316:See also: 268:elliptical 214:wine fault 172:physiology 119:winemaking 4863:Cedratine 4828:Sassolino 4798:Cointreau 4718:Brennivín 4694:Hazelnut 4658:Tia Maria 4617:Cinnamon 4601:Pertsivka 4566:Bierlikör 4369:Desi daru 4131:Bierbrand 4025:Tsikoudia 3988:Slivovitz 3888:Applejack 3766:Kabarawan 3742:Galangal 3696:Palm wine 3573:Rice wine 3553:Makgeolli 3488:Oshikundu 3407:Hippocras 3292:Rosé wine 3282:Port wine 3248:Bokbunja 3121:Lautering 2755:Noble rot 2659:M. Baldy 2519:23 August 2254:Jeff Cox 2170:from the 2071:microbial 2024:Kloeckera 1980:Wild non- 1949:anaerobic 1904:from the 1852:manganese 1773:Potassium 1755:Phosphate 1742:(such as 1736:magnesium 1734:(such as 1428:Kloeckera 1167:wines age 1130:sequenced 1117:In 1996, 1044:arabinose 1042:(such as 1024:raffinose 906:Kloeckera 783:Jura wine 781:from the 778:Vin jaune 751:and many 749:Champagne 723:mouthfeel 676:Champagne 649:bâtonnage 645:reductive 641:autolysis 633:fine lees 621:tartrates 482:propionic 401:bisulfite 392:Kloeckera 238:etymology 196:Zinfandel 150:Kloeckera 99:filtering 57:. In the 4888:Sloe gin 4883:Schnapps 4848:Licor 43 4844:Vanilla 4838:Charanda 4769:Juniper 4748:Drambuie 4683:Advocaat 4639:Coconut 4576:Patxaran 4515:Anisette 4494:Absinthe 4479:Amaretto 4464:Liqueurs 4329:Lambanog 4237:Sorghum 4176:Cheongju 4030:Tsipouro 3974:Juniper 3948:Armagnac 3893:Calvados 3836:Parakari 3816:Kombucha 3716:Tunggang 3691:Bahalina 3658:Huangjiu 3624:Sorghum 3578:Rượu cần 3543:Gwaha-ju 3478:Tesgüino 3402:Dubonnet 3397:Conditum 3387:Colonche 3302:Vermouth 3287:Red wine 3272:Mistelle 2770:Pressing 2693:Archived 2586:Archived 2281:D. Bird 2168:Riesling 2164:Burgundy 2140:pictured 2003:pictured 1960:. These 1929:cysteine 1920:arginine 1916:ammonium 1906:pressing 1840:chlorine 1821:Inositol 1783:proteins 1740:vitamins 1732:minerals 1646:and the 1640:Burgundy 1636:Bordeaux 1583:pyruvate 1505:Bordeaux 1475:macerate 1372:selected 1281:riddling 1273:UC-Davis 1214:Riesling 1202:cysteine 1190:Sémillon 1182:varietal 1040:pentoses 799:anamorph 791:taxonomy 711:tartrate 672:Muscadet 615:corking. 591:lactones 581:Various 546:rackings 541:sulfites 537:sulfates 527:oxidized 437:diacetyl 415:Methanol 370:glycerol 168:toxicity 4894:Walnut 4858:Campari 4803:Curaçao 4794:Orange 4778:Jenever 4763:Krupnik 4708:Aquavit 4649:Coffee 4621:Tentura 4582:Cherry 4556:Sambuca 4541:Mastika 4525:Hierbas 4499:Anisado 4475:Almond 4424:Horilka 4414:Akvavit 4364:Clairin 4359:Cachaça 4309:Tequila 4186:Lao-Lao 4171:Awamori 4107:Barley 4101:Cereals 4090:Schnaps 4080:Pálinka 4070:Nalewka 4035:Zivania 3968:Zivania 3909:Cherry 3866:Liquors 3732:Ginger 3568:Pangasi 3528:Choujiu 3508:Amazake 3484:Millet 3473:Tejuino 3449:Barley 3443:Cereals 3432:Sangria 3357:Tepache 3333:Lychee 3323:Longan 3238:Bignay 3223:Banana 3116:Malting 3111:Brewery 3106:Brewing 2954:Terroir 2912:Related 2760:Vintage 2742:Harvest 2227:ethanol 2059:Candida 2028:Candida 1999:Candida 1962:sterols 1897:ripasso 1884:Go-Ferm 1868:ripasso 1836:calcium 1817:itself. 1744:thiamin 1724:ammonia 1616:racking 1517:compost 1500:terroir 1471:pressed 1434:Candida 1364:natural 1356:ambient 1257:Épernay 1244:in the 1238:benzene 1206:enzymes 1142:strains 1032:ethanol 1028:maltose 1020:sucrose 1016:glucose 1013:ferment 951:budding 947:fission 868:Candida 861:Dekkera 844:brewing 828:Dekkera 731:tannins 699:mannose 653:sur lie 599:acetals 595:phenols 587:ketones 529:aromas. 474:butyric 449:acetoin 445:buttery 423:pectins 349:pentose 328:glucose 272:strains 220:History 156:Candida 145:harvest 87:dryness 71:alcohol 4930:Portal 4898:Nocino 4823:Pastis 4753:Glayva 4739:Honey 4733:Unicum 4728:Metaxa 4704:Herbs 4688:Eggnog 4664:Cream 4653:Kahlúa 4643:Malibu 4490:Anise 4434:Shōchū 4429:Poitín 4419:Bangla 4399:Tharra 4340:Dairy 4324:Laksoy 4319:Arrack 4304:Mezcal 4281:Whisky 4276:Shōchū 4266:Baijiu 4241:Baijiu 4231:Starka 4201:Shochu 4191:Lihing 4181:Cholai 4147:Maize 4045:Brandy 4010:Grappa 4005:Chacha 4000:Pomace 3958:Cognac 3953:Brandy 3939:Grape 3933:Boukha 3919:Dates 3884:Apple 3841:Ibwatu 3831:Chicha 3777:Sugar 3752:Honey 3722:Dairy 3681:Pulque 3628:Pendhā 3548:Hariya 3538:Dansul 3518:Beopju 3493:Tongba 3422:Nabidh 3297:Sherry 3258:Grape 3213:Apple 2919:Winery 2882:Solera 2667: 2621: 2567: 2544: 2499: 2458: 2411: 2350: 2289: 2262: 2180:vector 2079:Pichia 2063:Pichia 2055:oxygen 2044:thiols 2007:Pichia 2005:) and 1933:thiols 1902:pomace 1888:Ferm-K 1844:copper 1797:lipids 1779:Biotin 1738:) and 1601:barrel 1513:pomace 1459:pomace 1440:Pichia 1289:Sherry 1265:France 1236:, and 1218:Muscat 1194:thiols 1127:genome 874:Pichia 789:Yeast 637:settle 583:esters 522:Sherry 478:formic 452:aroma. 312:yeast. 285:sulfur 160:genera 107:brandy 67:sugars 4944:Drink 4633:Lakka 4562:Beer 4439:Vodka 4389:Sulai 4374:Guaro 4344:Arkhi 4334:Sotol 4299:Agave 4292:Other 4211:Sulai 4167:Rice 4127:Beer 4085:Rakia 4055:Geist 4020:Orujo 3993:Țuică 3984:Plum 3963:Pisco 3923:Araqi 3878:Fruit 3826:Cauim 3801:Palek 3796:Intus 3781:Kilju 3761:Byais 3746:Byais 3726:Kumis 3711:Tuhak 3669:Other 3618:Kvass 3608:Tapuy 3603:Tapai 3598:Sonti 3563:Mirin 3558:Mijiu 3533:Chuak 3503:Agkud 3499:Rice 3468:Pozol 3463:Mageu 3459:Corn 3427:Pruno 3412:Jabol 3377:Rimon 3363:Plum 3347:Perry 3343:Pear 3217:Cider 3207:Fruit 3131:Yeast 2869:Aging 2617:2006 2495:2006 2474:Yeast 2346:2006 2172:Mosel 2162:from 2010:wine. 1811:redox 1406:冶葛). 1394:cǎoqū 1348:flora 1287:. In 1154:bread 383:redox 293:aroma 200:Syrah 192:ports 105:with 63:yeast 55:juice 52:fruit 50:from 4917:List 4679:Egg 4551:Rakı 4546:Ouzo 4520:Arak 4394:Sura 4221:Korn 4217:Rye 4206:Soju 4075:Oghi 4015:Marc 3943:Arak 3929:Fig 3903:Feni 3807:Tea 3791:Basi 3771:Mead 3756:Bais 3706:Tubâ 3701:Tuak 3653:Boza 3648:Beer 3614:Rye 3588:Sato 3583:Sake 3523:Brem 3126:Wort 2665:ISBN 2619:ISBN 2565:ISBN 2542:ISBN 2521:2023 2497:ISBN 2456:ISBN 2409:ISBN 2348:ISBN 2287:ISBN 2260:ISBN 2209:and 2095:flor 2085:and 2061:and 2026:and 1956:and 1886:and 1856:zinc 1854:and 1848:iron 1746:and 1726:and 1612:lees 1610:and 1589:and 1543:rose 1532:non- 1457:and 1455:lees 1437:and 1404:yěgé 1323:and 1299:and 1297:fino 1293:flor 1283:and 1267:and 1246:must 1216:and 1188:and 1156:and 1150:beer 1136:and 1030:and 1022:and 963:Brix 921:and 895:and 842:and 721:and 719:body 713:and 625:lees 606:Lees 597:and 552:and 515:flor 480:and 394:and 387:body 320:and 204:Brix 198:and 164:must 153:and 73:and 48:wine 36:must 4773:Gin 4379:Rum 4256:Ara 3978:Gin 3643:Ara 3638:Ale 3513:Apo 2877:Oak 1392:" ( 1374:or 1362:or 1263:of 1251:In 1144:of 729:of 539:or 421:of 274:of 121:is 4956:: 2691:. 2602:^ 2552:^ 2529:^ 2419:^ 2358:^ 2297:^ 2270:^ 2241:^ 2050:. 2038:, 1850:, 1846:, 1842:, 1838:, 1785:, 1761:, 1642:, 1638:, 1618:, 1585:, 1507:, 1431:, 1370:, 1358:, 1329:. 1317:, 1311:, 1248:. 1232:, 1212:, 1169:. 1152:, 1018:, 889:, 883:, 877:, 733:. 686:, 670:, 593:, 589:, 585:, 476:, 255:. 184:, 133:pH 114:. 61:, 4932:: 4247:) 4243:( 3006:e 2999:t 2992:v 2726:e 2719:t 2712:v 2689:" 2685:" 2592:" 2581:" 2523:. 2476:" 2472:" 2138:( 2001:( 1831:. 1722:( 1659:" 1402:( 953:. 901:) 863:) 601:. 505:. 462:. 20:)

Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.

Index