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Corythosaurus

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beak; expanded section in front of the elongated nares; a small narial opening; a vertebral formula of 15 cervicals, 19 dorsals, 8 sacrals, and 61+ caudals; possession of dorsal spines of a medium height; high anterior caudal spines; long chevrons; long scapulae that possess a blade of medium width; a radius considerably longer than the humerus; comparatively short metacarpals, an anteriorly decurved ilium; a long ischium with a foot-like terminal expansion; a pubis with an anterior blade that is short and broadly expanded at the end; a femur that is longer than the tibia; the phalanges of pes are short; that the integument over the sides and tail composed of polygonal tuberculate scales without pattern, but graded in size in different parts of the body; and a belly with longitudinal rows of large conical limpet-like scales separated by uniformly large polygonal tubercles. Again, the presumed traits of the snout are incorrect because Brown confused the praemaxillae with the nasal bones and the nasal bones with the frontals. Most of the postcranial traits are today known to be shared with various other lambeosaurines.
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praemaxillae for the nasals. The snout is actually largely formed by them and they do separate the nasals. Brown also thought that, on the top and back of the crest, the whole external face is covered by the frontals. Again he made a mistake, as what he assumed to be the frontals are in fact the nasals. The nasals end at the back of the squamosals in a hooked, short process. The prefrontals also make up part of the crest. However, Brown mistook the lower upper branch of the praemaxilla for the prefrontal. The actual prefrontal, which is triangular in shape, is located at the side of the crest base. It was seen by Brown as a part of the frontal. The real frontals, which are largely internal to the crest base structure, are not visible from the side.
2122:, to have had surviving fossilized juveniles assigned to it. Juveniles are harder to assign to species because, at a young age, they lack the distinctive larger crests of adults. As they age, lambeosaurine crests tend to grow and become more prominent come maturity. In the Dinosaur Park Formation, over fifty articulated specimens have been found that come from many different genera. Among them, juveniles are hard to identify at the species level. Earlier, four genera and thirteen species were recognized from the formation's area when paleontologists used differences in size and crest shape to differentiate taxa. The smallest specimens were identified as 1084:
central chamber, and onward into the respiratory system. The skull of the type specimen has no dermal impressions on it. During preservation, it was compressed laterally and the width is now about two-thirds of what it would have been in real life. According to Brown, the compression also caused the nasals to shift where they pressed down on the premaxillaries. Because they were pressed on the premaxillaries, the nasals would have closed the nares. Apart from the compression, the skull appears to be normal. Contrary to what Brown assumed, the areas concerned were fully part of the praemaxillae.
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exhaled. The individual crests would produce different sounds, so it is likely that each species of lambeosaurine would have had a unique sound. However, even though the range for different lambeosaurine nasal passages vary greatly, they all probably made low-pitched sounds. This might be because low
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The lower jaw of the holotype is 66.9 centimetres (26.3 in) long and 10 centimetres (3.9 in) deep. The total length of the crest from the beak to the uppermost tip of the type specimen is 83.7 centimetres (33.0 in), its total length is 81.2 centimetres (32.0 in), and its height is
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is narrow. The praemaxillae each form two long folds that enclose air passages extending the narial passages to the front of the snout. There, they end in narrow openings, sometimes called "pseudonares", which are false bony nostrils. These were mistaken by Brown for the real nares or nostrils. These
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are shieldlike scales, arranged close together in rows. Ossified tendons are present on all the vertebrae, except for those in the cervical region. On no vertebrae do the tendons extend below the transverse processes. Each tendon is flattened at its origin, transversely ovoid in the central rod, and
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alone in later stages of the formation. Studies of the jaw anatomy and mechanics of these dinosaurs suggests they probably all occupied slightly different ecological niches in order to avoid direct competition for food in such a crowded eco-space. The only large predators known from the same levels
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When they were first described, crested hadrosaurs were thought to be aquatic, an assessment based incorrectly on webbing that is now known to be padding. The theory was that the animals could swim deep in the water and use the crest to store air to breath. However, it has now been proven that the
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In 1916, Brown expanded the character set to include even more features. In the revised version, these extra features include a comparatively short skull with a high helmet-like crest formed by nasals, prefrontals, and frontals; the nasals not being separated in front by premaxillaries; a narrow
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Over twenty skulls have been found from this dinosaur. As with other lambeosaurines, the animal bore a tall, elaborate, bony crest atop its skull that contained the elongate narial passages. The narial passages extended into the crest, first into separate pockets in the sides, then into a single
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The nasals make up most of the crest. Brown assumed that they extended from the beaks' tip to the highest spot along the crest and that, unlike those in other genera, the nasals meet in the center and are not separated in front by an ascending premaxillary process. However, Brown mistook the
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studied the differences between the skulls and crests of different species of lambeosaurine dinosaurs. He found that the differences in size and shape may have actually been related to the sex and age of the animal. Only one species is currently recognized for certain,
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of the province. The left or underside of the skeleton was preserved in carbonaceous clay, making it difficult to expose the skin to the elements. The skeleton was articulated and only missing about the last 0.61 metres (2.0 ft) of the tail and front legs. Both
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dinosaurs in the region. Large herbivores like the hadrosaurs living in North America during the Late Cretaceous had "remarkably small geographic ranges" despite their large body size and high mobility. This restricted distribution strongly contrasts with modern
760:). Apparently, the remaining front legs were weathered or eroded away. Impressions of the integument were preserved covering over a large part of the skeleton’s outlining and shows the form of the body. Another specimen, AMNH 5338, was found in 1914 by Brown and 1438:
as a base model to analyze which theory is more likely true. He found that, even though birds are thought of as saurischians, it is very plausible for them to have evolved their specific pelvic musculature and anatomy if they evolved from ornithschians like
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It was once thought that this dinosaur lived mostly in the water, due to the appearance of webbed hands and feet. However, it was later discovered that the so-called "webs" were in fact deflated padding, much like that found on many modern mammals.
1352: 2096: 838: 3199:"Testing Size–frequency Distributions As a Method of Ontogenetic Aging: A Life-history Assessment of Hadrosaurid Dinosaurs from the Dinosaur Park Formation of Alberta, Canada, with Implications for Hadrosaurid Paleoecology" 3276:
Dodson, Peter & Britt, Brooks & Carpenter, Kenneth & Forster, Catherine A. & Gillette, David D. & Norell, Mark A. & Olshevsky, George & Parrish, J. Michael & Weishampel, David B. (1994).
1567: 2958:, an indeterminate juvenile hadrosaurid from the Two Medicine Formation of Montana: implications for hadrosauroid diversity in the latest Cretaceous (Campanian-Maastrichtian) of western North America". 2204:
in that the sutures of the skull are sinuous, not smooth and straight. This feature helps to differentiate Parasaurolophini from Lambeosaurini. Generally, the crests of juveniles of lambeosaurines like
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and secured by him in the Fall of 1912. As well as an almost complete skeleton, the find was notable because impressions of much of the creature's skin had also survived. The specimen came from the
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proposed that the two orders of dinosaurs might have evolved separately and that birds, based on the shape and proportions of their ilia, might truly be specialized ornithischians. He used both
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Mallon, Jordan C.; Evans, David C.; Ryan, Michael J.; Anderson, Jason S. (2012). "Megaherbivorous dinosaur turnover in the Dinosaur Park Formation (upper Campanian) of Alberta, Canada".
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specimen AMNH 5240 was found to be 8.1 m (27 ft) long, with a weight close to 3.1 tonnes (3.0510 long tons; 3.4172 short tons). In 2016, Gregory S. Paul estimated that
630:. The most likely function of the crest is thought to be vocalization. As in a trombone, sound waves would travel through many chambers in the crest and then get amplified when 618:
found by Brown in 1911. The holotype skeleton is only missing the last section of the tail and part of the front legs, but was preserved with impressions of polygonal scales.
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bill, the two pseudonares unite into one. Because of his incorrect identification, Brown assumed that the holotype's inferior process of the premaxillary was shorter than in
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model used was modelled by Vincent Fusco, after a mounted skeleton, and supervised by Barnum Brown. After testing, it was concluded that the average weight of
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was shallow and delicate, concluding that it must have been used to feed upon soft vegetation. Based on the climate of the Late Cretaceous, they guessed that
2023:, implies the existence of a sensitive middle ear. It is possible that hadrosaurid ears are sensitive enough to detect as much sound as a modern crocodilian. 1151:, which is another difference from those genera. The praemaxilla actually does touch the lacrimal and extends to the rear until well behind the eye socket. 3498: 608:
has an estimated length of 7.7–9 metres (25–30 ft) and has a skull, including the crest, that is 70.8 centimetres (27.9 in; 2.32 ft) tall.
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crest did not have any holes in the end and the water pressure at even 3 metres (9.8 ft) would be too great for the lungs to be able to inflate.
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only possessing U-shaped tubes. Any vocalization would travel through these elaborate chambers and probably get amplified. Scientists speculate that
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is one of very few hadrosaurids which have preserved skin impressions on the hind limbs and feet. A study in 2013 showed that, amongst hadrosaurids,
6611: 6145: 1499:, they all share similar looking skulls and crests. However, research published in 2003 has suggested that even though it possesses a unique crest, 2893:"Diversity, Relationships, and Biogeography of the Lambeosaurine Dinosaurs from the European Archipelago, with Description of the New Aralosaurin 5230: 6711: 6140: 3632: 4217: 4155: 4128: 4101: 4074: 4003: 3744: 3173: 2766: 6741: 5244: 3567:"A New Basal Hadrosauroid Dinosaur (Dinosauria: Ornithopoda) with Transitional Features from the Late Cretaceous of Henan Province, China" 6716: 2658: 1030:
reached 7.7 metres (25 ft) in length and 2.5 metric tons (2.8 short tons) in weight. A "morphologically adult-sized specimen" of
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could make loud, low pitched cries "like a wind or brass instrument", such as a trombone. The sounds could serve to alert other
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from the deposits, lived about 77 to 76.5 mya. The holotype specimen was clearly a carcass that had floated up on a beach, as
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are quite complex, making possible a call that could be used as a warning or for attracting a mate. Nasal passageways of
6726: 1201: 1131:, the expanded portion of the premaxillary in front of the pseudonaris' opening is elongate. By comparison, the bill of 1107:. The two halves of the crest are separated by a median suture. In front of the orbit, the crest is made of thick bone. 142: 3952:, a New Crested Dinosaur from the Belly River Cretaceous, with Provisional Classification of the Family Trachodontidae" 2157: 1069: 6151: 1694: 1219: 961:
and the two species are not identical, which supported the separation of them in a 2009 study. The invalid species,
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to the presence of food or a potential threat from a predator. The nasal passages emit low-frequency sounds when
791:, was named by Barnum Brown in 1914, based on the first specimen collected by him in 1912. AMNH 5240 is thus the 3140: 4317: 2328: 1855: 1838: 3416:
Schmitz, L.; Motani, R. (2011). "Nocturnality in Dinosaurs Inferred from Scleral Ring and Orbit Morphology".
3329:"Standardized terminology and potential taxonomic utility for hadrosaurid skin impressions: A case study for 6706: 2339: 1281: 1213: 816: 658: 3565:
Xing, H.; Wang, D.; Han, F.; Sullivan, C.; Ma, Q.; He, Y.; Hone, D. W. E.; Yan, R.; Du, F.; Xu, X. (2014).
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specimen has even been preserved with its last meal in its chest cavity. Inside the cavity were remains of
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specimen has been preserved with its last meal in its chest cavity. Inside the cavity were remains of
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from Alberta. These include a comparatively short skull with a high helmet-like crest formed by the
931: 3679: 3131: 2480: 1711: 1275: 1103:, but instead of projecting backwards as a spine, they rise up to make the highest point above the 2080:) travel a set distance in any environment, while higher sounds (above 400 Hz) have a larger 761: 6602: 6258: 5869: 4454: 4045: 3671: 3663: 3449: 3228: 3144: 2831: 2778: 1099:. They are formed by a combination of the praemaxillae, nasals, prefrontals, and frontals, as in 396: 376: 137: 4066: 3736: 3724: 1171: 930:
has been recognized as valid in some studies. It is based on specimen ROM 776, a skull found by
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probably lived in a woodland forest and might have occasionally wandered into swampy areas.
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would have been a selective feeder, eating only the juiciest fruits and youngest leaves. A
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preserved the best and most complete foot and limb integument, although other species like
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reached 8 metres (26 ft) long and 2.8 metric tons (3.1 short tons) in weight and that
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Bramble, Katherine; Currie, Philip J.; Tanke, Darren H.; Torices, Angelica (August 2017).
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Bramble, Katherine; Currie, Philip J.; Tanke, Darren H.; Torices, Angelica (August 2017).
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Prieto-Márquez, A.; Vecchia, F.M.D.; Gaete, R.; Galobart, À. (2013). Dodson, Peter (ed.).
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Campione, N. E.; Brink, K. S.; Freedman, E. A.; McGarrity, C. T.; Evans, D. C. (2013). "
2912: 6501: 6496: 6434: 6371: 6244: 6093: 6056: 6038: 5983: 5615: 5431: 5424: 5353: 5275: 5262: 5190: 5178: 5147: 5122: 4986: 4908: 4869: 4855: 4732: 4643: 4565: 4543: 4536: 4277: 4086: 4059: 3792: 3761: 3633:"Acoustic Analysis of Vocalization of Lambeosaurine Dinosaurs (Reptilia: Ornithischia)" 3601: 3566: 3525:"A remarkable hollow-crested hadrosaur from Russia: an Asian origin for lambeosaurines" 3367: 3328: 2931: 2892: 2562: 2541: 2409: 1994: 1553: 1448: 1372: 1368: 1148: 940: 861: 671: 571: 404: 261: 219: 121: 3551: 3524: 2031: 6695: 6441: 6385: 6364: 6335: 6251: 6188: 6086: 6079: 6065: 5975: 5955: 5948: 5845: 5795: 5772: 5747: 5720: 5713: 5706: 5677: 5628: 5590: 5555: 5536: 5508: 5452: 5115: 5108: 5063: 4993: 4862: 4721: 4706: 4668: 4579: 4472: 4435: 4428: 4113: 4016:; Burns, M. E.; Sissons, R. L. (2009). "A redescription of the ankylosaurid dinosaur 3232: 2535: 1823: 1802: 1587: 1538: 1525: 1430: 1420: 1376: 1039: 757: 594: 582: 559: 274: 248: 72: 4049: 3675: 3453: 3148: 2323: 2228:
Work by Dodson (1975) recognized that there were many less taxa present in Alberta.
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is among many lambeosaurines that possess crests and it was the crest that lends
6624: 6596: 6538: 6427: 6392: 6230: 6072: 6045: 5997: 5990: 5823: 5809: 5781: 5761: 5740: 5684: 5663: 5656: 5649: 5583: 5576: 5529: 5494: 5466: 5402: 5395: 5164: 5014: 5000: 4972: 4950: 4892: 4755: 4586: 4550: 4514: 4410: 4397: 4356: 4226: 3729:(Ornithischia: Hadrosauridae) and the Identification of Juvenile Lambeosaurines" 2814:(1975). "Taxonomic implications of relative growth in lambeosaurine dinosaurs". 2439: 2427: 1744: 1649: 1632: 1601: 1466: 1452: 1381: 1321: 1057: 1051: 865: 689: 536: 235: 47: 4184: 2465:
is widespread throughout the lower unit of the Dinosaur Park Formation. In it,
2407:, are among the taxa. Also from the section of the formation are the theropods 744:
are preserved in position, but the rest of the front legs are gone (except for
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faunas whose large herbivores' ranges "typical ... span much of a continent."
2517: 2002: 1785: 1501: 1495: 1489: 1474: 1391: 1337: 1325: 600: 92: 57: 6587: 3861: 3183: 3112:"A New Method to Calculate Allometric Length-Mass Relationships of Dinosaurs" 3040: 6462: 6378: 6357: 6195: 5892: 5830: 5339: 5311: 5171: 5035: 4965: 4936: 4762: 4748: 4491: 3437: 3298:
Weishampel, D.B., Dodson, P., Osmólska, H., & Hilton, Richard P., 2004,
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A limited fauna is known from the upper section of the Oldman Formation and
2347: 1768: 1564:. The below cladogram is a simplified version including only Lambeosaurini. 1549: 1519: 1470: 1092: 1045: 954: 824: 623: 588: 555: 154: 97: 41: 34: 4142:
Discovering Dinosaurs: Evolution, Extinction, and the Lessons of Prehistory
3801: 3610: 3445: 3376: 2940: 2260:, although the holotype of the species was still found to be assignable to 707: 4172: 3760:
Farke, A. A.; Chok, D. J.; Herrero, A.; Scolieri, B.; Werning, S. (2013).
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Sander, P. Martin; Gee, Carole T.; Hummel, Jürgen; Clauss, Marcus (2010).
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lived alongside numerous other giant herbivores, such as the hadrosaurids
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are quite alike, although other features can be used to distinguish them.
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juveniles grew crests when they were only 25% as long as adults. Juvenile
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ROM 870, the skull of a subadult, originally named as a separate species,
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A set of characters were indicated by Barnum Brown in 1914 to distinguish
1061:. But, when including its crest, its superficial area is almost as large. 6581: 6342: 6011: 5899: 5699: 5670: 5597: 5522: 5487: 5388: 5374: 5332: 5156: 4957: 4922: 4828: 4814: 4776: 4286: 4259: 4250: 4041: 2663:
Plants in Mesozoic Time: Morphological Innovations, Phylogeny, Ecosystems
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started developing its crest when they were half the size of adults, but
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Fossil holotype specimen AMNH 5240 partially covered in skin impressions
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Parks, 1924 (Ornithischia: Ankylosauria) and a revision of the genus".
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lived from ~77–75.7 million years ago. In the Dinosaur Park Formation,
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based largely on biometric information. The only non-typic specimen of
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was 3.82 tonnes (3.76 long tons; 4.21 short tons). The total length of
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is derived from its apparent intermediate position according to Parks.
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is known from many skulls with tall crests that resemble those of the
5501: 5094: 4244: 3163: 2665:. Bloomington, Indiana: Indiana University Press. pp. 341, 343. 2343: 2304: 2016: 1127:
are actually situated inside the crest, above the eye sockets. As in
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thus means "Cassowary-like reptile, with a Corinthian helmet crest".
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is known from many complete specimens, including the nearly complete
164: 6558: 2827: 2011:, also seems to have been greatly developed because of an elongated 2100:
ROM 759, a juvenile skull, originally named as a separate species,
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is short and the pseudonares extend far forward. At the end of the
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Proportionally, the skull is much shorter and smaller than that of
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in 1912, were lost on December 6, 1916, while being carried by the
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Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed
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Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed
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Parks, W.A. (1923). "New species of crested trachodont dinosaur".
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sp. The stars represent the age at which crest development starts
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as the most derived lambeosaurine and being the sister taxon to
1371:(now Hadrosauridae). Inside Trachodontidae were the subfamilies 1310: 753: 6562: 6167: 5260: 4395: 4237: 4199: 4195: 3523:
Godefroit, Pascal; Bolotsky, Yuri; Alifanov, Vladimir (2003).
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does bear a greater superficial resemblance to birds than the
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word κόρυς, meaning "helmet", named and described in 1914 by
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There were formerly up to seven species described, including
1293:) share a fair amount of preserved tissue on those regions. 501: 489: 3141:
10.1671/0272-4634(2001)021[0051:ANMTCA]2.0.CO;2
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Tyrannosaurus rex, University of California Press, p. 143
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has an average length of 9 metres (30 ft). In 1962,
880:, when the ship transporting them was sunk by the German 815:), "Corinthian helmet", and means "helmeted lizard". The 486: 3197:
Wosik, M.; Chiba, K.; Therrien, F.; Evans, D.C. (2020).
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pp. 54–82 in Currie, P.J. & Koppelhus, E.B. (2005).
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was found to be closely associated with the ceratopsid
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living from 75.8 to 75.7 mya. In the Oldman Formation,
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was an ornithopod, therefore being a herbivore. Benson
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pp. 310–328 in Tanke, D.H. & Carpenter, K. (2001).
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pelvis does, birds are now known to be highly derived
4119:. Berkeley: University of California Press. pp.  3929: 3927: 3925: 3923: 2311:
probably fed on all of these, implying that it was a
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are both present in slightly different levels of the
507: 498: 480: 474: 2777:. Carnegie Museum of Natural History. Archived from 2475:. Their associating was found in the Dinosaur Park, 842:
Quarry with one of the specimens lost at sea in 1916
483: 354: 336: 313: 6571: 6472: 6323: 6303: 6178: 6055: 6021: 5965: 5924: 5868: 5844: 5771: 5730: 5614: 5553: 5273: 5189: 5146: 5080: 4906: 4838: 4794: 4731: 4695: 4654: 4490: 4453: 4408: 4355: 4328: 4301: 4146:. London: University of California Press. pp.  4111:Weishampel, D.B.; Dodson, P.; Osmolska, H. (2004). 1034:measured approximately 9 metres (30 ft) long. 495: 4139: 4085: 4058: 2001:and modern reptiles suggest that it may have been 3898:Palaeogeography, Palaeoclimatology, Palaeoecology 3891: 3889: 3766:(Hadrosauridae) and heterochrony in hadrosaurids" 3698: 3696: 3694: 3692: 3626: 3624: 3622: 3620: 2798:p. 495 in Tanke, D.H. & Carpenter, K. (2001). 1336:; the nasals not being separated in front by the 3998:. London: Dorling Kindersley. pp. 344–345. 3465: 3463: 3397: 3395: 3310: 3308: 3281:. Publications International, LTD. p. 137. 3254: 3252: 3250: 3248: 3246: 3244: 3242: 2874:University of Toronto Studies, Geological Series 1195:, extensive skin impressions have been found on 4138:Norrell, M.; Gaffney, E.S.; Dingus, L. (2000). 3723:Evans, D.C.; Forster, C.A.; Reiz, R.R. (2005). 2726: 2724: 2722: 2720: 2718: 2716: 2706: 2704: 2702: 2130:, and the largest skeletons were called either 1443:. However, even though the pelvic structure of 1251:have preserved polygonal scales. The scales on 3994:Benson, R.B.J.; Brussatte, M.; Xu, X. (2012). 3411: 3409: 3407: 3070: 1536:In 2014, a study including the description of 1199:and notable integument has also been found on 827:, a bird with a similar skull crest. The full 4211: 3731:. In Currie, P.J.; Koppelhus, Eva B. (eds.). 3068: 3066: 3064: 3062: 3060: 3058: 3056: 3054: 3052: 3050: 2851:Bulletin of the Geological Society of America 2577:, even though it is one of the most abundant 1231:, and on unidentified ornithopods. Of these, 1147:and that the process does not unite with the 1006:used models of specific dinosaurs, including 8: 3718: 3716: 3714: 3712: 2659:"14: Mesozoic Plants and Dinosaur Herbivory" 1175:Abdominal skin impressions from Brown (1916) 3983:American Museum of Natural History Bulletin 3959:American Museum of Natural History Bulletin 3322: 3320: 3168:. Princeton University Press. p. 344. 2694:, 2011 Barnum Brown: The Man Who Discovered 2573:hasn't been discovered outside of southern 2110:is one of a few lambeosaurines, along with 1577:Corythosaurus cf. intermedius cf. excavatus 516: 6702:Late Cretaceous dinosaurs of North America 6559: 6175: 6164: 5850: 5620: 5561: 5281: 5270: 5257: 5086: 4914: 4806: 4802: 4660: 4420: 4416: 4405: 4392: 4334: 4307: 4298: 4234: 4218: 4204: 4196: 120: 20: 3791: 3781: 3600: 3590: 3366: 3356: 3222: 3130: 2960:Palaeobiodiversity and Palaeoenvironments 2930: 2920: 2872:, a new species of trachodont dinosaur". 2527:in the earliest stages of the formation, 1399:in Trachodontinae, whereas he classified 3302:. University of California Press. p. 450 2989: 2987: 2985: 2983: 2981: 2979: 2977: 2975: 2973: 2156: 2043:The internal structures of the crest of 1548:. The study included an almost complete 945:earlier that year. The specific name of 3762:"Ontogeny in the tube-crested dinosaur 2806: 2804: 2622: 1473:Lambeosaurinae. It is related to other 775:Excavation of the holotype specimen of 764:. Both specimens are now housed in the 389: 4084:Currie, P.J.; Koppelhus, E.B. (2005). 3976:: Skeleton, Musculature and Epidermis" 3165:The Princeton Field Guide to Dinosaurs 1579:Parks 1935 at the Royal Ontario Museum 1414:Later, Brown revised the phylogeny of 953:lived at a slightly later time in the 4092:. Indiana University Press. pp.  4065:. Indiana University Press. pp.  3735:. Indiana University Press. pp.  2380:turtle were preserved all around it. 2334:Fossils have been found in the upper 2256:, was later found to be referable to 646:, twigs, and fruits, suggesting that 7: 2391:Herbivores of the Oldman Formation, 2346:. The Oldman Formation dates to the 2221:, and primitive lambeosaurines like 990:(right, yellow) compared to a human. 846:The two best preserved specimens of 4057:Tanke, D.H.; Carpenter, K. (2001). 2569:Thomas M. Lehman has observed that 1885: 1830: 1818: 1760: 1736: 1686: 1674: 1624: 1617: 1593: 1583: 811:is derived from the Greek κόρυθος,( 4022:Journal of Vertebrate Paleontology 3499:"The Ilium in Dinosaurs and Birds" 3119:Journal of Vertebrate Paleontology 2358:lived from 76.6 to 75.9 mya, with 2138:. An adult was even identified as 2035:Outdated 1916 restoration showing 1509:'s closest known relative. Benson 766:American Museum of Natural History 14: 6737:Campanian genus first appearances 2279:(2012) realized that the beak of 1155:70.8 centimetres (27.9 in). 1087:As aforementioned, the crests of 938:in 1923. Originally, he named it 874:British Museum of Natural History 6150: 6144: 6139: 6133: 6126: 5243: 5236: 5229: 4183: 4171: 3401:pp. 712–715 in Brown, B. (1916). 2596: 2376:shells, water-worn bones, and a 2144:. Small lambeosaurines from the 1010:, to estimate their weight. The 719:5240, was discovered in 1911 by 470: 141: 6507:List of informal dinosaur names 768:in their original death poses. 558:. Its name is derived from the 6517:Timeline of hadrosaur research 2993:pp. 1117–1135 in Arbour, V.M. 2636:Merriam-Webster.com Dictionary 2612:Timeline of hadrosaur research 1363:Originally, Brown referred to 1301:Distinguishing characteristics 650:probably fed on all of these. 1: 6712:Fossil taxa described in 1914 3854:10.1016/j.cretres.2017.04.006 3552:10.1016/S1631-0683(03)00017-4 3033:10.1016/j.cretres.2017.04.006 2692:Lowell Dingus and Mark Norell 2232:was found to be juveniles of 1542:was published in the journal 1465:is currently classified as a 1122:The mouth of the holotype of 661:. Both still co-existed with 3914:10.1016/j.palaeo.2012.06.024 3592:10.1371/journal.pone.0098821 3358:10.1371/journal.pone.0031295 2922:10.1371/journal.pone.0069835 1202:Brachylophosaurus canadensis 890:in the middle of the ocean. 449:Corythosaurus brevicristatus 6742:Campanian genus extinctions 4115:The Dinosauria (Second ed.) 3883:p. 709 in Brown, B. (1916). 3727:Tetragonosaurus erectofrons 3487:p. 565 in Brown, B. (1914). 3478:p. 564 in Brown, B. (1914). 3469:p. 710 in Brown, B. (1916). 3389:p. 711 in Brown, B. (1916). 3314:p. 563 in Brown, B. (1914). 3258:p. 561 in Brown, B. (1914). 2730:p. 560 in Brown, B. (1914). 2710:p. 559 in Brown, B. (1914). 2483:formations, as well as the 2126:, now seen as a synonym of 2102:Tetragonosaurus erectofrons 2084:in the distance travelled. 1695:Lambeosaurus magnicristatus 1220:Lambeosaurus magnicristatus 1167:Replica of skin impressions 807:its name. The generic name 6763: 6717:Taxa named by Barnum Brown 4018:Dyoplosaurus acutosquameus 2560:and an unnamed species of 2146:Horseshoe Canyon Formation 1367:as a member of the family 1270:Saurolophus angustirostris 1229:Saurolophus angustirsotris 1191:Aside from those found on 1114:Skull of the type specimen 872:, a paleontologist of the 868:. They were being sent to 16:Extinct genus of dinosaurs 6174: 6163: 6124: 5853: 5623: 5564: 5284: 5269: 5256: 5227: 5089: 4917: 4809: 4805: 4663: 4423: 4419: 4404: 4391: 4337: 4310: 4297: 4233: 3660:10.1017/S0094837300004036 3631:Weishampel, D.B. (1981). 3333:from Canada and Mongolia" 3162:Paul, Gregory S. (2016). 3097:American Museum Novitates 3090:"The Weight of Dinosaurs" 2870:Corythosaurus intermedius 2217:, parasaurolophines like 2015:. The presence of a thin 1911:Corythosaurus intermedius 1907: 1890: 1883: 1852: 1835: 1828: 1816: 1799: 1782: 1765: 1758: 1741: 1734: 1708: 1691: 1684: 1672: 1646: 1629: 1622: 1615: 1598: 1591: 1556:relationships, including 1188:ends at a rounded point. 440:Corythosaurus bicristatus 382: 375: 333: 328: 312: 305: 138:Scientific classification 136: 128: 119: 23: 4061:Mesozoic Vertebrate Life 2511:, and the ankylosaurids 2329:Dinosaur Provincial Park 2215:Hypacrosaurus stebingeri 2116:Hypacrosaurus stebingeri 1993:Comparisons between the 1856:Hypacrosaurus stebingeri 1839:Hypacrosaurus altispinus 1493:. With the exception of 6732:Paleontology in Alberta 3974:Corythosaurus casuarius 3950:Corythosaurus casuarius 3438:10.1126/science.1200043 2775:Dinosaurs in Their Time 2769:Corythosaurus casuarius 2554:are the tyrannosaurids 2545:in the middle age, and 2463:Corythosaurus casuarius 2443:, and the ceratopsians 2401:Corythosaurus casuarius 2340:Dinosaur Park Formation 2167:Corythosaurus casuarius 2108:Corythosaurus casuarius 1894:Corythosaurus casuarius 1517:was closely related to 1282:Edmontosaurus annectens 1214:Parasaurolophus walkeri 1197:Edmontosaurus annectens 1193:Corythosaurus casuarius 917:(Parks 1923). In 1975, 833:Corythosaurus casuarius 789:Corythosaurus casuarius 659:Dinosaur Park Formation 570:is now thought to be a 392:Corythosaurus excavatus 318:Corythosaurus casuarius 3725:"The Type Specimen of 3532:Comptes Rendus Palevol 3110:Seebacher, F. (2001). 3088:Colbert, E.H. (1962). 2752:p. 158 in Norrell, M. 2739:p. 159 in Norrell, M. 2396: 2366:, the only species of 2331: 2174: 2104: 2040: 1580: 1360: 1313: 1176: 1168: 1115: 1080: 998:(2012) estimated that 991: 934:in 1920, was named by 843: 780: 712: 6664:Paleobiology Database 6487:Basturs Poble bonebed 3814:p. 41 in Norrell, M. 3702:p. 35 in Norrell, M. 2868:Parks, W.A. (1923). " 2390: 2326: 2160: 2099: 2059:, are S-shaped, with 2034: 1570: 1426:Alfred Sherwood Romer 1355: 1308: 1208:Gryposaurus notabilis 1174: 1166: 1113: 1072: 981: 870:Arthur Smith Woodward 841: 779:by the Red Deer River 774: 710: 703:Discovery and species 556:western North America 550:, about 77–75.7  6747:Ornithischian genera 6509:(contains hadrosaur 5907:Probrachylophosaurus 4180:at Wikimedia Commons 4042:10.1671/039.029.0405 3900:. 350–352: 124–138. 3506:Bulletin of the AMNH 3497:Romer, A.S. (1923). 3327:Bell, P. R. (2012). 3279:The Age of Dinosaurs 2895:Canardia garonnensis 2557:Gorgosaurus libratus 2550:of the formation as 2472:Centrosaurus apertus 2200:, but dissimilar to 852:Charles H. Sternberg 715:The first specimen, 6727:Dinosaur Park fauna 4034:2009JVPal..29.1117A 3906:2012PPP...350..124M 3846:2017CrRes..76....7B 3834:Cretaceous Research 3652:1981Pbio....7..252W 3583:2014PLoSO...998821X 3544:2003CRPal...2..143G 3430:2011Sci...332..705S 3349:2012PLoSO...731295B 3215:2020Pbio...46..379W 3025:2017CrRes..76....7B 3013:Cretaceous Research 2956:Glishades ericksoni 2913:2013PLoSO...869835P 2437:, the ankylosaurid 2419:, the hadrosaurids 2112:Lambeosaurus lambei 1712:Lambeosaurus lambei 1379:. Brown classified 1287:Lambeosaurus lambei 1179:In the holotype of 1091:resemble that of a 653:The two species of 6259:Ornithopodichnites 5870:Brachylophosaurini 4455:Hypsilophodontidae 3970:Brown, B. (1916). 3946:Brown, B. (1914). 3224:10.1017/pab.2020.2 2816:Systematic Zoology 2503:, the ceratopsids 2397: 2332: 2175: 2105: 2076:sounds (below 400 2041: 1581: 1513:(2012) found that 1411:in Saurolophinae. 1361: 1314: 1177: 1169: 1116: 1081: 992: 913:(Parks 1935), and 844: 781: 748:and pieces of the 713: 574:, thus related to 6689: 6688: 6651:Open Tree of Life 6565:Taxon identifiers 6556: 6555: 6552: 6551: 6548: 6547: 6266:Shiraminesauropus 6159: 6158: 6122: 6121: 6118: 6117: 6114: 6113: 6110: 6109: 6005:Naashoibitosaurus 5926:Austrokritosauria 5886:Brachylophosaurus 5840: 5839: 5252: 5251: 5225: 5224: 5221: 5220: 5217: 5216: 5213: 5212: 5209: 5208: 4902: 4901: 4790: 4789: 4786: 4785: 4770:Transylvanosaurus 4656:Rhabdodontomorpha 4602:Notohypsilophodon 4573:Macrogryphosaurus 4387: 4386: 4383: 4382: 4379: 4378: 4375: 4374: 4176:Media related to 4157:978-0-520-22501-5 4130:978-0-520-24209-8 4103:978-0-253-34595-0 4076:978-0-253-33907-2 4005:978-0-7566-9910-9 3783:10.7717/peerj.182 3746:978-0-253-34595-0 3175:978-1-78684-190-2 3074:p. 345 in Benson 2639:. Merriam-Webster 2422:Brachylophosaurus 2416:Saurornitholestes 2395:in the background 2148:were referred to 1985: 1984: 1976: 1975: 1967: 1966: 1958: 1957: 1949: 1948: 1940: 1939: 1931: 1930: 1922: 1921: 1867: 1866: 1723: 1722: 1661: 1660: 1571:ROM 845, mounted 1358:C. brevicristatus 1263:are all similar. 1257:S. angustirostris 1249:S. angustirsotris 1097:Corinthian helmet 903:C. brevicristatus 733:Belly River Group 678:Brachylophosaurus 628:Corinthian helmet 554:, in what is now 552:million years ago 461: 460: 454: 445: 436: 424: 412: 400: 369: 351: 301: 6754: 6682: 6681: 6672: 6671: 6659: 6658: 6646: 6645: 6633: 6632: 6620: 6619: 6607: 6606: 6605: 6592: 6591: 6590: 6560: 6533:mummy S.M. R4036 6407:Microhadrosaurus 6400:Mandschurosaurus 6210:Camptosaurichnus 6203:Bonaparteichnium 6176: 6169:Related articles 6165: 6154: 6148: 6143: 6138: 6137: 6130: 5851: 5732:Parasaurolophini 5643:Angulomastacator 5621: 5562: 5481:Shuangmiaosaurus 5382:Huehuecanauhtlus 5368:Gongpoquansaurus 5282: 5271: 5258: 5247: 5241: 5240: 5233: 5087: 5082:Hadrosauriformes 5050:Ratchasimasaurus 4915: 4807: 4803: 4685:Muttaburrasaurus 4661: 4594:Muttaburrasaurus 4508:Atlascopcosaurus 4443:Thescelosauridae 4421: 4417: 4406: 4393: 4335: 4308: 4299: 4292: 4291: 4235: 4220: 4213: 4206: 4197: 4188:Data related to 4187: 4175: 4161: 4145: 4134: 4118: 4107: 4091: 4080: 4064: 4053: 4028:(4): 1117–1135. 4009: 3996:Prehistoric Life 3990: 3980: 3966: 3956: 3934: 3931: 3918: 3917: 3893: 3884: 3881: 3875: 3872: 3866: 3865: 3825: 3819: 3812: 3806: 3805: 3795: 3785: 3757: 3751: 3750: 3720: 3707: 3700: 3687: 3686: 3684: 3678:. Archived from 3637: 3628: 3615: 3614: 3604: 3594: 3562: 3556: 3555: 3529: 3520: 3514: 3513: 3503: 3494: 3488: 3485: 3479: 3476: 3470: 3467: 3458: 3457: 3413: 3402: 3399: 3390: 3387: 3381: 3380: 3370: 3360: 3324: 3315: 3312: 3303: 3296: 3290: 3274: 3259: 3256: 3237: 3236: 3226: 3194: 3188: 3187: 3159: 3153: 3152: 3134: 3116: 3107: 3101: 3100: 3094: 3085: 3079: 3072: 3045: 3044: 3004: 2998: 2991: 2968: 2967: 2951: 2945: 2944: 2934: 2924: 2888: 2882: 2881: 2865: 2859: 2858: 2846: 2840: 2839: 2808: 2799: 2796: 2790: 2789: 2787: 2786: 2763: 2757: 2750: 2744: 2737: 2731: 2728: 2711: 2708: 2697: 2690: 2684: 2683: 2681: 2679: 2654: 2648: 2647: 2645: 2644: 2627: 2606: 2604:Dinosaurs portal 2601: 2600: 2599: 2489:Wheatland County 2485:Wind River Basin 2336:Oldman Formation 2244:was assigned to 2161:Skull growth of 1886: 1831: 1819: 1761: 1737: 1687: 1675: 1625: 1618: 1594: 1584: 1291:L. clavinitialis 1074:Life restoration 1004:Edwin H. Colbert 986:(left, red) and 909:(Gilmore 1923), 823:, refers to the 530: 527: 524: 521: 518: 514: 513: 510: 509: 506: 503: 500: 497: 494: 491: 488: 485: 482: 479: 476: 452: 443: 434: 422: 410: 395: 363: 356: 345: 338: 315: 293: 286: 273: 260: 247: 234: 218: 202: 146: 145: 124: 114: 44: 29:Temporal range: 21: 6762: 6761: 6757: 6756: 6755: 6753: 6752: 6751: 6692: 6691: 6690: 6685: 6677: 6675: 6667: 6662: 6654: 6649: 6641: 6636: 6628: 6623: 6615: 6610: 6601: 6600: 6595: 6586: 6585: 6580: 6567: 6557: 6544: 6525:mummy AMNH 5060 6492:Dakota (fossil) 6468: 6319: 6299: 6287:Tetrapodosaurus 6238:Hadrosaurichnus 6170: 6155: 6132: 6106: 6101:Shantungosaurus 6051: 6032:Augustynolophus 6017: 5961: 5935:Bonapartesaurus 5920: 5864: 5836: 5767: 5755:Parasaurolophus 5726: 5610: 5558: 5549: 5474:Qianjiangsaurus 5460:Probactrosaurus 5439:Penelopognathus 5278: 5265: 5248: 5235: 5205: 5185: 5142: 5076: 5057:Riabininohadros 4930:Bayannurosaurus 4911: 4898: 4834: 4799: 4782: 4727: 4697:Tenontosauridae 4691: 4650: 4486: 4465:Gideonmantellia 4449: 4413: 4400: 4371: 4351: 4324: 4318:Avemetatarsalia 4303:Avemetatarsalia 4293: 4240: 4239: 4229: 4224: 4168: 4158: 4137: 4131: 4110: 4104: 4083: 4077: 4056: 4012: 4006: 3993: 3978: 3969: 3954: 3945: 3942: 3937: 3932: 3921: 3895: 3894: 3887: 3882: 3878: 3873: 3869: 3827: 3826: 3822: 3813: 3809: 3764:Parasaurolophus 3759: 3758: 3754: 3747: 3722: 3721: 3710: 3701: 3690: 3682: 3635: 3630: 3629: 3618: 3564: 3563: 3559: 3527: 3522: 3521: 3517: 3501: 3496: 3495: 3491: 3486: 3482: 3477: 3473: 3468: 3461: 3424:(6030): 705–8. 3415: 3414: 3405: 3400: 3393: 3388: 3384: 3326: 3325: 3318: 3313: 3306: 3297: 3293: 3275: 3262: 3257: 3240: 3196: 3195: 3191: 3176: 3161: 3160: 3156: 3114: 3109: 3108: 3104: 3092: 3087: 3086: 3082: 3073: 3048: 3006: 3005: 3001: 2992: 2971: 2953: 2952: 2948: 2890: 2889: 2885: 2867: 2866: 2862: 2848: 2847: 2843: 2828:10.2307/2412696 2810: 2809: 2802: 2797: 2793: 2784: 2782: 2765: 2764: 2760: 2751: 2747: 2738: 2734: 2729: 2714: 2709: 2700: 2691: 2687: 2677: 2675: 2673: 2656: 2655: 2651: 2642: 2640: 2631:"Corythosaurus" 2629: 2628: 2624: 2620: 2602: 2597: 2595: 2592: 2501:Parasaurolophus 2434:Parasaurolophus 2321: 2307:, meaning that 2270: 2250:Tetragonosaurus 2230:Tetragonosaurus 2219:Parasaurolophus 2202:Parasaurolophus 2192:are similar to 2182:Parasaurolophus 2163:Parasaurolophus 2141:Parasaurolophus 2128:Procheneosaurus 2124:Tetragonosaurus 2094: 2061:Parasaurolophus 2039:as semi-aquatic 2029: 1991: 1986: 1977: 1968: 1959: 1950: 1941: 1932: 1923: 1868: 1724: 1662: 1350: 1320:from all other 1303: 1161: 1067: 976: 971: 882:merchant raider 857:SS Mount Temple 705: 684:Parasaurolophus 545:Late Cretaceous 528: 526:helmeted lizard 525: 522: 519: 473: 469: 457: 451: 442: 433: 429:Tetragonosaurus 421: 417:Procheneosaurus 409: 394: 388: 387: 362: 344: 324: 321: 292: 284: 271: 258: 245: 232: 216: 200: 140: 131:Carnegie Museum 115: 113: 112: 111: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 39: 38: 31:Late Cretaceous 27: 17: 12: 11: 5: 6760: 6758: 6750: 6749: 6744: 6739: 6734: 6729: 6724: 6719: 6714: 6709: 6707:Lambeosaurines 6704: 6694: 6693: 6687: 6686: 6684: 6683: 6673: 6660: 6647: 6634: 6621: 6608: 6593: 6577: 6575: 6569: 6568: 6563: 6554: 6553: 6550: 6549: 6546: 6545: 6543: 6542: 6535: 6527: 6519: 6514: 6504: 6502:Hypsilophodont 6499: 6497:Hadrosaur diet 6494: 6489: 6484: 6476: 6474: 6473:Other articles 6470: 6469: 6467: 6466: 6459: 6452: 6445: 6438: 6435:Stephanosaurus 6431: 6424: 6417: 6410: 6403: 6396: 6389: 6382: 6375: 6372:Fulgurotherium 6368: 6361: 6354: 6346: 6339: 6331: 6329: 6321: 6320: 6318: 6317: 6309: 6307: 6301: 6300: 6298: 6297: 6290: 6283: 6276: 6269: 6262: 6255: 6248: 6245:Iguanodontipus 6241: 6234: 6227: 6220: 6213: 6206: 6199: 6192: 6184: 6182: 6172: 6171: 6168: 6161: 6160: 6157: 6156: 6125: 6123: 6120: 6119: 6116: 6115: 6112: 6111: 6108: 6107: 6105: 6104: 6097: 6094:Laiyangosaurus 6090: 6083: 6076: 6069: 6061: 6059: 6057:Edmontosaurini 6053: 6052: 6050: 6049: 6042: 6039:Prosaurolophus 6035: 6027: 6025: 6019: 6018: 6016: 6015: 6008: 6001: 5994: 5987: 5984:Coahuilasaurus 5980: 5971: 5969: 5963: 5962: 5960: 5959: 5952: 5945: 5938: 5930: 5928: 5922: 5921: 5919: 5918: 5910: 5903: 5896: 5889: 5882: 5874: 5872: 5866: 5865: 5863: 5862: 5854: 5848: 5842: 5841: 5838: 5837: 5835: 5834: 5827: 5820: 5813: 5806: 5799: 5792: 5785: 5777: 5775: 5769: 5768: 5766: 5765: 5758: 5751: 5744: 5736: 5734: 5728: 5727: 5725: 5724: 5717: 5710: 5703: 5696: 5688: 5681: 5674: 5667: 5660: 5653: 5646: 5639: 5632: 5624: 5618: 5616:Lambeosaurinae 5612: 5611: 5609: 5608: 5601: 5594: 5587: 5580: 5573: 5565: 5559: 5554: 5551: 5550: 5548: 5547: 5540: 5533: 5526: 5519: 5512: 5505: 5498: 5491: 5484: 5477: 5470: 5463: 5456: 5449: 5442: 5435: 5432:Nanyangosaurus 5428: 5425:Nanningosaurus 5421: 5414: 5407: 5399: 5392: 5385: 5378: 5371: 5364: 5357: 5354:Gilmoreosaurus 5350: 5343: 5336: 5329: 5322: 5315: 5308: 5300: 5293: 5285: 5279: 5276:Hadrosauroidea 5274: 5267: 5266: 5263:Hadrosauroidea 5261: 5254: 5253: 5250: 5249: 5228: 5226: 5223: 5222: 5219: 5218: 5215: 5214: 5211: 5210: 5207: 5206: 5204: 5203: 5202: 5201: 5195: 5193: 5191:Hadrosauroidea 5187: 5186: 5184: 5183: 5179:Mantellisaurus 5175: 5168: 5161: 5152: 5150: 5148:Iguanodontidae 5144: 5143: 5141: 5140: 5133: 5126: 5123:Mantellisaurus 5119: 5112: 5105: 5098: 5090: 5084: 5078: 5077: 5075: 5074: 5067: 5060: 5053: 5046: 5039: 5032: 5025: 5018: 5011: 5004: 4997: 4990: 4987:Iguanacolossus 4983: 4976: 4969: 4962: 4954: 4947: 4940: 4933: 4926: 4918: 4912: 4909:Ankylopollexia 4907: 4904: 4903: 4900: 4899: 4897: 4896: 4889: 4881: 4873: 4870:Eousdryosaurus 4866: 4859: 4856:Dysalotosaurus 4852: 4844: 4842: 4836: 4835: 4833: 4832: 4825: 4818: 4810: 4800: 4795: 4792: 4791: 4788: 4787: 4784: 4783: 4781: 4780: 4773: 4766: 4759: 4752: 4745: 4737: 4735: 4733:Rhabdodontidae 4729: 4728: 4726: 4725: 4718: 4711: 4702: 4700: 4693: 4692: 4690: 4689: 4681: 4673: 4664: 4658: 4652: 4651: 4649: 4648: 4644:Weewarrasaurus 4640: 4633: 4626: 4619: 4612: 4605: 4598: 4590: 4583: 4576: 4569: 4566:Leaellynasaura 4562: 4554: 4547: 4544:Gasparinisaura 4540: 4537:Galleonosaurus 4533: 4525: 4518: 4511: 4504: 4496: 4494: 4488: 4487: 4485: 4484: 4476: 4469: 4460: 4458: 4451: 4450: 4448: 4447: 4439: 4432: 4424: 4414: 4409: 4402: 4401: 4396: 4389: 4388: 4385: 4384: 4381: 4380: 4377: 4376: 4373: 4372: 4370: 4369: 4368: 4367: 4361: 4359: 4353: 4352: 4350: 4349: 4348: 4347: 4338: 4332: 4326: 4325: 4323: 4322: 4321: 4320: 4311: 4305: 4295: 4294: 4290: 4289: 4280: 4278:Neornithischia 4271: 4262: 4253: 4247: 4238: 4231: 4230: 4225: 4223: 4222: 4215: 4208: 4200: 4194: 4193: 4192:at Wikispecies 4181: 4167: 4166:External links 4164: 4163: 4162: 4156: 4135: 4129: 4108: 4102: 4081: 4075: 4054: 4010: 4004: 3991: 3967: 3941: 3938: 3936: 3935: 3919: 3885: 3876: 3867: 3820: 3807: 3752: 3745: 3708: 3688: 3685:on 2014-10-06. 3646:(2): 252–261. 3616: 3557: 3538:(2): 143–151. 3515: 3489: 3480: 3471: 3459: 3403: 3391: 3382: 3316: 3304: 3300:The Dinosauria 3291: 3260: 3238: 3209:(3): 379–404. 3189: 3174: 3154: 3132:10.1.1.462.255 3102: 3080: 3046: 2999: 2969: 2946: 2883: 2860: 2841: 2800: 2791: 2758: 2745: 2732: 2712: 2698: 2685: 2671: 2649: 2621: 2619: 2616: 2615: 2614: 2608: 2607: 2591: 2588: 2563:Daspletosaurus 2547:Euoplocephalus 2542:Euoplocephalus 2410:Daspletosaurus 2405:C. intermedius 2360:C. intermedius 2327:Skeleton from 2320: 2317: 2269: 2266: 2254:T. erectofrons 2252:, assigned to 2242:T. erectofrons 2093: 2090: 2028: 2027:Crest function 2025: 1990: 1987: 1983: 1982: 1979: 1978: 1974: 1973: 1970: 1969: 1965: 1964: 1961: 1960: 1956: 1955: 1952: 1951: 1947: 1946: 1943: 1942: 1938: 1937: 1934: 1933: 1929: 1928: 1925: 1924: 1920: 1919: 1916: 1915: 1906: 1903: 1902: 1899: 1898: 1889: 1884: 1882: 1874: 1873: 1870: 1869: 1865: 1864: 1861: 1860: 1851: 1848: 1847: 1844: 1843: 1834: 1829: 1827: 1817: 1815: 1812: 1811: 1808: 1807: 1798: 1795: 1794: 1791: 1790: 1781: 1778: 1777: 1774: 1773: 1764: 1759: 1757: 1754: 1753: 1750: 1749: 1740: 1735: 1733: 1730: 1729: 1726: 1725: 1721: 1720: 1717: 1716: 1707: 1704: 1703: 1700: 1699: 1690: 1685: 1683: 1673: 1671: 1668: 1667: 1664: 1663: 1659: 1658: 1655: 1654: 1645: 1642: 1641: 1638: 1637: 1628: 1623: 1621: 1616: 1614: 1611: 1610: 1607: 1606: 1597: 1592: 1590: 1582: 1449:ornithischians 1405:Stephanosaurus 1373:Trachodontinae 1369:Trachodontidae 1349: 1348:Classification 1346: 1338:premaxillaries 1302: 1299: 1160: 1157: 1066: 1063: 1028:C. intermedius 988:C. intermedius 975: 972: 970: 967: 951:C. intermedius 947:C. intermedius 941:Stephanosaurus 932:Levi Sternberg 928:C. intermedius 915:C. intermedius 905:(Parks 1935), 901:(Parks 1935), 899:C. bicristatus 862:United Kingdom 725:Red Deer River 704: 701: 672:Daspletosaurus 667:ornithischians 539:"duck-billed" 459: 458: 456: 455: 446: 437: 425: 423:(Parks, 1931 ) 413: 405:Stephanosaurus 401: 385: 384: 383: 380: 379: 373: 372: 371: 370: 359:C. intermedius 352: 331: 330: 326: 325: 322: 310: 309: 303: 302: 282: 278: 277: 269: 265: 264: 262:Lambeosaurinae 256: 252: 251: 243: 239: 238: 230: 223: 222: 220:Neornithischia 214: 207: 206: 198: 191: 190: 185: 178: 177: 172: 168: 167: 162: 158: 157: 152: 148: 147: 134: 133: 126: 125: 117: 116: 108: 107: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 45: 28: 15: 13: 10: 9: 6: 4: 3: 2: 6759: 6748: 6745: 6743: 6740: 6738: 6735: 6733: 6730: 6728: 6725: 6723: 6720: 6718: 6715: 6713: 6710: 6708: 6705: 6703: 6700: 6699: 6697: 6680: 6674: 6670: 6665: 6661: 6657: 6652: 6648: 6644: 6639: 6635: 6631: 6626: 6622: 6618: 6613: 6609: 6604: 6603:Corythosaurus 6598: 6594: 6589: 6583: 6579: 6578: 6576: 6574: 6573:Corythosaurus 6570: 6566: 6561: 6541: 6540: 6536: 6534: 6532: 6531:Edmontosaurus 6528: 6526: 6524: 6523:Edmontosaurus 6520: 6518: 6515: 6512: 6508: 6505: 6503: 6500: 6498: 6495: 6493: 6490: 6488: 6485: 6483: 6482: 6478: 6477: 6475: 6471: 6465: 6464: 6460: 6458: 6457: 6453: 6451: 6450: 6446: 6444: 6443: 6442:Syngonosaurus 6439: 6437: 6436: 6432: 6430: 6429: 6425: 6423: 6422: 6418: 6416: 6415: 6411: 6409: 6408: 6404: 6402: 6401: 6397: 6395: 6394: 6390: 6388: 6387: 6386:Koutalisaurus 6383: 6381: 6380: 6376: 6374: 6373: 6369: 6367: 6366: 6365:Eucercosaurus 6362: 6360: 6359: 6355: 6352: 6351: 6347: 6345: 6344: 6340: 6338: 6337: 6336:Arstanosaurus 6333: 6332: 6330: 6328: 6327: 6322: 6316: 6315: 6311: 6310: 6308: 6306: 6302: 6296: 6295: 6291: 6289: 6288: 6284: 6282: 6281: 6277: 6275: 6274: 6270: 6268: 6267: 6263: 6261: 6260: 6256: 6254: 6253: 6252:Orcauichnites 6249: 6247: 6246: 6242: 6240: 6239: 6235: 6233: 6232: 6228: 6226: 6225: 6221: 6219: 6218: 6214: 6212: 6211: 6207: 6205: 6204: 6200: 6198: 6197: 6193: 6191: 6190: 6189:Amblydactylus 6186: 6185: 6183: 6181: 6177: 6173: 6166: 6162: 6153: 6149: 6147: 6142: 6136: 6129: 6103: 6102: 6098: 6096: 6095: 6091: 6089: 6088: 6087:Kundurosaurus 6084: 6082: 6081: 6080:Kerberosaurus 6077: 6075: 6074: 6070: 6068: 6067: 6066:Edmontosaurus 6063: 6062: 6060: 6058: 6054: 6048: 6047: 6043: 6041: 6040: 6036: 6034: 6033: 6029: 6028: 6026: 6024: 6020: 6014: 6013: 6009: 6007: 6006: 6002: 6000: 5999: 5995: 5993: 5992: 5988: 5986: 5985: 5981: 5978: 5977: 5976:Anasazisaurus 5973: 5972: 5970: 5968: 5964: 5958: 5957: 5956:Secernosaurus 5953: 5951: 5950: 5949:Kelumapusaura 5946: 5944: 5943: 5939: 5937: 5936: 5932: 5931: 5929: 5927: 5923: 5916: 5915: 5911: 5909: 5908: 5904: 5902: 5901: 5897: 5895: 5894: 5890: 5888: 5887: 5883: 5881: 5880: 5876: 5875: 5873: 5871: 5867: 5861: 5860: 5856: 5855: 5852: 5849: 5847: 5846:Saurolophinae 5843: 5833: 5832: 5828: 5826: 5825: 5821: 5819: 5818: 5814: 5812: 5811: 5807: 5805: 5804: 5800: 5798: 5797: 5796:Hypacrosaurus 5793: 5791: 5790: 5789:Corythosaurus 5786: 5784: 5783: 5779: 5778: 5776: 5774: 5773:Lambeosaurini 5770: 5764: 5763: 5759: 5757: 5756: 5752: 5750: 5749: 5748:Charonosaurus 5745: 5743: 5742: 5738: 5737: 5735: 5733: 5729: 5723: 5722: 5721:Tsintaosaurus 5718: 5716: 5715: 5714:Pararhabdodon 5711: 5709: 5708: 5707:Nipponosaurus 5704: 5702: 5701: 5697: 5694: 5693: 5689: 5687: 5686: 5682: 5680: 5679: 5678:Jaxartosaurus 5675: 5673: 5672: 5668: 5666: 5665: 5661: 5659: 5658: 5654: 5652: 5651: 5647: 5645: 5644: 5640: 5638: 5637: 5633: 5631: 5630: 5629:Adynomosaurus 5626: 5625: 5622: 5619: 5617: 5613: 5607: 5606: 5602: 5600: 5599: 5595: 5593: 5592: 5591:Lapampasaurus 5588: 5586: 5585: 5581: 5579: 5578: 5574: 5572: 5571: 5567: 5566: 5563: 5560: 5557: 5556:Hadrosauridae 5552: 5546: 5545: 5541: 5539: 5538: 5537:Zhanghenglong 5534: 5532: 5531: 5527: 5525: 5524: 5520: 5518: 5517: 5513: 5511: 5510: 5509:Telmatosaurus 5506: 5504: 5503: 5499: 5497: 5496: 5492: 5490: 5489: 5485: 5483: 5482: 5478: 5476: 5475: 5471: 5469: 5468: 5464: 5462: 5461: 5457: 5455: 5454: 5453:Portellsaurus 5450: 5448: 5447: 5443: 5441: 5440: 5436: 5434: 5433: 5429: 5427: 5426: 5422: 5420: 5419: 5415: 5413: 5412: 5408: 5405: 5404: 5400: 5398: 5397: 5393: 5391: 5390: 5386: 5384: 5383: 5379: 5377: 5376: 5372: 5370: 5369: 5365: 5363: 5362: 5358: 5356: 5355: 5351: 5349: 5348: 5344: 5342: 5341: 5337: 5335: 5334: 5330: 5328: 5327: 5323: 5321: 5320: 5316: 5314: 5313: 5309: 5306: 5305: 5301: 5299: 5298: 5294: 5292: 5291: 5287: 5286: 5283: 5280: 5277: 5272: 5268: 5264: 5259: 5255: 5246: 5242: 5239: 5232: 5199: 5198: 5197: 5196: 5194: 5192: 5188: 5181: 5180: 5176: 5174: 5173: 5169: 5167: 5166: 5162: 5159: 5158: 5154: 5153: 5151: 5149: 5145: 5139: 5138: 5134: 5132: 5131: 5127: 5125: 5124: 5120: 5118: 5117: 5116:Jinzhousaurus 5113: 5111: 5110: 5109:Hypselospinus 5106: 5104: 5103: 5099: 5097: 5096: 5092: 5091: 5088: 5085: 5083: 5079: 5073: 5072: 5068: 5066: 5065: 5064:Theiophytalia 5061: 5059: 5058: 5054: 5052: 5051: 5047: 5045: 5044: 5040: 5038: 5037: 5033: 5031: 5030: 5026: 5024: 5023: 5019: 5017: 5016: 5012: 5010: 5009: 5005: 5003: 5002: 4998: 4996: 4995: 4994:Lanzhousaurus 4991: 4989: 4988: 4984: 4982: 4981: 4977: 4975: 4974: 4970: 4968: 4967: 4963: 4960: 4959: 4955: 4953: 4952: 4948: 4946: 4945: 4941: 4939: 4938: 4934: 4932: 4931: 4927: 4925: 4924: 4920: 4919: 4916: 4913: 4910: 4905: 4895: 4894: 4890: 4887: 4886: 4882: 4879: 4878: 4874: 4872: 4871: 4867: 4865: 4864: 4863:Elrhazosaurus 4860: 4858: 4857: 4853: 4851: 4850: 4846: 4845: 4843: 4841: 4837: 4831: 4830: 4826: 4824: 4823: 4819: 4817: 4816: 4812: 4811: 4808: 4804: 4801: 4798: 4793: 4779: 4778: 4774: 4772: 4771: 4767: 4765: 4764: 4760: 4758: 4757: 4753: 4751: 4750: 4746: 4744: 4743: 4739: 4738: 4736: 4734: 4730: 4724: 4723: 4722:Tenontosaurus 4719: 4717: 4716: 4712: 4709: 4708: 4707:Convolosaurus 4704: 4703: 4701: 4698: 4694: 4687: 4686: 4682: 4679: 4678: 4674: 4671: 4670: 4669:Ampelognathus 4666: 4665: 4662: 4659: 4657: 4653: 4646: 4645: 4641: 4639: 4638: 4634: 4632: 4631: 4627: 4625: 4624: 4620: 4618: 4617: 4613: 4611: 4610: 4606: 4604: 4603: 4599: 4596: 4595: 4591: 4589: 4588: 4584: 4582: 4581: 4580:Mahuidacursor 4577: 4575: 4574: 4570: 4568: 4567: 4563: 4560: 4559: 4555: 4553: 4552: 4548: 4546: 4545: 4541: 4539: 4538: 4534: 4531: 4530: 4526: 4524: 4523: 4519: 4517: 4516: 4512: 4510: 4509: 4505: 4503: 4502: 4498: 4497: 4495: 4493: 4489: 4482: 4481: 4477: 4475: 4474: 4473:Hypsilophodon 4470: 4467: 4466: 4462: 4461: 4459: 4456: 4452: 4445: 4444: 4440: 4438: 4437: 4436:Callovosaurus 4433: 4431: 4430: 4429:Burianosaurus 4426: 4425: 4422: 4418: 4415: 4412: 4407: 4403: 4399: 4394: 4390: 4365: 4364: 4363: 4362: 4360: 4358: 4354: 4346: 4342: 4341: 4340: 4339: 4336: 4333: 4331: 4327: 4319: 4315: 4314: 4313: 4312: 4309: 4306: 4304: 4300: 4296: 4288: 4284: 4281: 4279: 4275: 4272: 4270: 4266: 4263: 4261: 4257: 4254: 4252: 4248: 4246: 4242: 4241: 4236: 4232: 4228: 4221: 4216: 4214: 4209: 4207: 4202: 4201: 4198: 4191: 4190:Corythosaurus 4186: 4182: 4179: 4178:Corythosaurus 4174: 4170: 4169: 4165: 4159: 4153: 4149: 4144: 4143: 4136: 4132: 4126: 4122: 4117: 4116: 4109: 4105: 4099: 4095: 4090: 4089: 4082: 4078: 4072: 4068: 4063: 4062: 4055: 4051: 4047: 4043: 4039: 4035: 4031: 4027: 4023: 4019: 4015: 4011: 4007: 4001: 3997: 3992: 3988: 3984: 3977: 3975: 3968: 3964: 3960: 3953: 3951: 3944: 3943: 3939: 3930: 3928: 3926: 3924: 3920: 3915: 3911: 3907: 3903: 3899: 3892: 3890: 3886: 3880: 3877: 3871: 3868: 3863: 3859: 3855: 3851: 3847: 3843: 3839: 3835: 3831: 3824: 3821: 3817: 3811: 3808: 3803: 3799: 3794: 3789: 3784: 3779: 3775: 3771: 3767: 3765: 3756: 3753: 3748: 3742: 3738: 3734: 3730: 3728: 3719: 3717: 3715: 3713: 3709: 3705: 3699: 3697: 3695: 3693: 3689: 3681: 3677: 3673: 3669: 3665: 3661: 3657: 3653: 3649: 3645: 3641: 3634: 3627: 3625: 3623: 3621: 3617: 3612: 3608: 3603: 3598: 3593: 3588: 3584: 3580: 3577:(6): e98821. 3576: 3572: 3568: 3561: 3558: 3553: 3549: 3545: 3541: 3537: 3533: 3526: 3519: 3516: 3512:(5): 141–145. 3511: 3507: 3500: 3493: 3490: 3484: 3481: 3475: 3472: 3466: 3464: 3460: 3455: 3451: 3447: 3443: 3439: 3435: 3431: 3427: 3423: 3419: 3412: 3410: 3408: 3404: 3398: 3396: 3392: 3386: 3383: 3378: 3374: 3369: 3364: 3359: 3354: 3350: 3346: 3343:(2): e31295. 3342: 3338: 3334: 3332: 3323: 3321: 3317: 3311: 3309: 3305: 3301: 3295: 3292: 3288: 3287:0-7853-0443-6 3284: 3280: 3273: 3271: 3269: 3267: 3265: 3261: 3255: 3253: 3251: 3249: 3247: 3245: 3243: 3239: 3234: 3230: 3225: 3220: 3216: 3212: 3208: 3204: 3200: 3193: 3190: 3185: 3181: 3177: 3171: 3167: 3166: 3158: 3155: 3150: 3146: 3142: 3138: 3133: 3128: 3124: 3120: 3113: 3106: 3103: 3099:(2076): 1–16. 3098: 3091: 3084: 3081: 3077: 3071: 3069: 3067: 3065: 3063: 3061: 3059: 3057: 3055: 3053: 3051: 3047: 3042: 3038: 3034: 3030: 3026: 3022: 3018: 3014: 3010: 3003: 3000: 2996: 2990: 2988: 2986: 2984: 2982: 2980: 2978: 2976: 2974: 2970: 2965: 2961: 2957: 2950: 2947: 2942: 2938: 2933: 2928: 2923: 2918: 2914: 2910: 2907:(7): e69835. 2906: 2902: 2898: 2896: 2887: 2884: 2879: 2875: 2871: 2864: 2861: 2856: 2852: 2845: 2842: 2837: 2833: 2829: 2825: 2821: 2817: 2813: 2807: 2805: 2801: 2795: 2792: 2781:on 2014-08-10 2780: 2776: 2772: 2770: 2762: 2759: 2755: 2749: 2746: 2742: 2736: 2733: 2727: 2725: 2723: 2721: 2719: 2717: 2713: 2707: 2705: 2703: 2699: 2695: 2689: 2686: 2674: 2672:9780253354563 2668: 2664: 2660: 2653: 2650: 2638: 2637: 2632: 2626: 2623: 2617: 2613: 2610: 2609: 2605: 2594: 2589: 2587: 2585: 2580: 2576: 2572: 2571:Corythosaurus 2567: 2565: 2564: 2559: 2558: 2553: 2552:Corythosaurus 2548: 2544: 2543: 2538: 2537: 2536:Panoplosaurus 2532: 2531: 2526: 2525: 2520: 2519: 2514: 2510: 2506: 2502: 2498: 2494: 2493:Corythosaurus 2490: 2486: 2482: 2478: 2474: 2473: 2468: 2467:Corythosaurus 2464: 2460: 2458: 2457:Corythosaurus 2454: 2453: 2448: 2447: 2442: 2441: 2436: 2435: 2430: 2429: 2424: 2423: 2418: 2417: 2412: 2411: 2406: 2403:, as well as 2402: 2394: 2393:Corythosaurus 2389: 2385: 2383: 2382:Corythosaurus 2379: 2375: 2374: 2369: 2368:Corythosaurus 2365: 2361: 2357: 2353: 2352:Corythosaurus 2349: 2345: 2341: 2337: 2330: 2325: 2318: 2316: 2314: 2310: 2309:Corythosaurus 2306: 2302: 2298: 2294: 2290: 2289:Corythosaurus 2286: 2285:Corythosaurus 2282: 2281:Corythosaurus 2278: 2274: 2273:Corythosaurus 2267: 2265: 2263: 2262:Corythosaurus 2259: 2258:Hypacrosaurus 2255: 2251: 2247: 2246:Corythosaurus 2243: 2239: 2235: 2234:Corythosaurus 2231: 2226: 2224: 2220: 2216: 2212: 2208: 2207:Corythosaurus 2203: 2199: 2198:Hypacrosaurus 2195: 2191: 2190:Corythosaurus 2187: 2186:Corythosaurus 2183: 2179: 2178:Corythosaurus 2172: 2168: 2164: 2159: 2155: 2153: 2152: 2147: 2143: 2142: 2137: 2133: 2132:Corythosaurus 2129: 2125: 2121: 2120:H. altispinus 2117: 2113: 2109: 2103: 2098: 2091: 2089: 2085: 2083: 2079: 2074: 2073:Corythosaurus 2070: 2069:Corythosaurus 2066: 2065:Corythosaurus 2062: 2058: 2054: 2053:Hypacrosaurus 2051:, as well as 2050: 2049:Corythosaurus 2046: 2045:Corythosaurus 2038: 2033: 2026: 2024: 2022: 2018: 2014: 2010: 2009: 2004: 2000: 1999:Corythosaurus 1996: 1995:scleral rings 1988: 1981: 1980: 1972: 1971: 1963: 1962: 1954: 1953: 1945: 1944: 1936: 1935: 1927: 1926: 1918: 1917: 1914: 1913: 1912: 1905: 1904: 1901: 1900: 1897: 1896: 1895: 1888: 1887: 1881: 1880: 1879:Corythosaurus 1876: 1875: 1872: 1871: 1863: 1862: 1859: 1858: 1857: 1850: 1849: 1846: 1845: 1842: 1841: 1840: 1833: 1832: 1826: 1825: 1824:Hypacrosaurus 1821: 1820: 1814: 1813: 1810: 1809: 1806: 1805: 1804: 1803:Nipponosaurus 1797: 1796: 1793: 1792: 1789: 1788: 1787: 1780: 1779: 1776: 1775: 1772: 1771: 1770: 1763: 1762: 1756: 1755: 1752: 1751: 1748: 1747: 1746: 1739: 1738: 1732: 1731: 1728: 1727: 1719: 1718: 1715: 1714: 1713: 1706: 1705: 1702: 1701: 1698: 1697: 1696: 1689: 1688: 1682: 1681: 1677: 1676: 1670: 1669: 1666: 1665: 1657: 1656: 1653: 1652: 1651: 1644: 1643: 1640: 1639: 1636: 1635: 1634: 1627: 1626: 1620: 1619: 1613: 1612: 1609: 1608: 1605: 1604: 1603: 1596: 1595: 1589: 1588:Lambeosaurini 1586: 1585: 1578: 1574: 1569: 1565: 1563: 1562:Hypacrosaurus 1559: 1558:Corythosaurus 1555: 1551: 1547: 1546: 1541: 1540: 1539:Zhanghenglong 1534: 1532: 1531:Hypacrosaurus 1528: 1527: 1526:Nipponosaurus 1522: 1521: 1516: 1515:Corythosaurus 1512: 1508: 1507:Corythosaurus 1504: 1503: 1498: 1497: 1492: 1491: 1486: 1485: 1480: 1479:Hypacrosaurus 1476: 1472: 1468: 1464: 1463:Corythosaurus 1460: 1458: 1454: 1450: 1446: 1445:Corythosaurus 1442: 1441:Corythosaurus 1437: 1436:Corythosaurus 1433: 1432: 1431:Tyrannosaurus 1427: 1423: 1422: 1421:Hypacrosaurus 1417: 1416:Corythosaurus 1412: 1410: 1406: 1402: 1401:Corythosaurus 1398: 1394: 1393: 1388: 1384: 1383: 1378: 1377:Saurolophinae 1374: 1370: 1366: 1365:Corythosaurus 1359: 1354: 1347: 1345: 1341: 1339: 1335: 1331: 1327: 1323: 1319: 1318:Corythosaurus 1312: 1307: 1300: 1298: 1294: 1292: 1288: 1284: 1283: 1278: 1277: 1272: 1271: 1266: 1265:Corythosaurus 1262: 1258: 1254: 1250: 1246: 1242: 1238: 1234: 1230: 1226: 1222: 1221: 1216: 1215: 1210: 1209: 1204: 1203: 1198: 1194: 1189: 1186: 1182: 1173: 1165: 1158: 1156: 1152: 1150: 1146: 1142: 1138: 1137:Corythosaurus 1134: 1130: 1125: 1124:Corythosaurus 1120: 1112: 1108: 1106: 1102: 1098: 1094: 1090: 1089:Corythosaurus 1085: 1079: 1075: 1071: 1064: 1062: 1060: 1059: 1054: 1053: 1048: 1047: 1042: 1041: 1040:Edmontosaurus 1035: 1033: 1029: 1025: 1021: 1020:Corythosaurus 1017: 1016:Corythosaurus 1013: 1012:Corythosaurus 1009: 1008:Corythosaurus 1005: 1001: 1000:Corythosaurus 997: 989: 985: 980: 973: 968: 966: 964: 960: 956: 952: 948: 944: 942: 937: 936:William Parks 933: 929: 925: 920: 916: 912: 908: 904: 900: 896: 891: 889: 888: 883: 879: 875: 871: 867: 863: 859: 858: 853: 849: 848:Corythosaurus 840: 836: 834: 830: 826: 822: 818: 817:specific name 814: 810: 809:Corythosaurus 806: 805:Corythosaurus 802: 801:Corythosaurus 798: 794: 790: 786: 778: 773: 769: 767: 763: 759: 755: 751: 747: 743: 739: 734: 730: 726: 722: 718: 709: 702: 700: 698: 697: 692: 691: 686: 685: 680: 679: 674: 673: 668: 664: 660: 656: 655:Corythosaurus 651: 649: 648:Corythosaurus 645: 641: 637: 636:Corythosaurus 634:exhaled. One 633: 632:Corythosaurus 629: 625: 621: 620:Corythosaurus 617: 613: 612:Corythosaurus 609: 607: 606:Corythosaurus 603: 602: 597: 596: 595:Hypacrosaurus 591: 590: 585: 584: 583:Nipponosaurus 579: 578: 573: 572:lambeosaurine 569: 568:Corythosaurus 565: 561: 557: 553: 549: 546: 542: 538: 534: 512: 467: 466: 465:Corythosaurus 450: 447: 441: 438: 432: 430: 426: 420: 418: 414: 408: 406: 402: 398: 393: 390: 381: 378: 374: 367: 361: 360: 353: 349: 346:Brown, 1914 ( 343: 342: 335: 334: 332: 327: 320: 319: 311: 308: 304: 300: 296: 291: 290: 289:Corythosaurus 283: 280: 279: 276: 275:Lambeosaurini 270: 267: 266: 263: 257: 254: 253: 250: 249:Hadrosauridae 244: 241: 240: 237: 231: 228: 225: 224: 221: 215: 212: 209: 208: 205: 199: 196: 193: 192: 189: 186: 183: 180: 179: 176: 173: 170: 169: 166: 163: 160: 159: 156: 153: 150: 149: 144: 139: 135: 132: 127: 123: 118: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 43: 40:77–75.7  36: 32: 26: 25:Corythosaurus 22: 19: 6722:Oldman fauna 6572: 6537: 6530: 6522: 6510: 6479: 6461: 6454: 6447: 6440: 6433: 6426: 6419: 6412: 6405: 6398: 6391: 6384: 6377: 6370: 6363: 6356: 6350:Claorhynchus 6348: 6341: 6334: 6326:Nomina dubia 6324: 6314:Guegoolithus 6312: 6292: 6285: 6280:Staurichnium 6278: 6273:Sousaichnium 6271: 6264: 6257: 6250: 6243: 6236: 6229: 6224:Dakotasaurus 6222: 6217:Caririchnium 6215: 6208: 6201: 6194: 6187: 6131: 6099: 6092: 6085: 6078: 6071: 6064: 6044: 6037: 6030: 6023:Saurolophini 6010: 6003: 5996: 5989: 5982: 5974: 5967:Kritosaurini 5954: 5947: 5942:Huallasaurus 5940: 5933: 5914:Wulagasaurus 5912: 5905: 5898: 5891: 5884: 5877: 5857: 5829: 5822: 5815: 5808: 5803:Lambeosaurus 5801: 5794: 5788: 5787: 5780: 5760: 5753: 5746: 5739: 5719: 5712: 5705: 5698: 5690: 5683: 5676: 5669: 5662: 5655: 5648: 5641: 5634: 5627: 5605:Yamatosaurus 5603: 5596: 5589: 5582: 5575: 5570:Aquilarhinus 5568: 5542: 5535: 5528: 5521: 5516:Tethyshadros 5514: 5507: 5500: 5493: 5486: 5479: 5472: 5465: 5458: 5451: 5446:Plesiohadros 5444: 5437: 5430: 5423: 5418:Lophorhothon 5416: 5409: 5401: 5394: 5387: 5380: 5373: 5366: 5359: 5352: 5345: 5338: 5331: 5324: 5317: 5310: 5304:Batyrosaurus 5302: 5297:Bactrosaurus 5295: 5288: 5234: 5177: 5170: 5163: 5155: 5137:Ouranosaurus 5135: 5128: 5121: 5114: 5107: 5102:Brighstoneus 5100: 5093: 5069: 5062: 5055: 5048: 5041: 5034: 5029:Osmakasaurus 5027: 5022:Oblitosaurus 5020: 5013: 5006: 4999: 4992: 4985: 4978: 4971: 4964: 4956: 4949: 4944:Camptosaurus 4942: 4935: 4928: 4921: 4891: 4885:Kangnasaurus 4883: 4875: 4868: 4861: 4854: 4847: 4840:Dryosauridae 4827: 4820: 4813: 4775: 4768: 4761: 4754: 4747: 4742:Matheronodon 4740: 4720: 4713: 4705: 4683: 4675: 4667: 4642: 4635: 4628: 4621: 4616:Sektensaurus 4614: 4609:Qantassaurus 4607: 4600: 4592: 4585: 4578: 4571: 4564: 4558:Kangnasaurus 4556: 4549: 4542: 4535: 4527: 4522:Diluvicursor 4520: 4513: 4506: 4499: 4480:Vectidromeus 4478: 4471: 4463: 4441: 4434: 4427: 4345:Ornithischia 4330:Ornithischia 4282: 4273: 4269:Ornithischia 4264: 4255: 4141: 4114: 4087: 4060: 4025: 4021: 4017: 4014:Arbour, V.M. 3995: 3986: 3982: 3973: 3962: 3958: 3949: 3897: 3879: 3870: 3837: 3833: 3823: 3815: 3810: 3773: 3769: 3763: 3755: 3732: 3726: 3703: 3680:the original 3643: 3640:Paleobiology 3639: 3574: 3570: 3560: 3535: 3531: 3518: 3509: 3505: 3492: 3483: 3474: 3421: 3417: 3385: 3340: 3336: 3330: 3299: 3294: 3278: 3206: 3203:Paleobiology 3202: 3192: 3164: 3157: 3125:(1): 51–60. 3122: 3118: 3105: 3096: 3083: 3075: 3016: 3012: 3002: 2994: 2963: 2959: 2955: 2949: 2904: 2900: 2894: 2886: 2877: 2873: 2869: 2863: 2854: 2850: 2844: 2822:(1): 37–54. 2819: 2815: 2794: 2783:. Retrieved 2779:the original 2774: 2768: 2767:"DinoGuide: 2761: 2753: 2748: 2740: 2735: 2693: 2688: 2676:. Retrieved 2662: 2652: 2641:. Retrieved 2634: 2625: 2570: 2568: 2561: 2555: 2551: 2546: 2540: 2534: 2530:Dyoplosaurus 2528: 2524:Dyoplosaurus 2522: 2516: 2512: 2509:Chasmosaurus 2508: 2505:Centrosaurus 2504: 2500: 2496: 2492: 2477:Judith River 2470: 2466: 2462: 2461: 2456: 2452:Chasmosaurus 2450: 2446:Coronosaurus 2444: 2438: 2432: 2426: 2420: 2414: 2408: 2404: 2400: 2398: 2392: 2381: 2371: 2367: 2364:C. casuarius 2363: 2359: 2356:C. casuarius 2355: 2351: 2333: 2319:Paleoecology 2308: 2288: 2284: 2280: 2276: 2272: 2271: 2261: 2257: 2253: 2249: 2245: 2241: 2238:Lambeosaurus 2237: 2233: 2229: 2227: 2222: 2218: 2214: 2211:Lambeosaurus 2210: 2206: 2201: 2197: 2194:Lambeosaurus 2193: 2189: 2185: 2181: 2177: 2176: 2170: 2166: 2162: 2151:Cheneosaurus 2149: 2139: 2136:Lambeosaurus 2135: 2131: 2127: 2123: 2119: 2115: 2111: 2107: 2106: 2101: 2086: 2072: 2068: 2064: 2060: 2057:Lambeosaurus 2056: 2052: 2048: 2044: 2042: 2037:C. casuarius 2036: 2008:Lophorhothon 2006: 1998: 1992: 1989:Paleobiology 1910: 1909: 1908: 1893: 1892: 1891: 1878: 1877: 1854: 1853: 1837: 1836: 1822: 1801: 1800: 1784: 1783: 1767: 1766: 1743: 1742: 1710: 1709: 1693: 1692: 1680:Lambeosaurus 1678: 1648: 1647: 1631: 1630: 1600: 1599: 1576: 1561: 1557: 1554:hadrosauroid 1543: 1537: 1535: 1530: 1524: 1518: 1514: 1510: 1506: 1500: 1494: 1488: 1484:Lambeosaurus 1482: 1478: 1462: 1461: 1457:maniraptoran 1444: 1440: 1435: 1429: 1419: 1415: 1413: 1408: 1404: 1400: 1396: 1390: 1386: 1380: 1364: 1362: 1357: 1342: 1322:hadrosaurids 1317: 1315: 1309:Skeleton at 1295: 1290: 1286: 1280: 1274: 1268: 1264: 1261:C. casuarius 1260: 1256: 1252: 1248: 1244: 1241:G. notabilis 1240: 1237:C. casuarius 1236: 1232: 1228: 1224: 1218: 1212: 1206: 1200: 1196: 1192: 1190: 1185:C. casuarius 1184: 1181:C. casuarius 1180: 1178: 1153: 1144: 1140: 1136: 1132: 1128: 1123: 1121: 1117: 1100: 1088: 1086: 1082: 1078:C. casuarius 1077: 1056: 1050: 1044: 1038: 1036: 1032:C. casuarius 1031: 1027: 1024:C. casuarius 1023: 1019: 1015: 1011: 1007: 999: 995: 993: 987: 984:C. casuarius 983: 963:C. excavatus 962: 959:C. casuarius 958: 950: 946: 939: 927: 924:C. casuarius 923: 919:Peter Dodson 914: 911:C. frontalis 910: 907:C. excavatus 906: 902: 898: 895:C. casuarius 894: 892: 886: 855: 847: 845: 832: 820: 812: 808: 804: 800: 788: 785:type species 782: 777:C. casuarius 776: 762:Peter Kaisen 721:Barnum Brown 714: 696:Chasmosaurus 694: 688: 682: 676: 670: 654: 652: 647: 635: 631: 619: 611: 610: 605: 599: 593: 587: 581: 577:Lambeosaurus 575: 567: 564:Barnum Brown 464: 463: 462: 448: 439: 427: 415: 403: 391: 358: 357: 341:C. casuarius 340: 339: 317: 316: 307:Type species 288: 287: 226: 210: 204:Ornithischia 194: 181: 129:Skeleton in 24: 18: 6625:iNaturalist 6597:Wikispecies 6539:Willinakaqe 6511:nomina nuda 6428:Siluosaurus 6393:Loncosaurus 6231:Dinehichnus 6073:Kamuysaurus 6046:Saurolophus 5998:Kritosaurus 5991:Gryposaurus 5824:Sahaliyania 5810:Magnapaulia 5782:Amurosaurus 5762:Tlatolophus 5741:Adelolophus 5685:Kazaklambia 5664:Blasisaurus 5657:Arenysaurus 5650:Aralosaurus 5584:Hadrosaurus 5577:Eotrachodon 5530:Yunganglong 5495:Sirindhorna 5467:Protohadros 5403:Koshisaurus 5396:Jintasaurus 5165:Comptonatus 5015:Napaisaurus 5001:Lurdusaurus 4973:Fukuisaurus 4951:Cedrorestes 4893:Valdosaurus 4756:Pareisactus 4587:Morrosaurus 4551:Isasicursor 4515:Chakisaurus 4411:Ornithopoda 4398:Ornithopoda 4357:Ornithopoda 4227:Ornithopoda 4148:35, 158–159 3331:Saurolophus 2513:Scolosaurus 2497:Gryposaurus 2440:Scolosaurus 2428:Gryposaurus 2223:Kazaklambia 1745:Magnapaulia 1650:Amurosaurus 1633:Sahaliyania 1602:Arenysaurus 1467:hadrosaurid 1459:theropods. 1453:saurischian 1409:Saurolophus 1397:Kritosaurus 1382:Hadrosaurus 1330:prefrontals 1159:Soft tissue 1145:Saurolophus 1141:Kritosaurus 1133:Kritosaurus 1129:Saurolophus 1101:Saurolophus 1058:Saurolophus 1052:Kritosaurus 969:Description 943:intermedius 926:, although 866:World War I 850:, found by 690:Scolosaurus 537:hadrosaurid 453:Parks, 1935 444:Parks, 1935 435:Parks, 1931 431:erectofrons 419:erectofrons 411:Parks, 1923 407:intermedius 323:Brown, 1914 255:Subfamily: 236:Ornithopoda 6696:Categories 6481:Arkharavia 6456:Tichosteus 6449:Thespesius 6421:Pteropelyx 6414:Orthomerus 6294:Wintonopus 5879:Acristavus 5859:Barsboldia 5817:Olorotitan 5692:Latirhinus 5544:Zuoyunlong 5411:Levnesovia 5361:Gobihadros 5326:Datonglong 5319:Claosaurus 5290:Altirhinus 5200:see below↓ 5130:Morelladon 5008:Magnamanus 4980:Hippodraco 4849:Dryosaurus 4822:Hesperonyx 4797:Dryomorpha 4637:Trinisaura 4630:Tietasaura 4623:Talenkauen 4501:Anabisetia 4366:see below↓ 4260:Dinosauria 3989:: 709–715. 3965:: 559–565. 3940:References 2812:Dodson, P. 2785:2014-07-27 2678:13 October 2643:2022-09-27 2518:Edmontonia 2338:and lower 2003:cathemeral 1786:Olorotitan 1502:Olorotitan 1496:Olorotitan 1490:Olorotitan 1475:hadrosaurs 1447:and other 1392:Claosaurus 1276:S. osborni 1245:P. walkeri 1043:(formerly 797:plesiotype 665:and other 601:Olorotitan 188:Dinosauria 6463:Trachodon 6379:Glishades 6358:Diclonius 6196:Anomoepus 6180:Ichnotaxa 5893:Maiasaura 5831:Velafrons 5340:Equijubus 5312:Choyrodon 5172:Iguanodon 5036:Planicoxa 4966:Dakotadon 4937:Calvarius 4763:Rhabdodon 4749:Mochlodon 4492:Elasmaria 4243:Kingdom: 3862:0195-6671 3818:. (2000). 3233:221666530 3184:985402380 3127:CiteSeerX 3041:0195-6671 2618:Footnotes 2584:mammalian 2579:Judithian 2481:Mesaverde 2348:Campanian 2295:needles, 2171:Casuarius 1769:Velafrons 1550:cladogram 1520:Velafrons 1471:subfamily 1387:Trachodon 1253:L. lambei 1233:L. lambei 1225:L. lambei 1093:cassowary 1046:Trachodon 955:Campanian 885:SMS  825:cassowary 821:casuarius 746:phalanges 742:coracoids 663:theropods 642:needles, 624:cassowary 589:Velafrons 543:from the 161:Kingdom: 155:Eukaryota 35:Campanian 6582:Wikidata 6343:Cionodon 6012:Rhinorex 5900:Ornatops 5700:Minqaria 5671:Canardia 5598:Malefica 5523:Xuwulong 5488:Siamodon 5389:Jeyawati 5375:Gonkoken 5333:Eolambia 5157:Barilium 4958:Cumnoria 4923:Barilium 4829:Owenodon 4815:Draconyx 4777:Zalmoxes 4677:Fostoria 4529:Fostoria 4287:Cerapoda 4251:Chordata 4249:Phylum: 4245:Animalia 4050:85665879 3840:: 7–18. 3802:24167777 3776:: e182. 3676:89109302 3611:24901454 3571:PLOS ONE 3454:33253407 3446:21493820 3377:22319623 3337:PLOS ONE 3149:53446536 3019:: 7–18. 2966:: 65–75. 2941:23922815 2901:PLOS ONE 2590:See also 2487:and the 1573:skeleton 1545:PLOS ONE 1477:such as 1334:frontals 1149:lacrimal 982:Size of 829:binomial 813:korythos 793:holotype 738:scapulae 616:holotype 541:dinosaur 377:Synonyms 368:, 1923 ) 329:Species 242:Family: 175:Chordata 171:Phylum: 165:Animalia 151:Domain: 6679:4126538 6656:4128523 6643:1065845 6617:4823234 6588:Q131078 5636:Ajnabia 5071:Uteodon 4067:280–328 4030:Bibcode 3902:Bibcode 3842:Bibcode 3793:3807589 3737:349–363 3706:(2000). 3668:2400478 3648:Bibcode 3602:4047018 3579:Bibcode 3540:Bibcode 3426:Bibcode 3418:Science 3368:3272031 3345:Bibcode 3211:Bibcode 3078:(2012). 3021:Bibcode 2997:(2009). 2932:3724916 2909:Bibcode 2880:: 5–57. 2836:2412696 2756:(2000). 2743:(2000). 2575:Alberta 2539:, and 2313:browser 2293:conifer 2021:eardrum 1469:in the 994:Benson 878:England 864:during 860:to the 729:Alberta 669:, like 640:conifer 531:) is a 520:  397:Gilmore 281:Genus: 268:Tribe: 109:↓ 6676:uBio: 6630:553031 6305:Ootaxa 5502:Tanius 5095:Bolong 4154:  4127:  4100:  4094:54–435 4073:  4048:  4002:  3860:  3800:  3790:  3743:  3704:et al. 3674:  3666:  3609:  3599:  3452:  3444:  3375:  3365:  3285:  3231:  3182:  3172:  3147:  3129:  3076:et al. 3039:  2995:et al. 2939:  2929:  2857:: 130. 2834:  2754:et al. 2741:et al. 2669:  2521:, and 2491:area. 2479:, and 2431:, and 2378:baenid 2344:Canada 2305:fruits 2303:, and 2277:et al. 2118:, and 2092:Growth 2082:spread 2017:stapes 2013:lagena 1529:, and 1511:et al. 1487:, and 1407:, and 1395:, and 1332:, and 1326:nasals 1285:, and 1259:, and 1247:, and 996:et al. 756:, and 750:humeri 693:, and 626:and a 598:, and 548:period 399:, 1923 6669:38760 6638:IRMNG 5347:Fylax 4877:Iyuku 4283:Clade 4274:Clade 4265:Clade 4256:Clade 4046:S2CID 3979:(PDF) 3955:(PDF) 3816:et al 3770:PeerJ 3683:(PDF) 3672:S2CID 3664:JSTOR 3636:(PDF) 3528:(PDF) 3502:(PDF) 3450:S2CID 3229:S2CID 3145:S2CID 3115:(PDF) 3093:(PDF) 2832:JSTOR 2301:twigs 2297:seeds 2165:sp., 1105:orbit 1095:or a 1065:Skull 1055:, or 957:than 758:radii 754:ulnae 644:seeds 560:Greek 533:genus 366:Parks 295:Brown 227:Clade 211:Clade 195:Clade 182:Clade 6612:GBIF 5043:Proa 4715:Iani 4343:see 4316:see 4152:ISBN 4125:ISBN 4098:ISBN 4071:ISBN 4000:ISBN 3858:ISSN 3798:PMID 3741:ISBN 3607:PMID 3442:PMID 3373:PMID 3283:ISBN 3180:OCLC 3170:ISBN 3037:ISSN 2937:PMID 2680:2020 2667:ISBN 2507:and 2499:and 2449:and 2413:and 2373:Unio 2268:Diet 2196:and 2169:and 2055:and 1434:and 1375:and 1311:AMNH 1143:and 974:Size 887:Möwe 783:The 740:and 717:AMNH 517:lit. 386:List 348:type 299:1914 48:PreꞒ 4121:834 4038:doi 3910:doi 3850:doi 3788:PMC 3778:doi 3656:doi 3597:PMC 3587:doi 3548:doi 3434:doi 3422:332 3363:PMC 3353:doi 3219:doi 3137:doi 3029:doi 2927:PMC 2917:doi 2824:doi 2342:of 2236:or 2134:or 1997:of 1575:of 1552:of 1505:is 1289:(= 1076:of 1049:), 876:in 831:of 727:of 723:in 580:, 535:of 502:ɔːr 37:), 6698:: 6666:: 6653:: 6640:: 6627:: 6614:: 6599:: 6584:: 4285:: 4276:: 4267:: 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Index

Late Cretaceous
Campanian
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Carnegie Museum
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Ornithischia
Neornithischia
Ornithopoda
Hadrosauridae
Lambeosaurinae
Lambeosaurini
Corythosaurus

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