1331:
1337:
417:
1887:
137:
2038:
119:
1917:
1893:
505:
diversity of the
Leptoceratopsidae. As the third leptoceratopsid from Asia, this find exhibits the coexistence and radiation of two closely related clades, whose differences in jaw and tooth adaptation may represent different feeding strategies.
449:. It is known from a partial articulated skeleton including vertebrae, ribs, teeth, and parts of the skull and mandibles. The fossils were recovered from the
2108:
2079:
1997:
542:"A New Leptoceratopsid (Ornithischia: Ceratopsia) from the Upper Cretaceous of Shandong, China and Its Implications for Neoceratopsian Evolution"
2128:
1330:
610:
652:
1336:
2123:
1910:
515:
2133:
2072:
136:
2138:
2143:
1886:
497:
had a particularly massive and deep 50 cm-long mandible that is also thin transversely. This and a number of other
2103:
493:
likely represents an adult, measuring 2.5 m (8.2 ft) in length and 175 kg (386 lb) in body mass.
2118:
2113:
2098:
2065:
743:
645:
861:
1937:
416:
2010:
1920:
770:
352:
290:
1975:
1892:
1984:
553:
458:
267:
1026:
638:
1746:
1696:
1128:
502:
131:
2015:
457:. This genus was named by Xing Xu, Kebai Wang, Xijin Zhao, Corwin Sullivan and Shuqing Chen in
1829:
1767:
1730:
1204:
1091:
616:
606:
581:
428:
333:
242:
2049:
1859:
1657:
1482:
879:
571:
561:
501:
unique to the genus lend it significance for increasing the morphological disparity and the
438:
340:
2037:
1753:
1705:
1664:
1600:
1503:
1454:
1156:
1080:
1035:
909:
895:
854:
728:
712:
454:
435:
374:
557:
384:
shares many features with
Leptoceratopsidae as well as other ceratopsian groups such as
1760:
1719:
1650:
1637:
1608:
1593:
1563:
1524:
1510:
1427:
1388:
1267:
1177:
1142:
1135:
1015:
965:
888:
868:
576:
541:
498:
213:
2092:
1838:
1817:
1787:
1778:
1678:
1572:
1517:
1496:
1468:
1461:
1447:
1440:
1246:
1235:
1228:
1170:
1163:
1149:
1107:
1042:
1008:
979:
932:
474:
450:
404:
390:
372:, which is of Late Cretaceous age, and most fossils are only disarticulated bones of
369:
69:
30:
1852:
1803:
1796:
1712:
1685:
1671:
1579:
1552:
1545:
1538:
1489:
1475:
1367:
1354:
1285:
1274:
1259:
1221:
1213:
1195:
1100:
1066:
1049:
1001:
994:
950:
755:
703:
385:
197:
118:
566:
2045:
1969:
1845:
1739:
1622:
1586:
1409:
1121:
986:
431:
44:
1615:
1531:
1381:
1114:
902:
836:
823:
782:
661:
336:
229:
89:
54:
1960:
620:
148:
94:
38:
585:
1954:
1416:
1402:
1395:
958:
694:
685:
466:
442:
365:
360:. The name is derived from the location of Zhucheng, the Latinized-Greek
181:
168:
84:
79:
64:
59:
49:
2002:
1810:
972:
916:
540:
Xing Xu; Kebai Wang; Xijin Zhao; Corwin
Sullivan; Shuqing Chen (2010).
330:
99:
74:
679:
600:
477:
158:
1931:
465:. The genus name was chosen for the location of Zhucheng, where the
408:, although internal relationships of this triplet were unresolved.
486:
to refer to the unexpected discovery of the articulated skeleton.
470:
446:
415:
344:
326:
1989:
364:, or "horned face", and the unexpected articulated nature of the
1935:
1352:
821:
672:
634:
630:
2053:
482:, meaning "horned face". They chose the species name
1944:
1828:
1786:
1777:
1729:
1695:
1636:
1562:
1426:
1365:
1284:
1245:
1203:
1194:
1090:
1064:
1025:
930:
878:
834:
781:
754:
727:
535:
533:
531:
388:. The overall size of the taxon was similar to
2073:
646:
8:
605:. Princeton University Press. p. 279.
2080:
2066:
1932:
1783:
1642:
1432:
1373:
1362:
1349:
1251:
1200:
1072:
942:
938:
846:
842:
831:
818:
760:
733:
724:
669:
653:
639:
631:
117:
20:
575:
565:
347:. It was first described in 2010, by Xu
527:
602:The Princeton Field Guide to Dinosaurs
7:
2034:
2032:
398:was analyzed to be in a group with
2052:. You can help Knowledge (XXG) by
14:
2109:Late Cretaceous dinosaurs of Asia
2036:
1916:
1915:
1891:
1885:
1335:
1329:
516:Timeline of ceratopsian research
368:. The skeleton was found in the
135:
1:
2129:Fossil taxa described in 2010
567:10.1371/journal.pone.0013835
478:
394:, although slightly larger.
463:Zhuchengceratops inexpectus
357:Zhuchengceratops inexpectus
303:Zhuchengceratops inexpectus
2160:
2031:
489:The recovered specimen of
461:, and the type species is
1905:
1883:
1645:
1435:
1376:
1361:
1348:
1326:
1254:
1075:
945:
941:
849:
845:
830:
817:
763:
736:
723:
668:
599:Paul, Gregory S. (2016).
420:Fossil skull and mandible
296:
289:
132:Scientific classification
130:
125:
116:
23:
16:Extinct genus of reptiles
2124:Paleontology in Shandong
2044:This article related to
862:Micropachycephalosaurus
434:which lived during the
441:in what is now Kugou,
421:
339:that lived during the
2134:Taxa named by Xu Xing
2011:Paleobiology Database
419:
2139:Ornithischian genera
412:Discovery and naming
2144:Ornithischian stubs
1027:Archaeoceratopsidae
558:2010PLoSO...513835X
469:was found, and the
2104:Maastrichtian life
1747:Crittendenceratops
1697:Albertaceratopsini
422:
351:, who created the
282:Z. inexpectus
2061:
2060:
2026:
2025:
1938:Taxon identifiers
1929:
1928:
1901:
1900:
1881:
1880:
1877:
1876:
1873:
1872:
1869:
1868:
1830:Pachyrhinosaurini
1768:Yehuecauhceratops
1731:Nasutoceratopsini
1632:
1631:
1344:
1343:
1324:
1323:
1320:
1319:
1316:
1315:
1312:
1311:
1308:
1307:
1304:
1303:
1205:Protoceratopsidae
1092:Leptoceratopsidae
1060:
1059:
926:
925:
813:
812:
809:
808:
805:
804:
801:
800:
612:978-1-78684-190-2
318:
317:
270:
243:Leptoceratopsidae
126:Life restoration
2151:
2119:Fossils of China
2114:Cretaceous China
2099:Leptoceratopsids
2082:
2075:
2068:
2040:
2033:
2019:
2018:
2006:
2005:
1993:
1992:
1980:
1979:
1978:
1976:Zhuchengceratops
1965:
1964:
1963:
1946:Zhuchengceratops
1933:
1919:
1918:
1896:
1895:
1889:
1860:Pachyrhinosaurus
1784:
1658:Machairoceratops
1643:
1483:Coahuilaceratops
1433:
1374:
1363:
1350:
1339:
1334:
1333:
1252:
1201:
1185:Zhuchengceratops
1073:
943:
939:
880:Chaoyangsauridae
847:
843:
832:
819:
761:
734:
725:
718:
717:
670:
655:
648:
641:
632:
625:
624:
596:
590:
589:
579:
569:
537:
495:Zhuchengceratops
491:Zhuchengceratops
481:
425:Zhuchengceratops
396:Zhuchengceratops
382:Zhuchengceratops
341:Upper Cretaceous
322:Zhuchengceratops
305:
301:
261:
257:Zhuchengceratops
254:
241:
228:
212:
196:
140:
139:
121:
111:
41:
29:Temporal range:
25:Zhuchengceratops
21:
2159:
2158:
2154:
2153:
2152:
2150:
2149:
2148:
2089:
2088:
2087:
2086:
2048:dinosaurs is a
2029:
2027:
2022:
2014:
2009:
2001:
1996:
1988:
1983:
1974:
1973:
1968:
1959:
1958:
1953:
1940:
1930:
1925:
1897:
1890:
1865:
1824:
1773:
1754:Furcatoceratops
1725:
1706:Albertaceratops
1691:
1665:Menefeeceratops
1628:
1601:Tatankaceratops
1558:
1504:Mercuriceratops
1455:Arrhinoceratops
1422:
1370:
1357:
1340:
1328:
1300:
1280:
1241:
1190:
1157:Montanoceratops
1086:
1081:Graciliceratops
1069:
1056:
1036:Archaeoceratops
1021:
935:
922:
910:Xuanhuaceratops
896:Hualianceratops
874:
855:Albalophosaurus
839:
826:
797:
777:
750:
744:Avemetatarsalia
729:Avemetatarsalia
719:
713:Marginocephalia
675:
674:
664:
659:
629:
628:
613:
598:
597:
593:
539:
538:
529:
524:
512:
455:Late Cretaceous
443:Zhucheng County
436:Late Cretaceous
429:leptoceratopsid
414:
375:Shantungosaurus
334:leptoceratopsid
314:
307:
299:
298:
285:
260:
252:
239:
226:
210:
194:
134:
112:
110:
109:
108:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
36:
35:
33:
27:
17:
12:
11:
5:
2157:
2155:
2147:
2146:
2141:
2136:
2131:
2126:
2121:
2116:
2111:
2106:
2101:
2091:
2090:
2085:
2084:
2077:
2070:
2062:
2059:
2058:
2041:
2024:
2023:
2021:
2020:
2007:
1994:
1981:
1966:
1950:
1948:
1942:
1941:
1936:
1927:
1926:
1924:
1923:
1913:
1906:
1903:
1902:
1899:
1898:
1884:
1882:
1879:
1878:
1875:
1874:
1871:
1870:
1867:
1866:
1864:
1863:
1856:
1849:
1842:
1834:
1832:
1826:
1825:
1823:
1822:
1814:
1807:
1800:
1792:
1790:
1781:
1775:
1774:
1772:
1771:
1764:
1761:Nasutoceratops
1757:
1750:
1743:
1735:
1733:
1727:
1726:
1724:
1723:
1720:Medusaceratops
1716:
1709:
1701:
1699:
1693:
1692:
1690:
1689:
1682:
1675:
1668:
1661:
1654:
1651:Diabloceratops
1646:
1640:
1638:Centrosaurinae
1634:
1633:
1630:
1629:
1627:
1626:
1619:
1612:
1609:Titanoceratops
1605:
1597:
1594:Regaliceratops
1590:
1583:
1576:
1568:
1566:
1564:Triceratopsini
1560:
1559:
1557:
1556:
1549:
1542:
1535:
1528:
1525:Sierraceratops
1521:
1514:
1511:Navajoceratops
1507:
1500:
1493:
1486:
1479:
1472:
1465:
1458:
1451:
1444:
1436:
1430:
1428:Chasmosaurinae
1424:
1423:
1421:
1420:
1413:
1406:
1399:
1392:
1389:Brachyceratops
1385:
1377:
1371:
1366:
1359:
1358:
1353:
1346:
1345:
1342:
1341:
1327:
1325:
1322:
1321:
1318:
1317:
1314:
1313:
1310:
1309:
1306:
1305:
1302:
1301:
1299:
1298:
1297:
1296:
1290:
1288:
1282:
1281:
1279:
1278:
1271:
1268:Turanoceratops
1264:
1255:
1249:
1243:
1242:
1240:
1239:
1232:
1225:
1218:
1209:
1207:
1198:
1192:
1191:
1189:
1188:
1181:
1178:Unescoceratops
1174:
1167:
1160:
1153:
1146:
1143:Ischioceratops
1139:
1136:Gryphoceratops
1132:
1125:
1118:
1111:
1104:
1096:
1094:
1088:
1087:
1085:
1084:
1076:
1070:
1065:
1062:
1061:
1058:
1057:
1055:
1054:
1046:
1039:
1031:
1029:
1023:
1022:
1020:
1019:
1016:Sasayamagnomus
1012:
1005:
998:
991:
983:
976:
969:
966:Auroraceratops
962:
955:
946:
936:
931:
928:
927:
924:
923:
921:
920:
913:
906:
899:
892:
889:Chaoyangsaurus
884:
882:
876:
875:
873:
872:
869:Psittacosaurus
865:
858:
850:
840:
835:
828:
827:
822:
815:
814:
811:
810:
807:
806:
803:
802:
799:
798:
796:
795:
794:
793:
787:
785:
779:
778:
776:
775:
774:
773:
764:
758:
752:
751:
749:
748:
747:
746:
737:
731:
721:
720:
716:
715:
706:
697:
688:
682:
673:
666:
665:
660:
658:
657:
650:
643:
635:
627:
626:
611:
591:
552:(11): e13835.
526:
525:
523:
520:
519:
518:
511:
508:
499:autapomorphies
413:
410:
343:of modern-day
316:
315:
308:
294:
293:
287:
286:
278:
276:
272:
271:
250:
246:
245:
237:
233:
232:
224:
217:
216:
214:Neornithischia
208:
201:
200:
192:
185:
184:
179:
172:
171:
166:
162:
161:
156:
152:
151:
146:
142:
141:
128:
127:
123:
122:
114:
113:
105:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
42:
28:
15:
13:
10:
9:
6:
4:
3:
2:
2156:
2145:
2142:
2140:
2137:
2135:
2132:
2130:
2127:
2125:
2122:
2120:
2117:
2115:
2112:
2110:
2107:
2105:
2102:
2100:
2097:
2096:
2094:
2083:
2078:
2076:
2071:
2069:
2064:
2063:
2057:
2055:
2051:
2047:
2042:
2039:
2035:
2030:
2017:
2012:
2008:
2004:
1999:
1995:
1991:
1986:
1982:
1977:
1971:
1967:
1962:
1956:
1952:
1951:
1949:
1947:
1943:
1939:
1934:
1922:
1914:
1912:
1908:
1907:
1904:
1894:
1888:
1862:
1861:
1857:
1855:
1854:
1850:
1848:
1847:
1843:
1841:
1840:
1839:Achelousaurus
1836:
1835:
1833:
1831:
1827:
1820:
1819:
1818:Styracosaurus
1815:
1813:
1812:
1808:
1806:
1805:
1801:
1799:
1798:
1794:
1793:
1791:
1789:
1788:Centrosaurini
1785:
1782:
1780:
1779:Eucentrosaura
1776:
1770:
1769:
1765:
1763:
1762:
1758:
1756:
1755:
1751:
1749:
1748:
1744:
1742:
1741:
1737:
1736:
1734:
1732:
1728:
1722:
1721:
1717:
1715:
1714:
1710:
1708:
1707:
1703:
1702:
1700:
1698:
1694:
1688:
1687:
1683:
1681:
1680:
1679:Wendiceratops
1676:
1674:
1673:
1669:
1667:
1666:
1662:
1660:
1659:
1655:
1653:
1652:
1648:
1647:
1644:
1641:
1639:
1635:
1625:
1624:
1620:
1618:
1617:
1613:
1611:
1610:
1606:
1603:
1602:
1598:
1596:
1595:
1591:
1589:
1588:
1584:
1582:
1581:
1577:
1575:
1574:
1573:Eotriceratops
1570:
1569:
1567:
1565:
1561:
1555:
1554:
1550:
1548:
1547:
1543:
1541:
1540:
1536:
1534:
1533:
1529:
1527:
1526:
1522:
1520:
1519:
1518:Pentaceratops
1515:
1513:
1512:
1508:
1506:
1505:
1501:
1499:
1498:
1497:Kosmoceratops
1494:
1492:
1491:
1487:
1485:
1484:
1480:
1478:
1477:
1473:
1471:
1470:
1469:Bravoceratops
1466:
1464:
1463:
1462:Bisticeratops
1459:
1457:
1456:
1452:
1450:
1449:
1448:Anchiceratops
1445:
1443:
1442:
1441:Agujaceratops
1438:
1437:
1434:
1431:
1429:
1425:
1419:
1418:
1414:
1412:
1411:
1407:
1405:
1404:
1400:
1398:
1397:
1393:
1391:
1390:
1386:
1384:
1383:
1379:
1378:
1375:
1372:
1369:
1364:
1360:
1356:
1351:
1347:
1338:
1332:
1294:
1293:
1292:
1291:
1289:
1287:
1283:
1277:
1276:
1272:
1270:
1269:
1265:
1262:
1261:
1257:
1256:
1253:
1250:
1248:
1247:Ceratopsoidea
1244:
1238:
1237:
1236:Protoceratops
1233:
1231:
1230:
1229:Breviceratops
1226:
1224:
1223:
1219:
1216:
1215:
1211:
1210:
1208:
1206:
1202:
1199:
1197:
1193:
1187:
1186:
1182:
1180:
1179:
1175:
1173:
1172:
1171:Udanoceratops
1168:
1166:
1165:
1164:Prenoceratops
1161:
1159:
1158:
1154:
1152:
1151:
1150:Leptoceratops
1147:
1145:
1144:
1140:
1138:
1137:
1133:
1131:
1130:
1126:
1124:
1123:
1119:
1117:
1116:
1112:
1110:
1109:
1108:Bainoceratops
1105:
1103:
1102:
1098:
1097:
1095:
1093:
1089:
1083:
1082:
1078:
1077:
1074:
1071:
1068:
1063:
1052:
1051:
1047:
1045:
1044:
1043:Helioceratops
1040:
1038:
1037:
1033:
1032:
1030:
1028:
1024:
1018:
1017:
1013:
1011:
1010:
1009:Mosaiceratops
1006:
1004:
1003:
999:
997:
996:
992:
989:
988:
984:
982:
981:
980:Koreaceratops
977:
975:
974:
970:
968:
967:
963:
961:
960:
956:
953:
952:
948:
947:
944:
940:
937:
934:
933:Neoceratopsia
929:
919:
918:
914:
912:
911:
907:
905:
904:
900:
898:
897:
893:
891:
890:
886:
885:
883:
881:
877:
871:
870:
866:
864:
863:
859:
857:
856:
852:
851:
848:
844:
841:
838:
833:
829:
825:
820:
816:
791:
790:
789:
788:
786:
784:
780:
772:
768:
767:
766:
765:
762:
759:
757:
753:
745:
741:
740:
739:
738:
735:
732:
730:
726:
722:
714:
710:
707:
705:
701:
698:
696:
692:
689:
687:
683:
681:
677:
676:
671:
667:
663:
656:
651:
649:
644:
642:
637:
636:
633:
622:
618:
614:
608:
604:
603:
595:
592:
587:
583:
578:
573:
568:
563:
559:
555:
551:
547:
543:
536:
534:
532:
528:
521:
517:
514:
513:
509:
507:
504:
500:
496:
492:
487:
485:
480:
476:
472:
468:
464:
460:
456:
452:
451:Wangshi Group
448:
444:
440:
437:
433:
430:
427:is a derived
426:
418:
411:
409:
407:
406:
405:Udanoceratops
401:
400:Leptoceratops
397:
393:
392:
391:Leptoceratops
387:
383:
379:
377:
376:
371:
370:Wangshi Group
367:
363:
359:
358:
354:
350:
346:
342:
338:
335:
332:
328:
324:
323:
312:
306:
304:
295:
292:
291:Binomial name
288:
284:
283:
277:
274:
273:
269:
265:
259:
258:
251:
248:
247:
244:
238:
235:
234:
231:
225:
222:
219:
218:
215:
209:
206:
203:
202:
199:
193:
190:
187:
186:
183:
180:
177:
174:
173:
170:
167:
164:
163:
160:
157:
154:
153:
150:
147:
144:
143:
138:
133:
129:
124:
120:
115:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
40:
32:
31:Maastrichtian
26:
22:
19:
2054:expanding it
2043:
2028:
1945:
1858:
1853:Stellasaurus
1851:
1844:
1837:
1816:
1809:
1804:Coronosaurus
1802:
1797:Centrosaurus
1795:
1766:
1759:
1752:
1745:
1738:
1718:
1713:Lokiceratops
1711:
1704:
1686:Xenoceratops
1684:
1677:
1672:Sinoceratops
1670:
1663:
1656:
1649:
1621:
1614:
1607:
1599:
1592:
1585:
1580:Nedoceratops
1578:
1571:
1553:Vagaceratops
1551:
1546:Utahceratops
1544:
1539:Terminocavus
1537:
1530:
1523:
1516:
1509:
1502:
1495:
1490:Judiceratops
1488:
1481:
1476:Chasmosaurus
1474:
1467:
1460:
1453:
1446:
1439:
1415:
1408:
1401:
1394:
1387:
1380:
1368:Ceratopsidae
1355:Ceratopsidae
1286:Ceratopsidae
1275:Zuniceratops
1273:
1266:
1260:Ajkaceratops
1258:
1234:
1227:
1222:Bagaceratops
1220:
1214:Ajkaceratops
1212:
1196:Coronosauria
1184:
1183:
1176:
1169:
1162:
1155:
1148:
1141:
1134:
1127:
1120:
1113:
1106:
1101:Asiaceratops
1099:
1079:
1067:Euceratopsia
1050:Yamaceratops
1048:
1041:
1034:
1014:
1007:
1002:Microceratus
1000:
995:Liaoceratops
993:
985:
978:
971:
964:
957:
951:Ajkaceratops
949:
915:
908:
901:
894:
887:
867:
860:
853:
771:Ornithischia
756:Ornithischia
708:
704:Ornithischia
699:
690:
601:
594:
549:
545:
494:
490:
488:
483:
462:
424:
423:
403:
399:
395:
389:
386:Ceratopsidae
381:
380:
373:
361:
356:
355:
348:
321:
320:
319:
310:
302:
297:
281:
280:
263:
256:
255:
220:
204:
198:Ornithischia
188:
175:
24:
18:
2046:ceratopsian
1970:Wikispecies
1846:Einiosaurus
1740:Avaceratops
1623:Triceratops
1587:Ojoceratops
1410:Monoclonius
1122:Ferrisaurus
987:Kulceratops
432:ceratopsian
337:ceratopsian
2093:Categories
1909:See also:
1616:Torosaurus
1532:Spiclypeus
1382:Agathaumas
1295:see below↓
1115:Cerasinops
903:Stenopelix
837:Ceratopsia
824:Ceratopsia
792:see below↓
783:Ceratopsia
695:Dinosauria
662:Ceratopsia
522:References
484:inexpectus
230:Ceratopsia
182:Dinosauria
678:Kingdom:
621:985402380
503:taxonomic
479:ceraptops
471:Latinized
453:, of the
275:Species:
155:Kingdom:
149:Eukaryota
1990:47459989
1955:Wikidata
1921:Category
1911:Timeline
1417:Polyonax
1403:Dysganus
1396:Ceratops
959:Aquilops
686:Chordata
684:Phylum:
680:Animalia
586:21079798
546:PLOS ONE
510:See also
467:holotype
366:holotype
362:ceratops
353:binomial
236:Family:
169:Chordata
165:Phylum:
159:Animalia
145:Domain:
37:70
2003:8463228
1961:Q170121
1811:Spinops
1129:Gremlin
917:Yinlong
577:2973951
554:Bibcode
331:extinct
300:†
279:†
249:Genus:
106:↓
2016:172379
619:
609:
584:
574:
439:period
349:et al.
313:, 2010
311:et al.
264:et al.
709:Clade
700:Clade
691:Clade
475:Greek
447:China
345:China
327:genus
325:is a
221:Clade
205:Clade
189:Clade
176:Clade
2050:stub
1998:GBIF
769:see
742:see
617:OCLC
607:ISBN
582:PMID
459:2010
402:and
268:2010
45:PreꞒ
1985:EoL
973:Beg
572:PMC
562:doi
329:of
309:Xu
262:Xu
2095::
2013::
2000::
1987::
1972::
1957::
711::
702::
693::
615:.
580:.
570:.
560:.
548:.
544:.
530:^
445:,
378:.
266:,
223::
207::
191::
178::
95:Pg
39:Ma
2081:e
2074:t
2067:v
2056:.
1821:?
1604:?
1263:?
1217:?
1053:?
990:?
954:?
654:e
647:t
640:v
623:.
588:.
564::
556::
550:5
473:-
253:†
240:†
227:†
211:†
195:†
100:N
90:K
85:J
80:T
75:P
70:C
65:D
60:S
55:O
50:Ꞓ
34:~
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