Knowledge (XXG)

Conodont

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606: 124: 4396: 426:, with Conodontophorida as a subordinate order for "true conodonts". All three ranks were attributed to Eichenberg, and Paraconodontida was also included as an order under Conodonta. This approach was criticized by FĂĄhraeus (1983), who argued that it overlooked Pander's historical relevance as a founder and primary figure in conodontology. FĂĄhraeus proposed to retain Conodonta as a phylum (attributed to Pander), with the single class Conodontata (Pander) and the single order Conodontophorida (Eichenberg). Subsequent authors continued to regard Conodonta as a phylum with an ever-increasing number of subgroups. 621: 527: 4426: 633: 4437: 386:, Conodontes (a Greek translation), which Huddle (1934) altered to the Latin spelling Conodonta. A few years earlier, Eichenberg (1930) established another name for the animals responsible for conodont fossils: Conodontophorida ("conodont bearers"). A few other scientific names were rarely and inconsistently applied to conodonts and their proposed close relatives during 20th century, such as Conodontophoridia, Conodontophora, Conodontochordata, Conodontiformes, and Conodontomorpha. 2124: 510: 142: 4421: 4403: 4431: 572: 4415: 796: 9560: 589: 9550: 4409: 9570: 333:. They are primarily known from their hard, mineralised tooth-like structures called "conodont elements" that in life were present in the oral cavity and used to process food. Rare soft tissue remains suggest that they had elongate eel-like bodies with large eyes. Conodonts were a long-lasting group with over 300 million years of existence from the 1241: 809:
The "teeth" of some conodonts have been interpreted as filter-feeding apparatuses, filtering plankton from the water and passing it down the throat. Others have been interpreted as a "grasping and crushing array". Wear on some conodont elements suggests that they functioned like teeth, with both wear
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is composed of an axial Sa element at the front, flanked by two groups of four close-set elongate Sb and Sc elements which were inclined obliquely inwards and forwards. Above these elements lay a pair of arched and inward pointing (makellate) M elements. Behind the S-M array lay transversely oriented
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For many years, conodonts were known only from enigmatic tooth-like microfossils (200 micrometers to 5 millimeters in length), which occur commonly, but not always, in isolation and were not associated with any other fossil. Until the early 1980s, conodont teeth had not been found in association with
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Diversity continued to decline during the Middle and Late Triassic, culminating in their extinction soon after the Triassic-Jurassic boundary. Much of their diversity during the Paleozoic was likely controlled by sea levels and temperature, with the major declines during the Late Ordovician and Late
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The earliest fossils of conodonts are known from the Cambrian period. Conodonts extensively diversified during the early Ordovician, reaching their apex of diversity during the middle part of the period, and experienced a sharp decline during the late Ordovician and Silurian, before reaching another
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The element array constituted a feeding apparatus that is radically different from the jaws of modern animals. They are now termed "conodont elements" to avoid confusion. The three forms of teeth, i.e., coniform cones, ramiform bars, and pectiniform platforms, probably performed different functions.
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Individual conodont elements are difficult to classify in a consistent manner, but an increasing number of conodont species are now known from multi-element assemblages, which offer more data to infer how different conodont lineages are related to each other. The following is a simplified cladogram
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Figures 1, 2. Conodonts from the Deer Valley Member of the Mauch Chunk Formation, Keystone quarry, Pa. This collection (93RS–79c) is from the lower 10 cm of the Deer Valley Member. Note the nonabraded, although slightly broken, conodont elements of the high-energy oolitic marine facies of the Deer
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It was only in the early 1980s that the first fossil evidence of the rest of the animal was found (see below). In the 1990s exquisite fossils were found in South Africa in which the soft tissue had been converted to clay, preserving even muscle fibres. The presence of muscles for rotating the eyes
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With increasingly strong evidence that conodonts lie within the phylum Chordata, more recent studies generally refer to "true conodonts" as the class Conodonta, containing multiple smaller orders. Paraconodonts are typically excluded from the group, though still regarded as close relatives. In
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3–9. Conodonts from the uppermost Loyalhanna Limestone Member of the Mauch Chunk Formation, Keystone quarry, Pa. This collection (93RS–79b) is from the upper 10 cm of the Loyalhanna Member. Note the highly abraded and reworked aeolian forms.
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Through their history of study, "conodont" is a term which has been applied to both the individual fossils and to the animals to which they belonged. The original German term used by Pander was "conodonten", which was subsequently
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Although conodont elements are abundant in the fossil record, fossils preserving soft tissues of conodont animals are known from only a few deposits in the world. One of the first possible body fossils of a conodont were those of
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10–14. Conodonts from the basal 20 cm of the Loyalhanna Limestone Member of the Mauch Chunk Formation, Keystone quarry, Pa. (93RS–79a), and Westernport, Md. (93RS–67), note the highly abraded and reworked aeolian forms
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Miyashita, Tetsuto; Coates, Michael I.; Farrar, Robert; Larson, Peter; Manning, Phillip L.; Wogelius, Roy A.; Edwards, Nicholas P.; Anné, Jennifer; Bergmann, Uwe; Palmer, A. Richard; Currie, Philip J. (2019-02-05).
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has been puzzling for more than a century. It has been hypothesized that the first mechanism of chordate tissue mineralization began either in the oral skeleton of conodonts or the dermal skeleton of early
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Bergström, S. M.; Carnes, J. B.; Ethington, R. L.; Votaw, R. B.; Wigley, P. B. (1974). "Appalachignathus, a New Multielement Conodont Genus from the Middle Ordovician of North America".
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at least) were efficient cruisers, but incapable of bursts of speed. Based on isotopic evidence, some Devonian conodonts have been proposed to have been low-level consumers that fed on
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Turner, S., Burrow, C.J., Schultze, H.P., Blieck, A., Reif, W.E., Rexroad, C.B., Bultynck, P., Nowlan, G.S.; Burrow; Schultze; Blieck; Reif; Rexroad; Bultynck; Nowlan (2010).
632: 851:/Ca ratios of a population of conodonts from a carbonate platform from the Silurian of Sweden found that the different conodont species and genera likely occupied different 1232:. One of the broadest studies of this nature was the analysis of Donoghue et al. (2008), which focused on "complex" conodonts (Prioniodontida and other descendant groups): 544:
The conodont apparatus may comprise a number of discrete elements, including the spathognathiform, ozarkodiniform, trichonodelliform, neoprioniodiform, and other forms.
482:, a proxy for thermal alteration in the host rock, because under higher temperatures, the phosphate undergoes predictable and permanent color changes, measured with the 3685: 9660: 4978: 2137: 3285:"Natural assemblages of the conodont Clarkina in lowermost Triassic deep-sea black claystone from northeastern Japan, with probable soft-tissue impressions" 4250: 3630: 1384:. Only a handful of conodont genera were present during the Permian, though diversity increased after the P-T extinction during the Early Triassic. 4395: 3238:"Eramosa Lagerstätte—Exceptionally preserved soft-bodied biotas with shallow-marine shelly and bioturbating organisms (Silurian, Ontario, Canada)" 509: 3193:
Gabbott, S.E.; R. J. Aldridge; J. N. Theron (1995). "A giant conodont with preserved muscle tissue from the Upper Ordovician of South Africa".
734:. There are other examples of conodont animals that only preserve the head region, including eyes, of the animals known from the Silurian aged 464:. This is because the conodont animal was soft-bodied, thus everything but the teeth was unsuited for preservation under normal circumstances. 2928: 389:
Conodonta and Conodontophorida are by far the most common scientific names used to refer to conodonts, though inconsistencies regarding their
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has been published. Among other things, it demonstrates that at least this taxon had short lifespans lasting around a month. A study
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still persist. Bengtson (1976)'s research on conodont evolution identified three morphological tiers of early conodont-like fossils:
4057: 2365: 3702:"Hagfish from the Cretaceous Tethys Sea and a reconciliation of the morphological–molecular conflict in early vertebrate phylogeny" 4425: 571: 401:, and "true conodonts" (euconodonts). Further investigations revealed that protoconodonts were probably more closely related to 4243: 2961:
Barnes, Christopher R. (1967). "A Questionable Natural Conodont Assemblage from Middle Ordovician Limestone, Ottawa, Canada".
3931: 934:, because they lack the main characteristics of these groups. More recently it has been proposed that conodonts may be stem- 405:(arrow worms) rather than true conodonts. On the other hand, paraconodonts are still considered a likely ancestral stock or 4988: 3769:"The middle–late Tournaisian crisis in conodont diversity: a comparison between Northeast Laurussia and Northeast Siberia" 4995: 9729: 443:
Conodont elements consist of mineralised teeth-like structures of varying morphology and complexity. The evolution of
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Shi, Yukun; Wang, Xiangdong; Fan, Junxuan; Huang, Hao; Xu, Huiqing; Zhao, Yingying; Shen, Shuzhong (September 2021).
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Quinton, Page C. (2016). "Effects of extraction protocols on the oxygen isotope composition of conodont elements".
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than either of these groups. However, this analysis comes with one caveat: the earliest conodont-like fossils, the
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In the 1930s, the concept of conodont assemblages was described by Hermann Schmidt and by Harold W. Scott in 1934.
141: 4430: 4963: 2157: 483: 3467:Świś, Przemysław (2019). "Population dynamics of the Late Devonian conodont Alternognathus calibrated in days". 3390: 4973: 3420:"Sr/Ca and Ba/Ca ratios support trophic partitioning within a Silurian conodont community from Gotland, Sweden" 2152: 959:
based on Sweet and Donoghue (2001), which summarized previous work by Sweet (1988) and Donoghue et al. (2000):
475: 358: 4121: 2818: 2537: 2360:. Boulder, Colorado; Lawrence, Kansas: Geological Society of America; University of Kansas. pp. 111–180. 2244: 9691: 4420: 4402: 4102: 2518: 2309:"The structure of some Middle Cambrian conodonts, and the early evolution of conodont structure and function" 9553: 5054: 5021: 4414: 4091:
The Conodonta: morphology, taxonomy, paleoecology, and evolutionary history of a long-extinct animal phylum.
2427:"The Conodonta: morphology, taxonomy, paleoecology and evolutionary history of a long-extinct animal phylum" 2147: 1381: 3418:
Terrill, David F.; Jarochowska, Emilia; Henderson, Charles M.; Shirley, Bryan; Bremer, Oskar (2022-04-08).
9590: 7930: 6144: 3768: 3141:"Panderodus from the Waukesha Lagerstätte of Wisconsin, USA: a primitive macrophagous vertebrate predator" 1576: 430:
practice, Conodonta, Conodontophorida, and Euconodonta are equivalent terms and are used interchangeably.
354: 201: 6903: 5962: 5826: 3880:"Global climate changes account for the main trends of conodont diversity but not for their final demise" 3815:"Carboniferous-earliest Permian marine biodiversification event (CPBE) during the Late Paleozoic Ice Age" 3661: 2659:
Trotter, Julie A. (2006). "Chemical systematics of conodont apatite determined by laser ablation ICPMS".
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Donoghue, P. C. J.; Forey, P. L.; Aldridge, R. J. (2000). "Conodont affinity and chordate phylogeny".
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Jeppsson, Lennart; Anehus, Rikard (1995). "A Buffered Formic Acid Technique for Conodont Extraction".
2383:"A critical look at the Treatise family-group classification of Conodonta: an exercise in eclecticism" 1987: 1929: 1650: 1641: 1584: 1412:
Conodonta taxonomy based on Sweet (1988), Sweet & Donoghue (2001), and Mikko's Phylogeny Archive.
822:(swimming above the sediment surface) niches. The preserved musculature suggests that some conodonts ( 747: 9647: 9449: 9089: 9056: 8957: 8887: 8866: 8615: 8262: 8158: 8047: 8033: 7967: 7842: 7732: 7711: 7526: 7507: 7336: 7278: 7158: 7114: 6874: 6833: 6584: 6239: 6232: 6045: 5835: 5650: 5597: 5557: 5016: 4968: 4655: 3891: 3826: 3713: 3540: 3378: 3296: 3249: 3202: 3054: 3001: 2779: 2668: 2013: 1996: 1902: 1745: 1737: 1623: 1568: 1543: 1460: 680:. As of 2023, there are only three described species of conodonts that have preserved trunk fossils: 657: 9496: 9428: 9386: 9273: 8859: 8580: 8566: 8429: 8332: 8325: 8241: 7871: 7821: 7801: 7787: 7766: 7750: 7681: 7514: 7486: 7441: 7144: 7032: 6812: 6744: 6675: 6668: 6661: 6515: 6331: 6247: 6125: 6118: 6104: 6040: 5993: 5885: 5727: 5671: 5283: 5177: 4065: 2052: 1833: 1824: 1686: 1551: 1347:, Pa element, upper view, reworked Late Devonian(?) through Mississippian morphotype, 93RS–67, X140 919:. Protoconodonts are probably not relatives of true conodonts, but likely represent a stem group to 239: 9512: 9470: 9456: 9393: 9372: 9365: 9316: 9282: 8902: 8762: 8721: 8707: 8601: 8531: 8475: 8353: 8100: 8088: 8081: 7960: 7946: 7919: 7905: 7828: 7647: 7638: 7612: 7584: 7477: 7378: 7357: 7322: 7240: 7190: 7169: 7064: 7023: 6972: 6935: 6889: 6854: 6789: 6709: 6449: 6268: 6076: 5929: 5922: 5915: 5400: 5275: 5261: 2940: 2347: 2129: 2020: 1956: 1938: 1920: 1893: 1805: 1797: 1753: 1721: 1713: 1677: 1668: 1401: 1332:, Pa element, upper view, reworked Late Devonian to Early Mississippian morphotype, 93RS–79a, X140 444: 323: 9526: 9435: 9414: 9351: 9261: 8946: 8932: 8797: 8776: 8741: 8651: 8538: 8395: 8297: 8137: 8130: 7835: 7739: 7718: 7690: 7670: 7598: 7577: 7554: 7389: 7315: 7308: 7073: 6979: 6958: 6944: 6798: 6771: 6755: 6633: 6626: 6522: 6375: 6275: 6261: 6225: 6211: 6197: 6132: 6111: 6097: 6083: 5894: 5865: 5858: 5749: 5586: 5533: 5389: 5375: 5304: 5290: 5268: 5254: 5141: 2308: 1911: 1761: 1513: 1452: 1338:, Pa element, upper view, reworked Late Devonian to Early Mississippian morphotype, 93RS–67, X140 1059: 801: 664:. This possible identification was based on the presence of conodont elements with the fossils of 471:(a phosphatic mineral). The conodont elements can be extracted from rock using adequate solvents. 9714: 9563: 9442: 9358: 8852: 8831: 8804: 8769: 8755: 8679: 8587: 8573: 8559: 8484: 8440: 8415: 8381: 8339: 8234: 8149: 8019: 8012: 7812: 7773: 7759: 7704: 7605: 7542: 7371: 7350: 7329: 7301: 7294: 7199: 7130: 7000: 6914: 6847: 6730: 6591: 6500: 6483: 6361: 6303: 6029: 6022: 5873: 5549: 5521: 5354: 5347: 5239: 5047: 4704: 4552: 4133: 4037: 3909: 3788: 3653: 3480: 3449: 3320: 3218: 3168: 2970: 2909: 2865: 2830: 2822: 2719: 2711: 2541: 2492: 2286: 2196: 2074: 1947: 1659: 1614: 885: 691: 579: 276: 260: 136: 9421: 9379: 9337: 9330: 9309: 9206: 9098: 8824: 8594: 8367: 8318: 8123: 8109: 8040: 7912: 7856: 7849: 7663: 7626: 7619: 7213: 7206: 7179: 7151: 7103: 7039: 6993: 6965: 6840: 6764: 6605: 6471: 6464: 6282: 6254: 6218: 6182: 6175: 6168: 6153: 6069: 5985: 5978: 5936: 5851: 5814: 5798: 5782: 5692: 5615: 5607: 5496: 5247: 5208: 5200: 5170: 5148: 3038: 1443: 9678: 9519: 9289: 8966: 8783: 8608: 8360: 8192: 8116: 7864: 7697: 7563: 7399: 7287: 7247: 7081: 6737: 6368: 6204: 6007: 5943: 5901: 5790: 5769: 5741: 5542: 5513: 5361: 5333: 5297: 5162: 1526: 1023: 9484: 9477: 9198: 8994: 8693: 8213: 8199: 7953: 7137: 6986: 6924: 6536: 6529: 6055: 6015: 5719: 5579: 5572: 4878: 4826: 4783: 4777: 4740: 4728: 4698: 4353: 4259: 4125: 4053: 4029: 4000: 3749: 3731: 3583: 3556: 3441: 3345: 3312: 3265: 3160: 3116: 2750: 2592: 2484: 2434: 2402: 2361: 2328: 2142: 1397: 1393: 766: 668:. This claim was disproved, however, as the conodont elements were actually in the creature's 9344: 8845: 8748: 7898: 7725: 7591: 7570: 7407: 6778: 6612: 5806: 5506: 5185: 2571:
Donoghue, Philip C. J.; Purnell, Mark A.; Aldridge, Richard J.; Zhang, Shunxin (2008-01-01).
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volume on the conodonts (Part W revised, supplement 2) lists Conodonta as the name of both a
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von Bitter, Peter H.; Purnell, Mark A.; Tetreault, Denis K.; Stott, Christopher A. (2007).
2990:"Architecture and functional morphology of the skeletal apparatus of ozarkodinid conodonts" 2464: 2351: 783:
is estimated to reach 40 cm (16 in) in length, if it had the same proportions as
9134: 9019: 8987: 8346: 8276: 8065: 7053: 6686: 5968: 5662: 5226: 4891: 4854: 4734: 4685: 4506: 4308: 4094: 3604: 3367:"Feeding mechanisms in conodonts and the function of the earliest vertebrate hard tissues" 2236: 2098: 2058: 1858: 1789: 1632: 1505: 1480: 976: 912: 468: 423: 398: 383: 342: 327: 41: 9628: 1311:, Pa element, upper view, reworked Late Devonian to Early Mississippian morphotype, X140 4184: 3895: 3830: 3717: 3544: 3382: 3300: 3253: 3206: 3140: 3085:"A possible home for a bizarre Carboniferous animal: Is Typhloesus a pelagic gastropod?" 3058: 3043:(Melton and Scott, 1973), a bizarre metazoan from the Carboniferous of Montana, U. S. A" 3005: 2783: 2672: 2382: 9300: 9254: 9168: 9077: 8939: 8732: 8642: 8522: 8504: 8463: 8406: 8269: 8248: 8074: 7493: 7089: 7046: 6654: 6563: 6439: 6426: 6397: 6296: 5630: 4866: 4844: 4789: 4771: 4722: 4714: 4691: 4649: 4625: 4370: 4165: 4025: 3744: 3701: 3111: 3084: 3022: 2989: 2645: 2480: 2398: 2324: 2090: 2082: 2066: 2044: 1850: 1842: 1605: 1472: 1372:
peak of diversity during the mid-late Devonian. Conodont diversity declined during the
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envision them as vertebrates similar in appearance to modern hagfish and lampreys, and
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Briggs, D. E. G.; Clarkson, E. N. K.; Aldridge, R. J. (1983). "The conodont animal".
2240: 2215: 2200: 2036: 2028: 2005: 1874: 1816: 1773: 1389: 1373: 1141: 1119: 947: 908: 852: 687: 534: 394: 244: 81: 4185:"An oblique anterior view of a model of the apparatus of the Pennsylvanian conodont 4137: 4041: 3904: 3879: 3838: 3657: 3531:
Briggs, D. (May 1992). "Conodonts: a major extinct group added to the vertebrates".
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Scott, Harold W. (1942). "Conodont Assemblages from the Heath Formation, Montana".
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Only a few studies approach the question of conodont ingroup relationships from a
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Philosophical Transactions of the Royal Society of London. B, Biological Sciences
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Preserved articulated association of conodont elements belonging to the species
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have been speculated to be venomous, based on grooves found on some elements.
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with other elements. Studies have concluded that conodonts taxa occupied both
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name was provided for several decades. MacFarlane (1923) described them as an
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Schmidt, Hermann (1934). "Conodonten-Funde in ursprĂĽnglichem Zusammenhang".
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A size comparison of the three conodont species with preserved body fossils.
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and bilaterally opposed (pectiniform, i.e. comb-shaped) Pb and Pa elements.
449: 153: 106: 9639: 4033: 4004: 3975: 3753: 3120: 3101: 3066: 3013: 2488: 3962:; Clarkson, E. N. K.; Clark, N. D. L. (1993). "The anatomy of conodonts". 3560: 3516: 3499: 757:
According to these fossils, conodonts had large eyes, fins with fin rays,
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Lindström, Maurits (1970). "A suprageneric taxonomy of the conodonts".
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Sepkoski, J. J. (2002). "A compendium of fossil marine animal genera".
3631:"False teeth: conodont-vertebrate phylogenetic relationships revisited" 3436: 3419: 3283:
Takahashi, Satoshi; Yamakita, Satoshi; Suzuki, Noritoshi (2019-06-15).
3237: 2974: 2929:"Late Mississippian conodonts from the Bird Spring Formation in Nevada" 2913: 2887: 2861: 2826: 2737:
Green, Owen R. (2001). "Extraction Techniques for Phosphatic Fossils".
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Aldridge, R. J.; Purnell, M. A. (1996). "The conodont controversies".
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Philosophical Transactions of the Royal Society B: Biological Sciences
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A Manual of Practical Laboratory and Field Techniques in Palaeobiology
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Murdock, Duncan J. E.; Smith, M. Paul (2021). Sansom, Robert (ed.).
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Philosophical Transactions of the Royal Society of London, Series B
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and a prolonged period of significant loss of diversity during the
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Harold W. Scott, "The Zoological Relationships of the Conodonts.
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showed definitively that the animals were primitive vertebrates.
4270: 3500:"The Earliest Known Venomous Animals Recognized Among Conodonts" 353:
The teeth-like fossils of the conodont were first discovered by
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Zhuravlev, Andrey V.; Plotitsyn, Artem N. (18 January 2022).
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Eichenberg, W. (1930). "Conodonten aus dem Culm des Harzes".
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Historical Biology: An International Journal of Paleobiology
2573:"The interrelationships of 'complex' conodonts (Vertebrata)" 1400:
area. However, their final demise is more likely related to
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The Great Fossil Enigma: The Search for the Conodont Animal
3605:"New evidence for the protoconodont origin of chaetognaths" 3391:
10.1130/0091-7613(1993)021<0375:FMICAT>2.3.CO;2
644:, at one time considered the first conodont body fossil. 2267:"Phylum Conodonta Pander, 1856 and Nomenclatural Priority" 926:
Moreover, some analyses do not regard conodonts as either
4197:"'The Jaws That Catch': an Introduction to the Conodonta" 4122:
10.1666/0022-3360(2001)075<1174:CPPF>2.0.CO;2
3582:. Victoria, Australia: Blackwell Publishing. p. 88. 2819:
10.1666/0022-3360(2001)075<1174:CPPF>2.0.CO;2
2538:
10.1666/0022-3360(2001)075<1174:CPPF>2.0.CO;2
777:
had lengths only reaching 4–5 cm (1.6–2.0 in),
345:
as indicative of particular periods of geological time.
3341:
Cambrian Ocean World: Ancient Sea Life of North America
2463:
Donoghue, P.C.J.; Forey, P.L.; Aldridge, R.J. (2000).
3078: 3076: 1388:
Carboniferous due to cooler temperatures, especially
1244:
Conodont elements from the Deer Valley Member of the
877:
As of 2012, scientists classify the conodonts in the
337:(over 500 million years ago) to the beginning of the 4919: 4874: 4841: 4823: 4673: 4595: 4522: 4489: 4377: 4366: 4357: 4350: 4335: 4313: 4296: 2214:
Sweet, Walter C.; Cooper, Barry J. (December 2008).
1376:, with an extinction event at the end of the middle 989: 283: 269: 9597: 9494: 9299: 9271: 9216: 9167: 9133: 9087: 9066: 9045: 9018: 8956: 8901: 8814: 8731: 8641: 8632: 8548: 8521: 8494: 8473: 8439: 8405: 8168: 8147: 8098: 8064: 8001: 7981: 7929: 7881: 7811: 7749: 7680: 7646: 7636: 7552: 7524: 7476: 7417: 7388: 7277: 7265: 7223: 7189: 7168: 7113: 7063: 7022: 7010: 6934: 6913: 6864: 6822: 6788: 6754: 6719: 6685: 6546: 6482: 6437: 6396: 6341: 6313: 6143: 6039: 5961: 5884: 5834: 5825: 5768: 5709: 5661: 5629: 5596: 5532: 5495: 5485: 5476: 5427: 5399: 5323: 5314: 5225: 5118: 5004: 4956: 4865: 4814: 4805: 4713: 4672: 4594: 4585: 4521: 4488: 4458: 4448: 4284: 3878:Ginot, Samuel; Goudemand, Nicolas (December 2020). 1293:, alpha morphotype, Pa element, lateral view, X140 1272:, gamma morphotype, Pa element, lateral view, X140 4048:Gould, Stephen Jay (1985). "Reducing Riddles". In 3083:Conway Morris, Simon; Caron, Jean-Bernard (2022). 3289:Palaeogeography, Palaeoclimatology, Palaeoecology 2174: 2172: 1248:in Pennsylvania, Maryland, and West Virginia, US 923:, an unrelated phylum that includes arrow worms. 911:, appear to form a distinct clade from the later 2566: 2564: 2562: 672:area. That animal is now regarded as a possible 4052:, 245-260. New York, W.W. Norton and Company. 3706:Proceedings of the National Academy of Sciences 3413: 3411: 3409: 3407: 3188: 3186: 3184: 3182: 3134: 3132: 3130: 2458: 2456: 2454: 2452: 2450: 2302: 2300: 2260: 2258: 2216:"C.H. Pander's introduction to conodonts, 1856" 1637:Bassler, 1925 emend. Ulrich & Bassler, 1926 490:where they are known, in rocks dating from the 3344:. Indiana University Press. pp. 300–301. 2350:. In Moore, Raymond C.; Robison, R.A. (eds.). 1404:, perhaps competition with new Mesozoic taxa. 5055: 4244: 2886:, Vol. 8, No. 4 (Dec., 1934), pages 448-455 ( 2512: 2510: 2508: 2506: 8: 3684:: CS1 maint: multiple names: authors list ( 2420: 2418: 2416: 2353:Part W, Miscellanea, Supplement 2: Conodonta 2138:Timeline of the evolutionary history of life 2431:Oxford Monographs on Geology and Geophysics 810:marks likely created by food as well as by 9585: 9549: 9130: 9117: 8638: 7643: 7473: 7460: 6434: 6421: 5831: 5774: 5635: 5492: 5482: 5473: 5460: 5320: 5231: 5115: 5104: 5062: 5048: 5040: 4811: 4591: 4455: 4251: 4237: 4229: 4086:(Indiana University Press; 2012) 440 pages 2465:"Conodont affinity and chordate phylogeny" 1356:, M element, lateral views, 93RS–67, X140 884:on the basis of their fins with fin rays, 761:and axial line, which were interpreted as 122: 31: 16:Extinct agnathan chordates resembling eels 4101:Sweet, W. C.; Donoghue, P. C. J. (2001). 3903: 3743: 3725: 3515: 3435: 3110: 3100: 3021: 2517:Sweet, W. C.; Donoghue, P. C. J. (2001). 1365:, Pa element, lateral view, 93RS–67, X140 2988:Purnell, M. A.; Donoghue, P. C. (1997). 1239: 349:Discovery and understanding of conodonts 2168: 567: 505: 3677: 1934:Bardashev, Weddige & Ziegler, 2002 1265:, Sa element, posterior view, X140 2. 833:A study on the population dynamics of 4103:"Conodonts: past, present and future" 2358:Treatise on Invertebrate Paleontology 656:, an enigmatic animal known from the 515:Complete element set of the conodont 478:. Conodont elements are also used as 415:Treatise on Invertebrate Paleontology 7: 9692:32A17EFD-164F-401D-8813-80A784E6A79E 9569: 3498:Szaniawski, Hubert (December 2009). 2348:"Chapter 3: Systematic Descriptions" 3958:Aldridge, R. J.; Briggs, D. E. G.; 3037:Conway Morris, Simon (1990-04-12). 2577:Journal of Systematic Palaeontology 1134: 1112: 1088: 1037: 1030: 999: 969: 962: 460:fossils of the host organism, in a 4166:10.1111/j.1502-3931.1970.tb00834.x 4026:10.1111/j.1469-185X.1999.tb00045.x 3861:Bulletins of American Paleontology 2646:10.1111/j.1502-3931.1983.tb01993.x 2519:"Conodonts: Past, Present, Future" 2481:10.1111/j.1469-185X.1999.tb00045.x 2399:10.1111/j.1502-3931.1984.tb00675.x 2325:10.1111/j.1502-3931.1976.tb00966.x 1456:Khodalevich & Tschernich, 1973 1284:, Sa element, lateral views, X140 14: 3365:Purnell, Mark A. (1 April 1993). 1733:Lindström, 1970 sensu Sweet, 1988 1302:, Pa element, lateral view, X140 378:as "conodonts", though no formal 9568: 9559: 9558: 9548: 4435: 4429: 4424: 4419: 4413: 4407: 4401: 4394: 4147:from the original on 2022-10-30. 2555:from the original on 2022-10-30. 2122: 2078:Harris & Hollingsworth, 1933 1992:Shen, Yuan & Henderson, 2015 903:analysis suggests they are more 631: 619: 604: 587: 570: 525: 508: 486:. This has made them useful for 140: 3985:Trends in Ecology and Evolution 3905:10.1016/j.gloplacha.2020.103325 3839:10.1016/j.earscirev.2021.103699 1320:, Pa element, upper view, X140 467:These microfossils are made of 369:Nomenclature and taxonomic rank 1701:Repetski & Ethington, 1983 1530:Pander, 1856 non Branson, 1938 1: 3612:Acta Palaeontologica Polonica 3504:Acta Palaeontologica Polonica 3477:10.1080/08912963.2018.1427088 2792:10.1016/j.chemgeo.2016.03.023 2681:10.1016/j.chemgeo.2006.03.004 357:and the results published in 4996:List of transitional fossils 4070:The tree of life web project 3997:10.1016/0169-5347(96)10048-3 3785:10.1016/j.palwor.2022.01.001 3309:10.1016/j.palaeo.2019.03.034 2850:Paläontologische Zeitschrift 2747:10.1007/978-94-017-0581-3_27 2181:Paläontologische Zeitschrift 1228:perspective, as informed by 868:Classification and phylogeny 9720:Furongian first appearances 3884:Global and Planetary Change 3338:Foster, John (2014-06-06). 1448:FĂĄhraeus & Hunter, 1985 539:(Ordovician, North America) 9746: 4276:Timeline of fish evolution 4214:Jim Davison (2002-10-15). 3926:Mikko's Phylogeny Archive 2381:FĂĄhraeus, Lars E. (1984). 2265:FĂĄhraeus, Lars E. (1983). 2009:Klapper & Philip, 1971 938:, more closely related to 858:Some species of the genus 18: 9725:Late Triassic extinctions 9544: 9129: 9116: 7472: 7459: 6433: 6420: 5777: 5638: 5472: 5459: 5234: 5114: 5103: 5077: 5030: 4964:Lists of prehistoric fish 4392: 4266: 2708:10.1017/s0022336000035319 2589:10.1017/S1477201907002234 2307:Bengtson, Stefan (1976). 2158:Conodont alteration index 2094:Austin & Rhodes, 1981 2086:Austin & Rhodes, 1981 2062:Austin & Rhodes, 1981 1809:MĂĽller & MĂĽller, 1957 1646:Wang & Aldridge, 2010 1628:Stouge & Bagnoli 1999 1501:Wang & Aldridge, 2010 1154: 1139: 1132: 1117: 1110: 1093: 1086: 1057: 1042: 1035: 1028: 1004: 997: 974: 967: 683:Clydagnathus windsorensis 555:The feeding apparatus of 484:conodont alteration index 266: 259: 214: 209: 137:Scientific classification 135: 130: 121: 34: 2346:Clark, David L. (1981). 2245:"The quick and the dead" 2153:Conodont biostratigraphy 1983:Branson & Mehl, 1944 1925:Branson & Mehl, 1944 1846:Branson & Mehl, 1944 551:Elements of ozarkodinids 359:Saint Petersburg, Russia 19:Not to be confused with 5022:Vertebrate paleontology 4110:Journal of Paleontology 3930:Haaramo, Mikko (2007). 3727:10.1073/pnas.1814794116 3603:Szaniawski, H. (2002). 3578:(2004). "Vertebrates". 3553:10.1126/science.1598571 3145:Papers in Palaeontology 2963:Journal of Paleontology 2933:Journal of Paleontology 2927:Dunn, David L. (1965). 2902:Journal of Paleontology 2884:Journal of Paleontology 2807:Journal of Paleontology 2696:Journal of Paleontology 2526:Journal of Paleontology 2148:List of conodont genera 476:used in biostratigraphy 4216:"Ordovician conodonts" 4089:Sweet, Walter (1988). 3976:10.1098/rstb.1993.0082 3102:10.1098/rsbl.2022.0179 3067:10.1098/rstb.1990.0102 3014:10.1098/rstb.1997.0141 1664:Pomešano-Cherchi, 1967 1368: 806: 759:chevron-shaped muscles 719:Panderodus unicostatus 613:Panderodus unicostatus 597:Panderodus unicostatus 355:Heinz Christian Pander 9674:Paleobiology Database 5109:Conodont-like fossils 3934:Conodonta - conodonts 3819:Earth-Science Reviews 3517:10.4202/app.2009.0045 2425:Sweet, W. C. (1988). 1246:Mauch Chunk Formation 1243: 1230:phylogenetic analyses 802:Manticolepis subrecta 799:Model of elements of 798: 488:petroleum exploration 462:konservat lagerstätte 8958:Spathognathodontidae 7931:Plectospathodontinae 6145:Protopanderodontidae 5017:Transitional fossils 4656:Pseudopetalichthyida 4050:The Flamingo's Smile 2741:. pp. 318–330. 2014:Spathognathodontidae 1577:Protopanderodontidae 1236:Evolutionary history 888:-shaped muscles and 658:Bear Gulch limestone 577:Life restoration of 326:, classified in the 8903:Pterospathodontidae 7478:Anchignathodontidae 7024:Paracordylodontidae 5963:Drepanoistodontidae 5827:Protopanderodontida 5401:Westergaardodinidae 4064:Janvier, P (1997). 3896:2020GPC...19503325G 3831:2021ESRv..22003699S 3718:2019PNAS..116.2146M 3580:Fossils at a Glance 3545:1992Sci...256.1285B 3539:(5061): 1285–1286. 3383:1993Geo....21..375P 3301:2019PPP...524..212T 3254:2007Geo....35..879V 3207:1995Natur.374..800G 3059:1990RSPTB.327..595M 3006:1997RSPTB.352.1545P 3000:(1361): 1545–1564. 2939:: 6. Archived from 2784:2016ChGeo.431...36Q 2673:2006ChGeo.233..196T 2130:Paleontology portal 2021:Pterospathodontidae 1894:Anchignathodontidae 1669:Paracordylodontidae 1560:Drepanoistodontidae 1536:Protopanderodontida 1402:biotic interactions 1045:Protopanderodontida 564:The conodont animal 445:mineralized tissues 324:jawless vertebrates 235:Protopanderodontida 9730:Cambrian chordates 9401:Neoplectospathodus 8551:Novispathodontidae 8441:Idiognathodontidae 8150:Gladigondolellidae 7883:Neoprioniodontinae 7435:Bergstroemognathus 7225:Polyplacognathidae 6866:Multioistodontidae 6343:Strachanognathidae 5952:Paraserratognathus 5438:"true conodonts" ( 4705:Ischnacanthiformes 4553:Cephalaspidomorphi 4014:Biological Reviews 3437:10.1017/pab.2022.9 2862:10.1007/BF03041668 2469:Biological Reviews 2271:Systematic Zoology 2193:10.1007/BF03044446 2075:Idiognathodontidae 1970:Novispathodontidae 1948:Gladigondolellidae 1782:Polyplacognathidae 1706:Multioistodontidae 1594:Strachanognathidae 1394:marine regressions 1369: 807: 701:Promissum pulchrum 692:Granton Shrimp Bed 580:Promissum pulchrum 131:Conodont elements 9702: 9701: 9591:Taxon identifiers 9582: 9581: 9540: 9539: 9536: 9535: 9112: 9111: 9108: 9107: 9048:Trucherognathidae 8912:Astropentagnathus 8897: 8896: 8694:Pseudopolygnathus 8673:Ferganaegnathodus 8200:Carnepigondolella 7977: 7976: 7940:Anastrophognathus 7892:Leptochirognathus 7455: 7454: 7451: 7450: 7344:Guizhouprioniodus 7234:Baltoplacognathus 7124:Acanthocordylodus 6882:Leptochirognathus 6416: 6415: 6412: 6411: 6408: 6407: 6392: 6391: 6347:(Cornuodontidae)? 6325:Serratognathoides 6001:Laurentoscandodus 5764: 5763: 5760: 5759: 5625: 5624: 5565:Drepanodistacodus 5455: 5454: 5451: 5450: 5423: 5422: 5419: 5418: 5369:Problematoconites 5134: 5037: 5036: 4952: 4951: 4948: 4947: 4879:Palaeonisciformes 4827:Onychodontiformes 4784:Petalodontiformes 4778:Iniopterygiformes 4741:Phoebodontiformes 4729:Ctenacanthiformes 4699:Diplacanthiformes 4581: 4580: 4354:Myllokunmingiidae 4260:Evolution of fish 3970:(1294): 405–421. 3650:10.5252/g2010n4a1 3589:978-0-632-06047-4 3351:978-0-253-01188-6 3262:10.1130/g23894a.1 3201:(6525): 800–803. 3157:10.1002/spp2.1389 3053:(1242): 595–624. 3041:Typhloesus wellsi 2756:978-90-481-4013-8 2440:978-0-19-504352-5 2143:Micropaleontology 2103: 2095: 2087: 2079: 2071: 2063: 2049: 2041: 2033: 2025: 2010: 2002: 1993: 1984: 1979:Trucherognathidae 1975: 1966: 1953: 1944: 1935: 1926: 1917: 1908: 1899: 1890: 1879: 1871: 1863: 1855: 1847: 1839: 1830: 1821: 1810: 1802: 1794: 1786: 1778: 1767: 1758: 1750: 1742: 1734: 1726: 1718: 1710: 1702: 1692: 1683: 1674: 1665: 1656: 1647: 1638: 1629: 1620: 1610: 1599: 1590: 1589:Zhen et al., 2009 1581: 1573: 1565: 1556: 1548: 1540: 1531: 1519: 1510: 1502: 1497:Pseudooneotodidae 1493: 1485: 1477: 1466: 1457: 1449: 1440: 1430: 1422: 1398:continental shelf 1221: 1220: 1212: 1211: 1203: 1202: 1194: 1193: 1185: 1184: 1176: 1175: 1167: 1166: 1070: 1069: 954:Ingroup relations 948:jawed vertebrates 818:(open ocean) and 767:dorsal nerve cord 611:A body fossil of 480:paleothermometers 434:Conodont elements 294: 293: 280: 205: 46:Earliest Jurassic 9737: 9695: 9694: 9682: 9681: 9669: 9668: 9656: 9655: 9643: 9642: 9633: 9632: 9631: 9618: 9617: 9616: 9586: 9572: 9571: 9562: 9561: 9552: 9551: 9248:Parapachycladina 9131: 9118: 9069:Vjalovognathidae 8881:Streptognathodus 8839:Ctenopolygnathus 8701:Rhachistognathus 8639: 8623:Triassospathodus 8389:Scythogondolella 8375:Pseudofurnishius 8186:Ancyrogondolella 8004:Eognathodontidae 7644: 7474: 7461: 7428:Appalachignathus 7419:Rhipidognathidae 7256:Polyplacognathus 6897:Neomultioistodus 6824:Jablonnodontidae 6721:Gamachignathidae 6557:Aldridgeognathus 6435: 6422: 6383:Strachanognathus 6315:Serratognathidae 6191:Molloydenticulus 5909:Eoserratognathus 5832: 5775: 5711:Fryxellodontidae 5636: 5493: 5483: 5474: 5461: 5434:Conodontophorida 5410:Westergaardodina 5383:Prosagittodontus 5321: 5232: 5124: 5120:Protoconodontida 5116: 5105: 5064: 5057: 5050: 5041: 5012:Prehistoric life 4923:Pycnodontiformes 4921: 4906:Acipenseriformes 4876: 4843: 4825: 4812: 4675: 4620:Brindabellaspida 4597: 4592: 4535:Pteraspidomorphi 4524: 4491: 4456: 4440: 4439: 4433: 4428: 4423: 4418: 4417: 4411: 4406: 4405: 4398: 4379: 4368: 4359: 4352: 4337: 4315: 4298: 4253: 4246: 4239: 4230: 4225: 4223: 4222: 4210: 4208: 4207: 4192: 4169: 4148: 4146: 4116:(6): 1174–1184. 4107: 4082:Knell, Simon J. 4079: 4077: 4076: 4045: 4008: 3979: 3946: 3945: 3943: 3942: 3924: 3918: 3917: 3907: 3875: 3869: 3868: 3856: 3850: 3849: 3847: 3845: 3810: 3804: 3803: 3801: 3799: 3764: 3758: 3757: 3747: 3729: 3712:(6): 2146–2151. 3696: 3690: 3689: 3683: 3675: 3673: 3672: 3666: 3660:. Archived from 3635: 3626: 3620: 3619: 3609: 3600: 3594: 3593: 3571: 3565: 3564: 3528: 3522: 3521: 3519: 3495: 3489: 3488: 3464: 3458: 3457: 3439: 3415: 3402: 3401: 3399: 3397: 3362: 3356: 3355: 3335: 3329: 3328: 3280: 3274: 3273: 3233: 3227: 3226: 3215:10.1038/374800a0 3190: 3177: 3176: 3151:(4): 1977–1993. 3136: 3125: 3124: 3114: 3104: 3080: 3071: 3070: 3034: 3028: 3027: 3025: 2985: 2979: 2978: 2969:(6): 1557–1560. 2958: 2952: 2951: 2949: 2948: 2924: 2918: 2917: 2897: 2891: 2880: 2874: 2873: 2845: 2839: 2838: 2802: 2796: 2795: 2772:Chemical Geology 2767: 2761: 2760: 2734: 2728: 2727: 2691: 2685: 2684: 2667:(3–4): 196–216. 2661:Chemical Geology 2656: 2650: 2649: 2629: 2623: 2622: 2620: 2618: 2607: 2601: 2600: 2568: 2557: 2556: 2554: 2532:(6): 1174–1184. 2523: 2514: 2501: 2500: 2460: 2445: 2444: 2422: 2411: 2410: 2378: 2372: 2371: 2343: 2337: 2336: 2304: 2295: 2294: 2262: 2253: 2252: 2233: 2227: 2226: 2224: 2222: 2211: 2205: 2204: 2187:(3–4): 177–182. 2176: 2132: 2127: 2126: 2125: 2101: 2093: 2085: 2077: 2069: 2061: 2047: 2039: 2031: 2023: 2016: 2008: 2000: 1991: 1988:Vjalovognathidae 1982: 1973: 1960: 1951: 1942: 1933: 1930:Eognathodontidae 1924: 1915: 1906: 1897: 1888: 1877: 1869: 1861: 1853: 1845: 1837: 1828: 1819: 1808: 1800: 1792: 1784: 1776: 1765: 1756: 1748: 1740: 1732: 1730:Rhipidognathidae 1724: 1716: 1708: 1700: 1690: 1681: 1672: 1663: 1654: 1651:Jablonnodontidae 1645: 1642:Gamachignathidae 1636: 1627: 1618: 1608: 1597: 1588: 1585:Serratognathidae 1579: 1571: 1563: 1554: 1546: 1538: 1529: 1517: 1508: 1500: 1491: 1489:Fryxellodontidae 1483: 1475: 1464: 1455: 1447: 1438: 1428: 1420: 1135: 1113: 1089: 1038: 1031: 1000: 991: 970: 963: 748:Akkamori section 635: 623: 608: 591: 574: 529: 518:Hindeodus parvus 512: 474:They are widely 409:to euconodonts. 285: 275: 273:Conodontophorida 271: 200: 195: 145: 144: 126: 116: 53: 38:Temporal range: 32: 9745: 9744: 9740: 9739: 9738: 9736: 9735: 9734: 9705: 9704: 9703: 9698: 9690: 9685: 9677: 9672: 9664: 9659: 9651: 9646: 9638: 9636: 9627: 9626: 9621: 9612: 9611: 9606: 9593: 9583: 9578: 9532: 9490: 9450:Prionognathodus 9295: 9267: 9212: 9163: 9135:Bactrognathidae 9125: 9104: 9090:Wapitiodontidae 9083: 9062: 9057:Trucherognathus 9041: 9020:Sweetognathidae 9014: 8988:Spathognathodus 8952: 8893: 8888:Synclydagnathus 8867:Parapolygnathus 8810: 8727: 8628: 8616:Sweetospathodus 8544: 8517: 8490: 8469: 8435: 8401: 8347:Parvigondolella 8277:Metapolygnathus 8164: 8159:Gladigondolella 8143: 8094: 8066:Elictognathidae 8060: 8048:Schmidtognathus 8034:Pseudogondwania 7997: 7973: 7968:Plectospathodus 7925: 7877: 7843:Hindeodelloides 7807: 7745: 7733:Tetraprioniodus 7712:Hibbardelloides 7676: 7632: 7548: 7527:Archeognathidae 7520: 7508:Pseudohindeodus 7468: 7447: 7413: 7384: 7337:Eoneoprioniodus 7279:Prioniodontidae 7273: 7261: 7219: 7185: 7164: 7159:Protophragmodus 7115:Phragmodontidae 7109: 7059: 7054:Protoprioniodus 7018: 7006: 6930: 6909: 6875:Dischidognathus 6860: 6834:Cononcfiodontus 6818: 6784: 6750: 6715: 6687:Distomodontidae 6681: 6585:Barrandegnathus 6542: 6478: 6429: 6404: 6388: 6346: 6337: 6309: 6240:Pseudooneotodus 6233:Protopanderodus 6139: 6046:Panderodontidae 6043: 6035: 5969:Distacodontidae 5966: 5957: 5880: 5836:Acanthodontidae 5821: 5756: 5705: 5663:Cordylodontidae 5657: 5651:Pseudooneotodus 5621: 5598:Dapsilodontidae 5592: 5558:Coelocerodontus 5528: 5468: 5447: 5437: 5432: 5415: 5395: 5310: 5227:Paraconodontida 5221: 5123: 5110: 5099: 5073: 5068: 5038: 5033: 5026: 5000: 4944: 4892:Polypteriformes 4861: 4855:Tetrapodomorpha 4801: 4735:Hybodontiformes 4709: 4686:Acanthodiformes 4668: 4577: 4517: 4507:Paraconodontida 4484: 4452: 4444: 4443: 4442: 4434: 4412: 4400: 4390: 4309:Cephalochordate 4280: 4262: 4257: 4220: 4218: 4213: 4205: 4203: 4195: 4179: 4176: 4151: 4144: 4105: 4100: 4095:Clarendon Press 4074: 4072: 4063: 4011: 3991:(11): 463–468. 3982: 3957: 3954: 3952:Further reading 3949: 3940: 3938: 3929: 3925: 3921: 3877: 3876: 3872: 3858: 3857: 3853: 3843: 3841: 3812: 3811: 3807: 3797: 3795: 3766: 3765: 3761: 3698: 3697: 3693: 3676: 3670: 3668: 3664: 3633: 3628: 3627: 3623: 3607: 3602: 3601: 3597: 3590: 3574:Milsom, Clare; 3573: 3572: 3568: 3530: 3529: 3525: 3497: 3496: 3492: 3466: 3465: 3461: 3417: 3416: 3405: 3395: 3393: 3364: 3363: 3359: 3352: 3337: 3336: 3332: 3282: 3281: 3277: 3235: 3234: 3230: 3192: 3191: 3180: 3138: 3137: 3128: 3089:Biology Letters 3082: 3081: 3074: 3036: 3035: 3031: 2987: 2986: 2982: 2960: 2959: 2955: 2946: 2944: 2926: 2925: 2921: 2899: 2898: 2894: 2881: 2877: 2847: 2846: 2842: 2804: 2803: 2799: 2769: 2768: 2764: 2757: 2736: 2735: 2731: 2693: 2692: 2688: 2658: 2657: 2653: 2631: 2630: 2626: 2616: 2614: 2609: 2608: 2604: 2570: 2569: 2560: 2552: 2521: 2516: 2515: 2504: 2462: 2461: 2448: 2441: 2424: 2423: 2414: 2380: 2379: 2375: 2368: 2345: 2344: 2340: 2306: 2305: 2298: 2283:10.2307/2413175 2264: 2263: 2256: 2243:(Jun 5, 1999). 2237:Jan Zalasiewicz 2235: 2234: 2230: 2220: 2218: 2213: 2212: 2208: 2178: 2177: 2170: 2166: 2128: 2123: 2121: 2118: 2099:Sweetognathidae 2059:Elictognathidae 1997:Wapitiodontidae 1974:(Orchard, 2005) 1903:Archeognathidae 1878:Lindström, 1970 1862:Lindström, 1970 1859:Bactrognathidae 1829:Bergström, 1981 1790:Distomodontidae 1785:Bergström, 1981 1766:Bergstrom, 1981 1749:Bergström, 1981 1746:Phragmodontidae 1738:Prioniodontidae 1725:Lindström, 1970 1717:Lindström, 1970 1691:Bergström, 1981 1673:Bergström, 1981 1633:Distacodontidae 1624:Cahabagnathidae 1611:(paraphyletic) 1598:Bergström, 1981 1580:Lindström, 1970 1572:Lindström, 1970 1569:Panderodontidae 1555:Lindström, 1970 1547:Lindström, 1970 1544:Acanthodontidae 1518:Bergstrom, 1981 1509:Lindström, 1970 1506:Proconodontidae 1484:Lindström, 1970 1481:Cordylodontidae 1461:Dapsilodontidae 1410: 1392:and associated 1367: 1366: 1357: 1348: 1339: 1336:Polygnathus sp. 1333: 1324: 1321: 1312: 1303: 1294: 1285: 1276: 1273: 1257: 1253: 1249: 1238: 1222: 1213: 1204: 1195: 1186: 1177: 1168: 1071: 995: 977:Paraconodontida 956: 875: 870: 793: 645: 636: 627: 624: 615: 609: 600: 594:Restoration of 592: 583: 575: 566: 553: 540: 530: 521: 513: 504: 469:hydroxylapatite 441: 436: 371: 351: 343:biostratigraphy 199: 193: 139: 117: 115: 114: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 42:Middle Cambrian 40: 39: 36: 28: 17: 12: 11: 5: 9743: 9741: 9733: 9732: 9727: 9722: 9717: 9707: 9706: 9700: 9699: 9697: 9696: 9683: 9670: 9657: 9644: 9634: 9619: 9603: 9601: 9595: 9594: 9589: 9580: 9579: 9577: 9576: 9566: 9556: 9545: 9542: 9541: 9538: 9537: 9534: 9533: 9531: 9530: 9523: 9516: 9509: 9501: 9499: 9497:Xaniognathidae 9492: 9491: 9489: 9488: 9481: 9474: 9467: 9460: 9453: 9446: 9439: 9432: 9429:Polygnathellus 9425: 9418: 9411: 9404: 9397: 9390: 9387:Metalonchodina 9383: 9376: 9369: 9362: 9355: 9348: 9341: 9334: 9327: 9320: 9313: 9305: 9303: 9301:Prioniodinidae 9297: 9296: 9294: 9293: 9286: 9278: 9276: 9274:Oepikodontidae 9269: 9268: 9266: 9265: 9258: 9255:Parafurnishius 9251: 9244: 9237: 9230: 9222: 9220: 9214: 9213: 9211: 9210: 9203: 9195: 9188: 9181: 9173: 9171: 9169:Chirognathidae 9165: 9164: 9162: 9161: 9154: 9147: 9139: 9137: 9127: 9126: 9121: 9114: 9113: 9110: 9109: 9106: 9105: 9103: 9102: 9094: 9092: 9085: 9084: 9082: 9081: 9078:Vjalovognathus 9073: 9071: 9064: 9063: 9061: 9060: 9052: 9050: 9043: 9042: 9040: 9039: 9032: 9024: 9022: 9016: 9015: 9013: 9012: 9005: 8998: 8991: 8984: 8977: 8970: 8962: 8960: 8954: 8953: 8951: 8950: 8943: 8940:Pterospathodus 8936: 8929: 8922: 8915: 8907: 8905: 8899: 8898: 8895: 8894: 8892: 8891: 8884: 8877: 8870: 8863: 8860:Pandorinellina 8856: 8849: 8842: 8835: 8828: 8820: 8818: 8812: 8811: 8809: 8808: 8801: 8794: 8787: 8780: 8773: 8766: 8759: 8752: 8745: 8737: 8735: 8733:Palmatolepidae 8729: 8728: 8726: 8725: 8718: 8711: 8704: 8697: 8690: 8683: 8676: 8669: 8662: 8655: 8647: 8645: 8643:Cavusgnathidae 8636: 8630: 8629: 8627: 8626: 8619: 8612: 8605: 8598: 8591: 8584: 8581:Icriospathodus 8577: 8570: 8567:Gongolelloides 8563: 8555: 8553: 8546: 8545: 8543: 8542: 8535: 8527: 8525: 8523:Mestognathidae 8519: 8518: 8516: 8515: 8508: 8505:Ctenognathodus 8500: 8498: 8492: 8491: 8489: 8488: 8480: 8478: 8471: 8470: 8468: 8467: 8464:Protognathodus 8460: 8453: 8445: 8443: 8437: 8436: 8434: 8433: 8430:Peudognathodus 8426: 8419: 8411: 8409: 8407:Gnathodontidae 8403: 8402: 8400: 8399: 8392: 8385: 8378: 8371: 8364: 8357: 8350: 8343: 8336: 8333:Paragondolella 8329: 8326:Norigondolella 8322: 8315: 8308: 8301: 8294: 8287: 8280: 8273: 8270:Mesogondolella 8266: 8259: 8252: 8249:Jinogondolella 8245: 8242:Icriospathodus 8238: 8231: 8224: 8217: 8210: 8203: 8196: 8189: 8182: 8174: 8172: 8166: 8165: 8163: 8162: 8154: 8152: 8145: 8144: 8142: 8141: 8134: 8127: 8120: 8113: 8105: 8103: 8096: 8095: 8093: 8092: 8085: 8078: 8075:Alternognathus 8070: 8068: 8062: 8061: 8059: 8058: 8051: 8044: 8037: 8030: 8023: 8016: 8008: 8006: 7999: 7998: 7996: 7995: 7987: 7985: 7979: 7978: 7975: 7974: 7972: 7971: 7964: 7957: 7950: 7943: 7935: 7933: 7927: 7926: 7924: 7923: 7916: 7909: 7902: 7895: 7887: 7885: 7879: 7878: 7876: 7875: 7872:Synprioniodina 7868: 7861: 7853: 7846: 7839: 7832: 7825: 7822:Euprioniodella 7817: 7815: 7809: 7808: 7806: 7805: 7802:Tasmanognathus 7798: 7791: 7788:Metaprioniodus 7784: 7777: 7770: 7767:Cervicornoides 7763: 7755: 7753: 7751:Hindeodellinae 7747: 7746: 7744: 7743: 7736: 7729: 7722: 7715: 7708: 7701: 7694: 7686: 7684: 7682:Hibbardellinae 7678: 7677: 7675: 7674: 7667: 7660: 7652: 7650: 7641: 7634: 7633: 7631: 7630: 7623: 7616: 7609: 7602: 7595: 7588: 7581: 7574: 7567: 7559: 7557: 7550: 7549: 7547: 7546: 7539: 7531: 7529: 7522: 7521: 7519: 7518: 7515:Synprioniodina 7511: 7504: 7497: 7494:Diplognathodus 7490: 7487:Anchignathodus 7482: 7480: 7470: 7469: 7464: 7457: 7456: 7453: 7452: 7449: 7448: 7446: 7445: 7442:Rhipidognathus 7438: 7431: 7423: 7421: 7415: 7414: 7412: 7411: 7404: 7395: 7393: 7386: 7385: 7383: 7382: 7375: 7368: 7361: 7354: 7347: 7340: 7333: 7326: 7319: 7312: 7305: 7298: 7291: 7283: 7281: 7275: 7274: 7271: 7269: 7263: 7262: 7260: 7259: 7252: 7244: 7237: 7229: 7227: 7221: 7220: 7218: 7217: 7210: 7203: 7195: 7193: 7187: 7186: 7184: 7183: 7175: 7173: 7166: 7165: 7163: 7162: 7155: 7148: 7145:Paraprioniodus 7141: 7134: 7127: 7119: 7117: 7111: 7110: 7108: 7107: 7100: 7093: 7090:Microzarkodina 7086: 7078: 7069: 7067: 7061: 7060: 7058: 7057: 7050: 7047:Paracordylodus 7043: 7036: 7033:Cooperignathus 7028: 7026: 7020: 7019: 7016: 7014: 7008: 7007: 7005: 7004: 6997: 6990: 6983: 6976: 6969: 6962: 6955: 6948: 6940: 6938: 6932: 6931: 6929: 6928: 6920: 6918: 6911: 6910: 6908: 6907: 6900: 6893: 6886: 6878: 6870: 6868: 6862: 6861: 6859: 6858: 6851: 6844: 6837: 6829: 6827: 6820: 6819: 6817: 6816: 6813:Pelekysgnathus 6809: 6802: 6794: 6792: 6786: 6785: 6783: 6782: 6775: 6768: 6760: 6758: 6752: 6751: 6749: 6748: 6745:Gamachignathus 6741: 6734: 6726: 6724: 6717: 6716: 6714: 6713: 6706: 6699: 6691: 6689: 6683: 6682: 6680: 6679: 6676:Trapezognathus 6672: 6669:Sagittodontina 6665: 6662:Rhodesognathus 6658: 6655:Pterospathodus 6651: 6644: 6637: 6630: 6623: 6616: 6609: 6602: 6595: 6588: 6581: 6574: 6567: 6564:Amorphognathus 6560: 6552: 6550: 6544: 6543: 6541: 6540: 6533: 6526: 6519: 6516:Pteracontiodus 6512: 6504: 6497: 6489: 6487: 6480: 6479: 6477: 6476: 6468: 6461: 6454: 6445: 6443: 6440:incertae sedis 6431: 6430: 6427:Prioniodontida 6425: 6418: 6417: 6414: 6413: 6410: 6409: 6406: 6405: 6402: 6400: 6398:Prioniodontida 6394: 6393: 6390: 6389: 6387: 6386: 6379: 6372: 6365: 6358: 6350: 6348: 6339: 6338: 6336: 6335: 6332:Serratognathus 6328: 6320: 6318: 6311: 6310: 6308: 6307: 6300: 6297:Variabiloconus 6293: 6286: 6279: 6272: 6265: 6258: 6251: 6248:Semiacontiodus 6244: 6236: 6229: 6222: 6215: 6208: 6201: 6194: 6187: 6179: 6172: 6165: 6158: 6149: 6147: 6141: 6140: 6138: 6137: 6129: 6126:Taoqupognathus 6122: 6119:Pseudobelodina 6115: 6108: 6105:Parapanderodus 6101: 6094: 6087: 6080: 6073: 6066: 6059: 6051: 6049: 6041:Panderodontida 6037: 6036: 6034: 6033: 6026: 6019: 6012: 6004: 5997: 5994:Drepanoistodus 5990: 5982: 5974: 5972: 5959: 5958: 5956: 5955: 5948: 5940: 5933: 5926: 5919: 5912: 5905: 5898: 5890: 5888: 5886:Clavohamulidae 5882: 5881: 5879: 5878: 5870: 5862: 5855: 5848: 5840: 5838: 5829: 5823: 5822: 5820: 5819: 5811: 5803: 5795: 5787: 5778: 5772: 5766: 5765: 5762: 5761: 5758: 5757: 5755: 5754: 5746: 5738: 5731: 5728:Fryxellodontus 5724: 5715: 5713: 5707: 5706: 5704: 5703: 5696: 5689: 5682: 5675: 5672:Cambrooistodus 5667: 5665: 5659: 5658: 5656: 5655: 5647: 5639: 5633: 5631:Proconodontida 5627: 5626: 5623: 5622: 5620: 5619: 5612: 5603: 5601: 5594: 5593: 5591: 5590: 5583: 5576: 5569: 5561: 5554: 5546: 5538: 5536: 5530: 5529: 5527: 5526: 5518: 5510: 5502: 5500: 5490: 5480: 5470: 5469: 5464: 5457: 5456: 5453: 5452: 5449: 5448: 5445: 5443: 5425: 5424: 5421: 5420: 5417: 5416: 5414: 5413: 5405: 5403: 5397: 5396: 5394: 5393: 5386: 5379: 5372: 5365: 5358: 5351: 5344: 5337: 5329: 5327: 5318: 5312: 5311: 5309: 5308: 5301: 5294: 5287: 5284:Serratocambria 5280: 5272: 5265: 5258: 5251: 5244: 5235: 5229: 5223: 5222: 5220: 5219: 5212: 5205: 5197: 5190: 5182: 5178:Huayuanodontus 5174: 5167: 5159: 5152: 5145: 5137: 5135: 5112: 5111: 5108: 5101: 5100: 5098: 5097: 5091: 5085: 5078: 5075: 5074: 5069: 5067: 5066: 5059: 5052: 5044: 5035: 5034: 5031: 5028: 5027: 5025: 5024: 5019: 5014: 5008: 5006: 5002: 5001: 4999: 4998: 4993: 4992: 4991: 4986: 4981: 4976: 4971: 4960: 4958: 4954: 4953: 4950: 4949: 4946: 4945: 4943: 4942: 4941: 4940: 4935: 4930: 4925: 4911: 4910: 4909: 4908: 4897: 4896: 4895: 4894: 4883: 4882: 4871: 4869: 4863: 4862: 4860: 4859: 4858: 4857: 4852: 4847: 4845:Porolepiformes 4834: 4829: 4820: 4818: 4809: 4803: 4802: 4800: 4799: 4798: 4797: 4792: 4790:Symmoriiformes 4786: 4780: 4774: 4772:Eugeneodontida 4762: 4761: 4760: 4759: 4754: 4749: 4743: 4737: 4731: 4723:Elasmobranchii 4719: 4717: 4711: 4710: 4708: 4707: 4701: 4695: 4692:Climatiiformes 4688: 4681: 4679: 4670: 4669: 4667: 4666: 4659: 4652: 4650:Acanthothoraci 4646: 4640: 4634: 4628: 4626:Petalichthyida 4622: 4616: 4610: 4603: 4601: 4589: 4583: 4582: 4579: 4578: 4576: 4575: 4574: 4573: 4567: 4561: 4549: 4543: 4537: 4530: 4528: 4519: 4518: 4516: 4515: 4509: 4503: 4500:Protoconodonta 4495: 4493: 4486: 4485: 4483: 4482: 4481: 4480: 4470: 4464: 4462: 4453: 4446: 4445: 4393: 4391: 4389: 4388: 4387: 4386: 4375: 4371:Myllokunmingia 4348: 4343: 4328: 4327: 4326: 4321: 4306: 4305: 4304: 4290: 4288: 4282: 4281: 4279: 4278: 4273: 4267: 4264: 4263: 4258: 4256: 4255: 4248: 4241: 4233: 4227: 4226: 4211: 4193: 4175: 4174:External links 4172: 4171: 4170: 4160:(4): 427–445. 4149: 4098: 4087: 4080: 4061: 4046: 4020:(2): 191–251. 4009: 3980: 3960:Smith, M. Paul 3953: 3950: 3948: 3947: 3919: 3870: 3851: 3805: 3779:(4): 633–645. 3759: 3691: 3644:(4): 545–594. 3621: 3595: 3588: 3566: 3523: 3510:(4): 669–676. 3490: 3459: 3430:(4): 601–621. 3403: 3377:(4): 375–377. 3357: 3350: 3330: 3275: 3228: 3178: 3126: 3072: 3029: 2980: 2953: 2919: 2908:(3): 293–300. 2892: 2875: 2856:(1–2): 76–85. 2840: 2813:(2): 227–235. 2797: 2762: 2755: 2729: 2702:(4): 790–794. 2686: 2651: 2624: 2602: 2583:(2): 119–153. 2558: 2502: 2475:(2): 191–251. 2446: 2439: 2412: 2393:(4): 293–305. 2373: 2366: 2338: 2319:(2): 185–206. 2296: 2277:(4): 455–459. 2254: 2228: 2206: 2167: 2165: 2162: 2161: 2160: 2155: 2150: 2145: 2140: 2134: 2133: 2117: 2114: 2113: 2112: 2111: 2110: 2109: 2108: 2107: 2106: 2105: 2104: 2096: 2091:Cavusgnathidae 2088: 2083:Mestognathidae 2080: 2072: 2067:Gnathodontidae 2064: 2056: 2053:Hindeodontidae 2050: 2045:Palmatolepidae 2042: 2034: 2026: 2018: 2011: 2003: 1994: 1985: 1976: 1967: 1954: 1952:(Hirsch, 1994) 1945: 1936: 1927: 1918: 1909: 1900: 1882: 1881: 1880: 1872: 1864: 1856: 1851:Prioniodinidae 1848: 1843:Chirognathidae 1840: 1834:Xaniognathidae 1831: 1825:Oepikodontidae 1813: 1812: 1811: 1803: 1795: 1787: 1779: 1770:Icriodontacea 1768: 1759: 1751: 1743: 1735: 1727: 1719: 1711: 1703: 1693: 1687:Ulrichodinidae 1684: 1675: 1666: 1657: 1648: 1639: 1630: 1621: 1606:Prioniodontida 1602: 1601: 1600: 1591: 1582: 1574: 1566: 1564:FĂĄhraeus, 1978 1557: 1552:Clavohamulidae 1549: 1524: 1523: 1522: 1521: 1520: 1511: 1503: 1494: 1486: 1473:Proconodontida 1469: 1468: 1467: 1458: 1450: 1409: 1406: 1396:which reduced 1390:glacial events 1256:Valley Member. 1254: 1251: 1250: 1237: 1234: 1219: 1218: 1215: 1214: 1210: 1209: 1206: 1205: 1201: 1200: 1197: 1196: 1192: 1191: 1188: 1187: 1183: 1182: 1179: 1178: 1174: 1173: 1170: 1169: 1165: 1164: 1161: 1160: 1153: 1150: 1149: 1146: 1145: 1138: 1133: 1131: 1128: 1127: 1124: 1123: 1116: 1111: 1109: 1106: 1105: 1102: 1101: 1097:Paracordylodus 1092: 1087: 1085: 1082:Prioniodontida 1077: 1076: 1073: 1072: 1068: 1067: 1064: 1063: 1056: 1053: 1052: 1049: 1048: 1041: 1036: 1034: 1029: 1027: 1019: 1018: 1015: 1014: 1011:Proconodontida 1003: 998: 996: 988: 985: 984: 981: 980: 973: 968: 966: 961: 955: 952: 909:protoconodonts 874: 871: 869: 866: 853:trophic niches 836:Alternognathus 792: 789: 728:Waukesha Biota 647: 646: 637: 630: 628: 625: 618: 616: 610: 603: 601: 593: 586: 584: 576: 569: 565: 562: 552: 549: 542: 541: 531: 524: 522: 514: 507: 503: 502:Full apparatus 500: 440: 437: 435: 432: 395:protoconodonts 391:taxonomic rank 370: 367: 350: 347: 292: 291: 290: 289: 281: 264: 263: 257: 256: 255: 254: 253: 252: 247: 242: 240:Panderodontida 237: 227: 226: 225: 223:Proconodontida 212: 211: 207: 206: 191: 187: 186: 181: 177: 176: 171: 167: 166: 161: 157: 156: 151: 147: 146: 133: 132: 128: 127: 119: 118: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 54: 37: 15: 13: 10: 9: 6: 4: 3: 2: 9742: 9731: 9728: 9726: 9723: 9721: 9718: 9716: 9713: 9712: 9710: 9693: 9688: 9684: 9680: 9675: 9671: 9667: 9662: 9658: 9654: 9649: 9645: 9641: 9635: 9630: 9624: 9620: 9615: 9609: 9605: 9604: 9602: 9600: 9596: 9592: 9587: 9575: 9567: 9565: 9557: 9555: 9547: 9546: 9543: 9529: 9528: 9524: 9522: 9521: 9517: 9515: 9514: 9513:Crypridodella 9510: 9508: 9507: 9503: 9502: 9500: 9498: 9493: 9487: 9486: 9482: 9480: 9479: 9475: 9473: 9472: 9471:Subbryantodus 9468: 9466: 9465: 9461: 9459: 9458: 9457:Pristognathus 9454: 9452: 9451: 9447: 9445: 9444: 9440: 9438: 9437: 9433: 9431: 9430: 9426: 9424: 9423: 9419: 9417: 9416: 9412: 9410: 9409: 9405: 9403: 9402: 9398: 9396: 9395: 9394:Multidentodus 9391: 9389: 9388: 9384: 9382: 9381: 9377: 9375: 9374: 9373:Kamuellerella 9370: 9368: 9367: 9366:Idioprionodus 9363: 9361: 9360: 9356: 9354: 9353: 9349: 9347: 9346: 9342: 9340: 9339: 9335: 9333: 9332: 9328: 9326: 9325: 9321: 9319: 9318: 9317:Camptognathus 9314: 9312: 9311: 9307: 9306: 9304: 9302: 9298: 9292: 9291: 9287: 9285: 9284: 9283:Lissoepikodus 9280: 9279: 9277: 9275: 9270: 9264: 9263: 9259: 9257: 9256: 9252: 9250: 9249: 9245: 9243: 9242: 9238: 9236: 9235: 9231: 9229: 9228: 9224: 9223: 9221: 9219: 9215: 9209: 9208: 9204: 9201: 9200: 9196: 9194: 9193: 9189: 9187: 9186: 9182: 9180: 9179: 9175: 9174: 9172: 9170: 9166: 9160: 9159: 9158:Staurognathus 9155: 9153: 9152: 9148: 9146: 9145: 9144:Bactrognathus 9141: 9140: 9138: 9136: 9132: 9128: 9124: 9123:Prioniodinida 9119: 9115: 9101: 9100: 9096: 9095: 9093: 9091: 9086: 9080: 9079: 9075: 9074: 9072: 9070: 9065: 9059: 9058: 9054: 9053: 9051: 9049: 9044: 9038: 9037: 9036:Sweetognathus 9033: 9031: 9030: 9026: 9025: 9023: 9021: 9017: 9011: 9010: 9006: 9004: 9003: 8999: 8997: 8996: 8992: 8990: 8989: 8985: 8983: 8982: 8978: 8976: 8975: 8971: 8969: 8968: 8964: 8963: 8961: 8959: 8955: 8949: 8948: 8944: 8942: 8941: 8937: 8935: 8934: 8930: 8928: 8927: 8923: 8921: 8920: 8916: 8914: 8913: 8909: 8908: 8906: 8904: 8900: 8890: 8889: 8885: 8883: 8882: 8878: 8876: 8875: 8871: 8869: 8868: 8864: 8862: 8861: 8857: 8855: 8854: 8850: 8848: 8847: 8843: 8841: 8840: 8836: 8834: 8833: 8829: 8827: 8826: 8822: 8821: 8819: 8817: 8816:Polygnathidae 8813: 8807: 8806: 8802: 8800: 8799: 8795: 8793: 8792: 8788: 8786: 8785: 8781: 8779: 8778: 8774: 8772: 8771: 8767: 8765: 8764: 8763:Klapperilepis 8760: 8758: 8757: 8753: 8751: 8750: 8746: 8744: 8743: 8739: 8738: 8736: 8734: 8730: 8724: 8723: 8722:Weyerognathus 8719: 8717: 8716: 8715:Taphrognathus 8712: 8710: 8709: 8708:Scaphignathus 8705: 8703: 8702: 8698: 8696: 8695: 8691: 8689: 8688: 8684: 8682: 8681: 8677: 8675: 8674: 8670: 8668: 8667: 8663: 8661: 8660: 8656: 8654: 8653: 8649: 8648: 8646: 8644: 8640: 8637: 8635: 8634:Polygnathacea 8631: 8625: 8624: 8620: 8618: 8617: 8613: 8611: 8610: 8606: 8604: 8603: 8602:Novispathodus 8599: 8597: 8596: 8592: 8590: 8589: 8585: 8583: 8582: 8578: 8576: 8575: 8571: 8569: 8568: 8564: 8562: 8561: 8557: 8556: 8554: 8552: 8547: 8541: 8540: 8536: 8534: 8533: 8532:Laterignathus 8529: 8528: 8526: 8524: 8520: 8514: 8513: 8509: 8507: 8506: 8502: 8501: 8499: 8497: 8496:Kockelellidae 8493: 8487: 8486: 8482: 8481: 8479: 8477: 8476:Iowagnathidae 8472: 8466: 8465: 8461: 8459: 8458: 8457:Idiognathodus 8454: 8452: 8451: 8447: 8446: 8444: 8442: 8438: 8432: 8431: 8427: 8425: 8424: 8420: 8418: 8417: 8413: 8412: 8410: 8408: 8404: 8398: 8397: 8393: 8391: 8390: 8386: 8384: 8383: 8379: 8377: 8376: 8372: 8370: 8369: 8365: 8363: 8362: 8358: 8356: 8355: 8354:Platyvillosus 8351: 8349: 8348: 8344: 8342: 8341: 8337: 8335: 8334: 8330: 8328: 8327: 8323: 8321: 8320: 8316: 8314: 8313: 8309: 8307: 8306: 8305:Neogondolella 8302: 8300: 8299: 8295: 8293: 8292: 8288: 8286: 8285: 8281: 8279: 8278: 8274: 8272: 8271: 8267: 8265: 8264: 8260: 8258: 8257: 8256:Kraussodontus 8253: 8251: 8250: 8246: 8244: 8243: 8239: 8237: 8236: 8232: 8230: 8229: 8225: 8223: 8222: 8221:Epigondolella 8218: 8216: 8215: 8211: 8209: 8208: 8204: 8202: 8201: 8197: 8195: 8194: 8190: 8188: 8187: 8183: 8181: 8180: 8176: 8175: 8173: 8171: 8170:Gondolellidae 8167: 8161: 8160: 8156: 8155: 8153: 8151: 8146: 8140: 8139: 8135: 8133: 8132: 8128: 8126: 8125: 8121: 8119: 8118: 8114: 8112: 8111: 8107: 8106: 8104: 8102: 8101:Francodinidae 8097: 8091: 8090: 8089:Pinacognathus 8086: 8084: 8083: 8082:Elictognathus 8079: 8077: 8076: 8072: 8071: 8069: 8067: 8063: 8057: 8056: 8052: 8050: 8049: 8045: 8043: 8042: 8038: 8036: 8035: 8031: 8029: 8028: 8024: 8022: 8021: 8017: 8015: 8014: 8010: 8009: 8007: 8005: 8000: 7994: 7993: 7989: 7988: 7986: 7984: 7983:Cryptotaxidae 7980: 7970: 7969: 7965: 7963: 7962: 7961:Keislognathus 7958: 7956: 7955: 7951: 7949: 7948: 7947:Aphelognathus 7944: 7942: 7941: 7937: 7936: 7934: 7932: 7928: 7922: 7921: 7920:Subprioniodus 7917: 7915: 7914: 7910: 7908: 7907: 7906:Neoprioniodus 7903: 7901: 7900: 7896: 7894: 7893: 7889: 7888: 7886: 7884: 7880: 7874: 7873: 7869: 7867: 7866: 7862: 7859: 7858: 7854: 7852: 7851: 7847: 7845: 7844: 7840: 7838: 7837: 7833: 7831: 7830: 7829:Euprioniodina 7826: 7824: 7823: 7819: 7818: 7816: 7814: 7810: 7804: 7803: 7799: 7797: 7796: 7792: 7790: 7789: 7785: 7783: 7782: 7778: 7776: 7775: 7771: 7769: 7768: 7764: 7762: 7761: 7757: 7756: 7754: 7752: 7748: 7742: 7741: 7737: 7735: 7734: 7730: 7728: 7727: 7723: 7721: 7720: 7716: 7714: 7713: 7709: 7707: 7706: 7702: 7700: 7699: 7695: 7693: 7692: 7688: 7687: 7685: 7683: 7679: 7673: 7672: 7668: 7666: 7665: 7661: 7659: 7658: 7654: 7653: 7651: 7649: 7648:Coleodontinae 7645: 7642: 7640: 7639:Coleodontidae 7635: 7629: 7628: 7624: 7622: 7621: 7617: 7615: 7614: 7613:Microcoelodus 7610: 7608: 7607: 7603: 7601: 7600: 7596: 7594: 7593: 7589: 7587: 7586: 7585:Cambropustula 7582: 7580: 7579: 7575: 7573: 7572: 7568: 7566: 7565: 7561: 7560: 7558: 7556: 7551: 7545: 7544: 7540: 7538: 7537: 7536:Archeognathus 7533: 7532: 7530: 7528: 7523: 7517: 7516: 7512: 7510: 7509: 7505: 7503: 7502: 7498: 7496: 7495: 7491: 7489: 7488: 7484: 7483: 7481: 7479: 7475: 7471: 7467: 7462: 7458: 7444: 7443: 7439: 7437: 7436: 7432: 7430: 7429: 7425: 7424: 7422: 7420: 7416: 7410: 7409: 7405: 7402: 7401: 7397: 7396: 7394: 7391: 7387: 7381: 7380: 7379:Stiptognathus 7376: 7374: 7373: 7369: 7367: 7366: 7362: 7360: 7359: 7358:Lambdagnathus 7355: 7353: 7352: 7348: 7346: 7345: 7341: 7339: 7338: 7334: 7332: 7331: 7327: 7325: 7324: 7323:Embsaygnathus 7320: 7318: 7317: 7313: 7311: 7310: 7306: 7304: 7303: 7299: 7297: 7296: 7292: 7290: 7289: 7285: 7284: 7282: 7280: 7276: 7270: 7268: 7267:Prioniodinida 7264: 7258: 7257: 7253: 7251: 7249: 7245: 7243: 7242: 7241:Cahabagnathus 7238: 7236: 7235: 7231: 7230: 7228: 7226: 7222: 7216: 7215: 7211: 7209: 7208: 7204: 7202: 7201: 7197: 7196: 7194: 7192: 7191:Plectodinidae 7188: 7182: 7181: 7177: 7176: 7174: 7171: 7170:Playfordiidae 7167: 7161: 7160: 7156: 7154: 7153: 7149: 7147: 7146: 7142: 7140: 7139: 7135: 7133: 7132: 7128: 7126: 7125: 7121: 7120: 7118: 7116: 7112: 7106: 7105: 7101: 7099: 7098: 7094: 7092: 7091: 7087: 7084: 7083: 7079: 7076: 7075: 7071: 7070: 7068: 7066: 7065:Periodontidae 7062: 7056: 7055: 7051: 7049: 7048: 7044: 7042: 7041: 7037: 7035: 7034: 7030: 7029: 7027: 7025: 7021: 7015: 7013: 7009: 7003: 7002: 6998: 6996: 6995: 6991: 6989: 6988: 6984: 6982: 6981: 6977: 6975: 6974: 6973:Loxognathodus 6970: 6968: 6967: 6963: 6961: 6960: 6956: 6954: 6953: 6949: 6947: 6946: 6942: 6941: 6939: 6937: 6936:Oistodontidae 6933: 6927: 6926: 6922: 6921: 6919: 6916: 6912: 6906: 6905: 6901: 6899: 6898: 6894: 6892: 6891: 6890:Multioistodus 6887: 6884: 6883: 6879: 6877: 6876: 6872: 6871: 6869: 6867: 6863: 6857: 6856: 6855:Mitrellataxis 6852: 6850: 6849: 6845: 6843: 6842: 6838: 6836: 6835: 6831: 6830: 6828: 6825: 6821: 6815: 6814: 6810: 6808: 6807: 6803: 6801: 6800: 6796: 6795: 6793: 6791: 6790:Icriodontidae 6787: 6781: 6780: 6776: 6774: 6773: 6769: 6767: 6766: 6762: 6761: 6759: 6757: 6753: 6747: 6746: 6742: 6740: 6739: 6735: 6733: 6732: 6728: 6727: 6725: 6722: 6718: 6712: 6711: 6710:Moskalenkodus 6707: 6705: 6704: 6700: 6698: 6697: 6696:Anticostiodus 6693: 6692: 6690: 6688: 6684: 6678: 6677: 6673: 6671: 6670: 6666: 6664: 6663: 6659: 6657: 6656: 6652: 6650: 6649: 6645: 6643: 6642: 6638: 6636: 6635: 6631: 6629: 6628: 6624: 6622: 6621: 6617: 6615: 6614: 6610: 6608: 6607: 6603: 6601: 6600: 6596: 6594: 6593: 6589: 6587: 6586: 6582: 6580: 6579: 6575: 6573: 6572: 6571:Arianagnathus 6568: 6566: 6565: 6561: 6559: 6558: 6554: 6553: 6551: 6549: 6548:Balognathidae 6545: 6539: 6538: 6534: 6532: 6531: 6527: 6525: 6524: 6520: 6518: 6517: 6513: 6510: 6509: 6505: 6503: 6502: 6498: 6496: 6495: 6491: 6490: 6488: 6485: 6481: 6474: 6473: 6469: 6467: 6466: 6462: 6460: 6459: 6455: 6452: 6451: 6450:Multidentodus 6447: 6446: 6444: 6442: 6441: 6436: 6432: 6428: 6423: 6419: 6401: 6399: 6395: 6385: 6384: 6380: 6378: 6377: 6373: 6371: 6370: 6366: 6364: 6363: 6359: 6357: 6356: 6352: 6351: 6349: 6344: 6340: 6334: 6333: 6329: 6327: 6326: 6322: 6321: 6319: 6316: 6312: 6306: 6305: 6301: 6299: 6298: 6294: 6292: 6291: 6287: 6285: 6284: 6280: 6278: 6277: 6273: 6271: 6270: 6269:Striatodontus 6266: 6264: 6263: 6259: 6257: 6256: 6252: 6250: 6249: 6245: 6242: 6241: 6237: 6235: 6234: 6230: 6228: 6227: 6223: 6221: 6220: 6216: 6214: 6213: 6209: 6207: 6206: 6202: 6200: 6199: 6195: 6193: 6192: 6188: 6185: 6184: 6180: 6178: 6177: 6173: 6171: 6170: 6166: 6164: 6163: 6159: 6156: 6155: 6151: 6150: 6148: 6146: 6142: 6135: 6134: 6130: 6128: 6127: 6123: 6121: 6120: 6116: 6114: 6113: 6109: 6107: 6106: 6102: 6100: 6099: 6095: 6093: 6092: 6088: 6086: 6085: 6081: 6079: 6078: 6077:Neopanderodus 6074: 6072: 6071: 6067: 6065: 6064: 6060: 6058: 6057: 6053: 6052: 6050: 6047: 6042: 6038: 6032: 6031: 6027: 6025: 6024: 6020: 6018: 6017: 6013: 6010: 6009: 6005: 6003: 6002: 5998: 5996: 5995: 5991: 5988: 5987: 5983: 5981: 5980: 5976: 5975: 5973: 5970: 5964: 5960: 5954: 5953: 5949: 5946: 5945: 5941: 5939: 5938: 5934: 5932: 5931: 5930:Hispidodontus 5927: 5925: 5924: 5923:Hirsutodontus 5920: 5918: 5917: 5916:Granatodontus 5913: 5911: 5910: 5906: 5904: 5903: 5899: 5897: 5896: 5892: 5891: 5889: 5887: 5883: 5876: 5875: 5871: 5868: 5867: 5863: 5861: 5860: 5856: 5854: 5853: 5849: 5847: 5846: 5842: 5841: 5839: 5837: 5833: 5830: 5828: 5824: 5817: 5816: 5812: 5809: 5808: 5804: 5801: 5800: 5796: 5793: 5792: 5788: 5785: 5784: 5780: 5779: 5776: 5773: 5771: 5767: 5752: 5751: 5747: 5744: 5743: 5739: 5737: 5736: 5732: 5730: 5729: 5725: 5722: 5721: 5717: 5716: 5714: 5712: 5708: 5702: 5701: 5700:Iapetognathus 5697: 5695: 5694: 5690: 5688: 5687: 5683: 5681: 5680: 5676: 5674: 5673: 5669: 5668: 5666: 5664: 5660: 5653: 5652: 5648: 5646: 5645: 5644:Proconodontus 5641: 5640: 5637: 5634: 5632: 5628: 5618: 5617: 5613: 5610: 5609: 5605: 5604: 5602: 5599: 5595: 5589: 5588: 5584: 5582: 5581: 5577: 5575: 5574: 5570: 5567: 5566: 5562: 5560: 5559: 5555: 5552: 5551: 5547: 5545: 5544: 5540: 5539: 5537: 5535: 5531: 5524: 5523: 5519: 5516: 5515: 5511: 5509: 5508: 5504: 5503: 5501: 5498: 5494: 5491: 5488: 5484: 5481: 5479: 5475: 5471: 5467: 5462: 5458: 5444: 5441: 5435: 5430: 5426: 5412: 5411: 5407: 5406: 5404: 5402: 5398: 5392: 5391: 5387: 5385: 5384: 5380: 5378: 5377: 5373: 5371: 5370: 5366: 5364: 5363: 5359: 5357: 5356: 5352: 5350: 5349: 5345: 5343: 5342: 5338: 5336: 5335: 5331: 5330: 5328: 5326: 5325:Furnishinidae 5322: 5319: 5317: 5316:Furnishinacea 5313: 5307: 5306: 5302: 5300: 5299: 5295: 5293: 5292: 5288: 5286: 5285: 5281: 5278: 5277: 5276:Prodistacodus 5273: 5271: 5270: 5266: 5264: 5263: 5262:Hunanognathus 5259: 5257: 5256: 5252: 5250: 5249: 5245: 5242: 5241: 5237: 5236: 5233: 5230: 5228: 5224: 5218: 5217: 5216:Protohertzina 5213: 5211: 5210: 5206: 5203: 5202: 5198: 5196: 5195: 5191: 5188: 5187: 5183: 5180: 5179: 5175: 5173: 5172: 5168: 5165: 5164: 5160: 5158: 5157: 5153: 5151: 5150: 5146: 5144: 5143: 5139: 5138: 5136: 5132: 5128: 5121: 5117: 5113: 5106: 5102: 5096: 5092: 5090: 5086: 5084: 5080: 5079: 5076: 5072: 5065: 5060: 5058: 5053: 5051: 5046: 5045: 5042: 5029: 5023: 5020: 5018: 5015: 5013: 5010: 5009: 5007: 5003: 4997: 4994: 4990: 4987: 4985: 4982: 4980: 4979:cartilaginous 4977: 4975: 4972: 4970: 4967: 4966: 4965: 4962: 4961: 4959: 4955: 4939: 4936: 4934: 4931: 4929: 4926: 4924: 4918: 4917: 4916: 4913: 4912: 4907: 4904: 4903: 4902: 4899: 4898: 4893: 4890: 4889: 4888: 4885: 4884: 4880: 4873: 4872: 4870: 4868: 4864: 4856: 4853: 4851: 4848: 4846: 4840: 4839: 4838: 4835: 4833: 4830: 4828: 4822: 4821: 4819: 4817: 4813: 4810: 4808: 4804: 4796: 4793: 4791: 4787: 4785: 4781: 4779: 4775: 4773: 4769: 4768: 4767: 4764: 4763: 4758: 4755: 4753: 4750: 4748: 4744: 4742: 4738: 4736: 4732: 4730: 4726: 4725: 4724: 4721: 4720: 4718: 4716: 4715:Cartilaginous 4712: 4706: 4702: 4700: 4696: 4693: 4689: 4687: 4683: 4682: 4680: 4678: 4671: 4664: 4663:Stensioellida 4660: 4657: 4653: 4651: 4647: 4645: 4641: 4639: 4638:Ptyctodontida 4635: 4633: 4629: 4627: 4623: 4621: 4617: 4615: 4611: 4609: 4605: 4604: 4602: 4600: 4593: 4590: 4588: 4584: 4572: 4568: 4566: 4562: 4560: 4556: 4555: 4554: 4550: 4548: 4544: 4542: 4538: 4536: 4532: 4531: 4529: 4527: 4520: 4514: 4510: 4508: 4504: 4501: 4497: 4496: 4494: 4487: 4479: 4476: 4475: 4474: 4471: 4469: 4466: 4465: 4463: 4461: 4457: 4454: 4451: 4447: 4441: 4438: 4432: 4427: 4422: 4416: 4410: 4404: 4397: 4384: 4383: 4382:Zhongxiniscus 4376: 4373: 4372: 4364: 4363: 4362:Haikouichthys 4355: 4349: 4347: 4344: 4342: 4341: 4334: 4333: 4332: 4329: 4325: 4322: 4320: 4319: 4312: 4311: 4310: 4307: 4303: 4302: 4295: 4294: 4293:Basal member 4292: 4291: 4289: 4287: 4283: 4277: 4274: 4272: 4269: 4268: 4265: 4261: 4254: 4249: 4247: 4242: 4240: 4235: 4234: 4231: 4217: 4212: 4202: 4198: 4194: 4190: 4188: 4187:Idiognathodus 4182: 4178: 4177: 4173: 4167: 4163: 4159: 4155: 4150: 4143: 4139: 4135: 4131: 4127: 4123: 4119: 4115: 4111: 4104: 4099: 4096: 4092: 4088: 4085: 4081: 4071: 4067: 4066:"Euconodonta" 4062: 4059: 4058:0-393-30375-6 4055: 4051: 4047: 4043: 4039: 4035: 4031: 4027: 4023: 4019: 4015: 4010: 4006: 4002: 3998: 3994: 3990: 3986: 3981: 3977: 3973: 3969: 3965: 3961: 3956: 3955: 3951: 3937: 3935: 3928: 3923: 3920: 3915: 3911: 3906: 3901: 3897: 3893: 3889: 3885: 3881: 3874: 3871: 3866: 3862: 3855: 3852: 3840: 3836: 3832: 3828: 3824: 3820: 3816: 3809: 3806: 3794: 3790: 3786: 3782: 3778: 3774: 3770: 3763: 3760: 3755: 3751: 3746: 3741: 3737: 3733: 3728: 3723: 3719: 3715: 3711: 3707: 3703: 3695: 3692: 3687: 3681: 3667:on 2012-03-19 3663: 3659: 3655: 3651: 3647: 3643: 3639: 3638:Geodiversitas 3632: 3625: 3622: 3617: 3613: 3606: 3599: 3596: 3591: 3585: 3581: 3577: 3570: 3567: 3562: 3558: 3554: 3550: 3546: 3542: 3538: 3534: 3527: 3524: 3518: 3513: 3509: 3505: 3501: 3494: 3491: 3486: 3482: 3478: 3474: 3470: 3463: 3460: 3455: 3451: 3447: 3443: 3438: 3433: 3429: 3425: 3421: 3414: 3412: 3410: 3408: 3404: 3392: 3388: 3384: 3380: 3376: 3372: 3368: 3361: 3358: 3353: 3347: 3343: 3342: 3334: 3331: 3326: 3322: 3318: 3314: 3310: 3306: 3302: 3298: 3294: 3290: 3286: 3279: 3276: 3271: 3267: 3263: 3259: 3255: 3251: 3247: 3243: 3239: 3232: 3229: 3224: 3220: 3216: 3212: 3208: 3204: 3200: 3196: 3189: 3187: 3185: 3183: 3179: 3174: 3170: 3166: 3162: 3158: 3154: 3150: 3146: 3142: 3135: 3133: 3131: 3127: 3122: 3118: 3113: 3108: 3103: 3098: 3094: 3090: 3086: 3079: 3077: 3073: 3068: 3064: 3060: 3056: 3052: 3048: 3044: 3042: 3033: 3030: 3024: 3019: 3015: 3011: 3007: 3003: 2999: 2995: 2991: 2984: 2981: 2976: 2972: 2968: 2964: 2957: 2954: 2943:on 2016-08-18 2942: 2938: 2934: 2930: 2923: 2920: 2915: 2911: 2907: 2903: 2896: 2893: 2889: 2885: 2879: 2876: 2871: 2867: 2863: 2859: 2855: 2851: 2844: 2841: 2836: 2832: 2828: 2824: 2820: 2816: 2812: 2808: 2801: 2798: 2793: 2789: 2785: 2781: 2777: 2773: 2766: 2763: 2758: 2752: 2748: 2744: 2740: 2733: 2730: 2725: 2721: 2717: 2713: 2709: 2705: 2701: 2697: 2690: 2687: 2682: 2678: 2674: 2670: 2666: 2662: 2655: 2652: 2647: 2643: 2639: 2635: 2628: 2625: 2613: 2606: 2603: 2598: 2594: 2590: 2586: 2582: 2578: 2574: 2567: 2565: 2563: 2559: 2551: 2547: 2543: 2539: 2535: 2531: 2527: 2520: 2513: 2511: 2509: 2507: 2503: 2498: 2494: 2490: 2486: 2482: 2478: 2474: 2470: 2466: 2459: 2457: 2455: 2453: 2451: 2447: 2442: 2436: 2433:(10): 1–211. 2432: 2428: 2421: 2419: 2417: 2413: 2408: 2404: 2400: 2396: 2392: 2388: 2384: 2377: 2374: 2369: 2367:0-8137-3028-7 2363: 2359: 2355: 2354: 2349: 2342: 2339: 2334: 2330: 2326: 2322: 2318: 2314: 2310: 2303: 2301: 2297: 2292: 2288: 2284: 2280: 2276: 2272: 2268: 2261: 2259: 2255: 2250: 2249:New Scientist 2246: 2242: 2241:Sarah Gabbott 2238: 2232: 2229: 2217: 2210: 2207: 2202: 2198: 2194: 2190: 2186: 2182: 2175: 2173: 2169: 2163: 2159: 2156: 2154: 2151: 2149: 2146: 2144: 2141: 2139: 2136: 2135: 2131: 2120: 2115: 2100: 2097: 2092: 2089: 2084: 2081: 2076: 2073: 2068: 2065: 2060: 2057: 2054: 2051: 2046: 2043: 2040:Bassler, 1925 2038: 2037:Polygnathidae 2035: 2032:Klapper, 1981 2030: 2029:Kockelellidae 2027: 2022: 2019: 2015: 2012: 2007: 2006:Cryptotaxidae 2004: 2001:Orchard, 2005 1998: 1995: 1989: 1986: 1980: 1977: 1971: 1968: 1964: 1958: 1957:Iowagnathidae 1955: 1949: 1946: 1940: 1939:Francodinidae 1937: 1931: 1928: 1922: 1921:Coleodontidae 1919: 1913: 1910: 1904: 1901: 1895: 1892: 1891: 1887: 1883: 1876: 1875:Gondolellidae 1873: 1868: 1865: 1860: 1857: 1854:Bassler, 1925 1852: 1849: 1844: 1841: 1835: 1832: 1826: 1823: 1822: 1818: 1817:Prioniodinida 1814: 1807: 1806:Icriodontidae 1804: 1799: 1798:Icriodellidae 1796: 1793:Klapper, 1981 1791: 1788: 1783: 1780: 1775: 1774:Balognathidae 1772: 1771: 1769: 1763: 1760: 1755: 1754:Plectodinidae 1752: 1747: 1744: 1741:Bassler, 1925 1739: 1736: 1731: 1728: 1723: 1722:Periodontidae 1720: 1715: 1714:Oistodontidae 1712: 1707: 1704: 1699: 1698: 1694: 1688: 1685: 1679: 1678:Playfordiidae 1676: 1670: 1667: 1661: 1658: 1652: 1649: 1643: 1640: 1634: 1631: 1625: 1622: 1616: 1613: 1612: 1607: 1603: 1595: 1592: 1586: 1583: 1578: 1575: 1570: 1567: 1561: 1558: 1553: 1550: 1545: 1542: 1541: 1537: 1533: 1532: 1528: 1525: 1515: 1512: 1507: 1504: 1498: 1495: 1490: 1487: 1482: 1479: 1478: 1474: 1470: 1462: 1459: 1454: 1451: 1445: 1442: 1441: 1436: 1432: 1431: 1427: 1424: 1423: 1419: 1415: 1414: 1413: 1407: 1405: 1403: 1399: 1395: 1391: 1385: 1383: 1382:Pennsylvanian 1379: 1375: 1374:Carboniferous 1364: 1362: 1355: 1353: 1346: 1344: 1337: 1331: 1329: 1319: 1317: 1310: 1308: 1301: 1299: 1292: 1290: 1283: 1281: 1271: 1269: 1264: 1262: 1247: 1242: 1235: 1233: 1231: 1227: 1217: 1216: 1208: 1207: 1199: 1198: 1190: 1189: 1181: 1180: 1172: 1171: 1163: 1162: 1159: 1158: 1152: 1151: 1148: 1147: 1144: 1143: 1142:Prioniodinida 1137: 1136: 1130: 1129: 1126: 1125: 1122: 1121: 1120:Balognathidae 1115: 1114: 1108: 1107: 1104: 1103: 1100: 1099: 1098: 1091: 1090: 1083: 1079: 1078: 1075: 1074: 1066: 1065: 1062: 1061: 1055: 1054: 1051: 1050: 1047: 1046: 1040: 1039: 1033: 1032: 1025: 1021: 1020: 1017: 1016: 1013: 1012: 1008: 1002: 1001: 994: 987: 986: 983: 982: 979: 978: 972: 971: 965: 964: 960: 953: 951: 949: 945: 941: 937: 933: 929: 924: 922: 918: 914: 913:paraconodonts 910: 906: 902: 898: 893: 891: 887: 883: 880: 872: 867: 865: 863: 862: 856: 854: 850: 846: 842: 838: 837: 831: 829: 825: 821: 817: 813: 804: 803: 797: 790: 788: 786: 782: 781: 776: 772: 768: 764: 760: 755: 753: 749: 745: 741: 737: 733: 729: 725: 721: 720: 715: 711: 707: 703: 702: 697: 693: 689: 688:Carboniferous 685: 684: 679: 675: 671: 667: 663: 659: 655: 654: 643: 642: 634: 629: 622: 617: 614: 607: 602: 599: 598: 590: 585: 582: 581: 573: 568: 563: 561: 558: 550: 548: 545: 538: 536: 535:Archeognathus 528: 523: 520: 519: 511: 506: 501: 499: 497: 493: 489: 485: 481: 477: 472: 470: 465: 463: 457: 453: 451: 446: 439:Lone elements 438: 433: 431: 427: 425: 421: 417: 416: 410: 408: 404: 400: 399:paraconodonts 396: 392: 387: 385: 381: 377: 368: 366: 362: 360: 356: 348: 346: 344: 340: 336: 332: 329: 325: 321: 317: 313: 309: 305: 302: 298: 288: 282: 278: 274: 268: 267: 265: 262: 258: 251: 248: 246: 245:Prioniodinida 243: 241: 238: 236: 233: 232: 231: 228: 224: 221: 220: 219: 216: 215: 213: 208: 203: 198: 192: 189: 188: 185: 182: 179: 178: 175: 172: 169: 168: 165: 162: 159: 158: 155: 152: 149: 148: 143: 138: 134: 129: 125: 120: 113: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 51: 47: 43: 33: 30: 26: 22: 9598: 9527:Xaniognathus 9525: 9518: 9511: 9504: 9483: 9476: 9469: 9462: 9455: 9448: 9441: 9436:Prioniodella 9434: 9427: 9420: 9415:Palmatodella 9413: 9406: 9399: 9392: 9385: 9378: 9371: 9364: 9357: 9352:Guizhoudella 9350: 9343: 9336: 9329: 9322: 9315: 9308: 9288: 9281: 9262:Stepanovites 9260: 9253: 9246: 9241:Hadrodontina 9239: 9232: 9225: 9218:Ellisoniidae 9205: 9197: 9190: 9185:Curtognathus 9183: 9178:Chirognathus 9176: 9156: 9151:Doliognathus 9149: 9142: 9097: 9076: 9055: 9034: 9027: 9009:Zieglerodina 9007: 9000: 8993: 8986: 8979: 8972: 8965: 8947:Xainzadontus 8945: 8938: 8933:Pranognathus 8931: 8924: 8917: 8910: 8886: 8879: 8872: 8865: 8858: 8851: 8844: 8837: 8830: 8823: 8803: 8798:Panderolepis 8796: 8791:Palmatolepis 8789: 8782: 8777:Manticolepis 8775: 8768: 8761: 8754: 8747: 8742:Conditolepis 8740: 8720: 8713: 8706: 8699: 8692: 8687:Patrognathus 8685: 8678: 8671: 8666:Clydagnathus 8664: 8659:Cavusgnathus 8657: 8652:Adetognathus 8650: 8621: 8614: 8607: 8600: 8593: 8586: 8579: 8572: 8565: 8558: 8539:Mestognathus 8537: 8530: 8510: 8503: 8483: 8462: 8455: 8448: 8428: 8421: 8414: 8396:Sephardiella 8394: 8387: 8380: 8373: 8366: 8359: 8352: 8345: 8338: 8331: 8324: 8317: 8312:Neospathodus 8310: 8303: 8298:Neocavitella 8296: 8289: 8282: 8275: 8268: 8261: 8254: 8247: 8240: 8233: 8226: 8219: 8212: 8205: 8198: 8191: 8184: 8179:Acuminatella 8177: 8157: 8138:Vogelgnathus 8136: 8131:Urbanekodina 8129: 8122: 8115: 8108: 8087: 8080: 8073: 8055:Siphonodella 8053: 8046: 8039: 8032: 8025: 8018: 8011: 7990: 7966: 7959: 7952: 7945: 7938: 7918: 7911: 7904: 7897: 7890: 7870: 7863: 7855: 7848: 7841: 7836:Hamulosodina 7834: 7827: 7820: 7800: 7793: 7786: 7781:Kladognathus 7779: 7772: 7765: 7758: 7740:Zentagnathus 7738: 7731: 7724: 7719:Ptilognathus 7717: 7710: 7703: 7696: 7691:Diplododella 7689: 7671:Pravognathus 7669: 7662: 7655: 7625: 7618: 7611: 7604: 7599:Hunanganthus 7597: 7590: 7583: 7578:Bengtsonella 7576: 7569: 7562: 7555:Belodontidae 7541: 7534: 7513: 7506: 7499: 7492: 7485: 7466:Ozarkodinida 7440: 7433: 7426: 7406: 7398: 7390:Pygodontidae 7377: 7370: 7363: 7356: 7349: 7342: 7335: 7328: 7321: 7316:Drucognathus 7314: 7309:Dichognathus 7307: 7300: 7293: 7286: 7254: 7246: 7239: 7232: 7212: 7205: 7198: 7178: 7157: 7150: 7143: 7136: 7129: 7122: 7102: 7095: 7088: 7080: 7074:Fahraeusodus 7072: 7052: 7045: 7038: 7031: 7012:Ozarkodinida 6999: 6992: 6985: 6980:Loxodentatus 6978: 6971: 6964: 6959:Juanognathus 6957: 6950: 6945:Graciliconus 6943: 6923: 6902: 6895: 6888: 6880: 6873: 6853: 6846: 6839: 6832: 6811: 6804: 6799:Caudicriodus 6797: 6777: 6772:Icriognathus 6770: 6763: 6756:Icrodellidae 6743: 6736: 6729: 6708: 6701: 6694: 6674: 6667: 6660: 6653: 6646: 6639: 6634:Pranognathus 6632: 6627:Omanognathus 6625: 6618: 6611: 6604: 6597: 6590: 6583: 6576: 6569: 6562: 6555: 6535: 6528: 6523:Triangulodus 6521: 6514: 6506: 6499: 6492: 6470: 6463: 6456: 6448: 6438: 6381: 6376:Scabbardella 6374: 6367: 6360: 6353: 6330: 6323: 6302: 6295: 6288: 6281: 6276:Stultodontus 6274: 6267: 6262:Staufferella 6260: 6253: 6246: 6238: 6231: 6226:Polycostatus 6224: 6217: 6212:Parutahconus 6210: 6203: 6198:Monocostodus 6196: 6189: 6181: 6174: 6167: 6162:Colaptoconus 6160: 6152: 6133:Belodellidae 6131: 6124: 6117: 6112:Plegagnathus 6110: 6103: 6098:Parabelodina 6096: 6089: 6084:Nudibelodina 6082: 6075: 6068: 6061: 6054: 6028: 6021: 6014: 6006: 5999: 5992: 5984: 5977: 5950: 5942: 5935: 5928: 5921: 5914: 5907: 5900: 5895:Clavohamulus 5893: 5872: 5866:Scalpellodus 5864: 5859:Parapaltodus 5857: 5850: 5843: 5813: 5805: 5797: 5789: 5781: 5750:Pygodontidae 5748: 5740: 5733: 5726: 5718: 5698: 5691: 5686:Eoconodontus 5684: 5677: 5670: 5649: 5642: 5614: 5606: 5587:Walliserodus 5585: 5578: 5571: 5563: 5556: 5548: 5541: 5534:Belodellidae 5520: 5512: 5505: 5408: 5390:Proscandodus 5388: 5381: 5376:Prooneotodus 5374: 5367: 5360: 5353: 5346: 5339: 5332: 5305:Yongshunella 5303: 5296: 5291:Shandongodus 5289: 5282: 5274: 5269:Laiwugnathus 5267: 5260: 5255:Bengtsonella 5253: 5246: 5238: 5214: 5207: 5199: 5192: 5184: 5176: 5169: 5161: 5154: 5147: 5142:Amphigeisina 5140: 5131:chaetognaths 5070: 4969:spiny sharks 4928:Halecomorphi 4747:Xenacanthida 4632:Phyllolepida 4565:Pituriaspida 4526:Ostracoderms 4460:Cyclostomata 4450:Jawless fish 4399: 4380: 4369: 4360: 4338: 4316: 4299: 4219:. Retrieved 4204:. Retrieved 4200: 4186: 4181:Mark Purnell 4157: 4153: 4113: 4109: 4090: 4083: 4073:. Retrieved 4069: 4049: 4017: 4013: 3988: 3984: 3967: 3963: 3939:. Retrieved 3933: 3922: 3887: 3883: 3873: 3864: 3860: 3854: 3842:. Retrieved 3822: 3818: 3808: 3796:. Retrieved 3776: 3772: 3762: 3709: 3705: 3694: 3680:cite journal 3669:. Retrieved 3662:the original 3641: 3637: 3624: 3615: 3611: 3598: 3579: 3569: 3536: 3532: 3526: 3507: 3503: 3493: 3468: 3462: 3427: 3424:Paleobiology 3423: 3394:. Retrieved 3374: 3370: 3360: 3340: 3333: 3292: 3288: 3278: 3245: 3241: 3231: 3198: 3194: 3148: 3144: 3092: 3088: 3050: 3046: 3040: 3032: 2997: 2993: 2983: 2966: 2962: 2956: 2945:. Retrieved 2941:the original 2936: 2932: 2922: 2905: 2901: 2895: 2883: 2878: 2853: 2849: 2843: 2810: 2806: 2800: 2775: 2771: 2765: 2738: 2732: 2699: 2695: 2689: 2664: 2660: 2654: 2637: 2633: 2627: 2615:. Retrieved 2605: 2580: 2576: 2529: 2525: 2472: 2468: 2430: 2390: 2386: 2376: 2352: 2341: 2316: 2312: 2274: 2270: 2248: 2231: 2219:. Retrieved 2209: 2184: 2180: 2102:Ritter, 1986 2055:(Hass, 1959) 2024:Cooper, 1977 1962: 1916:Huddle, 1934 1912:Belodontidae 1907:Miller, 1969 1886:Ozarkodinida 1867:Ellisoniidae 1777:(Hass, 1959) 1762:Pygodontidae 1709:Harris, 1964 1695: 1514:Pygodontidae 1492:Miller, 1981 1453:Belodellidae 1421:Pander, 1856 1417: 1411: 1386: 1370: 1361:Cavusgnathus 1359: 1352:Kladognathus 1350: 1341: 1335: 1326: 1314: 1305: 1298:Cavusgnathus 1296: 1289:Cavusgnathus 1287: 1280:Kladognathus 1278: 1268:Cavusgnathus 1266: 1261:Kladognathus 1259: 1223: 1157:Ozarkodinida 1155: 1140: 1118: 1095: 1094: 1060:Panderontida 1058: 1043: 1005: 992: 975: 957: 925: 921:Chaetognatha 916: 901:phylogenetic 894: 876: 859: 857: 834: 832: 823: 820:nektobenthic 808: 800: 785:Clydagnathus 784: 778: 774: 771:Clydagnathus 770: 756: 718: 714:South Africa 700: 682: 665: 651: 648: 639: 612: 595: 578: 557:ozarkodinids 554: 546: 543: 533: 516: 494:to the Late 473: 466: 458: 454: 442: 428: 413: 411: 407:sister group 403:chaetognaths 388: 372: 363: 352: 330: 311: 303: 296: 295: 286: 272: 250:Ozarkodinida 229: 217: 196: 29: 9623:Wikispecies 9443:Prioniodina 9359:Gyrognathus 9192:Erraticodon 9029:Meiognathus 8926:Complexodus 8874:Polygnathus 8853:Nicollidina 8832:Bispathodus 8805:Tripodellus 8770:Lagovilepis 8756:Kielcelepis 8680:Neolochriea 8588:Kashmirella 8574:Guanxidella 8560:Discretella 8485:Iowagnathus 8416:FIiognathus 8382:Quadralella 8340:Parapetella 8235:Hayashiella 8020:Gigantholus 8013:Eognathodus 7992:Cryptotaxis 7813:Ligonodinae 7774:Hindeodella 7760:Arcugnathus 7705:Hibbardella 7606:Lepognathus 7543:Neocoleodus 7372:Reutterodus 7351:Kailidontus 7330:Eocarniodus 7302:Cardiodella 7295:Astognathus 7200:Bryantodina 7131:Acanthodina 7001:Tricostatus 6952:Histiodella 6915:Nurrellidae 6848:Jablonnodus 6731:Birksfeldia 6599:Complexodus 6592:Birksfeldia 6578:Baltoniodus 6501:Diaphorodus 6484:Acodontidae 6362:Decoriconus 6304:Ventricodus 6063:Culumbodina 6030:Venoistodus 6023:Paroistodus 5874:Ulrichodina 5735:Kallidontus 5550:Caenodontus 5522:Tokognathus 5487:Belodellida 5440:Euconodonta 5355:Nogamiconus 5348:Muellerodus 5240:Diaphanodus 5093:Subphylum: 4989:lobe-finned 4915:Neopterygii 4901:Chondrostei 4837:Rhipidistia 4816:Lobe-finned 4766:Holocephali 4571:Osteostraci 4473:Hyperoartia 4318:Cathaymyrus 4286:Forerunners 3844:4 September 3773:Palaeoworld 3295:: 212–229. 3248:(10): 879. 2640:(1): 1–14. 2612:"Conodonts" 2070:Sweet, 1988 2048:Sweet, 1988 2017:Hass, 1959 1898:Clark, 1972 1870:Clark, 1972 1838:Sweet, 1981 1820:Sweet, 1988 1801:Sweet, 1988 1757:Sweet, 1988 1660:Nurrellidae 1615:Acodontidae 1539:Sweet, 1988 1476:Sweet, 1988 1465:Sweet, 1988 1439:Sweet, 1988 1435:Belodellida 1429:Sweet, 1988 1378:Tournaisian 1328:Polygnathus 1307:Polygnathus 936:cyclostomes 928:vertebrates 917:euconodonts 895:Milsom and 828:zooplankton 676:related to 638:Fossils of 361:, in 1856. 287:Euconodonta 180:Subphylum: 9709:Categories 9506:Chirodella 9464:Scotlandia 9422:Pluckidina 9380:Lagovidina 9338:Dyminodina 9331:Cornuramia 9324:Chirodella 9310:Bryantodus 9207:Oistodella 9099:Wapitiodus 8981:Ozarkodina 8825:Avignathus 8595:Mosherella 8512:Kockelella 8368:Primatella 8319:Nicoraella 8228:Gondolella 8124:Sweetodina 8110:Francodina 8041:Rhodalepis 7913:Pachysomia 7857:Lonchodina 7850:Ligonodina 7664:Hindeodina 7627:Ptiloconus 7620:Paibiconus 7365:Prioniodus 7214:Scyphiodus 7207:Plectodina 7180:Playfordia 7152:Phragmodus 7104:Pohlerodus 7040:Oelandodus 6994:Oleandodus 6966:Jumudontus 6841:Fungulodus 6765:Icriodella 6703:Distomodus 6641:Prioniodus 6620:Notiodella 6606:Icriodella 6472:Ansellidae 6465:Schopfodus 6283:Teridontus 6255:Scolopodus 6219:Planusodus 6183:Filodontus 6176:Eucharodus 6169:Drepanodus 6154:Aloxoconus 6091:Panderodus 6070:Naimanodus 5986:Drepanodus 5979:Distacodus 5937:Laiwunodus 5852:Kirkupodus 5845:Acanthodus 5815:Thrincodus 5799:Dapsilodus 5783:Besselodus 5693:Eodentatus 5679:Cordylodus 5616:Dapsilodus 5608:Besselodus 5497:Ansellidae 5341:Furnishina 5248:Dolabrodus 5209:Phakelodus 5201:Paibiconus 5194:Mongolodus 5171:Hagionella 5156:Gapparodus 5149:Ganloudina 5125:(Probably 5095:Vertebrata 4974:placoderms 4933:Ginglymodi 4867:Ray-finned 4832:Coelacanth 4677:Acanthodii 4614:Arthrodira 4599:Placoderms 4587:Jawed fish 4559:Galeaspida 4541:Thelodonti 4340:Haikouella 4331:Olfactores 4221:2009-07-07 4206:2013-07-01 4075:2007-09-05 3941:2015-12-30 3890:: 103325. 3825:: 103699. 3798:16 October 3671:2011-02-11 3576:Rigby, Sue 2947:2016-07-15 2888:Stable URL 2164:References 1943:Dzik, 2006 1889:Dzik, 1976 1682:Dzik, 2002 1655:Dzik, 2006 1619:Dzik, 1993 1609:Dzik, 1976 1444:Ansellidae 873:Affinities 861:Panderodus 775:Panderodus 710:Soom Shale 706:Ordovician 678:gastropods 666:Typhloesus 653:Typhloesus 641:Typhloesus 376:anglicized 322:group of 318:") are an 277:Eichenberg 210:Subgroups 184:Vertebrata 50:Hettangian 9715:Conodonts 9629:Conodonta 9599:Conodonta 9520:Ketinella 9290:Oepikodus 9227:Ellisonia 9002:Wurmiella 8967:Flajsella 8919:Carniodus 8784:Mesotaxis 8609:Smithodus 8450:Gnathodus 8361:Pridaella 8284:Misikella 8263:Mazzaella 8193:Borinella 8117:Planadina 8027:Gondwania 7865:Lonchodus 7795:Oncodella 7698:Elsonella 7564:Albiconus 7501:Hindeodus 7400:Nericodus 7288:Angulodus 7272:see below 7248:Polonodus 7082:Hamarodus 7017:see below 6738:Galerodus 6648:Promissum 6508:Nealeodus 6403:see below 6369:Macerodus 6355:Cornuodus 6290:Utahconus 6205:Oneotodus 6008:Nordiodus 5944:Nericodus 5902:Dasytodus 5791:Cristodus 5770:Conodonti 5742:Polonodus 5543:Belodella 5514:Hamarodus 5478:Cavidonti 5466:Conodonta 5446:see below 5429:Conodonta 5362:Proacodus 5334:Albiconus 5298:Trolmenia 5163:Glauderia 5081:Kingdom: 5071:Conodonts 5032:† extinct 4887:Cladistia 4644:Rhenanida 4608:Antiarchi 4513:Conodonta 4492:Conodonts 4130:0022-3360 3914:225005180 3793:246060690 3736:0027-8424 3618:(3): 405. 3454:248062641 3446:0094-8373 3325:134664744 3317:0031-0182 3270:0091-7613 3173:237769553 3165:2056-2799 2870:128496416 2778:: 36–43. 2724:131850219 2617:26 August 2610:MIRACLE. 2597:1477-2019 2407:0024-1164 2333:0024-1164 2221:3 January 2201:129519805 1527:Conodonti 1426:Cavidonti 1418:Conodonta 1343:Gnathodus 1316:Gnathodus 1291:unicornis 1270:unicornis 1226:cladistic 1024:Conodonti 1007:Cavidonti 993:Conodonta 932:craniates 890:notochord 824:Promissum 812:occlusion 780:Promissum 763:notochord 732:Wisconsin 722:from the 704:from the 686:from the 670:digestive 450:agnathans 412:The 1981 380:latinized 331:Conodonta 297:Conodonts 230:Conodonti 218:Cavidonti 197:Conodonta 160:Kingdom: 154:Eukaryota 35:Conodonts 9653:42349189 9637:BioLib: 9608:Wikidata 9564:Category 9485:Veghella 9478:Uncadina 9234:Foliella 9199:Lumidens 8995:Tortodus 8423:Icriodus 8214:Conudina 8207:Clarkina 7954:Gothodus 7657:Coleodus 7138:Nordiora 7097:Periodon 6987:Oistodus 6925:Nurrella 6904:Wandelia 6806:Icriodus 6537:Tropodus 6530:Tripodus 6458:Rossodus 6056:Belodina 6016:Paltodus 5720:Dzikodus 5580:Stolodus 5573:Dvorakia 5089:Chordata 5087:Phylum: 5083:Animalia 4850:Lungfish 4795:Chimaera 4547:Anaspida 4346:Tunicate 4324:Lancelet 4142:Archived 4138:53395896 4093:Oxford, 4042:22803015 4034:10881388 4005:21237922 3867:: 1–560. 3754:30670644 3658:86599352 3485:89835464 3121:36126687 2835:53395896 2550:Archived 2546:53395896 2497:22803015 2489:10881388 2116:See also 1697:Rossodus 1408:Taxonomy 1318:texanus? 946:than to 944:lampreys 882:Chordata 769:. While 744:Triassic 738:site in 724:Silurian 696:Scotland 496:Triassic 492:Cambrian 339:Jurassic 335:Cambrian 261:Synonyms 174:Chordata 170:Phylum: 164:Animalia 150:Domain: 25:Cynodont 9687:ZooBank 9640:1159347 9614:Q586245 9574:Commons 9408:Oulodus 9345:Falodus 8846:Mehlina 8749:Dinodus 8291:Mockina 7899:Loxodus 7726:Roundya 7592:Gumella 7571:Belodus 7408:Pygodus 6779:Pedavis 6613:Lenodus 5807:Loxodus 5507:Ansella 5186:Kijacus 5005:Related 4938:Teleost 4478:Lamprey 4468:Hagfish 4154:Lethaia 3892:Bibcode 3827:Bibcode 3745:6369785 3714:Bibcode 3561:1598571 3541:Bibcode 3533:Science 3471:: 1–9. 3396:15 July 3379:Bibcode 3371:Geology 3297:Bibcode 3250:Bibcode 3242:Geology 3223:4342260 3203:Bibcode 3112:9489302 3055:Bibcode 3023:1692076 3002:Bibcode 2975:1302203 2914:1298905 2827:1303249 2780:Bibcode 2716:1306313 2669:Bibcode 2634:Lethaia 2387:Lethaia 2313:Lethaia 2291:2413175 1084:  1026:  940:hagfish 905:derived 886:chevron 816:pelagic 791:Ecology 765:or the 740:Ontario 736:Eramosa 674:mollusk 662:Montana 320:extinct 190:Class: 21:Conodon 6494:Acodus 4301:Pikaia 4201:Palæos 4136:  4128:  4056:  4040:  4032:  4003:  3912:  3791:  3752:  3742:  3734:  3656:  3586:  3559:  3483:  3452:  3444:  3348:  3323:  3315:  3268:  3221:  3195:Nature 3171:  3163:  3119:  3109:  3020:  2973:  2912:  2868:  2833:  2825:  2753:  2722:  2714:  2595:  2544:  2495:  2487:  2437:  2405:  2364:  2331:  2289:  2199:  1965:, 2017 1963:et al. 1884:Order 1815:Order 1604:Order 1534:Order 1471:Order 1433:Order 1416:Class 1295:6, 7. 1277:3, 4. 1252:detail 1080:  1022:  879:phylum 716:, and 537:primus 422:and a 420:phylum 279:, 1930 204:, 1856 202:Pander 9679:33816 9661:IRMNG 8974:Lanea 4957:Lists 4757:Shark 4145:(PDF) 4134:S2CID 4106:(PDF) 4038:S2CID 3910:S2CID 3789:S2CID 3665:(PDF) 3654:S2CID 3634:(PDF) 3608:(PDF) 3481:S2CID 3450:S2CID 3321:S2CID 3219:S2CID 3169:S2CID 3095:(9). 2971:JSTOR 2910:JSTOR 2866:S2CID 2831:S2CID 2823:JSTOR 2720:S2CID 2712:JSTOR 2553:(PDF) 2542:S2CID 2522:(PDF) 2493:S2CID 2287:JSTOR 2197:S2CID 897:Rigby 752:Japan 746:aged 726:aged 708:aged 690:aged 424:class 384:order 328:class 316:tooth 312:odont 310:", + 304:kĹŤnos 301:Greek 9666:1062 9554:List 5127:stem 4984:bony 4807:Bony 4271:Fish 4126:ISSN 4054:ISBN 4030:PMID 4001:PMID 3846:2022 3800:2022 3750:PMID 3732:ISSN 3686:link 3584:ISBN 3557:PMID 3442:ISSN 3398:2021 3346:ISBN 3313:ISSN 3266:ISSN 3161:ISSN 3117:PMID 2751:ISBN 2619:2014 2593:ISSN 2485:PMID 2435:ISBN 2403:ISSN 2362:ISBN 2329:ISSN 2239:and 2223:2019 1961:Liu 1358:14. 1349:13. 1340:12. 1334:11. 1325:10. 942:and 915:and 847:and 773:and 742:and 308:cone 57:Preęž’ 9648:EoL 4752:Ray 4356:? 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Index

Conodon
Cynodont
Middle Cambrian
Earliest Jurassic
Hettangian
Preęž’
ęž’
O
S
D
C
P
T
J
K
Pg
N

Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Vertebrata
Conodonta
Pander
Proconodontida
Protopanderodontida
Panderodontida
Prioniodinida

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