344:
hypertrophied. The ulna is slightly longer than the humerus. The ulnar olecranon process has a mediolaterally pinched posterior margin. The ulnar distal end bends anteriorly and is strongly expanded laterally. The proximal end of metacarpal II has a strong ventrolateral extension. Metacarpal III is laterally bowed and adorned with a longitudinal ventral groove. Phalanx II-2 is without a proximodorsal lip and lacks strong dorsal arching. The femoral head is stout and lower than the trochanteric crest. The medial condyle of tibiotarsus distal end has a prominent distal extension. The foot shows the
1485:
1474:
1480:
139:
1491:
118:
1514:
321:
403:
are preserved as well. The caudal series of vertebrae are represented by 26 articulated caudal vertebrae, probably missing only the first caudal vertebra, likely making a full count of 27 caudal vertebrae in the tail. The sternum is represented by what is probably the left sternal plate. Five left sternal ribs are preserved, more than are seen in the related genera
402:
The specimen's remains preserve the four most posterior cervical vertebrae (likely 7–10 in regards to placement). Seven dorsal vertebrae are known, while eight dorsal ribs are known from the left side of the specimen in comparison to the five preserved for the right side. Five left uncinate processes
343:
is present with slender and posteriorly curved clavicular rami. The humeral proximal end is strongly offset medially from the humeral shaft. The internal tuberosity of the humeral proximal end is short. A large fenestra perforates the humeral deltopectoral crest. The humeral ulnar condyle is
355:
The articulated tail has elongate rod-like extensions of the prezygaphophyses and chevrons characteristic of most dromaeosaurids, and these reach almost to the most anterior caudals. The sternum is large, with a maximum anteroposterior length 56% of the femoral length and is long axially in
460:. However, a full understanding of niche differentiation of Jehol dromaeosaurids probably will require a full dataset of the tempo-spatial distribution of the many Jehol dromaeosaurid fossils and a better understanding of their ecology before such conclusions can be fully drawn.
448:. The varying size of the different dromaeosaurids likely indicates that they focused on different types of prey and that many were specific about what type of prey they consumed. Niche partitioning may also have been aided by differentiation in habitat preferences.
356:
proportion. Bone fusion in the scapula, vertebrae, coracoid, and tarsals suggest that the specimen is a mature adult weighing 0.31 kg (0.68 lb), making it one of the smallest known non-avian theropods thus far discovered.
419:
are represented by both humeri, the left ulna, the left radius, and a partial left manus. The specimen's hindlimbs meanwhile, are represented by both femora, both tibiotarsi, and portions of both feet.
392:
367:
was first reported in 2009 when a new species based on a specimen consisting of a partial postcranial articulated skeleton was recovered from the lake deposits in
352:
hand as the second, third and fourth, whereas most researchers see them as the first, second and third so that "metacarpal II" would become metacarpal I etc.
1594:
667:
511:"A new tiny dromaeosaurid dinosaur from the Lower Cretaceous Jehol Group of western Liaoning and niche differentiation among the Jehol dromaeosaurids"
432:
in the
Dromaeosauridae, and specifically the Microraptoria, based on its synapomorphies. Some studies offer an alternate position as an avialan.
1609:
308:, specifically the middle of the Jehol Biota, it is the smallest known microraptorine thus far discovered and one of the smallest non-avian
1507:
469:
1484:
474:
1473:
138:
660:
391:, China's founder of vertebrate paleontology. The holotype specimen, labelled as IVPP V 22775, is currently housed in the
1490:
1604:
1479:
510:
444:
in the Yixian
Formation, in addition to eight other dromaeosaurids suggests that there was a sufficient amount of
1599:
764:
653:
384:
258:
543:
Agnolin, Federico L.; Motta, Matias J.; Brissón Egli, Federico; Lo Coco, Gastón; Novas, Fernando E. (2019).
1534:
415:. The furcula is preserved as well, as are the scapulae and coracoids, which are fused. The forelimbs of
1566:
1517:
348:
condition. Metatarsal II is without ginglymi on its distal end. The authors identify the fingers in the
818:
372:
1235:
972:
791:
525:
445:
133:
1571:
339:. Proportionally long ossified uncinate processes are fused to the dorsal ribs. A widely arched
1432:
1323:
1259:
1155:
948:
631:
576:
368:
345:
1356:
1302:
1140:
621:
611:
566:
556:
411:
305:
30:
1416:
1270:
1173:
886:
864:
830:
749:
676:
222:
1074:
626:
596:"The phylogenetic affinities and morphological peculiarities of the bird-like dinosaur
595:
388:
336:
117:
1588:
1395:
1309:
1201:
1186:
1125:
1118:
1111:
1101:
1036:
1029:
958:
301:
297:
238:
68:
529:
1454:
1365:
1286:
1279:
1244:
1223:
1208:
1147:
1014:
1007:
1000:
990:
937:
922:
349:
267:
1557:
1409:
1387:
1338:
1294:
1251:
1216:
1194:
1133:
1044:
979:
803:
405:
43:
320:
1402:
1379:
1331:
1082:
1051:
929:
907:
715:
706:
453:
196:
88:
53:
580:
561:
544:
1440:
1372:
1067:
1059:
965:
776:
724:
209:
150:
93:
37:
635:
1551:
1424:
1345:
1316:
1090:
900:
700:
457:
376:
309:
183:
170:
83:
78:
63:
58:
48:
616:
571:
1447:
733:
396:
340:
98:
73:
1022:
914:
694:
160:
1528:
645:
545:"Paravian Phylogeny and the Dinosaur-Bird Transition: An Overview"
380:
319:
1532:
687:
649:
335:
is distinguishable from other microraptorines in the following
393:
Institute of
Vertebrate Paleontology and Paleoanthropology
387:
and Qin Zi-Chuan. The genus and specific name honor
274:
1541:
1355:
1269:
1234:
1172:
1100:
989:
947:
884:
829:
802:
775:
748:
661:
8:
504:
502:
500:
498:
496:
494:
492:
490:
1529:
1178:
892:
881:
808:
781:
754:
745:
684:
668:
654:
646:
116:
20:
625:
615:
570:
560:
452:was, in view of its small size, possibly
486:
600:from the Upper Cretaceous of Mongolia"
7:
1595:Early Cretaceous dinosaurs of Asia
470:Timeline of dromaeosaurid research
14:
1513:
1512:
1489:
1483:
1478:
1472:
509:Xu, Xing; Qin, Zi-Chuan (2017).
137:
475:2017 in archosaur paleontology
328:based on the holotype skeleton
125:An artist's reconstruction of
1:
1610:Fossil taxa described in 2017
428:Studies by Xu and Qin place
383:. It was first described by
304:. Believed to hail from the
1626:
549:Frontiers in Earth Science
1502:
1470:
1181:
895:
880:
861:
811:
784:
757:
744:
683:
273:
266:
134:Scientific classification
132:
124:
115:
23:
16:Extinct genus of reptiles
594:Cau A, Madzia D (2021).
562:10.3389/feart.2018.00252
324:Life reconstruction of
518:Vertebrata PalAsiatica
329:
326:Zhongjianosaurus yangi
279:Zhongjianosaurus yangi
127:Zhongjianosaurus yangi
1567:Paleobiology Database
598:Borogovia gracilicrus
323:
360:Discovery and naming
1605:Feathered dinosaurs
1236:Saurornitholestinae
617:10.7717/peerj.12640
446:niche partitioning
346:arctometatarsalian
330:
284:Xu & Qin, 2017
1582:
1581:
1535:Taxon identifiers
1526:
1525:
1498:
1497:
1468:
1467:
1464:
1463:
1433:Saurornitholestes
1324:Saurornitholestes
1260:Saurornitholestes
949:Halszkaraptorinae
876:
875:
872:
871:
857:
856:
853:
852:
849:
848:
440:The discovery of
300:belonging to the
289:
288:
262:
1617:
1575:
1574:
1562:
1561:
1560:
1543:Zhongjianosaurus
1530:
1516:
1515:
1493:
1487:
1482:
1476:
1357:Velociraptorinae
1303:Dromaeosauroides
1179:
1163:Zhongjianosaurus
1141:Sinornithosaurus
893:
882:
809:
782:
755:
746:
739:
738:
685:
670:
663:
656:
647:
640:
639:
629:
619:
591:
585:
584:
574:
564:
540:
534:
533:
515:
506:
450:Zhongjianosaurus
442:Zhongjianosaurus
430:Zhongjianosaurus
417:Zhongjianosaurus
412:Sinornithosaurus
365:Zhongjianosaurus
333:Zhongjianosaurus
306:Yixian Formation
293:Zhongjianosaurus
276:
257:
253:Zhongjianosaurus
250:
237:
221:
142:
141:
120:
110:
40:
31:Early Cretaceous
29:Temporal range:
25:Zhongjianosaurus
21:
1625:
1624:
1620:
1619:
1618:
1616:
1615:
1614:
1600:Microraptorians
1585:
1584:
1583:
1578:
1570:
1565:
1556:
1555:
1550:
1537:
1527:
1522:
1494:
1460:
1417:Luanchuanraptor
1351:
1271:Dromaeosaurinae
1265:
1230:
1174:Eudromaeosauria
1168:
1096:
985:
943:
889:
887:Dromaeosauridae
868:
865:Dromaeosauridae
845:
831:Dromaeosauridae
825:
798:
771:
765:Avemetatarsalia
750:Avemetatarsalia
740:
690:
689:
679:
677:Dromaeosauridae
674:
644:
643:
593:
592:
588:
542:
541:
537:
513:
508:
507:
488:
483:
466:
438:
426:
373:Lingyuan County
362:
318:
285:
282:
261:& Qin, 2017
256:
248:
235:
223:Dromaeosauridae
219:
136:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
27:
17:
12:
11:
5:
1623:
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1607:
1602:
1597:
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1563:
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1533:
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1510:
1503:
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1471:
1469:
1466:
1465:
1462:
1461:
1459:
1458:
1451:
1444:
1437:
1429:
1421:
1413:
1406:
1399:
1392:
1384:
1376:
1369:
1361:
1359:
1353:
1352:
1350:
1349:
1342:
1335:
1328:
1320:
1313:
1306:
1299:
1291:
1283:
1275:
1273:
1267:
1266:
1264:
1263:
1256:
1248:
1240:
1238:
1232:
1231:
1229:
1228:
1220:
1213:
1205:
1198:
1191:
1182:
1176:
1170:
1169:
1167:
1166:
1159:
1152:
1144:
1137:
1130:
1122:
1115:
1107:
1105:
1098:
1097:
1095:
1094:
1087:
1079:
1075:Unquillosaurus
1071:
1064:
1056:
1048:
1041:
1033:
1026:
1019:
1011:
1004:
996:
994:
987:
986:
984:
983:
976:
969:
962:
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952:
945:
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942:
941:
934:
926:
919:
911:
904:
896:
890:
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878:
877:
874:
873:
870:
869:
862:
859:
858:
855:
854:
851:
850:
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846:
844:
843:
842:
841:
835:
833:
827:
826:
824:
823:
822:
821:
812:
806:
800:
799:
797:
796:
795:
794:
785:
779:
773:
772:
770:
769:
768:
767:
758:
752:
742:
741:
737:
736:
727:
718:
709:
703:
697:
688:
681:
680:
675:
673:
672:
665:
658:
650:
642:
641:
586:
535:
524:(2): 129–144.
485:
484:
482:
479:
478:
477:
472:
465:
462:
437:
434:
425:
424:Classification
422:
389:Yang Zhongjian
361:
358:
337:autapomorphies
317:
314:
296:is a genus of
287:
286:
283:
271:
270:
264:
263:
246:
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241:
233:
226:
225:
217:
213:
212:
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174:
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130:
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104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
1622:
1611:
1608:
1606:
1603:
1601:
1598:
1596:
1593:
1592:
1590:
1573:
1568:
1564:
1559:
1553:
1549:
1548:
1546:
1544:
1540:
1536:
1531:
1519:
1511:
1509:
1505:
1504:
1501:
1492:
1488:
1486:
1481:
1475:
1457:
1456:
1452:
1450:
1449:
1445:
1443:
1442:
1438:
1435:
1434:
1430:
1427:
1426:
1422:
1419:
1418:
1414:
1412:
1411:
1407:
1405:
1404:
1400:
1398:
1397:
1396:Kansaignathus
1393:
1390:
1389:
1385:
1382:
1381:
1377:
1375:
1374:
1370:
1368:
1367:
1363:
1362:
1360:
1358:
1354:
1348:
1347:
1343:
1341:
1340:
1336:
1334:
1333:
1329:
1326:
1325:
1321:
1319:
1318:
1314:
1312:
1311:
1310:Dromaeosaurus
1307:
1305:
1304:
1300:
1297:
1296:
1292:
1289:
1288:
1284:
1282:
1281:
1277:
1276:
1274:
1272:
1268:
1262:
1261:
1257:
1254:
1253:
1249:
1247:
1246:
1242:
1241:
1239:
1237:
1233:
1226:
1225:
1221:
1219:
1218:
1214:
1211:
1210:
1206:
1204:
1203:
1202:Dineobellator
1199:
1197:
1196:
1192:
1189:
1188:
1184:
1183:
1180:
1177:
1175:
1171:
1165:
1164:
1160:
1158:
1157:
1153:
1150:
1149:
1145:
1143:
1142:
1138:
1136:
1135:
1131:
1128:
1127:
1126:Hesperonychus
1123:
1121:
1120:
1119:Graciliraptor
1116:
1114:
1113:
1112:Changyuraptor
1109:
1108:
1106:
1103:
1102:Microraptoria
1099:
1093:
1092:
1088:
1085:
1084:
1080:
1077:
1076:
1072:
1070:
1069:
1065:
1062:
1061:
1057:
1054:
1053:
1049:
1047:
1046:
1042:
1039:
1038:
1037:Ornithodesmus
1034:
1032:
1031:
1030:Neuquenraptor
1027:
1025:
1024:
1020:
1017:
1016:
1012:
1010:
1009:
1005:
1003:
1002:
998:
997:
995:
992:
988:
982:
981:
977:
975:
974:
970:
968:
967:
963:
961:
960:
959:Halszkaraptor
956:
955:
953:
950:
946:
940:
939:
935:
932:
931:
927:
925:
924:
920:
917:
916:
912:
910:
909:
905:
903:
902:
898:
897:
894:
891:
888:
883:
879:
867:
866:
860:
839:
838:
837:
836:
834:
832:
828:
820:
816:
815:
814:
813:
810:
807:
805:
801:
793:
789:
788:
787:
786:
783:
780:
778:
774:
766:
762:
761:
760:
759:
756:
753:
751:
747:
743:
735:
731:
728:
726:
722:
719:
717:
713:
710:
708:
704:
702:
698:
696:
692:
691:
686:
682:
678:
671:
666:
664:
659:
657:
652:
651:
648:
637:
633:
628:
623:
618:
613:
609:
605:
601:
599:
590:
587:
582:
578:
573:
568:
563:
558:
554:
550:
546:
539:
536:
531:
527:
523:
519:
512:
505:
503:
501:
499:
497:
495:
493:
491:
487:
480:
476:
473:
471:
468:
467:
463:
461:
459:
455:
451:
447:
443:
435:
433:
431:
423:
421:
418:
414:
413:
408:
407:
400:
398:
394:
390:
386:
382:
378:
375:, in western
374:
370:
366:
359:
357:
353:
351:
347:
342:
338:
334:
327:
322:
315:
313:
311:
307:
303:
302:Microraptoria
299:
298:dromaeosaurid
295:
294:
281:
280:
272:
269:
265:
260:
255:
254:
247:
244:
243:
240:
239:Microraptoria
234:
231:
228:
227:
224:
218:
215:
214:
211:
208:
205:
202:
201:
198:
195:
192:
189:
188:
185:
182:
179:
176:
175:
172:
169:
166:
165:
162:
159:
156:
155:
152:
149:
146:
145:
140:
135:
131:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
36:125–120
32:
26:
22:
19:
1542:
1477:
1455:Velociraptor
1453:
1446:
1439:
1431:
1423:
1415:
1408:
1401:
1394:
1386:
1378:
1371:
1366:Acheroraptor
1364:
1344:
1337:
1330:
1322:
1315:
1308:
1301:
1293:
1287:Dakotaraptor
1285:
1280:Achillobator
1278:
1258:
1250:
1245:Atrociraptor
1243:
1224:Zhenyuanlong
1222:
1215:
1209:Tianyuraptor
1207:
1200:
1193:
1185:
1162:
1161:
1154:
1148:Tianyuraptor
1146:
1139:
1132:
1124:
1117:
1110:
1089:
1081:
1073:
1066:
1058:
1050:
1043:
1035:
1028:
1021:
1015:Dakotaraptor
1013:
1008:Buitreraptor
1006:
1001:Austroraptor
999:
991:Unenlagiinae
978:
971:
964:
957:
938:Zhenyuanlong
936:
928:
923:Tianyuraptor
921:
913:
906:
899:
863:
729:
720:
711:
607:
603:
597:
589:
572:11336/130197
552:
548:
538:
521:
517:
449:
441:
439:
436:Paleobiology
429:
427:
416:
410:
404:
401:
364:
363:
354:
350:maniraptoran
332:
331:
325:
292:
291:
290:
278:
277:
268:Type species
252:
251:
229:
203:
190:
177:
126:
24:
18:
1410:Linheraptor
1388:Deinonychus
1339:Yurgovuchia
1295:Deinonychus
1252:Bambiraptor
1217:Vectiraptor
1195:Bambiraptor
1134:Microraptor
1045:Pamparaptor
980:Natovenator
819:Maniraptora
804:Maniraptora
406:Microraptor
316:Description
312:dinosaurs.
1589:Categories
1506:See also:
1380:Boreonykus
1332:Utahraptor
1083:Variraptor
1052:Pyroraptor
930:Variraptor
908:Pyroraptor
840:see below↓
716:Dinosauria
707:Sauropsida
610:: e12640.
481:References
454:omnivorous
197:Saurischia
184:Dinosauria
1558:Q29366656
1373:Adasaurus
1068:Unenlagia
1060:Rahonavis
966:Hulsanpes
792:Theropoda
777:Theropoda
725:Theropoda
693:Kingdom:
581:2296-6463
210:Theropoda
157:Kingdom:
151:Eukaryota
1552:Wikidata
1518:Category
1508:Timeline
1425:Nuthetes
1346:Zapsalis
1317:Itemirus
1091:Ypupiara
973:Mahakala
901:Daurlong
701:Chordata
699:Phylum:
695:Animalia
636:34963824
530:44178386
464:See also
458:arboreal
377:Liaoning
369:Sihedang
310:theropod
216:Family:
171:Chordata
167:Phylum:
161:Animalia
147:Domain:
1448:Tsaagan
734:Paraves
705:Class:
627:8656384
397:Beijing
385:Xu Xing
341:furcula
245:Genus:
105:↓
1572:391290
1187:Balaur
1156:Wulong
1023:Diuqin
915:Shanag
634:
624:
579:
528:
730:Clade
721:Clade
712:Clade
604:PeerJ
526:S2CID
514:(PDF)
395:, in
381:China
230:Clade
204:Clade
191:Clade
178:Clade
1441:Shri
1403:Kuru
817:see
790:see
763:see
632:PMID
577:ISSN
456:and
409:and
44:PreꞒ
622:PMC
612:doi
567:hdl
557:doi
1591::
1569::
1554::
732::
723::
714::
630:.
620:.
606:.
602:.
575:.
565:.
555:.
551:.
547:.
522:55
520:.
516:.
489:^
399:.
379:,
371:,
259:Xu
232::
206::
193::
180::
94:Pg
38:Ma
33:,
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669:e
662:t
655:v
638:.
614::
608:9
583:.
569::
559::
553:6
532:.
275:†
249:†
236:†
220:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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