261:
416:
111:
133:
880:
667:
went extinct at the end of the Eocene. A supported theory is that these mammals went extinct during the Eocene-Oligocene extinction event; A large faunal turnover at the end of the
783:, a new machaeroidine (Oxyaenidae) from the Santiago Formation (late Uintan) of southern California and the relationships of Machaeroidinae, the oldest group of sabertooth mammals"
260:
415:
3398:
1195:
862:
3439:
655:
451:
3454:
1162:
454:. This creature as well as its subfamily were some of the first predatory saber toothed mammals to have evolved, 30 million years before the
890:
1056:"Eocene-Oligocene mammalian faunal turnover in the Hampshire Basin, UK: calibration to the global time scale and the major cooling event"
407:. According to a paper on the creature, ''The present study highlights how poorly documented the machaeroidine fossil record remains''.
3444:
345:
306:
973:
384:
395:
until 2022 when the fossils were described and recognised as a new genus and species. This creature is so far the only
132:
1188:
1126:
974:"High-resolution (104 years) deep-sea foraminiferal stable isotope records of the Eocene-Oligocene climate transition"
841:
110:
1103:
3366:
1181:
1019:
Shackleton, N. J. (1986-10-01). "Boundaries and Events in the
Paleogene Paleogene stable isotope events".
365:
361:
338:
3411:
1109:
905:
450:. Its discovery has made paleontologists question whether this group's extinction was caused due to the
380:
1651:
1561:
2457:
2449:
2182:
1601:
1553:
1070:
1028:
985:
920:
2593:
2563:
2473:
2465:
1843:
1585:
3449:
3058:
3040:
2601:
2118:
2088:
2062:
1956:
1609:
1465:
885:
3293:
3171:
2780:
2489:
2441:
2221:
2002:
1940:
1869:
1742:
1704:
1635:
1521:
3309:
3187:
3163:
2979:
2663:
2633:
2547:
2306:
2166:
1732:
1593:
1577:
1545:
1489:
1086:
954:
591:
127:
3416:
3340:
3179:
3123:
2963:
2671:
2481:
2406:
2352:
2229:
2080:
2072:
1909:
1569:
1399:
1383:
2804:
2398:
2376:
2190:
2158:
2110:
1667:
1537:
1529:
1369:
1361:
3083:
3011:
2971:
2955:
2609:
2360:
2336:
2042:
2034:
1901:
1885:
1659:
1158:
1001:
946:
822:
804:
2150:
2126:
1805:
1758:
1720:
1473:
2853:
1770:
1078:
1036:
993:
936:
928:
812:
794:
463:
455:
2260:
2741:
2729:
2142:
1855:
1780:
1445:
1298:
676:
1121:
906:"Cooler winters as a possible cause of mass extinctions at the Eocene/Oligocene boundary"
1074:
1032:
989:
924:
3329:
2653:
2537:
2100:
1968:
1877:
1825:
1813:
1455:
1241:
817:
778:
722:). It seems once the Oxyaenids went extinct their ecological nice was filled in by the
692:
473:
435:
292:
213:
3433:
2987:
2916:
2326:
1994:
1750:
1625:
1497:
1289:
1277:
1205:
1150:
1090:
1055:
1040:
680:
672:
396:
64:
3389:
3283:
3271:
3261:
3233:
3199:
3149:
3139:
2995:
2908:
2679:
2571:
2368:
2026:
2018:
1948:
1893:
958:
711:
684:
518:
467:
424:
368:
239:
671:
was probably caused by climate change, like the freezing of
Antarctica disrupting
1173:
3091:
2885:
2861:
2764:
2641:
2625:
2134:
1712:
1505:
1481:
1428:
1391:
1319:
1256:
1235:
715:
688:
39:
423:
showing a missing tooth in SDSNH 38343, the paper has noted that this could be
3301:
3225:
3217:
3209:
3110:
3050:
3033:
2877:
2826:
2816:
2796:
2583:
2527:
2514:
2429:
2174:
2010:
1986:
1917:
1675:
1643:
1340:
1250:
727:
723:
699:
695:
420:
310:
296:
200:
187:
84:
49:
1005:
808:
387:
by paleontologist Brad Riney in 1988. The fossils were housed for over three
3249:
3241:
3022:
2924:
2788:
2701:
2691:
2388:
2344:
1978:
1835:
1513:
1082:
664:
447:
334:
144:
89:
33:
950:
826:
3383:
3003:
2932:
2869:
2283:
2237:
1223:
1149:
Werdelin, L.; Yamaguchi, N.; Johnson, W. E. & O'Brien, S. J. (2010).
941:
719:
707:
443:
439:
164:
79:
74:
59:
54:
44:
3403:
799:
2772:
2617:
2314:
459:
404:
388:
285:
94:
69:
997:
863:"Paleontologists discover new San Diego saber-toothed catlike species"
3099:
2414:
1229:
1217:
972:
Zachos, James C.; Quinn, Terrence M.; Salamy, Karen A. (1996-06-01).
932:
703:
392:
318:
314:
174:
154:
101:
3360:
1130:. Colombo, Sri Lanka: Associated Newspapers of Ceylon. July 16, 2012
462:. Due to the lack of remains, there is questioning to these animals
1262:
904:
Ivany, Linda C.; Patterson, William P.; Lohmann, Kyger C. (2000).
668:
414:
288:
842:"A new saber-toothed mammal was among the first hypercarnivores"
400:
3364:
2727:
2511:
1425:
1275:
1177:
726:, a family of saber-toothed mammals that belonged to the group
675:
and global temperature. Many of the primitive mammals like the
881:"Not all saber-toothed animals were predators, fossils reveal"
353:
777:
Zack, Shawn P.; Poust, Ashley W.; Wagner, Hugh (2022-03-15).
480:
is actually the latest surviving member of its subfamily.
1157:. Oxford, UK: Oxford University Press. pp. 59–82.
1153:. In Macdonald, D. W. & Loveridge, A. J. (eds.).
364:
in honor of her research on carnivorous mammals and
3373:
3327:
3281:
3259:
3197:
3147:
3137:
3109:
3074:
3048:
3032:
3021:
2942:
2899:
2836:
2814:
2751:
2740:
2689:
2651:
2581:
2535:
2525:
2428:
2386:
2324:
2293:
2270:
2247:
2208:
2098:
2060:
1966:
1927:
1853:
1823:
1788:
1778:
1768:
1730:
1689:
1623:
1453:
1443:
1359:
1338:
1317:
1297:
1288:
1054:Hooker, J.J.; Collinson, M.E.; Sille, N.P. (2004).
434:was small in stature, with a size comparable to a
1021:Palaeogeography, Palaeoclimatology, Palaeoecology
691:were replaced by more advanced mammals like the
291:of placental mammals from extinct the subfamily
452:large faunal turnover at the end of the Eocene
1189:
8:
1151:"Phylogeny and evolution of cats (Felidae)"
399:species of Machaeroidinae known outside of
3361:
3154:
3144:
3029:
2946:
2755:
2748:
2737:
2724:
2532:
2522:
2508:
2212:
1860:
1785:
1775:
1695:
1450:
1440:
1436:
1422:
1294:
1285:
1272:
1196:
1182:
1174:
427:or it could have happened well antemortem.
259:
109:
20:
940:
816:
798:
488:The phylogenetic relationships of genus
379:The holotype fossils were discovered in
273:as well as other machaeroidinid genera.
1155:Biology and Conservation of Wild Felids
738:
466:. However, there are good remains from
321:around 46.2 to 39.7 million years ago.
299:. This genus contains only one species
492:are shown in the following cladogram:
772:
770:
768:
766:
764:
762:
7:
857:
855:
760:
758:
756:
754:
752:
750:
748:
746:
744:
742:
458:(saber-toothed cats) evolved in the
442:fossils (SDSNH 38343) consists of a
583:
559:
510:
503:
495:
663:along with all the members of its
14:
1063:Journal of the Geological Society
656:Eocene–Oligocene extinction event
268:A map showing the fossil finds of
840:Sofia Quaglia (March 15, 2022).
131:
313:. This mammal lived during the
253:Zack, Poust, & Wagner, 2022
233:Zack, Poust, & Wagner, 2022
1:
3440:Fossil taxa described in 2022
568:Diegoaelurus vanvalkenburghae
432:Diegoaelurus vanvalkenburghae
358:Diegoaelurus vanvalkenburghae
302:Diegoaelurus vanvalkenburghae
248:Diegoaelurus vanvalkenburghae
120:Diegoaelurus vanvalkenburghae
3455:Prehistoric placental genera
1041:10.1016/0031-0182(86)90008-8
3471:
710:, and other more advanced
653:
344:
284:("San Diego's cat") is an
3445:Mammals described in 2022
3157:
2949:
2758:
2736:
2723:
2521:
2507:
2215:
1863:
1698:
1439:
1435:
1421:
1377:
1284:
1271:
1212:
606:
588:
581:
564:
557:
533:
515:
508:
501:
476:lifestyle for the group.
305:, which was found in the
267:
258:
245:
238:
128:Scientific classification
126:
117:
108:
23:
1083:10.1144/0016-764903-091
16:Extinct genus of mammal
889:. 2020. Archived from
428:
366:saber-toothed predator
362:Blaire Van Valkenburgh
295:within extinct family
3412:Paleobiology Database
3075:other representatives
2209:other representatives
1122:"Meet the Cat Family"
1110:Paleobiology Database
418:
360:was named after Dr.
1127:The Sunday Observer
1124:. Junior Observer.
1075:2004JGSoc.161..161H
1033:1986PPP....57...91S
990:1996PalOc..11..251Z
925:2000Natur.407..887I
886:National Geographic
800:10.7717/peerj.13032
536:Machaeroides eothen
478:D. vanvalkenburghae
469:Machaeroides eothen
446:and well preserved
319:Middle Eocene Epoch
271:D. vanvalkenburghae
1733:Indohyaenodontidae
429:
329:The name of genus
307:Santiago Formation
3427:
3426:
3367:Taxon identifiers
3358:
3357:
3354:
3353:
3350:
3349:
3323:
3322:
3319:
3318:
3133:
3132:
3070:
3069:
3066:
3065:
2895:
2894:
2719:
2718:
2715:
2714:
2711:
2710:
2503:
2502:
2499:
2498:
2424:
2423:
2204:
2203:
2200:
2199:
2056:
2055:
2052:
2051:
1685:
1684:
1652:Minimovellentodon
1619:
1618:
1562:Paracynohyaenodon
1417:
1416:
1413:
1412:
1409:
1408:
1204:Mammals of clade
1164:978-0-19-923445-5
998:10.1029/96PA00571
919:(6806): 887–890.
893:on March 3, 2021.
867:Los Angeles Times
646:
645:
637:
636:
628:
627:
619:
618:
546:
545:
464:ecological niches
277:
276:
234:
3462:
3420:
3419:
3407:
3406:
3394:
3393:
3392:
3362:
3155:
3145:
3030:
2947:
2756:
2749:
2738:
2725:
2533:
2523:
2509:
2213:
1861:
1786:
1776:
1771:Hyainailouroidea
1696:
1451:
1441:
1437:
1423:
1295:
1286:
1273:
1198:
1191:
1184:
1175:
1169:
1168:
1146:
1140:
1139:
1137:
1135:
1118:
1112:
1101:
1095:
1094:
1060:
1051:
1045:
1044:
1016:
1010:
1009:
978:Paleoceanography
969:
963:
962:
944:
933:10.1038/35038044
910:
901:
895:
894:
877:
871:
870:
859:
850:
849:
837:
831:
830:
820:
802:
774:
609:Apataelurus kayi
584:
560:
511:
504:
496:
472:which support a
456:Machairodontinae
385:San Diego county
348:
333:comes from city
263:
232:
225:
212:
199:
186:
136:
135:
122:, (SDSNH 38343)
113:
99:
36:
29:Temporal range:
21:
3470:
3469:
3465:
3464:
3463:
3461:
3460:
3459:
3430:
3429:
3428:
3423:
3415:
3410:
3402:
3397:
3388:
3387:
3382:
3369:
3359:
3346:
3315:
3277:
3255:
3193:
3129:
3115:Carnivoraformes
3114:
3105:
3062:
3044:
3025:
3017:
2938:
2891:
2832:
2810:
2744:
2742:Carnivoraformes
2732:
2730:Carnivoramorpha
2707:
2685:
2647:
2577:
2517:
2495:
2458:Sarcotherichnus
2450:Dischidodacylus
2432:
2420:
2382:
2320:
2289:
2266:
2243:
2196:
2183:Paratritemnodon
2143:Ekweeconfractus
2094:
2048:
1962:
1923:
1856:Hyainailourinae
1849:
1819:
1781:Hyainailouridae
1764:
1726:
1691:
1681:
1615:
1602:Protoproviverra
1554:Paenoxyaenoides
1446:Hyaenodontoidea
1431:
1405:
1373:
1355:
1334:
1313:
1308:Carnivoramorpha
1299:Carnivoramorpha
1280:
1267:
1208:
1202:
1172:
1165:
1148:
1147:
1143:
1133:
1131:
1120:
1119:
1115:
1102:
1098:
1058:
1053:
1052:
1048:
1018:
1017:
1013:
971:
970:
966:
908:
903:
902:
898:
879:
878:
874:
861:
860:
853:
839:
838:
834:
776:
775:
740:
736:
714:as well as the
677:Palaeotheriidae
658:
652:
647:
638:
629:
620:
547:
499:Machaeroidinae
486:
474:hypercarnivorus
413:
377:
327:
269:
254:
251:
231:
223:
210:
197:
184:
130:
104:
98:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
32:46.2–39.7
31:
30:
27:
17:
12:
11:
5:
3468:
3466:
3458:
3457:
3452:
3447:
3442:
3432:
3431:
3425:
3424:
3422:
3421:
3408:
3395:
3379:
3377:
3371:
3370:
3365:
3356:
3355:
3352:
3351:
3348:
3347:
3345:
3344:
3335:
3333:
3330:Incertae sedis
3325:
3324:
3321:
3320:
3317:
3316:
3314:
3313:
3305:
3297:
3288:
3286:
3279:
3278:
3276:
3275:
3266:
3264:
3262:Ictidopappinae
3257:
3256:
3254:
3253:
3245:
3237:
3229:
3221:
3213:
3204:
3202:
3195:
3194:
3192:
3191:
3183:
3175:
3167:
3158:
3152:
3142:
3135:
3134:
3131:
3130:
3128:
3127:
3118:
3116:
3107:
3106:
3104:
3103:
3095:
3087:
3078:
3076:
3072:
3071:
3068:
3067:
3064:
3063:
3056:
3054:
3046:
3045:
3038:
3036:
3027:
3019:
3018:
3016:
3015:
3007:
2999:
2991:
2983:
2975:
2967:
2959:
2950:
2944:
2940:
2939:
2937:
2936:
2928:
2920:
2912:
2903:
2901:
2897:
2896:
2893:
2892:
2890:
2889:
2881:
2873:
2865:
2857:
2848:
2846:
2834:
2833:
2831:
2830:
2821:
2819:
2812:
2811:
2809:
2808:
2800:
2792:
2784:
2776:
2768:
2759:
2753:
2746:
2734:
2733:
2728:
2721:
2720:
2717:
2716:
2713:
2712:
2709:
2708:
2706:
2705:
2696:
2694:
2687:
2686:
2684:
2683:
2675:
2667:
2658:
2656:
2654:Palaeonictinae
2649:
2648:
2646:
2645:
2637:
2629:
2621:
2613:
2605:
2597:
2594:Argillotherium
2588:
2586:
2579:
2578:
2576:
2575:
2567:
2564:Isphanatherium
2559:
2551:
2542:
2540:
2538:Machaeroidinae
2530:
2519:
2518:
2512:
2505:
2504:
2501:
2500:
2497:
2496:
2494:
2493:
2485:
2477:
2474:Hyaenodontipus
2469:
2466:Zanclonychopus
2461:
2453:
2445:
2436:
2434:
2433:of Hyaenodonta
2426:
2425:
2422:
2421:
2419:
2418:
2410:
2402:
2393:
2391:
2384:
2383:
2381:
2380:
2372:
2364:
2356:
2348:
2340:
2331:
2329:
2322:
2321:
2319:
2318:
2310:
2301:
2299:
2291:
2290:
2288:
2287:
2278:
2276:
2268:
2267:
2265:
2264:
2255:
2253:
2245:
2244:
2242:
2241:
2233:
2225:
2216:
2210:
2206:
2205:
2202:
2201:
2198:
2197:
2195:
2194:
2186:
2178:
2170:
2162:
2154:
2146:
2138:
2130:
2122:
2114:
2105:
2103:
2101:Teratodontinae
2096:
2095:
2093:
2092:
2084:
2076:
2067:
2065:
2058:
2057:
2054:
2053:
2050:
2049:
2047:
2046:
2038:
2030:
2022:
2014:
2006:
1998:
1990:
1982:
1973:
1971:
1969:Hyainailourini
1964:
1963:
1961:
1960:
1952:
1944:
1935:
1933:
1925:
1924:
1922:
1921:
1913:
1905:
1897:
1889:
1881:
1878:Megistotherium
1873:
1864:
1858:
1851:
1850:
1848:
1847:
1844:Quasiapterodon
1839:
1830:
1828:
1826:Apterodontinae
1821:
1820:
1818:
1817:
1814:Orienspterodon
1809:
1800:
1798:
1795:Orienspterodon
1783:
1773:
1766:
1765:
1763:
1762:
1754:
1746:
1737:
1735:
1728:
1727:
1725:
1724:
1716:
1708:
1699:
1693:
1687:
1686:
1683:
1682:
1680:
1679:
1671:
1663:
1655:
1647:
1639:
1630:
1628:
1621:
1620:
1617:
1616:
1614:
1613:
1605:
1597:
1589:
1586:Prodissopsalis
1581:
1573:
1565:
1557:
1549:
1541:
1533:
1525:
1517:
1509:
1501:
1493:
1485:
1477:
1469:
1460:
1458:
1456:Hyaenodontidae
1448:
1433:
1432:
1426:
1419:
1418:
1415:
1414:
1411:
1410:
1407:
1406:
1404:
1403:
1395:
1387:
1378:
1375:
1374:
1366:
1364:
1357:
1356:
1354:
1353:
1345:
1343:
1336:
1335:
1333:
1332:
1324:
1322:
1315:
1314:
1312:
1311:
1303:
1301:
1292:
1282:
1281:
1276:
1269:
1268:
1266:
1265:
1259:
1253:
1244:
1242:Laurasiatheria
1238:
1232:
1226:
1220:
1213:
1210:
1209:
1203:
1201:
1200:
1193:
1186:
1178:
1171:
1170:
1163:
1141:
1113:
1096:
1069:(2): 161–172.
1046:
1011:
984:(3): 251–266.
964:
896:
872:
851:
832:
737:
735:
732:
693:Rhinocerotidae
673:ocean currents
654:Main article:
651:
648:
644:
643:
640:
639:
635:
634:
631:
630:
626:
625:
622:
621:
617:
616:
613:
612:
605:
602:
601:
598:
597:
587:
582:
580:
577:
576:
573:
572:
563:
558:
556:
553:
552:
549:
548:
544:
543:
540:
539:
532:
529:
528:
525:
524:
514:
509:
507:
502:
500:
494:
485:
484:Classification
482:
412:
409:
397:North American
376:
373:
326:
323:
293:Machaeroidinae
275:
274:
265:
264:
256:
255:
252:
243:
242:
236:
235:
221:
217:
216:
214:Machaeroidinae
208:
204:
203:
195:
191:
190:
182:
178:
177:
172:
168:
167:
162:
158:
157:
152:
148:
147:
142:
138:
137:
124:
123:
115:
114:
106:
105:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
37:
28:
15:
13:
10:
9:
6:
4:
3:
2:
3467:
3456:
3453:
3451:
3448:
3446:
3443:
3441:
3438:
3437:
3435:
3418:
3413:
3409:
3405:
3400:
3396:
3391:
3385:
3381:
3380:
3378:
3376:
3372:
3368:
3363:
3343:
3342:
3337:
3336:
3334:
3332:
3331:
3326:
3312:
3311:
3306:
3304:
3303:
3298:
3296:
3295:
3290:
3289:
3287:
3285:
3280:
3274:
3273:
3268:
3267:
3265:
3263:
3258:
3252:
3251:
3246:
3244:
3243:
3238:
3236:
3235:
3230:
3228:
3227:
3222:
3220:
3219:
3214:
3212:
3211:
3206:
3205:
3203:
3201:
3196:
3190:
3189:
3184:
3182:
3181:
3176:
3174:
3173:
3168:
3166:
3165:
3160:
3159:
3156:
3153:
3151:
3146:
3143:
3141:
3140:Viverravoidea
3136:
3126:
3125:
3120:
3119:
3117:
3112:
3108:
3102:
3101:
3096:
3094:
3093:
3088:
3086:
3085:
3080:
3079:
3077:
3073:
3060:
3055:
3053:
3052:
3047:
3042:
3037:
3035:
3031:
3028:
3024:
3020:
3014:
3013:
3008:
3006:
3005:
3000:
2998:
2997:
2992:
2990:
2989:
2988:Procynodictis
2984:
2982:
2981:
2976:
2974:
2973:
2968:
2966:
2965:
2960:
2958:
2957:
2952:
2951:
2948:
2945:
2941:
2935:
2934:
2929:
2927:
2926:
2921:
2919:
2918:
2917:Palaearctonyx
2913:
2911:
2910:
2905:
2904:
2902:
2898:
2888:
2887:
2882:
2880:
2879:
2874:
2872:
2871:
2866:
2864:
2863:
2858:
2856:
2855:
2850:
2849:
2847:
2844:
2840:
2835:
2829:
2828:
2823:
2822:
2820:
2818:
2817:Quercygalidae
2813:
2807:
2806:
2801:
2799:
2798:
2793:
2791:
2790:
2785:
2783:
2782:
2777:
2775:
2774:
2769:
2767:
2766:
2761:
2760:
2757:
2754:
2750:
2747:
2743:
2739:
2735:
2731:
2726:
2722:
2704:
2703:
2698:
2697:
2695:
2693:
2688:
2682:
2681:
2676:
2674:
2673:
2668:
2666:
2665:
2660:
2659:
2657:
2655:
2650:
2644:
2643:
2638:
2636:
2635:
2630:
2628:
2627:
2622:
2620:
2619:
2614:
2612:
2611:
2606:
2604:
2603:
2602:Dipsalidictis
2598:
2596:
2595:
2590:
2589:
2587:
2585:
2580:
2574:
2573:
2568:
2566:
2565:
2560:
2558:
2557:
2552:
2550:
2549:
2544:
2543:
2541:
2539:
2534:
2531:
2529:
2524:
2520:
2516:
2510:
2506:
2492:
2491:
2486:
2484:
2483:
2478:
2476:
2475:
2470:
2468:
2467:
2462:
2460:
2459:
2454:
2452:
2451:
2446:
2444:
2443:
2438:
2437:
2435:
2431:
2427:
2417:
2416:
2411:
2409:
2408:
2403:
2401:
2400:
2395:
2394:
2392:
2390:
2385:
2379:
2378:
2373:
2371:
2370:
2365:
2363:
2362:
2357:
2355:
2354:
2349:
2347:
2346:
2341:
2339:
2338:
2333:
2332:
2330:
2328:
2327:Limnocyonidae
2323:
2317:
2316:
2311:
2309:
2308:
2303:
2302:
2300:
2297:
2292:
2286:
2285:
2280:
2279:
2277:
2274:
2269:
2263:
2262:
2257:
2256:
2254:
2251:
2246:
2240:
2239:
2234:
2232:
2231:
2226:
2224:
2223:
2218:
2217:
2214:
2211:
2207:
2193:
2192:
2187:
2185:
2184:
2179:
2177:
2176:
2171:
2169:
2168:
2163:
2161:
2160:
2155:
2153:
2152:
2147:
2145:
2144:
2139:
2137:
2136:
2131:
2129:
2128:
2123:
2121:
2120:
2119:Brychotherium
2115:
2113:
2112:
2107:
2106:
2104:
2102:
2097:
2091:
2090:
2089:Thereutherium
2085:
2083:
2082:
2077:
2075:
2074:
2069:
2068:
2066:
2064:
2063:Prionogalidae
2059:
2045:
2044:
2039:
2037:
2036:
2031:
2029:
2028:
2023:
2021:
2020:
2015:
2013:
2012:
2007:
2005:
2004:
1999:
1997:
1996:
1995:Hyainailouros
1991:
1989:
1988:
1983:
1981:
1980:
1975:
1974:
1972:
1970:
1965:
1959:
1958:
1957:Ischnognathus
1953:
1951:
1950:
1945:
1943:
1942:
1937:
1936:
1934:
1931:
1926:
1920:
1919:
1914:
1912:
1911:
1906:
1904:
1903:
1898:
1896:
1895:
1890:
1888:
1887:
1882:
1880:
1879:
1874:
1872:
1871:
1866:
1865:
1862:
1859:
1857:
1852:
1846:
1845:
1840:
1838:
1837:
1832:
1831:
1829:
1827:
1822:
1816:
1815:
1810:
1808:
1807:
1802:
1801:
1799:
1796:
1792:
1787:
1784:
1782:
1777:
1774:
1772:
1767:
1761:
1760:
1755:
1753:
1752:
1751:Indohyaenodon
1747:
1745:
1744:
1739:
1738:
1736:
1734:
1729:
1723:
1722:
1717:
1715:
1714:
1709:
1707:
1706:
1701:
1700:
1697:
1694:
1690:†Afro‑Arabian
1688:
1678:
1677:
1672:
1670:
1669:
1664:
1662:
1661:
1656:
1654:
1653:
1648:
1646:
1645:
1640:
1638:
1637:
1632:
1631:
1629:
1627:
1626:Proviverridae
1622:
1612:
1611:
1610:Quercytherium
1606:
1604:
1603:
1598:
1596:
1595:
1590:
1588:
1587:
1582:
1580:
1579:
1574:
1572:
1571:
1566:
1564:
1563:
1558:
1556:
1555:
1550:
1548:
1547:
1542:
1540:
1539:
1534:
1532:
1531:
1526:
1524:
1523:
1518:
1516:
1515:
1510:
1508:
1507:
1502:
1500:
1499:
1498:Cynohyaenodon
1494:
1492:
1491:
1486:
1484:
1483:
1478:
1476:
1475:
1470:
1468:
1467:
1466:Alienetherium
1462:
1461:
1459:
1457:
1452:
1449:
1447:
1442:
1438:
1434:
1430:
1424:
1420:
1402:
1401:
1396:
1394:
1393:
1388:
1386:
1385:
1380:
1379:
1376:
1372:
1371:
1365:
1363:
1358:
1351:
1347:
1346:
1344:
1342:
1337:
1330:
1326:
1325:
1323:
1321:
1316:
1309:
1305:
1304:
1302:
1300:
1296:
1293:
1291:
1290:Pan-Carnivora
1287:
1283:
1279:
1278:Pan-Carnivora
1274:
1270:
1264:
1260:
1258:
1254:
1252:
1248:
1245:
1243:
1239:
1237:
1233:
1231:
1227:
1225:
1221:
1219:
1215:
1214:
1211:
1207:
1206:Pan-Carnivora
1199:
1194:
1192:
1187:
1185:
1180:
1179:
1176:
1166:
1160:
1156:
1152:
1145:
1142:
1129:
1128:
1123:
1117:
1114:
1111:
1107:
1106:
1100:
1097:
1092:
1088:
1084:
1080:
1076:
1072:
1068:
1064:
1057:
1050:
1047:
1042:
1038:
1034:
1030:
1027:(1): 91–102.
1026:
1022:
1015:
1012:
1007:
1003:
999:
995:
991:
987:
983:
979:
975:
968:
965:
960:
956:
952:
948:
943:
942:2027.42/62707
938:
934:
930:
926:
922:
918:
914:
907:
900:
897:
892:
888:
887:
882:
876:
873:
868:
864:
858:
856:
852:
847:
843:
836:
833:
828:
824:
819:
814:
810:
806:
801:
796:
792:
788:
784:
782:
773:
771:
769:
767:
765:
763:
761:
759:
757:
755:
753:
751:
749:
747:
745:
743:
739:
733:
731:
729:
725:
721:
717:
713:
712:Artiodacytyls
709:
705:
701:
697:
694:
690:
686:
682:
681:Xiphodontidae
678:
674:
670:
666:
662:
657:
649:
642:
641:
633:
632:
624:
623:
615:
614:
611:
610:
604:
603:
600:
599:
596:
595:
594:pishigouensis
593:
586:
585:
579:
578:
575:
574:
571:
570:
569:
562:
561:
555:
554:
551:
550:
542:
541:
538:
537:
531:
530:
527:
526:
523:
522:
520:
513:
512:
506:
505:
498:
497:
493:
491:
483:
481:
479:
475:
471:
470:
465:
461:
457:
453:
449:
445:
441:
437:
433:
426:
422:
417:
410:
408:
406:
402:
398:
394:
390:
386:
382:
374:
372:
370:
367:
363:
359:
355:
351:
347:
343:
340:
339:Ancient Greek
337:and from
336:
332:
324:
322:
320:
317:stage of the
316:
312:
308:
304:
303:
298:
294:
290:
287:
283:
282:
272:
266:
262:
257:
250:
249:
244:
241:
237:
230:
229:
222:
219:
218:
215:
209:
206:
205:
202:
196:
193:
192:
189:
183:
180:
179:
176:
173:
170:
169:
166:
163:
160:
159:
156:
153:
150:
149:
146:
143:
140:
139:
134:
129:
125:
121:
116:
112:
107:
103:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
35:
26:
22:
19:
3375:Diegoaelurus
3374:
3339:
3328:
3308:
3300:
3294:Simpsonictis
3292:
3284:Viverravinae
3272:Ictidopappus
3270:
3248:
3240:
3234:Pristinictis
3232:
3224:
3216:
3208:
3200:Didymictinae
3186:
3178:
3172:Pappictidops
3170:
3162:
3150:Viverravidae
3122:
3098:
3090:
3082:
3049:
3010:
3002:
2996:Prodaphaenus
2994:
2986:
2978:
2970:
2962:
2954:
2931:
2923:
2915:
2909:Dormaalocyon
2907:
2884:
2876:
2868:
2860:
2852:
2842:
2838:
2825:
2803:
2795:
2787:
2781:Simamphicyon
2779:
2771:
2763:
2700:
2692:Tytthaeninae
2680:Palaeonictis
2678:
2670:
2662:
2640:
2632:
2624:
2616:
2608:
2600:
2592:
2572:Machaeroides
2570:
2562:
2556:Diegoaelurus
2555:
2554:
2546:
2515:Oxyaenodonta
2490:Sarjeantipes
2488:
2480:
2472:
2464:
2456:
2448:
2442:Creodontipus
2440:
2413:
2405:
2397:
2375:
2369:Prolimnocyon
2367:
2359:
2351:
2343:
2335:
2313:
2305:
2295:
2282:
2272:
2259:
2249:
2236:
2228:
2222:Eoproviverra
2220:
2189:
2181:
2173:
2165:
2157:
2149:
2141:
2133:
2125:
2117:
2109:
2087:
2079:
2071:
2041:
2033:
2027:Isohyaenodon
2025:
2019:Sivapterodon
2017:
2009:
2003:Parapterodon
2001:
1993:
1985:
1977:
1955:
1949:Hemipsalodon
1947:
1941:Akhnatenavus
1939:
1930:Akhnatenavus
1929:
1916:
1908:
1900:
1894:Metapterodon
1892:
1884:
1876:
1870:Leakitherium
1868:
1842:
1834:
1812:
1804:
1794:
1790:
1757:
1749:
1743:Glibzegdouia
1741:
1719:
1711:
1705:Parvavorodon
1703:
1674:
1666:
1658:
1650:
1642:
1636:Allopterodon
1634:
1608:
1600:
1592:
1584:
1576:
1568:
1560:
1552:
1544:
1536:
1528:
1522:Leonhardtina
1520:
1512:
1504:
1496:
1488:
1480:
1472:
1464:
1398:
1390:
1382:
1368:
1350:Oxyaenodonta
1349:
1341:Oxyaenodonta
1328:
1307:
1255:Grandorder:
1246:
1240:Superorder:
1234:Infraclass:
1154:
1144:
1132:. Retrieved
1125:
1116:
1105:Diegoaelurus
1104:
1099:
1066:
1062:
1049:
1024:
1020:
1014:
981:
977:
967:
916:
912:
899:
891:the original
884:
875:
866:
846:Science News
845:
835:
790:
786:
781:Diegoaelurus
780:
696:(true rhinos
685:Dichobunidae
661:Diegoaelurus
660:
659:
608:
607:
590:
589:
567:
566:
565:
535:
534:
519:Machaeroides
517:
516:
490:Diegoaelurus
489:
487:
477:
468:
431:
430:
425:pathological
378:
369:paleoecology
357:
349:
342:
331:Diegoaelurus
330:
328:
301:
300:
281:Diegoaelurus
280:
279:
278:
270:
247:
246:
240:Type species
228:Diegoaelurus
227:
226:
188:Oxyaenodonta
119:
118:Holotype of
25:Diegoaelurus
24:
18:
3310:Viverriscus
3188:Variviverra
3164:Orientictis
3092:Dawsonicyon
3026:(Clade "E")
2980:Neovulpavus
2886:Paroodectes
2862:Gracilocyon
2839:Gracilocyon
2765:Messelogale
2745:(Clade "A")
2664:Ambloctonus
2642:Sarkastodon
2634:Protopsalis
2626:Patriofelis
2548:Apataelurus
2307:Boualitomus
2167:Masrasector
2135:Dissopsalis
1713:Tritemnodon
1594:Propterodon
1578:Preregidens
1546:Oxyaenoides
1506:Eurotherium
1490:Consobrinus
1482:Cartierodon
1429:Hyaenodonta
1392:Altacreodus
1362:Wyolestidae
1329:Hyaenodonta
1320:Hyaenodonta
1257:Ferungulata
1236:Placentalia
592:Apataelurus
411:Description
350:(aílouros-)
207:Subfamily:
3450:Oxyaenidae
3434:Categories
3390:Q111251465
3341:Ravenictis
3302:Viverravus
3226:Intyrictis
3218:Didymictis
3210:Bryanictis
3180:Preonictis
3124:Falcatipes
3059:Feliformia
3051:Feliformia
3041:Caniformia
3034:Caniformia
2964:Harpalodon
2878:Paramiacis
2827:Quercygale
2797:Xinyuictis
2672:Dipsalodon
2584:Oxyaeninae
2528:Oxyaenidae
2482:Quiritipes
2407:Prototomus
2353:Oxyaenodon
2230:Gazinocyon
2175:Metasinopa
2081:Prionogale
2073:Namasector
2011:Sectisodon
1987:Falcatodon
1918:Simbakubwa
1910:Paroxyaena
1676:Proviverra
1644:Lesmesodon
1570:Praecodens
1400:Tinerhodon
1384:Simidectes
1261:Mirorder:
1251:Scrotifera
1134:January 2,
793:: e13032.
734:References
728:Feliformia
716:Erinaceids
700:Castoridae
650:Extinction
421:radiograph
311:California
297:Oxyaenidae
201:Oxyaenidae
3250:Raphictis
3242:Protictis
3111:ichnotaxa
3023:Carnivora
2943:Clade "D"
2925:Vassacyon
2900:Clade "C"
2805:Zodiocyon
2789:Uintacyon
2752:Clade "B"
2702:Tytthaena
2430:ichnotaxa
2399:Acarictis
2389:Sinopidae
2377:Thinocyon
2345:Limnocyon
2191:Teratodon
2159:Kyawdawia
2111:Anasinopa
1979:Exiguodon
1836:Apterodon
1668:Parvagula
1538:Neosinopa
1530:Matthodon
1514:Hyaenodon
1370:Wyolestes
1216:Kingdom:
1091:140576090
1006:1944-9186
809:2167-8359
724:Nimravids
720:hedgehogs
708:Oreodonts
665:subfamily
448:dentition
381:Oceanside
375:Discovery
335:San Diego
325:Etymology
151:Kingdom:
145:Eukaryota
3404:11652575
3384:Wikidata
3084:Africtis
3012:Walshius
3004:Tapocyon
2972:Lycarion
2956:Ceruttia
2933:Vulpavus
2870:Oodectes
2843:Oodectes
2610:Malfelis
2361:Prolaena
2337:Iridodon
2284:Galecyon
2273:Galecyon
2238:Pyrocyon
2043:Pterodon
2035:Kerberos
1902:Pakakali
1886:Mlanyama
1660:Morlodon
1230:Mammalia
1224:Chordata
1222:Phylum:
1218:Animalia
951:11057663
827:35310159
689:Adapidae
687:and the
521:simpsoni
444:mandible
440:holotype
346:αἴλουρος
194:Family:
175:Mammalia
165:Chordata
161:Phylum:
155:Animalia
141:Domain:
2773:Miocyon
2618:Oxyaena
2315:Lahimia
2296:Lahimia
2151:Furodon
2127:Buhakia
1806:Maocyon
1791:Maocyon
1759:Yarshea
1721:Koholia
1474:Boritia
1352:below ↓
1331:below ↓
1310:below ↓
1228:Class:
1108:in the
1071:Bibcode
1029:Bibcode
986:Bibcode
959:4408282
921:Bibcode
869:. 2022.
818:8932314
704:beavers
460:Miocene
405:Wyoming
389:decades
352: '
286:extinct
220:Genus:
181:Order:
171:Class:
100:middle
3417:453634
3100:Miacis
2854:Eogale
2415:Sinopa
1161:
1089:
1004:
957:
949:
913:Nature
825:
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807:
438:. The
393:museum
315:Uintan
102:Eocene
3057:(See
3039:(See
2845:clade
2298:clade
2275:clade
2261:Arfia
2252:clade
2250:Arfia
1932:clade
1797:clade
1692:clade
1263:Ferae
1247:Clade
1087:S2CID
1059:(PDF)
955:S2CID
909:(PDF)
787:PeerJ
669:epoch
436:fossa
391:in a
341:
289:genus
3399:GBIF
1348:See
1327:See
1306:See
1159:ISBN
1136:2013
1002:ISSN
947:PMID
823:PMID
805:ISSN
403:and
401:Utah
40:PreꞒ
1079:doi
1067:161
1037:doi
994:doi
937:hdl
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813:PMC
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