42:
37:
254:
305:, from July 1965 to June 1966. He has been senior investigator in the Department of Transfusion Medicine at the NIH from July 1969 to present; chief of infectious diseases section at the department of transfusion medicine in the Clinical Center NIH from December 1972 to present; associate director for research at the department of transfusion medicine at the Clinical Center at NIH from January 1987 to present. He was the recipient of 1992
27:
244:"The discovery of the Hepatitis C virus by this year’s Laureates laid the foundation for our current understanding about how the virus survives in its niche during the long chronic phase of the infection, and how liver disease develops. And importantly, it led to the development of highly effective anti-viral medicines that now cure the infection in almost all treated persons."
284:
degree in 1956. In 1960, Alter obtained a medical degree from
University of Rochester and began a residency at Strong Memorial. Alters's post graduate training includes a rotation as a clinical associate at the National Institutes of Health in Bethesda, Maryland, from December 1961 to June 1964; a
524:, Gitnick G.L., Redeker A.G., Purcell R.H., Miyamura T., Dienstag J.L., Alter C.E., Stevens C.E., Tegtmeier G.E., Bonino F., Colombo M., Lee W.S., Kuo C., Berger K., Shuster J.R., Overby L.R., Bradley D.W.,
999:
2798:
992:
553:
2542:
1008:
767:
2696:
985:
310:
740:
614:
286:
2500:
2489:
2474:
204:
797:
2692:
47:
390:
426:
314:
114:
2535:
2656:
1012:
220:
195:
257:
Seminal experiments by HJ Alter, M Houghton and CM Rice leading to the discovery of HCV as the causative agent of non-A, non-B hepatitis.
386:
2772:
2767:
2762:
2752:
2747:
2742:
2737:
2732:
2528:
549:
2372:
1565:
186:
2388:
1653:
360:
306:
2590:
1080:
298:
2291:
2104:
359:
where he is also director of the Li Ka Shing
Applied Virology Institute. He was a recipient of numerous prizes such as 1992
2675:
2463:
2315:
397:
in New York. During 2001–2018, he was the
Scientific and Executive Director at the Center for the Study of Hepatitis C at
326:
229:
158:
56:
2012:
2777:
1215:
1992:
1773:
588:
2586:
1733:
1549:
1301:
1277:
1072:
414:
2561:
1856:
336:
801:
2726:
1968:
1701:
1529:
1357:
654:
385:
Charles Moen Rice was born in 1952 in
Sacramento, California. He received his Ph.D. degree in 1981 from the
273:
389:
where he also trained as a postdoctoral fellow between 1981 and 1985. He established his research group at
2625:
2610:
2569:
1852:
1713:
1494:
1466:
1418:
398:
394:
364:
1629:
1140:
977:
1896:
1872:
1573:
1486:
1442:
1406:
1293:
1257:
619:
356:
348:
1685:
1478:
2803:
2617:
1996:
1980:
1924:
1785:
1545:
1541:
1533:
685:"The road not taken or how I learned to love the liver: A personal perspective on hepatitis history"
41:
2364:
2299:
2136:
2128:
2120:
2080:
2020:
1984:
1944:
1833:
1697:
1450:
1203:
1100:
1088:
1044:
640:
401:
where he remains active. He was the recipient of the 2007 M.W. Beijerinck
Virology Prize, the 2015
290:
277:
253:
908:
2448:
2444:
2187:
2076:
1956:
1916:
1900:
1817:
1801:
1729:
1490:
1474:
1345:
1313:
1273:
956:
722:
344:
2493:
36:
443:
Posttransfusion hepatitis after exclusion of commercial and hepatitis-B antigen-positive donors
331:
Michael James
Houghton was born in 1949 in the United Kingdom. He received his Ph.D. degree in
2606:
2504:
2482:
2331:
2327:
2283:
2207:
2108:
2064:
2044:
2028:
2016:
1972:
1940:
1888:
1813:
1681:
1669:
1617:
1426:
1365:
1329:
1211:
1116:
932:
879:
861:
714:
706:
529:
An essay for circulating antibodies to a major etiologic virus of human non-A, non-B hepatitis
402:
237:
67:
2711:
2440:
2428:
2420:
2404:
2392:
2303:
2152:
2116:
2088:
2032:
1908:
1725:
1597:
1569:
1458:
1285:
1249:
1223:
1028:
869:
853:
696:
410:
340:
302:
281:
574:
173:
46:
2681:
2669:
2639:
2575:
2467:
2459:
2416:
2412:
2376:
2348:
2307:
2275:
2223:
2100:
2004:
1964:
1864:
1829:
1825:
1793:
1645:
1609:
1414:
1265:
1108:
741:"Rochester graduate awarded 2020 Nobel Prize for 'landmark achievement' against hepatitis"
380:
267:
233:
225:
162:
154:
124:
60:
52:
2597:
1741:
240:." During the award ceremony on December 10, 2020, Prof. Gunilla Karlsson-Hedestam said:
920:
824:
491:
Isolation of a cDNA clone derived from a blood-borne non-A, non-B viral hepatitis genome
2645:
2360:
2287:
2267:
2243:
2219:
2203:
2164:
2160:
2092:
2068:
1932:
1928:
1884:
1868:
1781:
1765:
1749:
1745:
1521:
1373:
1321:
1199:
1132:
874:
841:
944:
896:
393:
in 1986 and became a full professor in 1995. Since 2001, he has been professor at the
2792:
2633:
2508:
2356:
2323:
2235:
2140:
2040:
1809:
1797:
1761:
1709:
1689:
1661:
1637:
1621:
1589:
1557:
1434:
1124:
1068:
441:, Holland P.V., Purcell R.H., Lander J.J., Feinstone S.M., Morrow A.G., Schmidt P.J.
135:
726:
313:
or his work leading to the discovery of the virus that causes hepatitis C, and 2013
2705:
2478:
2432:
2380:
2271:
2247:
2231:
2211:
2056:
1912:
1880:
1757:
1673:
1657:
1641:
1605:
1593:
1581:
1337:
1325:
1305:
1184:
1176:
1092:
406:
368:
332:
826:
Report of program activities : National
Institutes of Health. Clinical Center
857:
2552:
2344:
2259:
2255:
2168:
2144:
2052:
1717:
1513:
1430:
1060:
1052:
1036:
589:"Nobel Prize in Medicine Awarded to Scientists Who Discovered Hepatitis C Virus"
26:
2195:
1454:
1402:
1349:
425:
The following publications were the fundamental researches that motivated the
294:
2520:
865:
710:
2396:
1948:
508:
Transmission of hepatitis C by intrahepatic inoculation with transcribed RNA
96:
883:
718:
2191:
768:"UR graduate Harvey Alter awarded Nobel Prize in fight against Hepatitis"
352:
462:, Holland P.V., Morrow A.G., Purcell R.H., Feinstone S.M., Moritsugu Y.
503:
Kolykhalov A.A., Agapov E.V., Blight K.J., Mihalik K., Feinstone S.M.,
454:
Transfusion-associated hepatitis not due to viral hepatitis type A or B
701:
684:
464:
Clinical and serological analysis of transfusion-associated hepatitis
272:
Alter was born in New York City in a Jewish family. He attended the
641:
Award ceremony speech - 2020 Nobel Prize in
Physiology or Medicine
252:
2524:
981:
823:
National
Institutes of Health (U.S.). Clinical Center (1958).
615:"3 Scientists Win Nobel Prize For Hepatitis C Virus Discovery"
957:
Press release: The Nobel Prize in
Physiology or Medicine 2020
842:"The hazards of blood transfusion in historical perspective"
486:
Choo Q.L., Kuo G., Weiner A.J., Overby L.R., Bradley D.W.,
972:
655:"NIH Clinical Center scientist a Lasker Award recipient"
840:
Alter, Harvey J.; Klein, Harvey G. (October 1, 2008).
429:
to award the 2020 Prize to Alter, Houghton and Rice:
587:
Katherine J. Wu and Daniel Victor (5 October 2020).
2691:
2655:
2624:
2605:
2585:
2560:
2178:
1843:
1504:
1240:
1019:
168:
150:
142:
130:
120:
110:
102:
92:
77:
24:
792:
790:
788:
829:. NIH Library. Bethesda, Md. : The Center.
550:"Nobel Prize amount is raised by SEK 1 million"
242:
575:The Nobel Prize in Physiology or Medicine 2020
311:Lasker-DeBakey Clinical Medical Research Award
30:The 2020 Nobel Prize in Physiology or Medicine
2799:Nobel Prize in Physiology or Medicine by year
2536:
993:
474:Transmissible agent in non-A, non-B hepatitis
448:Feinstone S.M., Kapikian A.Z., Purcell R.H.,
293:, from July 1964 to June 1965; and work as a
8:
287:University of Washington School of Medicine
2543:
2529:
2521:
1000:
986:
978:
653:Henrichsen, Colleen (September 21, 2000).
363:(together with Harvey J. Alter), the 1994
174:2020 Nobel Prize in Physiology or Medicine
21:
873:
700:
973:Official website of the Nobel Foundation
472:, Purcell R.H., Holland P.V., Popper H.
391:Washington University School of Medicine
351:in 2010 and is currently a professor of
224:was awarded to the American virologists
766:Freile, Victoria E. (October 6, 2020).
541:
427:Nobel Assembly at Karolinska Institutet
315:Gairdner Foundation International Award
115:Nobel Assembly at Karolinska Institutet
1013:Nobel Prize in Physiology or Medicine
221:Nobel Prize in Physiology or Medicine
196:Nobel Prize in Physiology or Medicine
7:
678:
676:
556:from the original on 1 February 2018
445:. Ann Intern Med. 1972; 77:691-699.
347:in California. He relocated to the
798:"Harvey J. Alter Curriculum Vitae"
456:. N Engl J Med. 1975; 292:767-770.
387:California Institute of Technology
14:
285:year of residency in medicine at
236:(b. 1952) "for the discovery of
45:
40:
35:
25:
361:Karl Landsteiner Memorial Award
307:Karl Landsteiner Memorial Award
299:Georgetown University Hospital
1:
531:. Science. 1989; 244:362-364.
510:. Science. 1997; 277:570-574.
493:. Science. 1989; 244:359-362.
327:Michael Houghton (virologist)
858:10.1182/blood-2008-07-077370
613:Rob Stein (5 October 2020).
2105:Christiane NĂĽsslein-Volhard
2820:
476:. Lancet. 1978; 1:459-463.
466:. Lancet. 1975; 2:838-841.
378:
341:G. D. Searle & Company
324:
265:
2721:
1302:Charles Scott Sherrington
1278:Frederick Gowland Hopkins
683:Alter, Harvey J. (2014).
415:Ralf F. W. Bartenschlager
343:before moving in 1982 to
180:
34:
2292:Françoise Barré-Sinoussi
909:Michael Houghton – Facts
81:5 October 2020
933:Charles M. Rice – Facts
274:University of Rochester
2727:Nobel Prize recipients
2657:Physiology or Medicine
2570:Emmanuelle Charpentier
1073:Santiago RamĂłn y Cajal
772:Democrat and Chronicle
399:Rockefeller University
395:Rockefeller University
365:William Beaumont Prize
258:
246:
66:"for the discovery of
1574:Dickinson W. Richards
1258:Julius Wagner-Jauregg
620:National Public Radio
357:University of Alberta
349:University of Alberta
337:King's College London
256:
2618:World Food Programme
1997:Rita Levi-Montalcini
1358:Albert Szent-Györgyi
804:on September 4, 2005
2389:William C. Campbell
2300:Elizabeth Blackburn
2137:Robert F. Furchgott
2129:Stanley B. Prusiner
2121:Rolf M. Zinkernagel
2021:George H. Hitchings
1985:Joseph L. Goldstein
1945:Bengt I. Samuelsson
1774:Earl Sutherland Jr.
520:Kuo G., Choo Q.L.,
291:Seattle, Washington
278:Rochester, New York
2449:William Kaelin Jr.
2445:Peter J. Ratcliffe
2188:Leland H. Hartwell
2077:Richard J. Roberts
1957:Barbara McClintock
1901:Godfrey Hounsfield
1802:Nikolaas Tinbergen
1734:Marshall Nirenberg
1702:Charles B. Huggins
1491:Tadeusz Reichstein
1479:AntĂłnio Egas Moniz
1274:Christiaan Eijkman
593:The New York Times
552:. Nobelprize.org.
515:Collaborative work
345:Chiron Corporation
259:
2786:
2785:
2693:Economic Sciences
2518:
2517:
2483:Ardem Patapoutian
2332:Ralph M. Steinman
2328:Jules A. Hoffmann
2316:Robert G. Edwards
2284:Harald zur Hausen
2208:H. Robert Horvitz
2109:Eric F. Wieschaus
2045:E. Donnall Thomas
2029:J. Michael Bishop
2017:Gertrude B. Elion
1889:Hamilton O. Smith
1814:Christian de Duve
1550:Frederick Robbins
1427:Alexander Fleming
1366:Corneille Heymans
1212:Frederick Banting
1117:Allvar Gullstrand
747:. October 5, 2020
702:10.1002/hep.26787
403:Robert Koch Prize
238:Hepatitis C virus
215:
214:
211:
210:
68:Hepatitis C virus
2811:
2729:
2712:Robert B. Wilson
2678:(United Kingdom)
2676:Michael Houghton
2636:(United Kingdom)
2545:
2538:
2531:
2522:
2511:
2496:
2485:
2470:
2464:Michael Houghton
2451:
2441:Gregg L. Semenza
2435:
2429:James P. Allison
2423:
2421:Michael W. Young
2407:
2405:Yoshinori Ohsumi
2399:
2383:
2367:
2365:Thomas C. SĂĽdhof
2351:
2339:
2318:
2310:
2304:Carol W. Greider
2294:
2278:
2262:
2250:
2238:
2226:
2214:
2198:
2171:
2155:
2147:
2131:
2123:
2117:Peter C. Doherty
2111:
2095:
2089:Alfred G. Gilman
2083:
2071:
2059:
2047:
2035:
2033:Harold E. Varmus
2023:
2007:
1999:
1987:
1975:
1959:
1951:
1935:
1919:
1909:Baruj Benacerraf
1903:
1891:
1875:
1859:
1836:
1820:
1804:
1788:
1776:
1768:
1752:
1736:
1726:Robert W. Holley
1720:
1704:
1692:
1676:
1664:
1648:
1632:
1630:Georg von Békésy
1624:
1612:
1600:
1598:Joshua Lederberg
1584:
1576:
1570:Werner Forssmann
1560:
1552:
1536:
1524:
1516:
1497:
1481:
1469:
1461:
1459:Bernardo Houssay
1445:
1437:
1421:
1409:
1397:
1396:
1390:
1389:
1383:
1382:
1376:
1368:
1360:
1352:
1340:
1332:
1316:
1308:
1296:
1288:
1286:Karl Landsteiner
1280:
1268:
1260:
1252:
1250:Johannes Fibiger
1233:
1232:
1226:
1224:Willem Einthoven
1218:
1206:
1194:
1193:
1187:
1179:
1171:
1170:
1164:
1163:
1157:
1156:
1150:
1149:
1143:
1135:
1127:
1119:
1111:
1103:
1095:
1089:Élie Metchnikoff
1083:
1081:Alphonse Laveran
1075:
1063:
1055:
1047:
1039:
1031:
1002:
995:
988:
979:
960:
954:
948:
942:
936:
930:
924:
921:Michael Houghton
918:
912:
906:
900:
894:
888:
887:
877:
852:(7): 2617–2626.
837:
831:
830:
820:
814:
813:
811:
809:
800:. Archived from
794:
783:
782:
780:
778:
763:
757:
756:
754:
752:
737:
731:
730:
704:
680:
671:
670:
668:
666:
650:
644:
638:
632:
631:
629:
627:
610:
604:
603:
601:
599:
584:
578:
572:
566:
565:
563:
561:
546:
481:Michael Houghton
421:Key publications
411:Michael J. Sofia
339:. He joined the
321:Michael Houghton
303:Washington, D.C.
282:Bachelor of Arts
230:Michael Houghton
182:
181:
176:
159:Michael Houghton
88:
86:
49:
44:
39:
29:
22:
2819:
2818:
2814:
2813:
2812:
2810:
2809:
2808:
2789:
2788:
2787:
2782:
2725:
2717:
2714:(United States)
2708:(United States)
2687:
2684:(United States)
2682:Charles M. Rice
2672:(United States)
2670:Harvey J. Alter
2651:
2648:(United States)
2640:Reinhard Genzel
2620:
2601:
2600:(United States)
2581:
2578:(United States)
2576:Jennifer Doudna
2556:
2549:
2519:
2514:
2499:
2488:
2473:
2468:Charles M. Rice
2460:Harvey J. Alter
2454:
2438:
2426:
2417:Michael Rosbash
2413:Jeffrey C. Hall
2410:
2402:
2386:
2377:May-Britt Moser
2370:
2354:
2349:Shinya Yamanaka
2342:
2321:
2313:
2308:Jack W. Szostak
2297:
2281:
2276:Oliver Smithies
2265:
2253:
2241:
2229:
2224:Peter Mansfield
2217:
2212:John E. Sulston
2201:
2185:
2174:
2158:
2150:
2134:
2126:
2114:
2101:Edward B. Lewis
2098:
2086:
2074:
2062:
2050:
2038:
2026:
2010:
2005:Susumu Tonegawa
2002:
1990:
1978:
1962:
1954:
1938:
1922:
1906:
1894:
1878:
1865:Roger Guillemin
1862:
1857:Daniel Gajdusek
1853:Baruch Blumberg
1850:
1839:
1830:Renato Dulbecco
1826:David Baltimore
1823:
1807:
1794:Karl von Frisch
1791:
1779:
1771:
1755:
1739:
1723:
1714:Haldan Hartline
1707:
1695:
1679:
1667:
1651:
1646:Maurice Wilkins
1635:
1627:
1615:
1610:Arthur Kornberg
1603:
1587:
1579:
1563:
1555:
1539:
1527:
1519:
1511:
1500:
1484:
1472:
1464:
1448:
1440:
1424:
1415:Joseph Erlanger
1412:
1400:
1394:
1393:
1387:
1386:
1380:
1379:
1371:
1363:
1355:
1343:
1335:
1319:
1311:
1299:
1291:
1283:
1271:
1266:Charles Nicolle
1263:
1255:
1247:
1236:
1230:
1229:
1221:
1209:
1197:
1191:
1190:
1182:
1174:
1168:
1167:
1161:
1160:
1154:
1153:
1147:
1146:
1138:
1130:
1122:
1114:
1109:Albrecht Kossel
1106:
1098:
1086:
1078:
1066:
1058:
1050:
1042:
1034:
1026:
1015:
1006:
969:
964:
963:
955:
951:
945:Charles M. Rice
943:
939:
931:
927:
919:
915:
907:
903:
897:Harvey J. Alter
895:
891:
839:
838:
834:
822:
821:
817:
807:
805:
796:
795:
786:
776:
774:
765:
764:
760:
750:
748:
739:
738:
734:
682:
681:
674:
664:
662:
652:
651:
647:
639:
635:
625:
623:
612:
611:
607:
597:
595:
586:
585:
581:
573:
569:
559:
557:
548:
547:
543:
538:
517:
500:
498:Charles M. Rice
483:
452:, Holland P.V.
435:
433:Harvey J. Alter
423:
405:, and the 2016
383:
381:Charles M. Rice
377:
375:Charles M. Rice
367:, and the 2002
329:
323:
280:, and earned a
270:
268:Harvey J. Alter
264:
262:Harvey J. Alter
251:
234:Charles M. Rice
226:Harvey J. Alter
172:
163:Charles M. Rice
155:Harvey J. Alter
125:Thomas Perlmann
84:
82:
73:
20:
17:
12:
11:
5:
2817:
2815:
2807:
2806:
2801:
2791:
2790:
2784:
2783:
2781:
2780:
2775:
2770:
2765:
2760:
2755:
2750:
2745:
2740:
2735:
2730:
2722:
2719:
2718:
2716:
2715:
2709:
2702:
2700:
2689:
2688:
2686:
2685:
2679:
2673:
2666:
2664:
2653:
2652:
2650:
2649:
2646:Andrea M. Ghez
2643:
2637:
2630:
2628:
2622:
2621:
2616:
2614:
2603:
2602:
2596:
2594:
2583:
2582:
2580:
2579:
2573:
2566:
2564:
2558:
2557:
2550:
2548:
2547:
2540:
2533:
2525:
2516:
2515:
2513:
2512:
2505:Katalin KarikĂł
2497:
2486:
2471:
2452:
2436:
2424:
2408:
2400:
2384:
2368:
2361:Randy Schekman
2352:
2340:
2319:
2311:
2295:
2288:Luc Montagnier
2279:
2268:Mario Capecchi
2263:
2251:
2244:Barry Marshall
2239:
2227:
2220:Paul Lauterbur
2215:
2204:Sydney Brenner
2199:
2182:
2180:
2176:
2175:
2173:
2172:
2165:Paul Greengard
2161:Arvid Carlsson
2156:
2148:
2132:
2124:
2112:
2096:
2093:Martin Rodbell
2084:
2072:
2069:Edwin G. Krebs
2065:Edmond Fischer
2060:
2048:
2036:
2024:
2013:James W. Black
2008:
2000:
1988:
1976:
1973:CĂ©sar Milstein
1969:Georges Köhler
1960:
1952:
1941:Sune Bergström
1936:
1933:Torsten Wiesel
1929:David H. Hubel
1920:
1904:
1892:
1885:Daniel Nathans
1876:
1869:Andrew Schally
1860:
1847:
1845:
1841:
1840:
1838:
1837:
1821:
1805:
1789:
1782:Gerald Edelman
1777:
1769:
1766:Julius Axelrod
1753:
1750:Salvador Luria
1746:Alfred Hershey
1737:
1721:
1705:
1693:
1682:François Jacob
1677:
1665:
1649:
1633:
1625:
1613:
1601:
1585:
1577:
1566:André Cournand
1561:
1553:
1537:
1525:
1522:Selman Waksman
1517:
1508:
1506:
1502:
1501:
1499:
1498:
1487:Edward Kendall
1482:
1470:
1462:
1446:
1443:Hermann Muller
1438:
1422:
1419:Herbert Gasser
1410:
1398:
1391:
1384:
1377:
1374:Gerhard Domagk
1369:
1361:
1353:
1341:
1333:
1330:William Murphy
1322:George Whipple
1317:
1309:
1297:
1289:
1281:
1269:
1261:
1253:
1244:
1242:
1238:
1237:
1235:
1234:
1227:
1219:
1207:
1200:Archibald Hill
1195:
1188:
1180:
1172:
1165:
1158:
1151:
1144:
1136:
1133:Charles Richet
1128:
1120:
1112:
1104:
1096:
1084:
1076:
1064:
1056:
1048:
1040:
1032:
1023:
1021:
1017:
1016:
1007:
1005:
1004:
997:
990:
982:
976:
975:
968:
967:External links
965:
962:
961:
959:nobelprize.org
949:
947:britannica.com
937:
935:nobelprize.org
925:
923:britannica.com
913:
911:nobelprize.org
901:
899:nobelprize.org
889:
832:
815:
784:
758:
732:
672:
645:
643:nobelprize.org
633:
605:
579:
577:nobelprize.org
567:
540:
539:
537:
534:
533:
532:
516:
513:
512:
511:
499:
496:
495:
494:
482:
479:
478:
477:
467:
457:
446:
434:
431:
422:
419:
379:Main article:
376:
373:
325:Main article:
322:
319:
266:Main article:
263:
260:
250:
247:
232:(b. 1949) and
213:
212:
209:
208:
198:
193:
178:
177:
170:
166:
165:
152:
151:2020 laureates
148:
147:
144:
140:
139:
132:
128:
127:
122:
118:
117:
112:
108:
107:
104:
100:
99:
94:
90:
89:
79:
75:
74:
50:
32:
31:
18:
15:
13:
10:
9:
6:
4:
3:
2:
2816:
2805:
2802:
2800:
2797:
2796:
2794:
2779:
2776:
2774:
2771:
2769:
2766:
2764:
2761:
2759:
2756:
2754:
2751:
2749:
2746:
2744:
2741:
2739:
2736:
2734:
2731:
2728:
2724:
2723:
2720:
2713:
2710:
2707:
2704:
2703:
2701:
2698:
2694:
2690:
2683:
2680:
2677:
2674:
2671:
2668:
2667:
2665:
2662:
2658:
2654:
2647:
2644:
2641:
2638:
2635:
2634:Roger Penrose
2632:
2631:
2629:
2627:
2623:
2619:
2615:
2612:
2608:
2604:
2599:
2595:
2592:
2588:
2584:
2577:
2574:
2571:
2568:
2567:
2565:
2563:
2559:
2554:
2546:
2541:
2539:
2534:
2532:
2527:
2526:
2523:
2510:
2509:Drew Weissman
2506:
2502:
2498:
2495:
2491:
2487:
2484:
2480:
2476:
2472:
2469:
2465:
2461:
2457:
2453:
2450:
2446:
2442:
2437:
2434:
2430:
2425:
2422:
2418:
2414:
2409:
2406:
2401:
2398:
2394:
2393:Satoshi ĹŚmura
2390:
2385:
2382:
2378:
2374:
2369:
2366:
2362:
2358:
2357:James Rothman
2353:
2350:
2346:
2341:
2337:
2333:
2329:
2325:
2324:Bruce Beutler
2320:
2317:
2312:
2309:
2305:
2301:
2296:
2293:
2289:
2285:
2280:
2277:
2273:
2269:
2264:
2261:
2257:
2252:
2249:
2245:
2240:
2237:
2236:Linda B. Buck
2233:
2228:
2225:
2221:
2216:
2213:
2209:
2205:
2200:
2197:
2193:
2189:
2184:
2183:
2181:
2177:
2170:
2166:
2162:
2157:
2154:
2153:GĂĽnter Blobel
2149:
2146:
2142:
2141:Louis Ignarro
2138:
2133:
2130:
2125:
2122:
2118:
2113:
2110:
2106:
2102:
2097:
2094:
2090:
2085:
2082:
2081:Phillip Sharp
2078:
2073:
2070:
2066:
2061:
2058:
2054:
2049:
2046:
2042:
2041:Joseph Murray
2037:
2034:
2030:
2025:
2022:
2018:
2014:
2009:
2006:
2001:
1998:
1994:
1993:Stanley Cohen
1989:
1986:
1982:
1981:Michael Brown
1977:
1974:
1970:
1966:
1961:
1958:
1953:
1950:
1946:
1942:
1937:
1934:
1930:
1926:
1921:
1918:
1914:
1910:
1905:
1902:
1898:
1897:Allan Cormack
1893:
1890:
1886:
1882:
1877:
1874:
1873:Rosalyn Yalow
1870:
1866:
1861:
1858:
1854:
1849:
1848:
1846:
1842:
1835:
1831:
1827:
1822:
1819:
1818:George Palade
1815:
1811:
1810:Albert Claude
1806:
1803:
1799:
1798:Konrad Lorenz
1795:
1790:
1787:
1786:Rodney Porter
1783:
1778:
1775:
1770:
1767:
1763:
1762:Ulf von Euler
1759:
1754:
1751:
1747:
1743:
1738:
1735:
1731:
1727:
1722:
1719:
1715:
1711:
1710:Ragnar Granit
1706:
1703:
1699:
1694:
1691:
1690:Jacques Monod
1687:
1683:
1678:
1675:
1671:
1666:
1663:
1662:Andrew Huxley
1659:
1655:
1650:
1647:
1643:
1639:
1638:Francis Crick
1634:
1631:
1626:
1623:
1622:Peter Medawar
1619:
1614:
1611:
1607:
1602:
1599:
1595:
1591:
1590:George Beadle
1586:
1583:
1578:
1575:
1571:
1567:
1562:
1559:
1558:Hugo Theorell
1554:
1551:
1547:
1546:Thomas Weller
1543:
1538:
1535:
1534:Fritz Lipmann
1531:
1526:
1523:
1518:
1515:
1510:
1509:
1507:
1503:
1496:
1492:
1488:
1483:
1480:
1476:
1471:
1468:
1463:
1460:
1456:
1452:
1447:
1444:
1439:
1436:
1435:Howard Florey
1432:
1428:
1423:
1420:
1416:
1411:
1408:
1404:
1399:
1392:
1385:
1378:
1375:
1370:
1367:
1362:
1359:
1354:
1351:
1347:
1342:
1339:
1334:
1331:
1327:
1323:
1318:
1315:
1314:Thomas Morgan
1310:
1307:
1303:
1298:
1295:
1290:
1287:
1282:
1279:
1275:
1270:
1267:
1262:
1259:
1254:
1251:
1246:
1245:
1243:
1239:
1228:
1225:
1220:
1217:
1213:
1208:
1205:
1204:Otto Meyerhof
1201:
1196:
1189:
1186:
1181:
1178:
1173:
1166:
1159:
1152:
1145:
1142:
1141:Róbert Bárány
1137:
1134:
1129:
1126:
1125:Alexis Carrel
1121:
1118:
1113:
1110:
1105:
1102:
1097:
1094:
1090:
1085:
1082:
1077:
1074:
1070:
1069:Camillo Golgi
1065:
1062:
1057:
1054:
1049:
1046:
1041:
1038:
1033:
1030:
1025:
1024:
1022:
1018:
1014:
1010:
1003:
998:
996:
991:
989:
984:
983:
980:
974:
971:
970:
966:
958:
953:
950:
946:
941:
938:
934:
929:
926:
922:
917:
914:
910:
905:
902:
898:
893:
890:
885:
881:
876:
871:
867:
863:
859:
855:
851:
847:
843:
836:
833:
828:
827:
819:
816:
803:
799:
793:
791:
789:
785:
773:
769:
762:
759:
746:
742:
736:
733:
728:
724:
720:
716:
712:
708:
703:
698:
694:
690:
686:
679:
677:
673:
660:
659:Press release
656:
649:
646:
642:
637:
634:
622:
621:
616:
609:
606:
594:
590:
583:
580:
576:
571:
568:
555:
551:
545:
542:
535:
530:
527:
523:
519:
518:
514:
509:
506:
502:
501:
497:
492:
489:
485:
484:
480:
475:
471:
468:
465:
461:
458:
455:
451:
447:
444:
440:
437:
436:
432:
430:
428:
420:
418:
416:
412:
408:
404:
400:
396:
392:
388:
382:
374:
372:
370:
366:
362:
358:
354:
350:
346:
342:
338:
335:in 1977 from
334:
328:
320:
318:
316:
312:
308:
304:
300:
296:
292:
288:
283:
279:
275:
269:
261:
255:
248:
245:
241:
239:
235:
231:
227:
223:
222:
207: →
206:
202:
199:
197:
194:
192:
188:
185:←
184:
183:
179:
175:
171:
167:
164:
160:
156:
153:
149:
145:
143:First awarded
141:
137:
133:
129:
126:
123:
119:
116:
113:
109:
105:
101:
98:
95:
91:
80:
76:
72:
71:
69:
63:
62:
58:
54:
48:
43:
38:
33:
28:
23:
2757:
2706:Paul Milgrom
2660:
2598:Louise GlĂĽck
2494:Svante Pääbo
2479:David Julius
2455:
2433:Tasuku Honjo
2381:Edvard Moser
2373:John O'Keefe
2336:posthumously
2335:
2272:Martin Evans
2248:Robin Warren
2232:Richard Axel
2179:2001–present
2057:Bert Sakmann
1925:Roger Sperry
1917:George Snell
1913:Jean Dausset
1881:Werner Arber
1834:Howard Temin
1758:Bernard Katz
1742:Max DelbrĂĽck
1698:Francis Rous
1674:Feodor Lynen
1670:Konrad Bloch
1658:Alan Hodgkin
1642:James Watson
1618:Frank Burnet
1606:Severo Ochoa
1594:Edward Tatum
1582:Daniel Bovet
1495:Philip Hench
1407:Edward Doisy
1338:Hans Spemann
1326:George Minot
1306:Edgar Adrian
1294:Otto Warburg
1216:John Macleod
1185:August Krogh
1177:Jules Bordet
1093:Paul Ehrlich
1045:Niels Finsen
1029:Emil Behring
952:
940:
928:
916:
904:
892:
849:
845:
835:
825:
818:
806:. Retrieved
802:the original
775:. Retrieved
771:
761:
749:. Retrieved
744:
735:
692:
688:
663:. Retrieved
658:
648:
636:
624:. Retrieved
618:
608:
596:. Retrieved
592:
582:
570:
558:. Retrieved
544:
528:
525:
521:
507:
504:
490:
487:
473:
469:
463:
459:
453:
449:
442:
438:
424:
407:Lasker Award
384:
369:Lasker Award
333:biochemistry
330:
271:
243:
218:
216:
200:
190:
111:Presented by
65:
64:
51:
2804:2020 awards
2553:Nobel Prize
2345:John Gurdon
2260:Craig Mello
2256:Andrew Fire
2169:Eric Kandel
2145:Ferid Murad
2053:Erwin Neher
1965:Niels Jerne
1730:Har Khorana
1718:George Wald
1686:André Lwoff
1654:John Eccles
1542:John Enders
1514:Max Theiler
1475:Walter Hess
1467:Paul MĂĽller
1431:Ernst Chain
1101:Emil Kocher
1061:Robert Koch
1053:Ivan Pavlov
1037:Ronald Ross
695:(1): 4–12.
665:October 20,
526:Houghton M.
488:Houghton M.
309:, the 2002
228:(b. 1935),
2793:Categories
2587:Literature
2196:Paul Nurse
1530:Hans Krebs
1455:Gerty Cori
1403:Henrik Dam
1350:Otto Loewi
1346:Henry Dale
777:October 6,
751:October 5,
745:NewsCenter
689:Hepatology
536:References
522:Alter H.J.
470:Alter H.J.
460:Alter H.J.
450:Alter H.J.
439:Alter H.J.
297:fellow at
295:hematology
134:9 million
85:2020-10-05
2642:(Germany)
2562:Chemistry
2555:laureates
2397:Tu Youyou
1949:John Vane
1844:1976–2000
1505:1951–1975
1451:Carl Cori
1241:1926–1950
1020:1901–1925
1009:Laureates
866:0006-4971
808:April 21,
711:1527-3350
626:20 August
598:20 August
560:5 October
505:Rice C.M.
249:Laureates
131:Reward(s)
121:Hosted by
97:Stockholm
2572:(France)
2192:Tim Hunt
884:18809775
727:30040670
719:24123147
554:Archived
353:virology
93:Location
57:Houghton
2626:Physics
1011:of the
875:2962447
355:at the
169:Website
103:Country
83: (
2439:2019:
2427:2018:
2411:2017:
2403:2016:
2387:2015:
2371:2014:
2355:2013:
2343:2012:
2322:2011:
2314:2010:
2298:2009:
2282:2008:
2266:2007:
2254:2006:
2242:2005:
2230:2004:
2218:2003:
2202:2002:
2186:2001:
2159:2000:
2151:1999:
2135:1998:
2127:1997:
2115:1996:
2099:1995:
2087:1994:
2075:1993:
2063:1992:
2051:1991:
2039:1990:
2027:1989:
2011:1988:
2003:1987:
1991:1986:
1979:1985:
1963:1984:
1955:1983:
1939:1982:
1923:1981:
1907:1980:
1895:1979:
1879:1978:
1863:1977:
1851:1976:
1824:1975:
1808:1974:
1792:1973:
1780:1972:
1772:1971:
1756:1970:
1740:1969:
1724:1968:
1708:1967:
1696:1966:
1680:1965:
1668:1964:
1652:1963:
1636:1962:
1628:1961:
1616:1960:
1604:1959:
1588:1958:
1580:1957:
1564:1956:
1556:1955:
1540:1954:
1528:1953:
1520:1952:
1512:1951:
1485:1950:
1473:1949:
1465:1948:
1449:1947:
1441:1946:
1425:1945:
1413:1944:
1401:1943:
1372:1939:
1364:1938:
1356:1937:
1344:1936:
1336:1935:
1320:1934:
1312:1933:
1300:1932:
1292:1931:
1284:1930:
1272:1929:
1264:1928:
1256:1927:
1248:1926:
1222:1924:
1210:1923:
1198:1922:
1183:1920:
1175:1919:
1139:1914:
1131:1913:
1123:1912:
1115:1911:
1107:1910:
1099:1909:
1087:1908:
1079:1907:
1067:1906:
1059:1905:
1051:1904:
1043:1903:
1035:1902:
1027:1901:
882:
872:
864:
725:
717:
709:
409:(with
203:
189:
161:, and
138:(2017)
106:Sweden
2758:2020
2607:Peace
2551:2020
846:Blood
723:S2CID
661:. NIH
219:2020
53:Alter
19:Award
16:Award
2778:2024
2773:2023
2768:2022
2763:2021
2753:2019
2748:2018
2743:2017
2738:2016
2733:2015
2697:2020
2661:2020
2611:2020
2591:2020
2501:2023
2490:2022
2475:2021
2456:2020
1395:1942
1388:1941
1381:1940
1231:1925
1192:1921
1169:1918
1162:1917
1155:1916
1148:1915
880:PMID
862:ISSN
810:2006
779:2020
753:2020
715:PMID
707:ISSN
667:2009
628:2022
600:2022
562:2020
413:and
217:The
205:2021
187:2019
146:1901
78:Date
61:Rice
59:and
870:PMC
854:doi
850:112
697:doi
417:).
276:in
136:SEK
2795::
2507:/
2503::
2492::
2481:/
2477::
2466:/
2462:/
2458::
2447:/
2443:/
2431:/
2419:/
2415:/
2395:/
2391:/
2379:/
2375:/
2363:/
2359:/
2347:/
2330:/
2326:/
2306:/
2302:/
2290:/
2286:/
2274:/
2270:/
2258:/
2246:/
2234:/
2222:/
2210:/
2206:/
2194:/
2190:/
2167:/
2163:/
2143:/
2139:/
2119:/
2107:/
2103:/
2091:/
2079:/
2067:/
2055:/
2043:/
2031:/
2019:/
2015:/
1995:/
1983:/
1971:/
1967:/
1947:/
1943:/
1931:/
1927:/
1915:/
1911:/
1899:/
1887:/
1883:/
1871:/
1867:/
1855:/
1832:/
1828:/
1816:/
1812:/
1800:/
1796:/
1784:/
1764:/
1760:/
1748:/
1744:/
1732:/
1728:/
1716:/
1712:/
1700:/
1688:/
1684:/
1672:/
1660:/
1656:/
1644:/
1640:/
1620:/
1608:/
1596:/
1592:/
1572:/
1568:/
1548:/
1544:/
1532:/
1493:/
1489:/
1477:/
1457:/
1453:/
1433:/
1429:/
1417:/
1405:/
1348:/
1328:/
1324:/
1304:/
1276:/
1214:/
1202:/
1091:/
1071:/
878:.
868:.
860:.
848:.
844:.
787:^
770:.
743:.
721:.
713:.
705:.
693:59
691:.
687:.
675:^
657:.
617:.
591:.
371:.
317:.
301:,
289:,
157:,
70:."
55:,
2699:)
2695:(
2663:)
2659:(
2613:)
2609:(
2593:)
2589:(
2544:e
2537:t
2530:v
2338:)
2334:(
1001:e
994:t
987:v
886:.
856::
812:.
781:.
755:.
729:.
699::
669:.
630:.
602:.
564:.
201:·
191:·
87:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
