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Pseudosuchia

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crocodiles than to birds" (Gauthier and Padian), "Extant crocodiles and all extinct archosaurs that are closer to crocodiles than they are to birds" (Gauthier, 1986), and more recently "the most inclusive clade within Archosauria that includes Crocodylia but not Aves" (Senter, 2005). As a branch-based clade, Pseudosuchia is the
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Pseudosuchia was defined as a stem-based clade in 1985. It includes crocodiles and all archosaurs more closely related to crocodiles than to birds. A second clade with a similar definition, Crocodylotarsi, was named in 1988, possibly as a replacement for Pseudosuchia. The name Pseudosuchia, meaning
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placement of phytosaurs. In 2011, Sterling J. Nesbitt found phytosaurs to be the sister taxon of Archosauria, and therefore not crocodile-line archosaurs. Because phytosaurs are included in the definition of Crurotarsi, crurotarsans are not solely crocodile-line archosaurs, but also bird-line
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for all archosaurs more closely related to crocodilians than to birds. This made the name Pseudosuchia somewhat ironic because true crocodiles (i.e. members of Crocodylia) were now included in the group. Phylogenetic definitions of Pseudosuchia include "Crocodiles and all archosaurs closer to
697:'s 2011 analysis places one crurotarsan group, Phytosauria, outside Pseudosuchia. Since the definition of Crurotarsi relies on phytosaurs, their placement outside Pseudosuchia (and thus Archosauria) means that the clade Crurotarsi includes both pseudosuchians and avemetatarsalians. 1677:, the retention of phytosaurs within Pseudosuchia, and a close relation between aetosaurs and crocodylomorphs, replicate the results of older studies. However, the findings of Nesbitt (2011) have been more widely supported by pseudosuchian-focused analyses published since 2011. 689:. It is not equivalent to Pseudosuchia, which by definition must include all crocodilian-line archosaurs. For many years, Pseudosuchia and Crurotarsi have been considered partial synonyms because the latter clade encompasses all crocodilian-line archosaurs in most 815:
progressed, the Protosuchia gave rise to more typically crocodile-like forms. While dinosaurs were the dominant animals on land, the crocodiles flourished in rivers, swamps, and the oceans, with far greater diversity than they have today. With the
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Contrary to popular belief, crocodilians differ significantly from their ancestors and distant relatives, as Pseudosuchia contains a staggering diversity of reptiles with many different lifestyles. Early pseudosuchians were successful in the
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Ezcurra, Martín D.; Fiorelli, Lucas E.; Martinelli, Agustín G.; Rocher, Sebastián; von Baczko, M. Belén; Ezpeleta, Miguel; Taborda, Jeremías R.A.; Hechenleitner, E. Martín; Trotteyn, M. Jimena; Desojo, Julia B. (11 September 2017).
848:("true crocodiles") as well. Crocodylotarsi may have been named to remove confusion, but as a stem-based clade, it is synonymous with Pseudosuchia. Because Pseudosuchia was named first, it has precedence. A third group, 1799: 1093: 339:
and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs". Despite Pseudosuchia meaning "false crocodiles", the name is a
535:. The clade Pseudosuchia as originally defined could still be used to identify crocodilian-line archosaurs, and since many recent studies support Nesbitt's findings, Pseudosuchia is again commonly used. 857:
archosaurs and phytosaurs. Under this phylogeny, Crurotarsi includes phytosaurs, crocodiles, pterosaurs, and dinosaurs, while Pseudosuchia still contains only crocodile-line archosaurs. Below is a
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Gauthier, J. A.; Padian, K. (1985). "Phylogenetic, functional, and aerodynamic analyses of the origin of birds and their flight". In Hecht, M. K.; Ostrom, J. H.; Viohl, G.; Wellnhofer, P. (eds.).
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Interestingly, the Mesozoic range of cranial disparity is higher than the Triassic one, suggesting crocodylomorphs attained a high degree of diversification compared to Triassic pseudosuchians.
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The following cladogram is from a slightly older study, Brusatte, Benton, Desojo and Langer (2010). Bold terminal taxa are collapsed. Several results of the study, such as the retention of a
844:"false crocodiles", has been used for over a century, and traditionally included only non-crocodilians, but when defined as a clade, Pseudosuchia came to include the group 2656: 721:', although achieving it a different way. The body is often protected by two or more rows of armored plates. Many crurotarsans reached lengths of three meters or more. 6961: 852:, traditionally included the same archosaurs as Pseudosuchia, but as a node-based clade it is not synonymous. The scope of Crurotarsi has recently been changed by the 713:; the snout is narrow and tends to be elongated, the neck is short and strong, and the limb posture ranges from a typical reptilian sprawl to an erect stance like 604:
dinosaur-like bipeds — to be typical pseudosuchians. These small forms were assumed to be the ancestors of all later archosaurs. The name Pseudosuchia became a
516:(evolutionary grouping) of archosaurs encompassing most reptiles previously identified as pseudosuchians. By this time, Pseudosuchia had also been defined as a 2939:; Julia B. Desojo; Max C. Langer. 2010. The higher-level phylogeny of Archosauria (Tetrapoda: Diapsida). Journal of Systematic Palaeontology, 8: 1, 3 — 47pp. 2878:"Rapid Initial Morphospace Expansion and Delayed Morphological Disparity Peak in the First 100 Million Years of the Archosauromorph Evolutionary Radiation" 3042: 527:
proposed that Crurotarsi, as it was then defined, must include not only crocodilian-line archosaurs, but all other archosaurs including birds, non-avian
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and all of its descendants. Benton and Clark also named a group called Crocodylotarsi, which includes most taxa now considered pseudosuchians.
2985: 800:), the latter being the ancestors of modern-day crocodiles. A study published in 2010 postulates that there is significant evidence that 395:. As a result, Crurotarsi could be a much broader clade than Pseudosuchia. Other recent studies support a more traditional phylogeny. 5938: 5915: 5334: 2817:"Compound-specific carbon isotopes from Earth's largest flood basalt eruptions directly linked to the end-Triassic mass extinction" 476:
periods, although only a single subset of crocodylomorphs, the Crocodilia, survive to the present day. Living crocodilians include
820:, the dinosaurs became extinct, with the exception of the birds, while the crocodilians continued with little change. Today, the 6798: 5930: 681:
to supplant Pseudosuchia. However, Sereno defined Crurotarsi as a node-based clade, relying on the inclusion of groups such as
588:. Zittel's aetosaurs were placed in their own suborder, Aetosauria. Colbert considered small lightly built archosaurs, such as 566: 3035: 4400: 2815:
Whiteside, Jessica H.; Olsen, Paul E.; Eglinton, Timothy; Brookfield, Michael E.; Sambrotto, Raymond N. (22 March 2010).
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Butler, Richard J.; Brusatte, Stephen L.; Reich, Mike; Nesbitt, Sterling J.; Schoch, Rainer R.; Hornung, Jahn J. (2011).
5923: 1765: 987: 576: 3006: 161: 124: 620:, two enigmatic Triassic reptiles that bear little resemblance to archosaurs, have been regarded as pseudosuchians. 6784: 4746: 557:) that were superficially crocodilian-like, but were not actually crocodilian. Hence the name "false crocodiles". 4859: 4720: 3028: 581: 804:
changed the climate, causing a mass extinction that wiped out the dinosaurs' main competitors. This allowed the
5319: 4319: 4210: 3744: 3694: 647:. Avemetatarsalians are bird-line archosaurs, including pterosaurs and dinosaurs (the latter including birds). 4387: 468:, survived the major extinction. Crocodylomorphs themselves evolved a diverse array of lifestyles during the 6956: 6851: 6222: 3643: 383:, was not closely related to other traditional "crurotarsans", at least compared to "bird-line archosaurs" ( 4394: 3014: 2542:
from the latest Early Triassic of Germany and the timing and biogeography of the early archosaur radiation"
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were small, light-footed predators. A few groups acquired herbivorous diets, such as the heavily armored
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Senter, P. (2005). "Phylogenetic taxonomy and the names of the major archosaurian (Reptilia) clades".
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niches. Their heyday was the Late Triassic, during which time their ranks included erect-limbed
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into which all thecodonts that did not fit in the other three suborders could be placed. Even
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was originally given to a group of superficially crocodile-like prehistoric reptiles from the
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Gauthier, J. (1986). "Saurischian monophyly and the origin of birds". In Padian, K. (ed.).
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modified from Nesbitt (2011) showing the new changes (bold terminal taxa are collapsed).
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Brochu, C.A. (1997). "Synonymy, redundancy, and the name of the crocodile stem-group".
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period, but fell out of use in the late 20th century, especially after the name
430: 143: 82: 2705: 2689: 630:(1985) and Gauthier (1986) became the first to establish the name Pseudosuchia in a 6831: 6671: 6609: 6469: 6455: 6357: 6320: 6290: 6183: 6174: 6102: 5840: 5832: 5796: 5780: 5772: 5763: 5749: 5709: 5688: 5646: 5596: 5505: 5486: 5458: 5275: 5182: 5148: 5141: 5134: 5095: 4998: 4990: 4669: 4510: 4468: 4458: 4438: 4118: 4044: 3939: 3480: 3373: 3299: 3228: 2801: 2389: 2295: 2236: 2183: 2156: 1951: 1671: 1481: 1454: 1358: 1284: 853: 789: 778: 774: 710: 690: 663: 640: 631: 627: 450: 426: 421: 415: 138: 3017:
archives, for comments critical of applying "Pseudosuchia" in a cladistic context.
2922: 2706:"Deep faunistic turnovers preceded the rise of dinosaurs in southwestern Pangaea" 2652:"The early evolution of archosaurs: Relationships and the origin of major clades" 2566: 6900: 6836: 6705: 6650: 6520: 6477: 6462: 6431: 6424: 6401: 6385: 6378: 6350: 6031: 5803: 5729: 5698: 5666: 5656: 5586: 5555: 5548: 5494: 5431: 5382: 5261: 5118: 5084: 5074: 5052: 5012: 4956: 4674: 4653: 4623: 4603: 4583: 4573: 4563: 4553: 4533: 4488: 4108: 4067: 4051: 4008: 3918: 3896: 3812: 3658: 3587: 3517: 3459: 2332: 1833: 1779: 1390: 1072: 1053: 682: 674: 616: 585: 454: 445: 409: 336: 332: 263: 133: 57: 6891: 2369: 1427: 6841: 6826: 6816: 6751: 6722: 6693: 6530: 6123: 5847: 5818: 5788: 5636: 5616: 5521: 5416: 4817: 4792: 4593: 4523: 4354: 4227: 4185: 4171: 4148: 4098: 4022: 3978: 3971: 3911: 3843: 3832: 3822: 3701: 3524: 3510: 3473: 3359: 3324: 3249: 3124: 3081: 2944: 2902: 2877: 2729: 2094: 1933: 1877: 1814: 1730: 1674: 1243: 1034: 1014: 950: 849: 829: 762: 678: 509: 489: 473: 439: 388: 352: 148: 102: 67: 17: 2747: 6734: 6664: 6062: 4966: 4801: 4634: 4498: 4263: 4240: 4088: 4030: 3947: 3618: 3570: 3396: 3003: 2843: 2046: 1223: 858: 821: 766: 758: 746: 706: 532: 481: 477: 392: 380: 216: 173: 107: 51: 2862: 2755: 2585: 1703: 923: 2241: 2189: 460:
Many of these Triassic pseudosuchian groups went extinct at or before the
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to succeed them as the dominant terrestrial carnivores and herbivores.
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caused the extinction of all the pseudosuchians with the exception of
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Foth, Christian; Sookias, Roland B.; Ezcurra, Martín D. (2021).
2821: 709:. The skull is often massively built, especially in contrast to 705:
Pseudosuchia is one of the two primary "daughter" clades of the
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as true crocodilians are now defined as a subset of the group.
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definition: "all taxa the least inclusive clade containing
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The Dinosaur Book: The ruling reptiles and their relatives
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Origin, decline, and re-use of the clade name Pseudosuchia
2613:. Vol. 1. Oxford: Clarendon Press. pp. 295–338. 457:
were both bipedal and herbivorous, with toothless beaks.
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Colbert, Edwin Harris; Knight, Charles Robert (1951).
757:(late Middle Triassic) they dominated the terrestrial 6702: 6593: 6577: 6567: 6556: 6546: 6536: 6526: 6516: 6506: 6443: 6412: 6331: 6316: 6306: 6296: 6286: 6276: 6266: 6256: 6208: 6171: 6134: 6119: 6108: 6098: 6088: 6078: 6068: 6058: 6048: 6038: 6027: 6017: 6007: 5997: 5987: 5760: 5745: 5735: 5725: 5715: 5705: 5694: 5684: 5673: 5662: 5652: 5642: 5632: 5622: 5612: 5602: 5592: 5582: 5393: 5378: 5234: 5166: 5106: 5091: 5080: 5070: 5059: 5048: 4978: 4962: 4952: 4942: 4619: 4609: 4599: 4589: 4579: 4569: 4559: 4549: 4539: 4529: 4519: 4494: 4484: 4474: 4464: 4454: 4444: 4434: 4293: 4282: 4237: 4196: 4159: 4144: 4134: 4124: 4114: 4104: 4094: 4084: 3989: 3959: 3884: 3869: 3859: 3849: 3839: 3828: 3818: 3808: 3681: 3655: 3629: 3614: 3604: 3593: 3583: 3321: 3305: 3295: 3285: 3275: 3265: 3255: 3245: 3234: 3224: 3214: 2385: 2360: 2328: 2291: 2232: 2212: 2179: 2152: 2108: 2062: 2037: 1947: 1910: 1873: 1829: 1758: 1726: 1694: 1600:(crocodilian‑lineage of archosaurs)  1477: 1450: 1418: 1386: 1354: 1317: 1280: 1239: 1214: 1182: 1157: 1120: 1049: 1010: 978: 946: 914: 882: 269: 260: 250: 6875: 6733: 6701: 6692: 6640: 6592: 6491: 6442: 6411: 6330: 6237: 6207: 6170: 6133: 5968: 5881: 5759: 5569: 5531: 5504: 5441: 5392: 5363: 5285: 5233: 5206: 5165: 5105: 5033: 4977: 4923: 4870: 4843: 4816: 4662: 4633: 4508: 4427: 4336: 4307: 4271: 4262: 4236: 4225: 4195: 4158: 4065: 3988: 3958: 3883: 3789: 3711: 3680: 3654: 3628: 3568: 3449: 3421: 3349: 3319: 3199: 3150: 3123: 832:are the surviving representatives of this lineage. 2657:Bulletin of the American Museum of Natural History 600:— both of which were at the time reconstructed as 371:(Laurenti, 1768)." However, a major 2011 study of 331:, "crocodile") is one of two major divisions of 2963:The Origin of Birds and the Evolution of Flight 2921:. International Archaeopteryx Conference 1984. 781:crocodilians, and a few other assorted groups. 425:. Ornithosuchids were large scavengers, while 4747: 3036: 2925:: Freunde des Jura-Museums. pp. 185–197. 2611:Phylogeny and Classification of the Tetrapods 520:, but it was not widely embraced until 2011. 8: 6863: 6698: 6601: 6499: 6249: 6245: 5980: 5976: 5965: 5952: 5575: 5401: 5371: 5360: 5347: 5174: 5041: 4935: 4931: 4920: 4907: 4849: 4822: 4813: 4770: 4754: 4740: 4732: 4424: 4413: 4268: 4233: 4077: 4073: 3801: 3797: 3786: 3773: 3576: 3330: 3207: 3196: 3183: 3129: 3120: 3059: 3043: 3029: 3021: 351:Pseudosuchia is potentially equivalent to 123: 31: 2901: 2852: 2842: 2737: 2679: 2669: 2630:. New York, NY: McGraw-Hill. p. 153. 2575: 2565: 403:period. They included giant, quadrupedal 745:Pseudosuchians appeared during the late 650:A different definition was suggested by 547:in 1887–1890 to include three taxa (two 355:even though the latter has a different, 6962:Extant Early Triassic first appearances 2775: 2773: 2645: 2643: 2641: 2639: 2637: 2528: 2597: 2595: 580:, Pseudosuchia constitutes one of the 512:was established in 1990 to label the 7: 560:In mid-20th century textbooks, like 543:The name Pseudosuchia was coined by 437:, and several were bipedal, such as 2353: 2321: 2284: 2205: 2172: 2145: 2138: 2101: 2030: 2023: 1940: 1903: 1866: 1859: 1822: 1786: 1751: 1719: 1687: 1680: 1470: 1443: 1411: 1379: 1347: 1310: 1273: 1207: 1150: 1113: 1079: 1042: 1003: 971: 939: 907: 875: 864: 643:of another branch-based clade, the 3004:Re: Even more last papers for 2005 2782:Journal of Vertebrate Paleontology 462:Triassic–Jurassic extinction event 375:archosaur relations proposed that 25: 733:Mounted skeleton of the Triassic 6847: 6846: 6797: 6790: 6783: 5937: 5929: 5922: 5914: 5333: 5326: 5318: 4716: 4715: 4399: 4393: 4386: 3758: 3750: 3743: 2392: 2368: 2336: 2299: 2240: 2188: 2116: 2070: 2045: 1978: 1955: 1918: 1881: 1837: 1798: 1764: 1734: 1702: 1508: 1485: 1426: 1394: 1362: 1325: 1288: 1247: 1222: 1190: 1165: 1128: 1092: 1057: 1018: 986: 954: 922: 890: 160: 2609:. In Benton, Michael J. (ed.). 2802:10.1080/02724634.1997.10010992 2710:Nature Ecology & Evolution 1: 3013:and follow up posts, on the 2986:Stem Archosauria—TaxonSearch 2567:10.1371/journal.pone.0025693 1091:(bird-lineage of archosaurs) 577:Evolution of the Vertebrates 2999:Taxon Search – Pseudosuchia 584:of the now-abandoned order 6978: 2882:Frontiers in Earth Science 2605:; Clark, James M. (1988). 464:. However, one group, the 6811: 6780: 6604: 6502: 6252: 6248: 5983: 5979: 5964: 5951: 5911: 5578: 5404: 5374: 5359: 5346: 5315: 5177: 5044: 4938: 4934: 4919: 4906: 4852: 4825: 4812: 4769: 4711: 4423: 4412: 4383: 4080: 4076: 3804: 3800: 3785: 3772: 3741: 3579: 3333: 3210: 3195: 3182: 3132: 3119: 3058: 2945:10.1080/14772010903537732 2903:10.3389/feart.2021.723973 2730:10.1038/s41559-017-0305-5 2538:"The sail-backed reptile 2383: 2358: 2351: 2326: 2319: 2289: 2282: 2230: 2210: 2203: 2177: 2170: 2150: 2143: 2136: 2106: 2099: 2060: 2035: 2028: 2021: 1970: 1945: 1938: 1908: 1901: 1871: 1864: 1857: 1827: 1820: 1791: 1784: 1756: 1749: 1724: 1717: 1692: 1685: 1500: 1475: 1468: 1448: 1441: 1416: 1409: 1384: 1377: 1352: 1345: 1315: 1308: 1278: 1271: 1237: 1212: 1205: 1180: 1155: 1148: 1118: 1111: 1084: 1077: 1047: 1040: 1008: 1001: 976: 969: 944: 937: 912: 905: 880: 873: 818:end-Cretaceous extinction 325: 314: 292: 285: 247: 242: 157:Scientific classification 155: 131: 122: 34: 297:Benton & Clark, 1988 2919:The Beginnings of Birds 2844:10.1073/pnas.1001706107 786:end-Triassic extinction 634:context, using it as a 567:Vertebrate Paleontology 523:In 2011 paleontologist 5410:Benggwigwishingasuchus 2650:Nesbitt, S.J. (2011). 773:, the large predatory 742: 545:Karl Alfred von Zittel 6929:Paleobiology Database 3015:Dinosaur Mailing List 3009:12 April 2016 at the 2935:Stephen L. Brusatte; 732: 654:and Clark, 1988: the 562:Alfred Sherwood Romer 362:Rutiodon carolinensis 2984:Sereno, P. C. 2005. 660:last common ancestor 368:Crocodylus niloticus 365:(Emmons, 1856), and 5366:Paracrocodylomorpha 5353:Paracrocodylomorpha 5287:Paracrocodylomorpha 2894:2021FrEaS...9..763F 2835:2010PNAS..107.6721W 2794:1997JVPal..17..448B 2722:2017NatEE...1.1477E 2558:2011PLoSO...625693B 1897:Paracrocodylomorpha 1304:Paracrocodylomorpha 739:Saurosuchus galilei 335:, including living 6083:Pseudhesperosuchus 4685:Ichthyosauromorpha 4675:Drepanosauromorpha 2603:Benton, Michael J. 802:volcanic eruptions 777:, the small agile 743: 685:, Aetosauria, and 677:erected the clade 636:branch-based taxon 6944: 6943: 6869:Taxon identifiers 6860: 6859: 6807: 6806: 6778: 6777: 6774: 6773: 6770: 6769: 6766: 6765: 6762: 6761: 6688: 6687: 6597:Shartegosuchoidea 6561:Pachycheilosuchus 6487: 6486: 6233: 6232: 6073:Phyllodontosuchus 5947: 5946: 5909: 5908: 5905: 5904: 5901: 5900: 5565: 5564: 5342: 5341: 5313: 5312: 5309: 5308: 5305: 5304: 5229: 5228: 5029: 5028: 4902: 4901: 4898: 4897: 4894: 4893: 4890: 4889: 4729: 4728: 4707: 4706: 4703: 4702: 4408: 4407: 4381: 4380: 4377: 4376: 4373: 4372: 4369: 4368: 4365: 4364: 4221: 4220: 4061: 4060: 3768: 3767: 3739: 3738: 3735: 3734: 3731: 3730: 3564: 3563: 3554:Sharovipterygidae 3423:Sharovipterygidae 3367:Dinocephalosaurus 3178: 3177: 3174: 3173: 3170: 3169: 2937:Michael J. Benton 2716:(10): 1477–1483. 2519: 2518: 2510: 2509: 2501: 2500: 2492: 2491: 2483: 2482: 2469: 2468: 2460: 2459: 2451: 2450: 2442: 2441: 2433: 2432: 2424: 2423: 2415: 2414: 2406: 2405: 2271: 2270: 2262: 2261: 2253: 2252: 2083: 2082: 2010: 2009: 2001: 2000: 1992: 1991: 1762:Proterochampsidae 1667: 1666: 1658: 1657: 1649: 1648: 1640: 1639: 1631: 1630: 1622: 1621: 1613: 1612: 1604: 1603: 1594: 1593: 1585: 1584: 1576: 1575: 1567: 1566: 1558: 1557: 1549: 1548: 1540: 1539: 1531: 1530: 1522: 1521: 1260: 1259: 606:wastebasket taxon 539:Taxonomic history 385:Avemetatarsalians 304: 303: 298: 238: 27:Clade of reptiles 16:(Redirected from 6969: 6937: 6936: 6924: 6923: 6911: 6910: 6909: 6896: 6895: 6894: 6864: 6850: 6849: 6802: 6801: 6794: 6788: 6787: 6704: 6699: 6642:Shartegosuchidae 6602: 6595: 6579: 6569: 6558: 6548: 6538: 6528: 6518: 6511:Dianchungosaurus 6508: 6500: 6494:Mesoeucrocodylia 6445: 6414: 6333: 6318: 6308: 6298: 6288: 6278: 6271:Eopneumatosuchus 6268: 6258: 6250: 6246: 6210: 6173: 6136: 6121: 6110: 6100: 6090: 6080: 6070: 6060: 6050: 6040: 6029: 6019: 6009: 5999: 5989: 5981: 5977: 5966: 5953: 5942: 5941: 5934: 5933: 5927: 5926: 5919: 5918: 5762: 5747: 5737: 5727: 5717: 5707: 5696: 5686: 5675: 5664: 5654: 5644: 5634: 5624: 5614: 5604: 5594: 5584: 5576: 5443:Ctenosauriscidae 5402: 5395: 5380: 5372: 5361: 5348: 5337: 5331: 5330: 5323: 5322: 5236: 5175: 5168: 5108: 5093: 5082: 5072: 5061: 5050: 5042: 4980: 4964: 4954: 4944: 4936: 4932: 4921: 4908: 4860:Archosauromorpha 4850: 4845:Archosauromorpha 4823: 4814: 4807: 4806: 4771: 4756: 4749: 4742: 4733: 4719: 4718: 4663:Possible members 4621: 4611: 4601: 4591: 4581: 4571: 4561: 4551: 4541: 4531: 4521: 4496: 4486: 4476: 4466: 4456: 4446: 4436: 4425: 4414: 4403: 4397: 4391: 4390: 4295: 4284: 4269: 4239: 4234: 4198: 4161: 4146: 4136: 4126: 4116: 4106: 4096: 4086: 4078: 4074: 3991: 3961: 3886: 3871: 3861: 3851: 3841: 3830: 3820: 3810: 3802: 3798: 3792:Archosauriformes 3787: 3779:Archosauriformes 3774: 3763: 3762: 3755: 3754: 3747: 3713:Archosauriformes 3683: 3657: 3631: 3616: 3606: 3598:Cryptovaranoides 3595: 3585: 3577: 3504:Langobardisaurus 3390:Protanystropheus 3351:Trachelosauridae 3331: 3323: 3310:Vritramimosaurus 3307: 3297: 3287: 3277: 3270:Mecistotrachelos 3267: 3257: 3247: 3236: 3226: 3219:Aenigmastropheus 3216: 3208: 3202:Archosauromorpha 3197: 3189:Archosauromorpha 3184: 3152:Archosauromorpha 3130: 3121: 3114: 3113: 3060: 3052:Archosauromorpha 3045: 3038: 3031: 3022: 2981: 2966: 2947: 2933: 2927: 2926: 2914: 2908: 2907: 2905: 2873: 2867: 2866: 2856: 2846: 2812: 2806: 2805: 2777: 2768: 2767: 2741: 2700: 2694: 2693: 2683: 2673: 2647: 2632: 2631: 2621: 2615: 2614: 2599: 2590: 2589: 2579: 2569: 2533: 2396: 2387: 2372: 2362: 2354: 2340: 2330: 2322: 2303: 2293: 2285: 2244: 2234: 2214: 2206: 2192: 2181: 2173: 2154: 2146: 2139: 2120: 2110: 2102: 2074: 2064: 2049: 2039: 2031: 2024: 1983: 1982: 1959: 1949: 1941: 1922: 1912: 1904: 1885: 1875: 1867: 1860: 1841: 1831: 1823: 1802: 1787: 1768: 1760: 1752: 1738: 1728: 1720: 1706: 1696: 1688: 1681: 1513: 1512: 1489: 1479: 1471: 1452: 1444: 1430: 1420: 1412: 1398: 1388: 1380: 1366: 1356: 1348: 1329: 1319: 1311: 1292: 1282: 1274: 1251: 1241: 1226: 1216: 1208: 1194: 1184: 1169: 1159: 1151: 1132: 1122: 1114: 1096: 1080: 1061: 1051: 1043: 1022: 1012: 1004: 990: 980: 972: 958: 948: 940: 926: 916: 908: 894: 884: 876: 869:Archosauriformes 865: 695:Sterling Nesbitt 656:node-based taxon 572:Edwin H. Colbert 525:Sterling Nesbitt 330: 328: 319: 317: 296: 271: 262: 252: 233: 165: 164: 127: 117: 54: 38:Temporal range: 32: 21: 6977: 6976: 6972: 6971: 6970: 6968: 6967: 6966: 6947: 6946: 6945: 6940: 6932: 6927: 6919: 6914: 6905: 6904: 6899: 6890: 6889: 6884: 6871: 6861: 6856: 6803: 6796: 6782: 6758: 6729: 6684: 6636: 6617:Shantungosuchus 6588: 6496: 6483: 6438: 6407: 6372:Hemiprotosuchus 6326: 6242: 6240:Crocodyliformes 6229: 6203: 6166: 6161:Terrestrisuchus 6138:Saltoposuchidae 6129: 6113:Syntomiprosopus 5973: 5971:Crocodylomorpha 5960: 5958:Crocodylomorpha 5943: 5936: 5928: 5921: 5913: 5897: 5883:Crocodylomorpha 5877: 5871:Vytshegdosuchus 5863:Vjushkovisaurus 5755: 5561: 5527: 5515:Dolichobrachium 5500: 5437: 5388: 5368: 5355: 5338: 5325: 5317: 5301: 5281: 5238:Gracilisuchidae 5225: 5202: 5170:Aetosauriformes 5161: 5101: 5038: 5025: 4982:Ornithosuchidae 4973: 4928: 4915: 4886: 4866: 4839: 4808: 4776: 4775: 4765: 4760: 4730: 4725: 4699: 4680:Kuehneosauridae 4658: 4629: 4614:Vjushkovisaurus 4504: 4479:Protecovasaurus 4419: 4404: 4385: 4361: 4332: 4320:Avemetatarsalia 4309:Avemetatarsalia 4303: 4258: 4230: 4217: 4211:Proterochampsia 4200:Proterochampsia 4191: 4154: 4070: 4057: 4002:Bharitalasuchus 3993:Erythrosuchidae 3984: 3954: 3888:Proterosuchidae 3879: 3874:Syntomiprosopus 3794: 3781: 3764: 3757: 3749: 3727: 3707: 3676: 3650: 3624: 3573: 3560: 3532:Sclerostropheus 3467:Augustaburiania 3451:Tanystropheidae 3445: 3417: 3404:Sclerostropheus 3345: 3327: 3315: 3239:Elachistosuchus 3204: 3191: 3166: 3146: 3115: 3065: 3064: 3054: 3049: 3011:Wayback Machine 2995: 2990: 2969: 2960: 2956: 2951: 2950: 2934: 2930: 2916: 2915: 2911: 2875: 2874: 2870: 2814: 2813: 2809: 2779: 2778: 2771: 2702: 2701: 2697: 2649: 2648: 2635: 2623: 2622: 2618: 2601: 2600: 2593: 2535: 2534: 2530: 2525: 2520: 2511: 2502: 2493: 2484: 2470: 2461: 2452: 2443: 2434: 2425: 2416: 2407: 2272: 2263: 2254: 2084: 2067:Ornithosuchidae 2011: 2002: 1993: 1977: 1974:Crocodylomorpha 1795:Avemetatarsalia 1668: 1659: 1650: 1641: 1632: 1623: 1614: 1605: 1595: 1586: 1577: 1568: 1559: 1550: 1541: 1532: 1523: 1507: 1504:Crocodylomorpha 1261: 1125:Ornithosuchidae 1088:Avemetatarsalia 983:Proterochampsia 887:Proterosuchidae 841: 798:Crocodylomorpha 794:Crocodyliformes 727: 703: 687:Crocodylomorpha 645:Avemetatarsalia 541: 498: 466:crocodylomorphs 449:. The bizarre, 326: 315: 295:Crocodylotarsi 273:Ornithosuchidae 232: 159: 118: 116: 115: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 49: 48: 39: 36: 28: 23: 22: 15: 12: 11: 5: 6975: 6973: 6965: 6964: 6959: 6957:Pseudosuchians 6949: 6948: 6942: 6941: 6939: 6938: 6925: 6912: 6897: 6881: 6879: 6873: 6872: 6867: 6858: 6857: 6855: 6854: 6844: 6839: 6834: 6829: 6824: 6822:Prestosuchidae 6819: 6812: 6809: 6808: 6805: 6804: 6781: 6779: 6776: 6775: 6772: 6771: 6768: 6767: 6764: 6763: 6760: 6759: 6757: 6756: 6755: 6754: 6748: 6739: 6737: 6731: 6730: 6728: 6727: 6726: 6725: 6719: 6710: 6708: 6696: 6690: 6689: 6686: 6685: 6683: 6682: 6679:Shartegosuchus 6675: 6668: 6661: 6654: 6646: 6644: 6638: 6637: 6635: 6634: 6627: 6624:Sichuanosuchus 6620: 6613: 6605: 6599: 6590: 6589: 6587: 6586: 6582:Thalattosuchia 6575: 6565: 6554: 6544: 6541:Lusitanisuchus 6534: 6524: 6514: 6503: 6497: 6492: 6489: 6488: 6485: 6484: 6482: 6481: 6474: 6466: 6459: 6451: 6449: 6440: 6439: 6437: 6436: 6428: 6420: 6418: 6416:Notochampsidae 6409: 6408: 6406: 6405: 6398: 6394:Pedeticosaurus 6390: 6382: 6375: 6368: 6365:Erythrochampsa 6361: 6354: 6347: 6344:Coloradisuchus 6339: 6337: 6328: 6327: 6325: 6324: 6314: 6304: 6294: 6284: 6274: 6264: 6253: 6243: 6238: 6235: 6234: 6231: 6230: 6228: 6227: 6226: 6225: 6223:Thalattosuchia 6216: 6214: 6212:Thalattosuchia 6205: 6204: 6202: 6201: 6194: 6187: 6179: 6177: 6168: 6167: 6165: 6164: 6157: 6150: 6142: 6140: 6131: 6130: 6128: 6127: 6117: 6106: 6096: 6093:Redondavenator 6086: 6076: 6066: 6056: 6046: 6036: 6025: 6015: 6005: 6002:Dibothrosuchus 5995: 5984: 5974: 5969: 5962: 5961: 5956: 5949: 5948: 5945: 5944: 5912: 5910: 5907: 5906: 5903: 5902: 5899: 5898: 5896: 5895: 5894: 5893: 5887: 5885: 5879: 5878: 5876: 5875: 5867: 5859: 5852: 5844: 5837: 5829: 5822: 5815: 5807: 5800: 5793: 5785: 5777: 5768: 5766: 5757: 5756: 5754: 5753: 5743: 5733: 5723: 5713: 5703: 5692: 5682: 5678:Incertovenator 5671: 5660: 5650: 5640: 5630: 5620: 5610: 5600: 5590: 5579: 5573: 5567: 5566: 5563: 5562: 5560: 5559: 5552: 5545: 5537: 5535: 5529: 5528: 5526: 5525: 5518: 5510: 5508: 5502: 5501: 5499: 5498: 5491: 5483: 5480:Hypselorhachis 5476: 5473:Ctenosauriscus 5469: 5466:Bystrowisuchus 5462: 5455: 5447: 5445: 5439: 5438: 5436: 5435: 5428: 5420: 5413: 5405: 5399: 5390: 5389: 5387: 5386: 5375: 5369: 5364: 5357: 5356: 5351: 5344: 5343: 5340: 5339: 5316: 5314: 5311: 5310: 5307: 5306: 5303: 5302: 5300: 5299: 5298: 5297: 5291: 5289: 5283: 5282: 5280: 5279: 5272: 5265: 5258: 5250: 5242: 5240: 5231: 5230: 5227: 5226: 5224: 5223: 5222: 5221: 5212: 5210: 5204: 5203: 5201: 5200: 5197:Revueltosaurus 5193: 5186: 5178: 5172: 5163: 5162: 5160: 5159: 5152: 5145: 5138: 5131: 5123: 5114: 5112: 5110:Erpetosuchidae 5103: 5102: 5100: 5099: 5089: 5078: 5068: 5057: 5045: 5039: 5034: 5031: 5030: 5027: 5026: 5024: 5023: 5020:Venaticosuchus 5016: 5009: 5002: 4995: 4986: 4984: 4975: 4974: 4972: 4971: 4960: 4950: 4947:Dasygnathoides 4939: 4929: 4924: 4917: 4916: 4911: 4904: 4903: 4900: 4899: 4896: 4895: 4892: 4891: 4888: 4887: 4885: 4884: 4883: 4882: 4876: 4874: 4868: 4867: 4865: 4864: 4863: 4862: 4853: 4847: 4841: 4840: 4838: 4837: 4836: 4835: 4826: 4820: 4810: 4809: 4805: 4804: 4795: 4789: 4783: 4774: 4767: 4766: 4761: 4759: 4758: 4751: 4744: 4736: 4727: 4726: 4724: 4723: 4712: 4709: 4708: 4705: 4704: 4701: 4700: 4698: 4697: 4695:Thalattosauria 4692: 4687: 4682: 4677: 4672: 4666: 4664: 4660: 4659: 4657: 4656: 4651: 4646: 4640: 4638: 4631: 4630: 4628: 4627: 4617: 4607: 4597: 4587: 4577: 4567: 4557: 4547: 4537: 4527: 4516: 4514: 4506: 4505: 4503: 4502: 4492: 4482: 4472: 4462: 4452: 4442: 4431: 4429: 4421: 4420: 4418:Related topics 4417: 4410: 4409: 4406: 4405: 4384: 4382: 4379: 4378: 4375: 4374: 4371: 4370: 4367: 4366: 4363: 4362: 4360: 4359: 4358: 4357: 4351: 4342: 4340: 4334: 4333: 4331: 4330: 4329: 4328: 4322: 4313: 4311: 4305: 4304: 4302: 4301: 4291: 4279: 4277: 4274:Incertae sedis 4266: 4260: 4259: 4257: 4256: 4255: 4254: 4245: 4243: 4231: 4226: 4223: 4222: 4219: 4218: 4216: 4215: 4214: 4213: 4204: 4202: 4193: 4192: 4190: 4189: 4182: 4179:Halazhaisuchus 4175: 4167: 4165: 4156: 4155: 4153: 4152: 4142: 4132: 4129:Marcianosuchus 4122: 4112: 4102: 4092: 4081: 4071: 4066: 4063: 4062: 4059: 4058: 4056: 4055: 4048: 4041: 4038:Guchengosuchus 4034: 4027: 4019: 4012: 4005: 3997: 3995: 3986: 3985: 3983: 3982: 3975: 3967: 3965: 3956: 3955: 3953: 3952: 3944: 3936: 3929: 3922: 3915: 3908: 3904:Chasmatosuchus 3900: 3892: 3890: 3881: 3880: 3878: 3877: 3867: 3857: 3847: 3837: 3826: 3816: 3805: 3795: 3790: 3783: 3782: 3777: 3770: 3769: 3766: 3765: 3742: 3740: 3737: 3736: 3733: 3732: 3729: 3728: 3726: 3725: 3724: 3723: 3717: 3715: 3709: 3708: 3706: 3705: 3698: 3690: 3688: 3678: 3677: 3675: 3674: 3673: 3672: 3663: 3661: 3652: 3651: 3649: 3648: 3647: 3646: 3644:Allokotosauria 3637: 3635: 3633:Allokotosauria 3626: 3625: 3623: 3622: 3612: 3609:Tasmaniosaurus 3602: 3591: 3580: 3574: 3569: 3566: 3565: 3562: 3561: 3559: 3558: 3550: 3543: 3536: 3528: 3521: 3514: 3507: 3500: 3497:Gwyneddosaurus 3493: 3485: 3477: 3470: 3463: 3455: 3453: 3447: 3446: 3444: 3443: 3436: 3428: 3426: 3419: 3418: 3416: 3415: 3412:Trachelosaurus 3408: 3400: 3393: 3386: 3378: 3370: 3363: 3355: 3353: 3347: 3346: 3344: 3343: 3334: 3328: 3320: 3317: 3316: 3314: 3313: 3303: 3293: 3283: 3280:Prolacertoides 3273: 3263: 3260:Malutinisuchus 3253: 3243: 3232: 3222: 3211: 3205: 3200: 3193: 3192: 3187: 3180: 3179: 3176: 3175: 3172: 3171: 3168: 3167: 3165: 3164: 3163: 3162: 3156: 3154: 3148: 3147: 3145: 3144: 3143: 3142: 3133: 3127: 3117: 3116: 3112: 3111: 3102: 3093: 3084: 3078: 3072: 3063: 3056: 3055: 3050: 3048: 3047: 3040: 3033: 3025: 3019: 3018: 3001: 2994: 2993:External links 2991: 2989: 2988: 2982: 2967: 2957: 2955: 2952: 2949: 2948: 2928: 2909: 2868: 2829:(15): 6721–5. 2807: 2788:(2): 448–449. 2769: 2695: 2633: 2616: 2591: 2552:(10): e25693. 2540:Ctenosauriscus 2527: 2526: 2524: 2521: 2517: 2516: 2513: 2512: 2508: 2507: 2504: 2503: 2499: 2498: 2495: 2494: 2490: 2489: 2486: 2485: 2481: 2480: 2472: 2471: 2467: 2466: 2463: 2462: 2458: 2457: 2454: 2453: 2449: 2448: 2445: 2444: 2440: 2439: 2436: 2435: 2431: 2430: 2427: 2426: 2422: 2421: 2418: 2417: 2413: 2412: 2409: 2408: 2404: 2403: 2400: 2399: 2382: 2379: 2378: 2375: 2374: 2357: 2352: 2350: 2347: 2346: 2343: 2342: 2325: 2320: 2318: 2315:Prestosuchidae 2310: 2309: 2306: 2305: 2288: 2283: 2281: 2278: 2277: 2274: 2273: 2269: 2268: 2265: 2264: 2260: 2259: 2256: 2255: 2251: 2250: 2247: 2246: 2229: 2226: 2225: 2222: 2221: 2209: 2204: 2202: 2199: 2198: 2195: 2194: 2176: 2171: 2169: 2166: 2165: 2162: 2161: 2149: 2144: 2142: 2137: 2135: 2127: 2126: 2123: 2122: 2105: 2100: 2098: 2090: 2089: 2086: 2085: 2081: 2080: 2077: 2076: 2059: 2056: 2055: 2052: 2051: 2042:Revueltosaurus 2034: 2029: 2027: 2022: 2020: 2017: 2016: 2013: 2012: 2008: 2007: 2004: 2003: 1999: 1998: 1995: 1994: 1990: 1989: 1986: 1985: 1969: 1966: 1965: 1962: 1961: 1944: 1939: 1937: 1929: 1928: 1925: 1924: 1907: 1902: 1900: 1892: 1891: 1888: 1887: 1870: 1865: 1863: 1858: 1856: 1848: 1847: 1844: 1843: 1826: 1821: 1819: 1809: 1808: 1805: 1804: 1790: 1785: 1783: 1775: 1774: 1771: 1770: 1755: 1750: 1748: 1745: 1744: 1741: 1740: 1723: 1718: 1716: 1713: 1712: 1709: 1708: 1691: 1686: 1684: 1679: 1665: 1664: 1661: 1660: 1656: 1655: 1652: 1651: 1647: 1646: 1643: 1642: 1638: 1637: 1634: 1633: 1629: 1628: 1625: 1624: 1620: 1619: 1616: 1615: 1611: 1610: 1607: 1606: 1602: 1601: 1597: 1596: 1592: 1591: 1588: 1587: 1583: 1582: 1579: 1578: 1574: 1573: 1570: 1569: 1565: 1564: 1561: 1560: 1556: 1555: 1552: 1551: 1547: 1546: 1543: 1542: 1538: 1537: 1534: 1533: 1529: 1528: 1525: 1524: 1520: 1519: 1516: 1515: 1499: 1496: 1495: 1492: 1491: 1474: 1469: 1467: 1464: 1463: 1460: 1459: 1447: 1442: 1440: 1437: 1436: 1433: 1432: 1415: 1410: 1408: 1405: 1404: 1401: 1400: 1383: 1378: 1376: 1373: 1372: 1369: 1368: 1351: 1346: 1344: 1336: 1335: 1332: 1331: 1314: 1309: 1307: 1299: 1298: 1295: 1294: 1277: 1272: 1270: 1267: 1266: 1263: 1262: 1258: 1257: 1254: 1253: 1236: 1233: 1232: 1229: 1228: 1219:Revueltosaurus 1211: 1206: 1204: 1201: 1200: 1197: 1196: 1179: 1176: 1175: 1172: 1171: 1154: 1149: 1147: 1139: 1138: 1135: 1134: 1117: 1112: 1110: 1103: 1102: 1099: 1098: 1083: 1078: 1076: 1068: 1067: 1064: 1063: 1046: 1041: 1039: 1029: 1028: 1025: 1024: 1007: 1002: 1000: 997: 996: 993: 992: 975: 970: 968: 965: 964: 961: 960: 943: 938: 936: 933: 932: 929: 928: 911: 906: 904: 901: 900: 897: 896: 879: 874: 872: 863: 840: 837: 726: 723: 702: 699: 658:including the 540: 537: 497: 494: 431:gracilisuchids 405:apex predators 320:, "false" and 302: 301: 300: 299: 290: 289: 283: 282: 281: 280: 275: 267: 258: 255:Dasygnathoides 245: 244: 240: 239: 227: 220: 219: 214: 207: 206: 201: 197: 196: 191: 187: 186: 181: 177: 176: 171: 167: 166: 153: 152: 129: 128: 120: 119: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 55: 41:Early Triassic 37: 35:Pseudosuchians 26: 24: 18:Crocodylotarsi 14: 13: 10: 9: 6: 4: 3: 2: 6974: 6963: 6960: 6958: 6955: 6954: 6952: 6935: 6930: 6926: 6922: 6917: 6913: 6908: 6902: 6898: 6893: 6887: 6883: 6882: 6880: 6878: 6874: 6870: 6865: 6853: 6845: 6843: 6840: 6838: 6835: 6833: 6830: 6828: 6825: 6823: 6820: 6818: 6814: 6813: 6810: 6800: 6795: 6793: 6786: 6753: 6749: 6747: 6743: 6742: 6741: 6740: 6738: 6736: 6732: 6724: 6720: 6718: 6714: 6713: 6712: 6711: 6709: 6707: 6700: 6697: 6695: 6691: 6681: 6680: 6676: 6674: 6673: 6669: 6667: 6666: 6662: 6660: 6659: 6658:Fruitachampsa 6655: 6653: 6652: 6648: 6647: 6645: 6643: 6639: 6633: 6632: 6628: 6626: 6625: 6621: 6619: 6618: 6614: 6612: 6611: 6607: 6606: 6603: 6600: 6598: 6591: 6584: 6583: 6576: 6574: 6573: 6572:Titanochampsa 6566: 6563: 6562: 6555: 6553: 6552: 6551:Neuquensuchus 6545: 6543: 6542: 6535: 6533: 6532: 6525: 6523: 6522: 6515: 6513: 6512: 6505: 6504: 6501: 6498: 6495: 6490: 6480: 6479: 6475: 6472: 6471: 6467: 6465: 6464: 6460: 6458: 6457: 6453: 6452: 6450: 6448: 6447:Gobiosuchidae 6441: 6434: 6433: 6429: 6427: 6426: 6422: 6421: 6419: 6417: 6410: 6404: 6403: 6399: 6396: 6395: 6391: 6388: 6387: 6383: 6381: 6380: 6376: 6374: 6373: 6369: 6367: 6366: 6362: 6360: 6359: 6355: 6353: 6352: 6348: 6346: 6345: 6341: 6340: 6338: 6336: 6335:Protosuchidae 6329: 6323: 6322: 6315: 6313: 6312: 6311:Stegomosuchus 6305: 6303: 6302: 6301:Platyognathus 6295: 6293: 6292: 6285: 6283: 6282: 6275: 6273: 6272: 6265: 6263: 6262: 6261:Entradasuchus 6255: 6254: 6251: 6247: 6244: 6241: 6236: 6224: 6220: 6219: 6218: 6217: 6215: 6213: 6206: 6200: 6199: 6198:Macelognathus 6195: 6193: 6192: 6188: 6186: 6185: 6181: 6180: 6178: 6176: 6169: 6163: 6162: 6158: 6156: 6155: 6154:Saltoposuchus 6151: 6149: 6148: 6147:Litargosuchus 6144: 6143: 6141: 6139: 6132: 6126: 6125: 6118: 6115: 6114: 6107: 6105: 6104: 6097: 6095: 6094: 6087: 6085: 6084: 6077: 6075: 6074: 6067: 6065: 6064: 6057: 6055: 6054: 6053:Kayentasuchus 6047: 6045: 6044: 6043:Junggarsuchus 6037: 6034: 6033: 6026: 6024: 6023: 6022:Hesperosuchus 6016: 6014: 6013: 6012:Dromicosuchus 6006: 6004: 6003: 5996: 5994: 5993: 5986: 5985: 5982: 5978: 5975: 5972: 5967: 5963: 5959: 5954: 5950: 5940: 5935: 5932: 5925: 5917: 5891: 5890: 5889: 5888: 5886: 5884: 5880: 5873: 5872: 5868: 5865: 5864: 5860: 5858: 5857: 5853: 5850: 5849: 5845: 5843: 5842: 5838: 5835: 5834: 5830: 5828: 5827: 5826:Scolotosuchus 5823: 5821: 5820: 5816: 5813: 5812: 5811:Procerosuchus 5808: 5806: 5805: 5801: 5799: 5798: 5794: 5791: 5790: 5786: 5783: 5782: 5778: 5775: 5774: 5770: 5769: 5767: 5765: 5758: 5752: 5751: 5744: 5742: 5741: 5740:Stagonosuchus 5734: 5732: 5731: 5724: 5722: 5721: 5720:Procerosuchus 5714: 5712: 5711: 5704: 5701: 5700: 5693: 5691: 5690: 5683: 5680: 5679: 5672: 5669: 5668: 5661: 5659: 5658: 5651: 5649: 5648: 5641: 5639: 5638: 5631: 5629: 5628: 5627:Decuriasuchus 5621: 5619: 5618: 5611: 5609: 5608: 5607:Batrachotomus 5601: 5599: 5598: 5591: 5589: 5588: 5581: 5580: 5577: 5574: 5572: 5568: 5558: 5557: 5553: 5551: 5550: 5546: 5544: 5543: 5539: 5538: 5536: 5534: 5533:Shuvosauridae 5530: 5524: 5523: 5519: 5517: 5516: 5512: 5511: 5509: 5507: 5503: 5497: 5496: 5492: 5489: 5488: 5484: 5482: 5481: 5477: 5475: 5474: 5470: 5468: 5467: 5463: 5461: 5460: 5456: 5454: 5453: 5452:Arizonasaurus 5449: 5448: 5446: 5444: 5440: 5434: 5433: 5429: 5426: 5425: 5424:Schultzsuchus 5421: 5419: 5418: 5414: 5412: 5411: 5407: 5406: 5403: 5400: 5398: 5397:Poposauroidea 5391: 5385: 5384: 5377: 5376: 5373: 5370: 5367: 5362: 5358: 5354: 5349: 5345: 5336: 5332: 5329: 5321: 5295: 5294: 5293: 5292: 5290: 5288: 5284: 5278: 5277: 5273: 5271: 5270: 5269:Turfanosuchus 5266: 5264: 5263: 5259: 5256: 5255: 5251: 5249: 5248: 5247:Gracilisuchus 5244: 5243: 5241: 5239: 5232: 5220: 5216: 5215: 5214: 5213: 5211: 5209: 5205: 5199: 5198: 5194: 5192: 5191: 5190:Euscolosuchus 5187: 5185: 5184: 5180: 5179: 5176: 5173: 5171: 5164: 5158: 5157: 5153: 5151: 5150: 5146: 5144: 5143: 5139: 5137: 5136: 5132: 5129: 5128: 5124: 5121: 5120: 5116: 5115: 5113: 5111: 5104: 5098: 5097: 5090: 5087: 5086: 5079: 5077: 5076: 5069: 5066: 5065: 5064:Koilamasuchus 5058: 5055: 5054: 5047: 5046: 5043: 5040: 5037: 5032: 5022: 5021: 5017: 5015: 5014: 5010: 5008: 5007: 5006:Ornithosuchus 5003: 5001: 5000: 4996: 4993: 4992: 4988: 4987: 4985: 4983: 4976: 4969: 4968: 4961: 4959: 4958: 4951: 4949: 4948: 4941: 4940: 4937: 4933: 4930: 4927: 4922: 4918: 4914: 4909: 4905: 4880: 4879: 4878: 4877: 4875: 4873: 4869: 4861: 4857: 4856: 4855: 4854: 4851: 4848: 4846: 4842: 4834: 4830: 4829: 4828: 4827: 4824: 4821: 4819: 4815: 4811: 4803: 4799: 4796: 4794: 4790: 4788: 4784: 4782: 4778: 4777: 4772: 4768: 4764: 4757: 4752: 4750: 4745: 4743: 4738: 4737: 4734: 4722: 4714: 4713: 4710: 4696: 4693: 4691: 4690:Sauropterygia 4688: 4686: 4683: 4681: 4678: 4676: 4673: 4671: 4668: 4667: 4665: 4661: 4655: 4652: 4650: 4649:Protorosauria 4647: 4645: 4644:Proterosuchia 4642: 4641: 4639: 4636: 4632: 4626: 4625: 4618: 4616: 4615: 4608: 4606: 4605: 4598: 4596: 4595: 4588: 4586: 4585: 4578: 4576: 4575: 4568: 4566: 4565: 4558: 4556: 4555: 4548: 4546: 4545: 4538: 4536: 4535: 4528: 4526: 4525: 4518: 4517: 4515: 4513: 4512: 4507: 4501: 4500: 4493: 4491: 4490: 4483: 4481: 4480: 4473: 4471: 4470: 4463: 4461: 4460: 4453: 4451: 4450: 4449:Lucianosaurus 4443: 4441: 4440: 4433: 4432: 4430: 4426: 4422: 4415: 4411: 4402: 4398: 4396: 4389: 4356: 4352: 4350: 4346: 4345: 4344: 4343: 4341: 4339: 4335: 4327: 4323: 4321: 4317: 4316: 4315: 4314: 4312: 4310: 4306: 4300: 4299: 4292: 4289: 4288: 4287:Sikannisuchus 4281: 4280: 4278: 4276: 4275: 4270: 4267: 4265: 4261: 4253: 4249: 4248: 4247: 4246: 4244: 4242: 4235: 4232: 4229: 4224: 4212: 4208: 4207: 4206: 4205: 4203: 4201: 4194: 4188: 4187: 4183: 4181: 4180: 4176: 4174: 4173: 4169: 4168: 4166: 4164: 4163:Euparkeriidae 4157: 4151: 4150: 4143: 4141: 4140: 4139:Polymorphodon 4133: 4131: 4130: 4123: 4121: 4120: 4113: 4111: 4110: 4103: 4101: 4100: 4093: 4091: 4090: 4083: 4082: 4079: 4075: 4072: 4069: 4064: 4054: 4053: 4049: 4047: 4046: 4042: 4040: 4039: 4035: 4033: 4032: 4028: 4025: 4024: 4020: 4018: 4017: 4016:Erythrosuchus 4013: 4011: 4010: 4006: 4004: 4003: 3999: 3998: 3996: 3994: 3987: 3981: 3980: 3976: 3974: 3973: 3969: 3968: 3966: 3964: 3963:Protopyknosia 3957: 3950: 3949: 3945: 3942: 3941: 3937: 3935: 3934: 3933:Samsarasuchus 3930: 3928: 3927: 3926:Proterosuchus 3923: 3921: 3920: 3916: 3914: 3913: 3909: 3906: 3905: 3901: 3899: 3898: 3894: 3893: 3891: 3889: 3882: 3876: 3875: 3868: 3866: 3865: 3864:Sarmatosuchus 3858: 3856: 3855: 3854:Koilamasuchus 3848: 3846: 3845: 3838: 3835: 3834: 3827: 3825: 3824: 3817: 3815: 3814: 3807: 3806: 3803: 3799: 3796: 3793: 3788: 3784: 3780: 3775: 3771: 3761: 3756: 3753: 3746: 3721: 3720: 3719: 3718: 3716: 3714: 3710: 3704: 3703: 3699: 3697: 3696: 3692: 3691: 3689: 3686: 3685:Prolacertidae 3679: 3671: 3670:Rhynchosauria 3667: 3666: 3665: 3664: 3662: 3660: 3659:Rhynchosauria 3653: 3645: 3641: 3640: 3639: 3638: 3636: 3634: 3627: 3621: 3620: 3613: 3611: 3610: 3603: 3600: 3599: 3592: 3590: 3589: 3582: 3581: 3578: 3575: 3572: 3567: 3556: 3555: 3551: 3549: 3548: 3547:Tanytrachelos 3544: 3542: 3541: 3540:Tanystropheus 3537: 3534: 3533: 3529: 3527: 3526: 3522: 3520: 3519: 3515: 3513: 3512: 3508: 3506: 3505: 3501: 3499: 3498: 3494: 3491: 3490: 3489:Gracilicollum 3486: 3483: 3482: 3478: 3476: 3475: 3471: 3469: 3468: 3464: 3462: 3461: 3457: 3456: 3454: 3452: 3448: 3442: 3441: 3440:Sharovipteryx 3437: 3435: 3434: 3430: 3429: 3427: 3424: 3420: 3414: 3413: 3409: 3406: 3405: 3401: 3399: 3398: 3394: 3392: 3391: 3387: 3384: 3383: 3382:Gracilicollum 3379: 3376: 3375: 3371: 3369: 3368: 3364: 3362: 3361: 3357: 3356: 3354: 3352: 3348: 3341: 3340: 3339:Jesairosaurus 3336: 3335: 3332: 3329: 3326: 3318: 3312: 3311: 3304: 3302: 3301: 3294: 3292: 3291: 3290:Protorosaurus 3284: 3282: 3281: 3274: 3272: 3271: 3264: 3262: 3261: 3254: 3252: 3251: 3244: 3241: 3240: 3233: 3231: 3230: 3223: 3221: 3220: 3213: 3212: 3209: 3206: 3203: 3198: 3194: 3190: 3185: 3181: 3160: 3159: 3158: 3157: 3155: 3153: 3149: 3141: 3137: 3136: 3135: 3134: 3131: 3128: 3126: 3122: 3118: 3110: 3109:Archelosauria 3106: 3103: 3101: 3097: 3094: 3092: 3088: 3085: 3083: 3079: 3077: 3073: 3071: 3067: 3066: 3061: 3057: 3053: 3046: 3041: 3039: 3034: 3032: 3027: 3026: 3023: 3016: 3012: 3008: 3005: 3002: 3000: 2997: 2996: 2992: 2987: 2983: 2979: 2975: 2974: 2968: 2964: 2959: 2958: 2953: 2946: 2942: 2938: 2932: 2929: 2924: 2920: 2913: 2910: 2904: 2899: 2895: 2891: 2887: 2883: 2879: 2872: 2869: 2864: 2860: 2855: 2850: 2845: 2840: 2836: 2832: 2828: 2824: 2823: 2818: 2811: 2808: 2803: 2799: 2795: 2791: 2787: 2783: 2776: 2774: 2770: 2765: 2761: 2757: 2753: 2749: 2745: 2740: 2735: 2731: 2727: 2723: 2719: 2715: 2711: 2707: 2699: 2696: 2691: 2687: 2682: 2677: 2672: 2671:10.1206/352.1 2667: 2663: 2659: 2658: 2653: 2646: 2644: 2642: 2640: 2638: 2634: 2629: 2628: 2620: 2617: 2612: 2608: 2604: 2598: 2596: 2592: 2587: 2583: 2578: 2573: 2568: 2563: 2559: 2555: 2551: 2547: 2543: 2541: 2532: 2529: 2522: 2515: 2514: 2506: 2505: 2497: 2496: 2488: 2487: 2478: 2474: 2473: 2465: 2464: 2456: 2455: 2447: 2446: 2438: 2437: 2429: 2428: 2420: 2419: 2411: 2410: 2402: 2401: 2398: 2397: 2395: 2391: 2381: 2380: 2377: 2376: 2373: 2371: 2367: 2366: 2365:Batrachotomus 2356: 2355: 2349: 2348: 2345: 2344: 2341: 2339: 2335: 2334: 2324: 2323: 2316: 2312: 2311: 2308: 2307: 2304: 2302: 2298: 2297: 2287: 2286: 2280: 2279: 2276: 2275: 2267: 2266: 2258: 2257: 2249: 2248: 2245: 2243: 2239: 2238: 2228: 2227: 2224: 2223: 2220: 2219: 2218: 2217:Stagonosuchus 2208: 2207: 2201: 2200: 2197: 2196: 2193: 2191: 2187: 2185: 2175: 2174: 2168: 2167: 2164: 2163: 2160: 2159: 2158: 2148: 2147: 2141: 2140: 2133: 2132:Rauisuchoidea 2129: 2128: 2125: 2124: 2121: 2119: 2115: 2114: 2113:Poposauroidea 2104: 2103: 2096: 2092: 2091: 2088: 2087: 2079: 2078: 2075: 2073: 2069: 2068: 2058: 2057: 2054: 2053: 2050: 2048: 2044: 2043: 2033: 2032: 2026: 2025: 2019: 2018: 2015: 2014: 2006: 2005: 1997: 1996: 1988: 1987: 1984: 1981: 1976: 1975: 1968: 1967: 1964: 1963: 1960: 1958: 1954: 1953: 1943: 1942: 1935: 1931: 1930: 1927: 1926: 1923: 1921: 1917: 1916: 1915:Gracilisuchus 1906: 1905: 1898: 1894: 1893: 1890: 1889: 1886: 1884: 1880: 1879: 1869: 1868: 1862: 1861: 1854: 1850: 1849: 1846: 1845: 1842: 1840: 1836: 1835: 1825: 1824: 1817: 1816: 1811: 1810: 1807: 1806: 1803: 1801: 1797: 1796: 1789: 1788: 1781: 1777: 1776: 1773: 1772: 1769: 1767: 1763: 1754: 1753: 1747: 1746: 1743: 1742: 1739: 1737: 1733: 1732: 1722: 1721: 1715: 1714: 1711: 1710: 1707: 1705: 1701: 1700: 1699:Erythrosuchus 1690: 1689: 1683: 1682: 1678: 1676: 1673: 1663: 1662: 1654: 1653: 1645: 1644: 1636: 1635: 1627: 1626: 1618: 1617: 1609: 1608: 1599: 1598: 1590: 1589: 1581: 1580: 1572: 1571: 1563: 1562: 1554: 1553: 1545: 1544: 1536: 1535: 1527: 1526: 1518: 1517: 1514: 1511: 1506: 1505: 1498: 1497: 1494: 1493: 1490: 1488: 1484: 1483: 1473: 1472: 1466: 1465: 1462: 1461: 1458: 1457: 1456: 1446: 1445: 1439: 1438: 1435: 1434: 1431: 1429: 1425: 1424: 1423:Batrachotomus 1414: 1413: 1407: 1406: 1403: 1402: 1399: 1397: 1393: 1392: 1382: 1381: 1375: 1374: 1371: 1370: 1367: 1365: 1361: 1360: 1350: 1349: 1342: 1338: 1337: 1334: 1333: 1330: 1328: 1324: 1323: 1322:Poposauroidea 1313: 1312: 1305: 1301: 1300: 1297: 1296: 1293: 1291: 1287: 1286: 1276: 1275: 1269: 1268: 1265: 1264: 1256: 1255: 1252: 1250: 1246: 1245: 1235: 1234: 1231: 1230: 1227: 1225: 1221: 1220: 1210: 1209: 1203: 1202: 1199: 1198: 1195: 1193: 1189: 1188: 1187:Turfanosuchus 1178: 1177: 1174: 1173: 1170: 1168: 1164: 1163: 1162:Gracilisuchus 1153: 1152: 1145: 1141: 1140: 1137: 1136: 1133: 1131: 1127: 1126: 1116: 1115: 1109: 1105: 1104: 1101: 1100: 1097: 1095: 1090: 1089: 1082: 1081: 1074: 1070: 1069: 1066: 1065: 1062: 1060: 1056: 1055: 1045: 1044: 1037: 1036: 1031: 1030: 1027: 1026: 1023: 1021: 1017: 1016: 1006: 1005: 999: 998: 995: 994: 991: 989: 985: 984: 974: 973: 967: 966: 963: 962: 959: 957: 953: 952: 942: 941: 935: 934: 931: 930: 927: 925: 921: 920: 919:Erythrosuchus 910: 909: 903: 902: 899: 898: 895: 893: 889: 888: 878: 877: 870: 867: 866: 862: 860: 855: 851: 847: 838: 836: 833: 831: 827: 823: 819: 814: 809: 807: 803: 799: 795: 791: 787: 782: 780: 779:sphenosuchian 776: 772: 768: 764: 760: 756: 752: 748: 741: 740: 736: 731: 724: 722: 720: 716: 712: 708: 700: 698: 696: 692: 688: 684: 680: 676: 671: 669: 665: 661: 657: 653: 648: 646: 642: 637: 633: 629: 625: 621: 619: 618: 613: 612: 611:Sharovipteryx 607: 603: 599: 598: 597:Hesperosuchus 593: 592: 591:Ornithosuchus 587: 583: 579: 578: 573: 569: 568: 563: 558: 556: 555: 550: 546: 538: 536: 534: 530: 526: 521: 519: 515: 511: 507: 503: 495: 493: 491: 487: 483: 479: 475: 471: 467: 463: 458: 456: 452: 448: 447: 442: 441: 436: 432: 428: 427:erpetosuchids 424: 423: 418: 417: 412: 411: 406: 402: 396: 394: 390: 386: 382: 378: 374: 370: 369: 364: 363: 358: 354: 350: 345: 343: 338: 334: 323: 312: 308: 294: 293: 291: 288: 284: 279: 276: 274: 268: 265: 259: 257: 256: 249: 248: 246: 241: 236: 231: 228: 225: 222: 221: 218: 215: 212: 209: 208: 205: 202: 199: 198: 195: 192: 189: 188: 185: 182: 179: 178: 175: 172: 169: 168: 163: 158: 154: 150: 146: 145: 144:Desmatosuchus 140: 136: 135: 130: 126: 121: 114: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 53: 46: 42: 33: 30: 19: 6907:Pseudosuchia 6877:Pseudosuchia 6876: 6832:Sphenosuchia 6789: 6752:crocodilians 6723:notosuchians 6677: 6672:Nominosuchus 6670: 6663: 6656: 6649: 6629: 6622: 6615: 6610:Hadongsuchus 6608: 6580: 6570: 6559: 6549: 6539: 6529: 6519: 6509: 6476: 6470:Lisboasaurus 6468: 6461: 6456:Cassissuchus 6454: 6430: 6423: 6400: 6392: 6384: 6377: 6370: 6363: 6358:Edentosuchus 6356: 6349: 6342: 6321:Tagarosuchus 6319: 6309: 6299: 6291:Microchampsa 6289: 6279: 6269: 6259: 6196: 6189: 6184:Almadasuchus 6182: 6175:Hallopodidae 6159: 6152: 6145: 6122: 6111: 6103:Sphenosuchus 6101: 6091: 6081: 6071: 6061: 6051: 6041: 6030: 6020: 6010: 6000: 5990: 5920: 5869: 5861: 5854: 5846: 5841:Teratosaurus 5839: 5833:Scythosuchus 5831: 5824: 5817: 5809: 5802: 5797:Polonosuchus 5795: 5787: 5781:Energosuchus 5779: 5773:Arganasuchus 5771: 5764:Rauisuchidae 5750:Youngosuchus 5748: 5738: 5728: 5718: 5710:Prestosuchus 5708: 5697: 5689:Luperosuchus 5687: 5676: 5665: 5655: 5647:Fasolasuchus 5645: 5635: 5625: 5615: 5605: 5597:Arganasuchus 5595: 5585: 5554: 5547: 5540: 5520: 5513: 5506:Poposauridae 5493: 5487:Tsylmosuchus 5485: 5478: 5471: 5464: 5459:Bromsgroveia 5457: 5450: 5430: 5422: 5415: 5408: 5381: 5324: 5276:Yonghesuchus 5274: 5267: 5260: 5252: 5245: 5195: 5188: 5183:Acaenasuchus 5181: 5154: 5149:Parringtonia 5147: 5142:Pagosvenator 5140: 5135:Erpetosuchus 5133: 5125: 5117: 5096:Ticinosuchus 5094: 5083: 5073: 5062: 5051: 5018: 5011: 5004: 4999:Dynamosuchus 4997: 4991:Aenigmaspina 4989: 4965: 4955: 4945: 4926:Pseudosuchia 4925: 4913:Pseudosuchia 4912: 4872:Pseudosuchia 4871: 4797: 4763:Pseudosuchia 4762: 4670:Choristodera 4635:Paraphyletic 4622: 4612: 4602: 4592: 4582: 4572: 4562: 4552: 4542: 4532: 4522: 4511:Nomina dubia 4509: 4497: 4487: 4477: 4469:Palaeosaurus 4467: 4459:Tecovasaurus 4457: 4447: 4439:Crosbysaurus 4437: 4392: 4355:crocodilians 4349:Pseudosuchia 4338:Pseudosuchia 4337: 4296: 4285: 4272: 4184: 4177: 4170: 4147: 4137: 4127: 4119:Litorosuchus 4117: 4107: 4097: 4087: 4050: 4045:Shansisuchus 4043: 4036: 4029: 4021: 4014: 4007: 4000: 3977: 3970: 3946: 3940:Tsylmosuchus 3938: 3931: 3924: 3917: 3910: 3902: 3895: 3872: 3862: 3852: 3842: 3831: 3821: 3811: 3748: 3700: 3693: 3617: 3607: 3596: 3586: 3552: 3545: 3538: 3530: 3523: 3516: 3509: 3502: 3495: 3487: 3481:Fuyuansaurus 3479: 3472: 3465: 3458: 3438: 3431: 3410: 3402: 3395: 3388: 3380: 3374:Fuyuansaurus 3372: 3365: 3358: 3337: 3308: 3300:Rhombopholis 3298: 3288: 3278: 3268: 3258: 3248: 3237: 3229:Czatkowiella 3227: 3217: 3104: 3095: 3086: 2977: 2971: 2962: 2931: 2918: 2912: 2885: 2881: 2871: 2826: 2820: 2810: 2785: 2781: 2713: 2709: 2698: 2661: 2655: 2626: 2619: 2610: 2549: 2545: 2539: 2531: 2477:Pseudosuchia 2476: 2390:Prestosuchus 2388: 2384: 2363: 2359: 2331: 2327: 2296:Rauisuchidae 2294: 2290: 2237:Ticinosuchus 2235: 2231: 2215: 2211: 2184:Fasolasuchus 2182: 2178: 2157:Arganasuchus 2155: 2151: 2111: 2107: 2065: 2061: 2040: 2036: 1972: 1971: 1952:Erpetosuchus 1950: 1946: 1913: 1909: 1876: 1872: 1832: 1828: 1813: 1793: 1792: 1757: 1729: 1725: 1697: 1693: 1672:monophyletic 1669: 1502: 1501: 1482:Rauisuchidae 1480: 1476: 1455:Fasolasuchus 1453: 1449: 1421: 1417: 1389: 1385: 1359:Prestosuchus 1357: 1353: 1320: 1316: 1285:Ticinosuchus 1283: 1279: 1242: 1238: 1217: 1213: 1185: 1181: 1160: 1156: 1123: 1119: 1108:Pseudosuchia 1107: 1086: 1085: 1052: 1048: 1033: 1013: 1009: 981: 977: 949: 945: 917: 913: 885: 881: 854:phylogenetic 842: 834: 810: 790:Sphenosuchia 783: 763:rauisuchians 744: 737: 711:ornithodires 704: 691:phylogenetic 672: 664:Rauisuchidae 649: 641:sister taxon 632:phylogenetic 622: 615: 609: 595: 589: 575: 565: 559: 552: 542: 522: 502:Pseudosuchia 501: 499: 459: 455:shuvosaurids 451:ornithomimid 444: 438: 422:Fasolasuchus 420: 416:Prestosuchus 414: 408: 397: 376: 366: 360: 346: 337:crocodilians 307:Pseudosuchia 306: 305: 253: 230:Pseudosuchia 229: 223: 210: 142: 139:Rauisuchidae 132: 29: 6901:Wikispecies 6837:Protosuchia 6717:Ziphosuchia 6706:Ziphosuchia 6651:Adzhosuchus 6521:Hsisosuchus 6478:Zaraasuchus 6463:Gobiosuchus 6432:Orthosuchus 6425:Notochampsa 6402:Protosuchus 6386:Orthosuchus 6379:Hoplosuchus 6351:Dianosuchus 6032:Heteropelta 5804:Postosuchus 5730:Saurosuchus 5699:Mandasuchus 5667:Heteropelta 5657:Heptasuchus 5587:Apatosuchus 5556:Sillosuchus 5549:Shuvosaurus 5495:Xilousuchus 5432:Qianosuchus 5383:Mambawakale 5262:Parvosuchus 5119:Archeopelta 5085:Nundasuchus 5075:Mandasuchus 5053:Heteropelta 5013:Riojasuchus 4967:Phytosauria 4957:Nundasuchus 4802:Archosauria 4654:Thecodontia 4624:Wangisuchus 4604:Microcnemus 4584:Fenhosuchus 4574:Exilisuchus 4564:Collilongus 4554:Blomosuchus 4534:Ankistrodon 4489:Uatchitodon 4264:Archosauria 4252:Phytosauria 4241:Phytosauria 4109:Heteropelta 4068:Eucrocopoda 4052:Uralosaurus 4009:Chalishevia 3919:Jaikosuchus 3897:Archosaurus 3813:Antarctanax 3588:Boreopricea 3518:Macrocnemus 3460:Amotosaurus 2739:11336/41466 2333:Saurosuchus 1834:Phytosauria 1780:Archosauria 1391:Saurosuchus 1073:Archosauria 1054:Phytosauria 767:herbivorous 707:Archosauria 701:Description 693:analyses. 683:Phytosauria 675:Paul Sereno 617:Longisquama 586:Thecodontia 446:Postosuchus 410:Saurosuchus 381:Phytosauria 333:Archosauria 264:Phytosauria 217:Archosauria 134:Postosuchus 50:248–0  6951:Categories 6842:Mesosuchia 6827:Bathyotica 6817:Rauisuchia 6815:See also: 6694:Metasuchia 6531:Induszalim 6124:Trialestes 5892:see below↓ 5848:Tikisuchus 5819:Rauisuchus 5789:Jushatyria 5637:Etjosuchus 5617:Dagasuchus 5522:Poposaurus 5417:Lotosaurus 5296:see below↓ 5219:Aetosauria 5208:Aetosauria 4881:see below↓ 4833:Sauropsida 4818:Sauropsida 4793:Sauropsida 4594:Megacnemus 4524:Albisaurus 4428:Tooth taxa 4228:Crurotarsi 4186:Osmolskina 4172:Euparkeria 4149:Vancleavea 4099:Dorosuchus 4023:Fugusuchus 3979:Triopticus 3972:Kranosaura 3912:Gamosaurus 3844:Kalisuchus 3833:Eorasaurus 3823:Cuyosuchus 3722:see below↓ 3702:Prolacerta 3695:Kadimakara 3525:Raibliania 3511:Luxisaurus 3474:Cosesaurus 3360:Austronaga 3325:Tanysauria 3250:Elessaurus 3161:see below↓ 3140:Sauropsida 3125:Sauropsida 3082:Sauropsida 2523:References 2095:Rauisuchia 1934:Bathyotica 1878:Aetosauria 1815:Crurotarsi 1731:Euparkeria 1675:Rauisuchia 1244:Aetosauria 1035:Crurotarsi 1015:Euparkeria 951:Vancleavea 850:Crurotarsi 826:alligators 822:crocodiles 753:); by the 679:Crurotarsi 533:pterosaurs 510:Crurotarsi 482:alligators 478:crocodiles 474:Cretaceous 440:Poposaurus 389:pterosaurs 387:) such as 379:'s group, 357:node-based 353:Crurotarsi 243:Subgroups 149:Aetosauria 6750:includes 6746:Neosuchia 6735:Neosuchia 6721:includes 6665:Kyasuchus 6063:Parrishia 4779:Kingdom: 4499:Zanclodon 4353:includes 4324:includes 4089:Asperoris 4031:Garjainia 3948:Vonhuenia 3619:Teyujagua 3571:Crocopoda 3397:Pectodens 3068:Kingdom: 2980:(2): 1–7. 2973:PaleoBios 2923:Eichstätt 2764:256707805 2748:2397-334X 2681:2246/6112 2664:: 1–292. 859:cladogram 839:Phylogeny 806:dinosaurs 775:poposaurs 771:aetosaurs 759:carnivore 747:Olenekian 735:loricatan 725:Evolution 715:dinosaurs 673:In 1990, 668:aetosaurs 582:suborders 549:aetosaurs 529:dinosaurs 500:The name 490:gavialids 435:aetosaurs 393:dinosaurs 329:(souchos) 318:(pseudos) 180:Kingdom: 174:Eukaryota 6921:42334946 6892:Q2742285 6886:Wikidata 6852:Category 6631:Zosuchus 6281:Kemkemia 6191:Hallopus 5992:Carnufex 5571:Loricata 5156:Tarjadia 4787:Chordata 4785:Phylum: 4781:Animalia 4721:Category 3091:Diapsida 3076:Chordata 3074:Phylum: 3070:Animalia 3007:Archived 2863:20308590 2756:29185518 2690:83493714 2586:22022431 2546:PLOS ONE 1341:Loricata 846:Eusuchia 830:gharials 813:Mesozoic 769:armored 755:Ladinian 751:Triassic 624:Gauthier 602:theropod 506:Triassic 470:Jurassic 407:such as 401:Triassic 377:Rutiodon 373:Triassic 342:misnomer 287:Synonyms 204:Reptilia 194:Chordata 190:Phylum: 184:Animalia 170:Domain: 5856:Vivaron 5542:Effigia 5254:Maehary 5127:Dyoplax 4791:Class: 3080:Class: 2954:Sources 2890:Bibcode 2888:: 763. 2854:2872409 2831:Bibcode 2790:Bibcode 2718:Bibcode 2577:3194824 2554:Bibcode 2479:  2317:  2134:  2097:  1936:  1899:  1855:  1818:  1782:  1343:  1306:  1146:  1075:  1038:  871:  811:As the 749:(early 719:mammals 554:Dyoplax 486:caimans 200:Class: 45:Present 5036:Suchia 4637:groups 4544:Avipes 3433:Ozimek 3100:Sauria 2861:  2851:  2762:  2754:  2746:  2688:  2584:  2574:  2475:  2313:  2130:  2093:  1932:  1895:  1853:Suchia 1851:  1812:  1778:  1339:  1302:  1144:Suchia 1142:  1106:  1071:  1032:  828:, and 796:(both 652:Benton 628:Padian 531:, and 488:, and 453:-like 419:, and 327:σούχος 316:ψεύδος 309:(from 278:Suchia 237:, 1887 235:Zittel 141:) and 6934:93706 4798:Clade 4326:birds 3105:Clade 3096:Clade 3087:Clade 2760:S2CID 2686:S2CID 717:' or 518:clade 514:clade 349:clade 322:Greek 311:Greek 224:Clade 211:Clade 6744:see 6715:see 6221:see 5217:see 4858:see 4831:see 4347:see 4318:see 4298:Smok 4250:see 4209:see 3668:see 3642:see 3138:see 2859:PMID 2822:PNAS 2752:PMID 2744:ISSN 2582:PMID 792:and 784:The 666:and 626:and 614:and 594:and 570:and 551:and 472:and 443:and 429:and 391:and 347:The 58:PreꞒ 6916:EoL 2941:doi 2898:doi 2849:PMC 2839:doi 2827:107 2798:doi 2734:hdl 2726:doi 2676:hdl 2666:doi 2662:352 2572:PMC 2562:doi 662:of 574:'s 564:'s 6953:: 6931:: 6918:: 6903:: 6888:: 4800:: 3107:: 3098:: 3089:: 2978:25 2976:. 2896:. 2884:. 2880:. 2857:. 2847:. 2837:. 2825:. 2819:. 2796:. 2786:17 2784:. 2772:^ 2758:. 2750:. 2742:. 2732:. 2724:. 2712:. 2708:. 2684:. 2674:. 2660:. 2654:. 2636:^ 2594:^ 2580:. 2570:. 2560:. 2548:. 2544:. 824:, 765:, 492:. 484:, 480:, 413:, 324:: 313:: 226:: 213:: 151:) 108:Pg 52:Ma 47:, 6703:† 6594:† 6585:? 6578:† 6568:† 6564:? 6557:† 6547:† 6537:† 6527:† 6517:† 6507:† 6473:? 6444:† 6435:? 6413:† 6397:? 6389:? 6332:† 6317:† 6307:† 6297:† 6287:† 6277:† 6267:† 6257:† 6209:† 6172:† 6135:† 6120:† 6116:? 6109:† 6099:† 6089:† 6079:† 6069:† 6059:† 6049:† 6039:† 6035:? 6028:† 6018:† 6008:† 5998:† 5988:† 5874:? 5866:? 5851:? 5836:? 5814:? 5792:? 5784:? 5776:? 5761:† 5746:† 5736:† 5726:† 5716:† 5706:† 5702:? 5695:† 5685:† 5681:? 5674:† 5670:? 5663:† 5653:† 5643:† 5633:† 5623:† 5613:† 5603:† 5593:† 5583:† 5490:? 5427:? 5394:† 5379:† 5257:? 5235:† 5167:† 5130:? 5122:? 5107:† 5092:† 5088:? 5081:† 5071:† 5067:? 5060:† 5056:? 5049:† 4994:? 4979:† 4970:? 4963:† 4953:† 4943:† 4755:e 4748:t 4741:v 4620:† 4610:† 4600:† 4590:† 4580:† 4570:† 4560:† 4550:† 4540:† 4530:† 4520:† 4495:† 4485:† 4475:† 4465:† 4455:† 4445:† 4435:† 4294:† 4290:? 4283:† 4238:† 4197:† 4160:† 4145:† 4135:† 4125:† 4115:† 4105:† 4095:† 4085:† 4026:? 3990:† 3960:† 3951:? 3943:? 3907:? 3885:† 3870:† 3860:† 3850:† 3840:† 3836:? 3829:† 3819:† 3809:† 3687:? 3682:† 3656:† 3630:† 3615:† 3605:† 3601:? 3594:† 3584:† 3557:? 3535:? 3492:? 3484:? 3425:? 3407:? 3385:? 3377:? 3342:? 3322:† 3306:† 3296:† 3286:† 3276:† 3266:† 3256:† 3246:† 3242:? 3235:† 3225:† 3215:† 3044:e 3037:t 3030:v 2943:: 2906:. 2900:: 2892:: 2886:9 2865:. 2841:: 2833:: 2804:. 2800:: 2792:: 2766:. 2736:: 2728:: 2720:: 2714:1 2692:. 2678:: 2668:: 2588:. 2564:: 2556:: 2550:6 2386:† 2361:† 2329:† 2292:† 2233:† 2213:† 2186:' 2180:† 2153:† 2109:† 2063:† 2038:† 1948:† 1911:† 1874:† 1830:† 1759:† 1727:† 1695:† 1478:† 1451:† 1419:† 1387:† 1355:† 1318:† 1281:† 1240:† 1215:† 1183:† 1158:† 1121:† 1050:† 1011:† 979:† 947:† 915:† 883:† 270:† 266:? 261:† 251:† 147:( 137:( 113:N 103:K 98:J 93:T 88:P 83:C 78:D 73:S 68:O 63:Ꞓ 43:– 20:)

Index

Crocodylotarsi
Early Triassic
Present
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Postosuchus
Rauisuchidae
Desmatosuchus
Aetosauria
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Reptilia
Archosauria
Pseudosuchia
Zittel

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