675:, the upper jaws, were 11.9 centimetres (4.7 in) long and contained about eighteen small teeth on each side. The outside of the maxilla was covered by a large loreal osteoderm, overlapping the snout edge. A large lacrimal plate is present behind it. The rim above the eye socket has two osteoderms, each forming a separate peak as with Asian ankylosaurids. The rear edge of the skull is ornamented by small osteoderms, oriented somewhat to the midline and gradually increasing in size towards the outside of the skull. The squamosal horns on the back corners of the skull were very high, thick and forward-curving. The shape of the cheek horns is unknown because of damage inflicted during the fossilisation process.
2665:
780:
731:
638:
457:
2660:
150:
129:
2688:
746:, scutes of bone that sit inside of the skin. Apart from the osteoderms that are part of the skull, two cervical halfrings were present to protect the upper side of the neck and rows of oval scutes probably ran along the back and sides of the torso; the latter are only known as detached specimens. All osteoderms found have a surface that is densely pitted but lacks veins.
628:
in several traits. The edge on the squamosal horns is sharper. The points of these horns are more curved to the front instead of to below. The osteoderm peaks above the eye sockets are sharper. The caputegulae are more irregular in form, at best lightly convex instead of cone-shaped, have a more
771:
ankylosaurian side spikes and was thus identified as a lateral rump osteoderm. Some small ossicles were discovered also, round scutes of between two and four centimetres in cross-section, with a conical or flat shape and featuring a pitted surface. Their original position cannot be determined.
705:
At the underside of the skull, the paired front praemaxillae form a bony secondary palate, with a concave surface. Each praemaxilla is pierced by two foramina. Both elements are at the snout tip separated by a gap, ending about thirty-five millimetres in front of the tip of the narrow and flat
754:
the segments consisted of osteoderms that were in-between an oval and a rectangular shape. These osteoderms were keeled, featuring a high cutting edge. The osteoderms were between 146 and 169 millimetres long, between 58 and 104 millimetres wide and their keels varied between 2.4 centimetres
770:
Several unconnected osteoderms have been discovered. One of these might originally have been part of a halfring. A second one was seen as originating from the pectoral, upper breast, region. A third, measuring 118 by 68 millimetres, had a narrow keel and the shark fin shape typical of the
749:
The cervical halfrings of ankylosaurids typically consisted of six fused segments each, that were paired per side: one at the top, a second segment at the upper side and a third one at the lower side. These segments were fused to an underlying continuous bone ring. In
762:
Specimen NMMNH P-66930, a first cervical halfring, shared several traits with its counterpart of the holotype: high narrow keels, interstitial osteoderms in the middle and a lower side osteoderm that does not wrap around the lower edge. It was therefore referred to
666:
of the snout is exceptionally large, covering about half of the snout width whereas in other species possessing such a structure this is at most 40%. It is also triangular instead of hexagonal, which is the usual shape. Behind this central plate rows of smaller
606:
had deep, anteriorly curved squamosal horns (measuring 10.2 centimetres (4.0 in) from the base to the tip). The rear base of the skull featured three deep grooves. A further possible distinguishing trait is the mixture of concave, flat and convex
653:
The skull has no constriction in front of the eye sockets and its widest point is formed by the squamosal horns on the top rear corners. The skull is rather flat and this is only to a small degree caused by compression of the fossil. The
755:(0.94 in) and 10.9 centimetres (4.3 in) tall. Where the middle segments touched each other on the midline, an irregular punctuated row of small trapezium-shaped "interstitial" osteoderms was present within the
1510:
Sullivan, Robert M.; Robinson, Robert F. (2015). "A complete first cervical half ring (Ankylosauridae, Dinosauria) from the Upper
Cretaceous Kirtland Formation (Hunter Wash Member) San Juan Basin, New Mexico".
436:, where the fossils were found. Multiple specimens have been described to date, though the fossils are mostly from the front part of the animal. Its closest relative appears to be either
759:. At the front of the suture of the first cervical halfring, a small cone-shaped interstitial osteoderm protruded. Such interstitial elements were absent with the side segments.
1447:"A New Ankylosaurid Dinosaur from the Upper Cretaceous (Kirtlandian) of New Mexico with Implications for Ankylosaurid Diversity in the Upper Cretaceous of Western North America"
837:
were forced to be sister species was just one evolutionary step longer, i.e. requiring the presence of but a single additional shared trait, and thus only slightly less likely.
579:
is known from a complete skull and specimens of first cervical half ring, so its head and cervical armor may have been separated from the body before the specimen was buried.
2819:
829:
which lived in the same time and area. The latter being recovered as a relative of East-Asian ankylosaurids. However, the authors indicated that an alternative tree in which
500:
NMMNH P-64484 found from the De-na-zin Member and consisting of a complete skull lacking the lower jaws, parts of the first two cervical half-rings, and a number of partial
698:. To the front the basioccipital contacts the triangular basisphenoid of the underside of the braincase. The two elements are not fully fused but in side view show a clear
662:
were broad and square, and covered in a rather flat cranial ornamentation. The nasal vestibules are oriented towards the front and shaped like flat ovals. The central
465:
1445:
Arbour, Victoria M.; Burns, Michael E.; Sullivan, Robert M.; Lucas, Spencer G.; Cantrell, Amanda K.; Fry, Joshua; Suazo, Thomas L. (24 September 2014).
2824:
2755:
2839:
2664:
1357:. Provincialism in the sense of a faunal separation between the northern and southern regions of Laramidia cannot be deduced from the fact that
2834:
1619:
694:
element. More to below this basioccipital shows three deep parallel grooves, bordered by four rims. The middle groove has the opening of the
2829:
2681:
1394:
408:
The genus is named after the Zia sun symbol, a stylized sun with four groups of rays, having religious significance to the
2844:
2659:
596:
504:; and a referred specimen NMMNH P-66930 from the older Hunter Wash Member, consisting of a first cervical half-ring. By
469:
508:
of ash layers the age of the holotype skull has been determined at between 72.98 and 72.62 million years old, the late
671:
run backwards; they have a diameter of about three centimetres each and are rectangular, pentagonal or hexagonal. The
565:
429:
1558:
Arbour, V. M.; Currie, P. J. (2015). "Systematics, phylogeny and palaeobiogeography of the ankylosaurid dinosaurs".
1612:
489:
678:
At the rear of the skull, the paroccipital processes are fused with the quadrates. The contact with the neck, the
2814:
149:
1710:
557:
539:
473:
425:
409:
2809:
2708:
1327:
2781:
1737:
1605:
845:
779:
730:
637:
456:
2691:
1567:
1464:
1306:, Arbour and Currie suggested that earlier North American ankylosaurids had gone extinct by the late
340:
308:
2097:
1994:
1530:
710:
at the midline. Behind the praemaxillae, the internal wings of the maxillae form the edges of the
505:
1833:
1583:
543:
485:
481:
374:
370:
144:
2786:
512:. It was initially prepared by Amanda K. Cantrell and Thomas L. Suazo; later by Larry Rinehart.
2768:
2590:
1854:
1492:
1318:. They propose that ankylosaurids subsequently recolonized North America from Asia during the
1092:
824:
679:
616:
444:
2773:
2055:
2002:
1840:
1575:
1482:
1472:
756:
699:
413:
378:
1298:
and other Late
Cretaceous North American ankylosaurids are grouped with Asian genera, in a
2520:
2484:
2424:
2390:
2383:
2276:
2217:
2089:
1986:
1902:
1862:
1695:
1299:
910:
516:
394:
333:
301:
1571:
1468:
686:, the bones of the upper rear skull sides, the exoccipitalia, do not contribute to this
602:(middle bone plate), of the snout was large, prominent and roughly triangular in shape.
2746:
2583:
2569:
2555:
2463:
2436:
2332:
2313:
2228:
2174:
2039:
2009:
1979:
1942:
1918:
1487:
1446:
1175:
1149:
855:
811:
797:
785:
591:
535:
433:
362:
268:
255:
128:
2803:
2546:
2456:
2449:
2401:
2345:
2292:
2258:
2159:
2105:
2080:
2062:
1965:
1911:
1884:
1876:
1587:
1303:
1030:
869:
715:
520:
281:
79:
17:
822:
also containing other
American ankylosaurids. However it was not closely related to
629:
rectangular outline, instead of a rounded one, and are separated by deeper grooves.
2576:
2562:
2491:
2375:
2299:
2244:
2195:
2167:
2152:
2139:
2112:
2031:
2016:
1972:
1952:
1819:
1801:
1779:
1749:
1722:
1670:
1628:
1371:
1132:
1044:
927:
524:
390:
316:
242:
210:
1579:
2731:
1477:
2740:
2611:
2528:
2477:
2417:
2368:
2210:
2188:
2120:
2069:
1869:
1847:
1679:
1377:
1199:
893:
802:
655:
597:
438:
226:
54:
614:
From the
Kirtland Formation a second ankylosaur is known from limited remains,
2604:
2597:
2535:
2410:
2361:
2353:
2306:
2251:
2237:
2203:
2181:
2047:
1891:
1826:
1315:
1311:
999:
743:
659:
386:
99:
64:
2618:
2498:
2285:
2265:
2127:
2024:
1926:
1354:
1319:
1075:
958:
841:
815:
691:
509:
501:
421:
382:
161:
104:
48:
41:
1496:
2725:
1661:
1652:
1451:
1323:
497:
194:
181:
136:
94:
89:
74:
69:
59:
2760:
2513:
2505:
1362:
982:
687:
672:
496:
was identified as a new species from a number of fossils including the
355:
109:
84:
818:
analysis that was part of the describing paper. It thus was part of a
552:
1646:
1307:
807:
711:
366:
171:
2702:
682:, is kidney-shaped with a smooth surface surrounded by a groove. In
484:
in 2011. The specimens were found in the Hunter Wash and De-na-zin
2470:
819:
778:
729:
707:
636:
547:
455:
398:
358:
2632:
1460:
2706:
1639:
1601:
1597:
722:
though a triangular depression is present at their undersides.
714:, the internal nostrils. To the rear of the vomers, triangular
27:
Genus of ankylosaurid dinosaur from the Late
Cretaceous period
690:
which is fully formed by the basioccipital, the lower rear
1553:
1551:
1513:
New Mexico Museum of
Natural History and Science Bulletin
523:, Amanda Cantrell, Joshua Fry and Thomas Suazo named the
556:("shield"), referring to ankylosaurids' osteoderms. The
1531:"New dinosaur from New Mexico has relatives in Alberta"
1353:
and Asian forms indicates a migration between Asia and
814:
in several of the evolutionary trees rendered by the
323:
2715:
2545:
2435:
2400:
2331:
2275:
2227:
2138:
2079:
1951:
1940:
1901:
1799:
1748:
1721:
1694:
389:. It was named in 2014, in a research paper led by
848:of Ankylosaurinae conducted by Arbour and Currie:
658:portion of the skull was lightly convex, and the
369:have been found in the Hunter Wash and De-na-zin
466:New Mexico Museum of Natural History and Science
1385:stems from younger layers than these animals.
1613:
448:, depending on what cladistic model is used.
420:(Latin), a small shield, in reference to the
8:
718:are located. These are not pierced by large
412:of New Mexico, and the iconic symbol on the
1440:
1438:
1436:
1434:
1432:
1430:
2820:Late Cretaceous dinosaurs of North America
2703:
2441:
2337:
2144:
1957:
1948:
1811:
1807:
1796:
1727:
1700:
1691:
1636:
1620:
1606:
1598:
1428:
1426:
1424:
1422:
1420:
1418:
1416:
1414:
1412:
1410:
127:
31:
1486:
1476:
1406:
1326:ages of the Late Cretaceous, and there
590:showed three unique derived traits or
1330:once more, leading to genera such as
783:Braincase (A, B) compared to that of
568:, where the fossils were discovered.
460:Location and stratigraphy of the find
7:
1560:Journal of Systematic Palaeontology
1191:
1124:
1117:
1067:
1060:
1036:
974:
950:
943:
885:
861:
851:
397:. There is a single species in the
519:, Michael Burns, Robert Sullivan,
25:
800:, as possibly closely related to
2825:Cretaceous geology of New Mexico
2687:
2686:
2663:
2658:
649:at the middle top and lower left
424:found on all ankylosaurids. The
148:
1395:Timeline of ankylosaur research
2840:Taxa named by Spencer G. Lucas
702:, obliquely running to below.
1:
2835:Fossil taxa described in 2014
1580:10.1080/14772019.2015.1059985
1478:10.1371/journal.pone.0108804
538:, a religious image for the
470:State Museum of Pennsylvania
2861:
2830:Paleontology in New Mexico
645:pattern in ankylosaurids;
490:Bisti/De-Na-Zin Wilderness
2676:
2656:
2444:
2340:
2147:
1960:
1814:
1810:
1795:
1776:
1730:
1703:
1690:
1635:
1302:which the authors dubbed
1213:
1196:
1189:
1172:
1146:
1129:
1122:
1115:
1089:
1072:
1065:
1058:
1041:
1034:
996:
979:
972:
955:
948:
941:
924:
907:
890:
883:
866:
859:
734:Three cervical half-rings
476:, discovered a number of
322:
315:
145:Scientific classification
143:
135:
126:
34:
1365:shared by its relatives
488:of the formation in the
1361:was not present in the
474:Robert Michael Sullivan
464:An expedition from the
790:
735:
696:foramen basioccipitale
650:
542:and an element of the
534:) was named after the
461:
2782:Paleobiology Database
1349:The relation between
846:phylogenetic analysis
782:
733:
640:
588:Ziapelta sanjuanensis
583:Distinguishing traits
528:Ziapelta sanjuanensis
517:Victoria Megan Arbour
459:
404:Ziapelta sanjuanensis
328:Ziapelta sanjuanensis
18:Ziapelta sanjuanensis
2845:Ornithischian genera
1572:2016JSPal..14..385A
1469:2014PLoSO...9j8804A
1314:ages of the Middle
844:is based on a 2015
428:is in reference to
416:of New Mexico, and
796:was placed in the
791:
736:
651:
611:on the skull top.
544:flag of New Mexico
482:Kirtland Formation
462:
375:Kirtland Formation
2797:
2796:
2769:Open Tree of Life
2709:Taxon identifiers
2700:
2699:
2672:
2671:
2654:
2653:
2650:
2649:
2646:
2645:
2642:
2641:
2591:Nodocephalosaurus
2327:
2326:
2323:
2322:
1936:
1935:
1855:Serendipaceratops
1791:
1790:
1787:
1786:
1772:
1771:
1768:
1767:
1351:Nodocephalosaurus
1291:
1290:
1282:
1281:
1273:
1272:
1264:
1263:
1255:
1254:
1246:
1245:
1237:
1236:
1228:
1227:
1161:
1160:
1104:
1103:
1093:Nodocephalosaurus
1020:
1019:
1011:
1010:
835:Nodocephalosaurus
825:Nodocephalosaurus
742:was protected by
680:occipital condyle
626:Nodocephalosaurus
617:Nodocephalosaurus
445:Nodocephalosaurus
347:
346:
311:
16:(Redirected from
2852:
2815:Campanian genera
2790:
2789:
2777:
2776:
2764:
2763:
2751:
2750:
2749:
2736:
2735:
2734:
2704:
2690:
2689:
2667:
2662:
2442:
2338:
2145:
2056:Propanoplosaurus
2003:Glyptodontopelta
1958:
1949:
1841:Priodontognathus
1812:
1808:
1797:
1728:
1701:
1692:
1685:
1684:
1637:
1622:
1615:
1608:
1599:
1592:
1591:
1555:
1546:
1545:
1543:
1541:
1527:
1521:
1520:
1507:
1501:
1500:
1490:
1480:
1442:
1192:
1125:
1118:
1068:
1061:
1037:
975:
951:
944:
886:
862:
852:
599:
379:Upper Cretaceous
325:
300:
293:
280:
267:
254:
241:
225:
209:
153:
152:
131:
121:
51:
40:Temporal range:
32:
21:
2860:
2859:
2855:
2854:
2853:
2851:
2850:
2849:
2800:
2799:
2798:
2793:
2785:
2780:
2772:
2767:
2759:
2754:
2745:
2744:
2739:
2730:
2729:
2724:
2711:
2701:
2696:
2668:
2638:
2541:
2521:Tianzhenosaurus
2485:Minotaurasaurus
2431:
2425:Zhongyuansaurus
2396:
2391:Zhejiangosaurus
2384:Liaoningosaurus
2319:
2277:Struthiosaurini
2271:
2223:
2218:Tatankacephalus
2134:
2090:Gargoyleosaurus
2075:
1987:Gargoyleosaurus
1945:
1932:
1903:Parankylosauria
1897:
1863:Sinankylosaurus
1804:
1783:
1764:
1744:
1717:
1711:Avemetatarsalia
1696:Avemetatarsalia
1686:
1642:
1641:
1631:
1626:
1596:
1595:
1557:
1556:
1549:
1539:
1537:
1529:
1528:
1524:
1509:
1508:
1504:
1444:
1443:
1408:
1403:
1391:
1347:
1292:
1283:
1274:
1265:
1256:
1247:
1238:
1229:
1162:
1105:
1021:
1012:
911:Zhejiangosaurus
777:
765:Z. sanjuanensis
728:
635:
601:
585:
574:
566:San Juan County
494:Z. sanjuanensis
492:in New Mexico.
480:fossils in the
478:Z. sanjuanensis
454:
430:San Juan County
395:Victoria Arbour
343:
331:
299:
291:
278:
265:
252:
239:
223:
207:
147:
122:
120:
119:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
47:74.5–72.6
46:
45:
38:
28:
23:
22:
15:
12:
11:
5:
2858:
2856:
2848:
2847:
2842:
2837:
2832:
2827:
2822:
2817:
2812:
2802:
2801:
2795:
2794:
2792:
2791:
2778:
2765:
2752:
2737:
2721:
2719:
2713:
2712:
2707:
2698:
2697:
2695:
2694:
2684:
2677:
2674:
2673:
2670:
2669:
2657:
2655:
2652:
2651:
2648:
2647:
2644:
2643:
2640:
2639:
2637:
2636:
2629:
2622:
2615:
2608:
2601:
2594:
2587:
2584:Euoplocephalus
2580:
2573:
2570:Anodontosaurus
2566:
2559:
2556:Akainacephalus
2551:
2549:
2543:
2542:
2540:
2539:
2532:
2525:
2517:
2510:
2502:
2495:
2488:
2481:
2474:
2467:
2464:Crichtonsaurus
2460:
2453:
2445:
2439:
2437:Ankylosaurinae
2433:
2432:
2430:
2429:
2421:
2414:
2406:
2404:
2398:
2397:
2395:
2394:
2387:
2380:
2372:
2365:
2358:
2350:
2341:
2335:
2333:Ankylosauridae
2329:
2328:
2325:
2324:
2321:
2320:
2318:
2317:
2314:Struthiosaurus
2310:
2303:
2296:
2289:
2281:
2279:
2273:
2272:
2270:
2269:
2262:
2255:
2248:
2241:
2233:
2231:
2229:Panoplosaurini
2225:
2224:
2222:
2221:
2214:
2207:
2200:
2192:
2185:
2178:
2175:Niobrarasaurus
2171:
2164:
2156:
2148:
2142:
2136:
2135:
2133:
2132:
2124:
2117:
2109:
2102:
2094:
2085:
2083:
2077:
2076:
2074:
2073:
2066:
2059:
2052:
2044:
2040:Polacanthoides
2036:
2028:
2021:
2013:
2010:Horshamosaurus
2006:
1999:
1991:
1983:
1980:Dongyangopelta
1976:
1969:
1961:
1955:
1946:
1943:Euankylosauria
1941:
1938:
1937:
1934:
1933:
1931:
1930:
1923:
1919:Kunbarrasaurus
1915:
1907:
1905:
1899:
1898:
1896:
1895:
1888:
1881:
1873:
1866:
1859:
1851:
1844:
1837:
1830:
1823:
1815:
1805:
1800:
1793:
1792:
1789:
1788:
1785:
1784:
1777:
1774:
1773:
1770:
1769:
1766:
1765:
1763:
1762:
1761:
1760:
1754:
1752:
1746:
1745:
1743:
1742:
1741:
1740:
1731:
1725:
1719:
1718:
1716:
1715:
1714:
1713:
1704:
1698:
1688:
1687:
1683:
1682:
1673:
1664:
1655:
1649:
1640:
1633:
1632:
1627:
1625:
1624:
1617:
1610:
1602:
1594:
1593:
1547:
1522:
1502:
1405:
1404:
1402:
1399:
1398:
1397:
1390:
1387:
1367:Euoplocephalus
1346:
1345:Paleogeography
1343:
1332:Euoplocephalus
1289:
1288:
1285:
1284:
1280:
1279:
1276:
1275:
1271:
1270:
1267:
1266:
1262:
1261:
1258:
1257:
1253:
1252:
1249:
1248:
1244:
1243:
1240:
1239:
1235:
1234:
1231:
1230:
1226:
1225:
1222:
1221:
1212:
1209:
1208:
1205:
1204:
1195:
1190:
1188:
1185:
1184:
1181:
1180:
1176:Euoplocephalus
1171:
1168:
1167:
1164:
1163:
1159:
1158:
1155:
1154:
1150:Anodontosaurus
1145:
1142:
1141:
1138:
1137:
1128:
1123:
1121:
1116:
1114:
1111:
1110:
1107:
1106:
1102:
1101:
1098:
1097:
1088:
1085:
1084:
1081:
1080:
1071:
1066:
1064:
1059:
1057:
1054:
1053:
1050:
1049:
1040:
1035:
1033:
1027:
1026:
1023:
1022:
1018:
1017:
1014:
1013:
1009:
1008:
1005:
1004:
995:
992:
991:
988:
987:
978:
973:
971:
968:
967:
964:
963:
954:
949:
947:
942:
940:
937:
936:
933:
932:
923:
920:
919:
916:
915:
906:
903:
902:
899:
898:
889:
884:
882:
879:
878:
875:
874:
865:
860:
858:
856:Ankylosaurinae
850:
840:The following
812:sister species
798:Ankylosauridae
786:Euoplocephalus
776:
775:Classification
773:
727:
724:
716:palatine bones
634:
631:
595:
592:autapomorphies
584:
581:
573:
570:
536:Zia sun symbol
453:
450:
434:San Juan basin
345:
344:
332:
320:
319:
313:
312:
289:
285:
284:
276:
272:
271:
269:Ankylosaurinae
263:
259:
258:
256:Ankylosauridae
250:
246:
245:
237:
230:
229:
221:
214:
213:
205:
198:
197:
192:
185:
184:
179:
175:
174:
169:
165:
164:
159:
155:
154:
141:
140:
133:
132:
124:
123:
115:
114:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
52:
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
2857:
2846:
2843:
2841:
2838:
2836:
2833:
2831:
2828:
2826:
2823:
2821:
2818:
2816:
2813:
2811:
2810:Ankylosaurids
2808:
2807:
2805:
2788:
2783:
2779:
2775:
2770:
2766:
2762:
2757:
2753:
2748:
2742:
2738:
2733:
2727:
2723:
2722:
2720:
2718:
2714:
2710:
2705:
2693:
2685:
2683:
2679:
2678:
2675:
2666:
2661:
2635:
2634:
2630:
2628:
2627:
2623:
2621:
2620:
2616:
2614:
2613:
2609:
2607:
2606:
2602:
2600:
2599:
2595:
2593:
2592:
2588:
2586:
2585:
2581:
2579:
2578:
2574:
2572:
2571:
2567:
2565:
2564:
2560:
2558:
2557:
2553:
2552:
2550:
2548:
2547:Ankylosaurini
2544:
2538:
2537:
2533:
2531:
2530:
2526:
2523:
2522:
2518:
2516:
2515:
2511:
2508:
2507:
2503:
2501:
2500:
2496:
2494:
2493:
2489:
2487:
2486:
2482:
2480:
2479:
2475:
2473:
2472:
2468:
2466:
2465:
2461:
2459:
2458:
2457:Crichtonpelta
2454:
2452:
2451:
2450:Bissektipelta
2447:
2446:
2443:
2440:
2438:
2434:
2427:
2426:
2422:
2420:
2419:
2415:
2413:
2412:
2408:
2407:
2405:
2403:
2402:Shamosaurinae
2399:
2393:
2392:
2388:
2386:
2385:
2381:
2378:
2377:
2373:
2371:
2370:
2366:
2364:
2363:
2359:
2356:
2355:
2351:
2348:
2347:
2346:Ahshislepelta
2343:
2342:
2339:
2336:
2334:
2330:
2316:
2315:
2311:
2309:
2308:
2304:
2302:
2301:
2297:
2295:
2294:
2293:Hungarosaurus
2290:
2288:
2287:
2283:
2282:
2280:
2278:
2274:
2268:
2267:
2263:
2261:
2260:
2259:Panoplosaurus
2256:
2254:
2253:
2249:
2247:
2246:
2242:
2240:
2239:
2235:
2234:
2232:
2230:
2226:
2220:
2219:
2215:
2213:
2212:
2208:
2206:
2205:
2201:
2198:
2197:
2193:
2191:
2190:
2186:
2184:
2183:
2179:
2177:
2176:
2172:
2170:
2169:
2165:
2162:
2161:
2160:Ahshislepelta
2157:
2155:
2154:
2150:
2149:
2146:
2143:
2141:
2137:
2130:
2129:
2125:
2123:
2122:
2118:
2115:
2114:
2110:
2108:
2107:
2106:Hoplitosaurus
2103:
2100:
2099:
2095:
2092:
2091:
2087:
2086:
2084:
2082:
2081:Polacanthinae
2078:
2072:
2071:
2067:
2065:
2064:
2063:Rhadinosaurus
2060:
2058:
2057:
2053:
2050:
2049:
2045:
2042:
2041:
2037:
2034:
2033:
2029:
2027:
2026:
2022:
2019:
2018:
2014:
2012:
2011:
2007:
2005:
2004:
2000:
1997:
1996:
1992:
1989:
1988:
1984:
1982:
1981:
1977:
1975:
1974:
1970:
1968:
1967:
1966:Acanthopholis
1963:
1962:
1959:
1956:
1954:
1950:
1947:
1944:
1939:
1929:
1928:
1924:
1921:
1920:
1916:
1914:
1913:
1912:Antarctopelta
1909:
1908:
1906:
1904:
1900:
1894:
1893:
1889:
1887:
1886:
1885:Tianchisaurus
1882:
1879:
1878:
1877:Stegosaurides
1874:
1872:
1871:
1867:
1865:
1864:
1860:
1857:
1856:
1852:
1850:
1849:
1845:
1843:
1842:
1838:
1836:
1835:
1831:
1829:
1828:
1824:
1822:
1821:
1817:
1816:
1813:
1809:
1806:
1803:
1798:
1794:
1782:
1781:
1775:
1758:
1757:
1756:
1755:
1753:
1751:
1747:
1739:
1735:
1734:
1733:
1732:
1729:
1726:
1724:
1720:
1712:
1708:
1707:
1706:
1705:
1702:
1699:
1697:
1693:
1689:
1681:
1677:
1674:
1672:
1668:
1665:
1663:
1659:
1656:
1654:
1650:
1648:
1644:
1643:
1638:
1634:
1630:
1623:
1618:
1616:
1611:
1609:
1604:
1603:
1600:
1589:
1585:
1581:
1577:
1573:
1569:
1565:
1561:
1554:
1552:
1548:
1536:
1532:
1526:
1523:
1518:
1514:
1506:
1503:
1498:
1494:
1489:
1484:
1479:
1474:
1470:
1466:
1462:
1458:
1454:
1453:
1448:
1441:
1439:
1437:
1435:
1433:
1431:
1429:
1427:
1425:
1423:
1421:
1419:
1417:
1415:
1413:
1411:
1407:
1400:
1396:
1393:
1392:
1388:
1386:
1384:
1380:
1379:
1374:
1373:
1368:
1364:
1360:
1356:
1352:
1344:
1342:
1341:
1337:
1333:
1329:
1325:
1321:
1317:
1313:
1309:
1305:
1304:Ankylosaurini
1301:
1297:
1287:
1286:
1278:
1277:
1269:
1268:
1260:
1259:
1251:
1250:
1242:
1241:
1233:
1232:
1224:
1223:
1220:
1219:
1218:
1211:
1210:
1207:
1206:
1203:
1202:
1201:
1194:
1193:
1187:
1186:
1183:
1182:
1179:
1178:
1177:
1170:
1169:
1166:
1165:
1157:
1156:
1153:
1152:
1151:
1144:
1143:
1140:
1139:
1136:
1135:
1134:
1127:
1126:
1120:
1119:
1113:
1112:
1109:
1108:
1100:
1099:
1096:
1095:
1094:
1087:
1086:
1083:
1082:
1079:
1078:
1077:
1070:
1069:
1063:
1062:
1056:
1055:
1052:
1051:
1048:
1047:
1046:
1039:
1038:
1032:
1031:Ankylosaurini
1029:
1028:
1025:
1024:
1016:
1015:
1007:
1006:
1003:
1002:
1001:
994:
993:
990:
989:
986:
985:
984:
977:
976:
970:
969:
966:
965:
962:
961:
960:
953:
952:
946:
945:
939:
938:
935:
934:
931:
930:
929:
922:
921:
918:
917:
914:
913:
912:
905:
904:
901:
900:
897:
896:
895:
888:
887:
881:
880:
877:
876:
873:
872:
871:
870:Crichtonpelta
864:
863:
857:
854:
853:
849:
847:
843:
838:
836:
832:
828:
826:
821:
817:
813:
809:
805:
804:
799:
795:
788:
787:
781:
774:
772:
768:
766:
760:
758:
753:
747:
745:
741:
732:
725:
723:
721:
717:
713:
709:
703:
701:
697:
693:
689:
685:
681:
676:
674:
670:
665:
661:
657:
648:
644:
639:
632:
630:
627:
624:differs from
623:
619:
618:
612:
610:
605:
600:
593:
589:
582:
580:
578:
571:
569:
567:
564:) comes from
563:
559:
558:specific name
555:
554:
549:
545:
541:
537:
533:
530:. The genus (
529:
526:
522:
521:Spencer Lucas
518:
513:
511:
507:
503:
499:
495:
491:
487:
483:
479:
475:
471:
467:
458:
451:
449:
447:
446:
441:
440:
435:
431:
427:
426:specific name
423:
419:
415:
411:
407:
405:
400:
396:
392:
388:
384:
380:
376:
372:
368:
364:
360:
357:
353:
352:
342:
338:
335:
330:
329:
321:
318:
314:
310:
306:
303:
298:
297:
290:
287:
286:
283:
282:Ankylosaurini
277:
274:
273:
270:
264:
261:
260:
257:
251:
248:
247:
244:
238:
235:
232:
231:
228:
222:
219:
216:
215:
212:
206:
203:
200:
199:
196:
193:
190:
187:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
151:
146:
142:
138:
134:
130:
125:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
50:
43:
37:
33:
30:
19:
2716:
2631:
2625:
2624:
2617:
2610:
2603:
2596:
2589:
2582:
2577:Dyoplosaurus
2575:
2568:
2563:Ankylosaurus
2561:
2554:
2534:
2527:
2519:
2512:
2504:
2497:
2492:Pinacosaurus
2490:
2483:
2476:
2469:
2462:
2455:
2448:
2423:
2416:
2409:
2389:
2382:
2376:Hylaeosaurus
2374:
2367:
2360:
2352:
2344:
2312:
2305:
2300:Pawpawsaurus
2298:
2291:
2284:
2264:
2257:
2250:
2245:Denversaurus
2243:
2236:
2216:
2209:
2202:
2196:Peloroplites
2194:
2187:
2180:
2173:
2168:Borealopelta
2166:
2158:
2153:Acantholipan
2151:
2140:Nodosaurinae
2126:
2119:
2113:Peloroplites
2111:
2104:
2096:
2088:
2068:
2061:
2054:
2046:
2038:
2032:Mymoorapelta
2030:
2023:
2017:Hylaeosaurus
2015:
2008:
2001:
1993:
1985:
1978:
1973:Anoplosaurus
1971:
1964:
1953:Nodosauridae
1925:
1917:
1910:
1890:
1883:
1875:
1868:
1861:
1853:
1846:
1839:
1832:
1825:
1820:Cryptosaurus
1818:
1802:Ankylosauria
1780:Ankylosauria
1778:
1750:Ankylosauria
1738:Ornithischia
1723:Ornithischia
1675:
1671:Ornithischia
1666:
1657:
1629:Ankylosauria
1563:
1559:
1540:26 September
1538:. Retrieved
1534:
1525:
1516:
1512:
1505:
1456:
1450:
1382:
1376:
1372:Dyoplosaurus
1370:
1366:
1358:
1350:
1348:
1339:
1335:
1331:
1295:
1293:
1216:
1215:
1214:
1198:
1197:
1174:
1173:
1148:
1147:
1133:Ankylosaurus
1131:
1130:
1091:
1090:
1074:
1073:
1045:Dyoplosaurus
1043:
1042:
998:
997:
981:
980:
957:
956:
928:Pinacosaurus
926:
925:
909:
908:
892:
891:
868:
867:
839:
834:
830:
823:
810:, being its
801:
793:
792:
784:
769:
764:
761:
751:
748:
739:
738:The body of
737:
719:
704:
695:
683:
677:
668:
663:
652:
646:
642:
625:
621:
615:
613:
608:
603:
587:
586:
576:
575:
562:sanjuanensis
561:
551:
531:
527:
525:type species
514:
506:argon dating
493:
477:
463:
443:
437:
417:
403:
402:
363:ankylosaurid
350:
349:
348:
336:
327:
326:
317:Type species
304:
295:
294:
243:Ankylosauria
233:
217:
211:Ornithischia
201:
188:
35:
29:
2741:Wikispecies
2612:Scolosaurus
2529:Tsagantegia
2478:Jinyunpelta
2418:Shamosaurus
2369:Chuanqilong
2211:Silvisaurus
2189:Patagopelta
2121:Polacanthus
2070:Sauroplites
1870:Spicomellus
1848:Sarcolestes
1680:Thyreophora
1566:(5): 1–60.
1463:: e108804.
1378:Scolosaurus
1336:Scolosaurus
1328:diversified
1200:Scolosaurus
894:Tsagantegia
803:Scolosaurus
669:caputegulae
664:caputegulae
660:premaxillae
643:caputegulae
609:caputegulae
598:caputegulum
572:Description
439:Scolosaurus
393:researcher
262:Subfamily:
227:Thyreophora
2804:Categories
2680:See also:
2605:Platypelta
2598:Oohkotokia
2536:Zaraapelta
2411:Gobisaurus
2362:Cedarpelta
2354:Aletopelta
2307:Stegopelta
2252:Edmontonia
2238:Animantarx
2204:Sauropelta
2182:Nodosaurus
2048:Priconodon
1892:Vectipelta
1827:Dracopelta
1759:see below↓
1662:Dinosauria
1519:: 289–296.
1401:References
1316:Cretaceous
1312:Cenomanian
1000:Zaraapelta
744:osteoderms
726:Osteoderms
720:fenestrae,
656:antorbital
546:, and the
540:Zia people
502:osteoderms
422:osteoderms
414:state flag
410:Zia people
391:ankylosaur
387:New Mexico
195:Dinosauria
2732:Q18123387
2619:Talarurus
2499:Saichania
2286:Europelta
2266:Texasetes
2128:Taohelong
2025:Invictarx
1927:Stegouros
1645:Kingdom:
1588:214625754
1355:Laramidia
1320:Campanian
1076:Talarurus
959:Saichania
842:cladogram
816:cladistic
692:braincase
510:Campanian
472:, led by
452:Discovery
383:Campanian
168:Kingdom:
162:Eukaryota
42:Campanian
2747:Ziapelta
2726:Wikidata
2717:Ziapelta
2692:Category
2682:Timeline
2626:Ziapelta
2098:Gastonia
1995:Gastonia
1653:Chordata
1651:Phylum:
1647:Animalia
1535:Eukalert
1497:25250819
1452:PLOS ONE
1389:See also
1383:Ziapelta
1381:because
1359:Ziapelta
1340:Ziapelta
1324:Turonian
1296:Ziapelta
1217:Ziapelta
831:Ziapelta
794:Ziapelta
752:Ziapelta
740:Ziapelta
684:Ziapelta
673:maxillae
647:Ziapelta
622:Ziapelta
604:Ziapelta
577:Ziapelta
532:Ziapelta
515:In 2014
498:holotype
468:and the
432:and the
351:Ziapelta
296:Ziapelta
249:Family:
182:Chordata
178:Phylum:
172:Animalia
158:Domain:
137:Holotype
36:Ziapelta
2774:6150250
2761:8686391
2514:Tarchia
2506:Shanxia
1568:Bibcode
1488:4177562
1465:Bibcode
1363:habitat
1294:Since,
983:Tarchia
712:choanae
688:condyle
486:Members
373:of the
371:members
367:fossils
356:extinct
288:Genus:
275:Tribe:
116:↓
2787:320257
1586:
1495:
1485:
1375:, and
1308:Albian
808:Canada
757:suture
708:vomers
700:suture
594:. The
365:. Its
354:is an
337:et al.
334:Arbour
305:et al.
302:Arbour
139:skull
2471:Datai
1834:Minmi
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