2011:. Osteoporotic bone is spongy and porous whereas pachyostotic involves an increase in skeletal mass. Spongy/porous bone such as osteoporotic is associated with faster swimming and better maneuverability in water because of the reduction in bone tissue, many extant cetaceans and marine turtles have osteoporotic bone which enables them to be good swimmers. Pachyostotic bone is a general/local increase in skeletal mass which can be caused by osteosclerosis (inner compaction of bone), pachyostosis (hyperplasy of compact cortices) or pachyeosclerosis (combination of the two). Research on dyrosaur bone performed by Rafael César Lima Pedroso de Andrade and Juliana Manso Sayao revealed that this family had osteoporotic bone tissue indicative of a fast-swimming
2132:
from the ocean. From the lower Eocene Oulad Abdoun Basin, there are very few juvenile dyrosaurids, but numerous similarly-sized adult specimens. This has furthered the assumption that juveniles lived in freshwater environments and adults lived in marine environments. Recently however, the large-bodied and fully mature dyrosaurids of the Cerrejón
Formation have shown that some dyrosaurids lived their entire lives in inland environments, never returning to the coast.
2039:. In the left tibia, the cortex had a lamellar zonal bone with five lines of arrested growth (LAGs) which were spaced 300 mm apart, well as a clear vascular networks of primary osteons that decreased in density towards the membrane (periostially). The right femur had double LAGs and an EFS later as well as secondary osteons occurring in the deep cortex and the spongiosa. This tissue growth pattern is a common characteristic of slow growing animals.
142:
519:. Dyrosaurid fossils are globally distributed, having been found in Africa, Asia, Europe, North America and South America. Over a dozen species are currently known, varying greatly in overall size and cranial shape. A majority were aquatic, some terrestrial and others fully marine (see locomotion below), with species inhabiting both freshwater and marine environments. Ocean-dwelling dyrosaurids were among the few marine reptiles to survive the
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1056:
979:
855:
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1996:
119:
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phosphaticus, this feature of deep occlusal pits become less developed and broad. Dyrosaurus teeth are homodont, conical, long, and slender, with asymmetrically labial and lingual surfaces. Posterior teeth are shorter and more compressed than anterior tooth indicating that the tooth size decreased anterior to posterior.
2047:
Dyrosaurids are found in transitional marine sediments from the Late
Cretaceous to lower Eocene. This family is known mainly from Maastrichthian deposits in New Jersey and the late Cretaceous to early Paleogene rock from the Tethys Sea in northern and western Africa. Fossils have also been found from
532:
estimated at 9 m (30 ft) in length. Based on bone tissue evidence, it has been hypothesized that they were slow-growing near-shore marine animals with interlocking closed jaws, able to swim as well as walk on land. External nostrils at the posterior end of its snout and an internal naris in
2019:
which is a component of pachyostotic bone tissue. Osteoporosis is associated with a fully aquatic lifestyle whereas pachyostotic is not fully aquatic but is associated with fast swimming ecology. Therefore, dyrosaurs are semi-aquatic fast swimmers as indicated by their bone structure. Other evidence
2131:
In 1978, it was proposed that dyrosaurids lived as adults in the ocean but reproduced in inland freshwater environments. Remains belonging to small-bodied dyrosaurids from
Pakistan were interpreted as juveniles. Their presence in inland deposits was viewed as evidence that dyrosaurids hatched far
647:
have the largest snout proportions of all dyrosaurids. Snout length was previously used it to establish dyrosaurid relationships, while considering the lengthening of the snout to be a ‘more evolved’ character. This was not congruent with Jouve's conclusion which was that the longest snout is
619:
pits, present particularly in the posterior region of the maxillae that get less pronounced anteriorly. The pits are indicative of an interlocking closed jaw for dyrosaurids because the pits give a way for the upper and lower jaws to alternate. Although they are still present in dyrosaurus
2048:
the
Paleocene and Eocene strata of Pakistan, as well as South America, Brazil, India, Southern Asia as well as coastal. Generally dyrosaurids are recovered from coastal and estuarine deposits through North Africa and the Middle East confirming their existence as semi-aquatic animals.
2825:
Jouve, Stephanie; Bardet, Nathalie; Jalil, Nour-Eddine; Superbiola, Xabier Pereda; Bouya, Baâdi and
Amagzaz, Mbarek. 2008. The oldest African crocodylian: phylogeny, paleobiogeography, and differential survivorship of marine reptiles through the Cretaceous–Tertiary boundary.
2068:, Late Cretaceous discoveries in Egypt. Later, by the Maastrichtian of North America, the record of dyrosaurids became more complete by establishing a widespread distribution that appears to be maintained through the Paleocene and Eocene.
2840:
Hastings, A.K.; Bloch, J.; Jaramillo, C.A. (2011). "A new longirostrine dyrosaurid (Crocodylomorpha, Mesoeucrocodylia) from the
Paleocene of north-eastern Colombia: biogeographic and behavioural implications for new-world dyrosauridae".
656:
Dyrosaurids were once considered an
African group, but discoveries made starting from the 2000s indicate they inhabited the majority of the continents. In fact, basal forms suggest that their cradle may have been North America.
2724:
Head, J. J.; Bloch, J. I.; Hastings, A. K.; Borque, J. R.; Cadena, E. A.; Herrera, F. A.; Polly, P. D.; Jaramillo, C. A. (2009). "Giant boid snake from the
Palaeocene neotropics reveals hotter past equatorial temperatures".
2059:
was able to obtain a greater and more widespread geographic distribution during the Late
Cretaceous. The earliest records of dyrosaurids are either in or close to Africa with fragmentary occurrences from the
596:
bones which separated by the single nasal bone. The last premaxilla and first maxilla are widely separated by a fourth dentary tooth. Alveoli are widely spaced anteriorly and the space between them decreases
2020:
for near shore, semi-aquatic lifestyle is where the fossils are found, often in transitional marine sediments. -using axial frequency swimming (that used primarily by extant crocodylians) with a greater
2604:
Khosla, Ashu; Sertich, Joseph J. W.; Prasad, Guntupalli V. R.; Verma, Omkar (2009-12-12). "Dyrosaurid remains from the
Intertrappean Beds of India and the Late Cretaceous distribution of Dyrosauridae".
554:
Dyrosaurids are known to have a very characteristic skull shape with a long and thin snout that is approximately 68% of the total skull length. The most anterior part of the skull and snout is the
2999:
2003:
Evidence for the semi-aquatic life of dyrosaurids comes from careful analysis of bone structure. There are two types of structural bone organization that can occur in aquatic tetrapods:
592:
with shallow pits that extend posterior to the third maxillary alveoli. There are two premaxillae that are narrow as compared to the maxillary bones and extend in the two long
2111:, which is thought to represent a transitional marine-freshwater environment surrounded by rainforest more inland than the estuarine environment of the Umm Himar Formation.
2968:
Jouve, S.; Iarochène, M.; Bouya, B.; Amaghzaz, M. (2005). "A new dyrosaurid crocodyliform from the Palaeocene of Morocco and a phylogenetic analysis of Dyrosauridae".
4612:
608:) and forms most of the lateral margin of the skull. According to Jouve and Barbosa and perhaps depending on the age of the animal, each maxilla bears 13–19 teeth.
2146:
suggest this genus was more terrestrial than other dyrosaurids, and also shows that modern crocodylians are not good functional analogues for Dyrosauridae.
4673:
526:
The dyrosaurids were a group of mostly marine, long jawed, crocodile-like quadrupeds up to 6 metres (20 ft) long. The largest dyrosaurid was probably
520:
573:
Typical in dyrosaurids is a single nasal element with a characteristic collection of small pits and a constant width until it widens to contact the
4678:
4658:
4599:
2463:"A Long-Snouted Dyrosaurid (Crocodyliformes, Mesoeucrocodylia) from the Paleocene of Morocco: Phylogenetic and Palaeobiogeographic Implications"
533:
its pterygoid indicated a habit of hunting while swimming with the top of the head above the water, enabling it to breathe while stalking prey.
2935:"A long-snouted dyrosaurid (Crocodyliformes, Mesoeucrocodylia) from the Paleocene of Morocco: phylogenetic and palaeogeographic implications"
2120:
3031:
4663:
2556:"From dinosaurs to dyrosaurids (Crocodyliformes): removal of the post-Cenomanian (Late Cretaceous) record of Ornithischia from Africa"
2269:
Martin, Jeremy E.; Antoine, Pierre-Olivier; Perrier, Vincent; Welcomme, Jean-Loup; Metais, Gregoire; Marivaux, Laurent (2019-07-04).
2079:
deposits, indicating that they lived in a river setting. Bones from indeterminate dyrosaurids have been found in inland deposits in
648:
actually the primitive condition so the shorter or longer snout appears independently at least four times in dyrosaurid evolution.
2342:"Paleohistology and Lifestyle Inferences of a Dyrosaurid (Archosauria: Crocodylomorpha) from Paraíba Basin (Northeastern Brazil)"
2656:
Buffetaut, E.; Bussert, R.; Brinkmann, W. (1990). "A new nonmarine vertebrate fauna in the Upper Cretaceous of northern Sudan".
2031:
Dyrosaurids have a tissue pattern that is indicative of a slow-growth animal that was determined by careful analysis of a right
4668:
2878:"The postcranial skeleton of Cerrejonisuchus improcerus (Crocodyliformes: Dyrosauridae) and the unusual anatomy of dyrosaurids"
1682:
128:
2777:
2990:
2270:
1782:
141:
4683:
4474:
2677:"A new small short-snouted dyrosaurid (Crocodylomorpha, Mesoeucrocodylia) from the Paleocene of northeastern Colombia"
1576:
1463:
2020 provide a comprehensive analysis of the relationships of dyrosaurids, shown below. Note the former dyrosaurids
1544:
2515:
Jouve, Stéphane; de Muizon, Christian; Cespedes-Paz, Ricardo; Sossa-Soruco, Víctor; Knoll, Stephane (2020-10-19).
2230:
Buffetaut, E. (1978). "Les Dyrosauridae (Crocodylia, Mesosuchia) des phosphates de l'Eocène inférieur de Tunisie:
4495:
2574:
1871:
1830:
1496:
2517:"The longirostrine crocodyliforms from Bolivia and their evolution through the Cretaceous–Palaeogene boundary"
3024:
1806:
1740:
1723:
1593:
1520:
865:
A long-snouted marine dyrosaurid with the longest snout length in proportion to body size of any dyrosaurid
4023:
4002:
577:
bones, then tapering off for a short distance until it meets the boundary of the frontals and prefrontals.
4539:
1431:
Posteromedial wing of the retroarticular process dorsally situated ventrally on the retroarticular process
3995:
2084:
4625:
3119:
1854:
1699:
1626:
616:
2104:
936:
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1651:
555:
4470:
3732:
1437:
4008:
3862:
3767:
3342:
3017:
2758:
2706:
2638:
2586:
2414:"New dyrosaurid crocodylomorph and evidences for faunal turnover at the K–P transition in Brazil"
2309:
755:
136:
2925:
Buffetaut, E. (1985). L'evolution des crocodiliens. Les animaux disparus-Pour la science, Paris
2021:
1303:
1202:
990:
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4577:
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2750:
2630:
2578:
2536:
2443:
2389:
2371:
2301:
2205:
1441:
4617:
1251:
A large-bodied, long-snouted marine dyrosaurid with blunt teeth and a spoon-shaped snout tip
4279:
4251:
4212:
4197:
4102:
3876:
3621:
3446:
2954:
2899:
2889:
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2742:
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2622:
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2528:
2482:
2433:
2425:
2412:
Barbosa, José Antonio; Kellner, Alexander Wilhelm Armin; Viana, Maria Somália Sales (2008).
2379:
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803:
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391:
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4143:
4092:
3795:
3753:
3686:
3593:
3488:
3439:
3412:
3302:
3269:
3248:
3238:
2904:
2877:
2555:
2438:
2413:
2384:
2341:
2016:
1452:
Quadratojugal participates largely to the cranial condyle for articulation with the jaw
1424:
1319:
765:
508:
431:
283:
2178:(Thomas, 1893) (Mesoeucrocodylia: Dyrosauridae) from the Lower Eocene of North Africa"
4647:
4411:
4355:
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4175:
4164:
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3885:
3739:
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3584:
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3384:
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3228:
3218:
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2642:
2554:
Lamanna, Matthew C.; Smith, Joshua B.; Attia, Yousry S.; Dodson, Peter (2004-09-10).
2516:
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2313:
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2065:
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1196:
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873:
838:
727:
574:
481:
471:
411:
401:
381:
313:
303:
69:
2710:
2590:
1201:
Five species have been named, the most of any dyrosaurid genus; the genus spans the
1125:
4434:
4112:
3948:
3941:
3906:
3899:
3855:
3831:
3809:
3802:
3725:
3702:
3665:
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3600:
3559:
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3356:
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2008:
2004:
1471:
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351:
209:
20:
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3817:
3788:
3781:
3774:
3760:
3746:
3695:
3628:
3614:
3544:
3523:
3481:
3405:
3391:
3335:
3317:
3291:
3198:
2025:
1005:
598:
559:
461:
421:
323:
248:
44:
1995:
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118:
4459:
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4153:
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3913:
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3607:
3516:
3495:
3280:
3259:
3208:
3088:
3070:
2701:
2061:
1330:
1134:
1039:
900:
A long-snouted, long-armed dyrosaurid with the longest arms of any dyrosaurid
589:
567:
547:
451:
441:
89:
54:
4562:
2987:
2778:"UF researchers: Ancient crocodile relative likely food source for Titanoboa"
2634:
2582:
2540:
2375:
2305:
2209:
4265:
3363:
3181:
3168:
3131:
3040:
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917:
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774:
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612:
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505:
196:
153:
105:
94:
38:
2913:
2754:
2447:
2429:
2393:
2119:
lived in a neotropical setting during a time when global temperatures were
1082:
541:
2340:
de Andrade, Rafael César Lima Pedroso; Sayão, Juliana Manso (2014-07-17).
4556:
4397:
4221:
4055:
4042:
3966:
3064:
2807:
Buffetaut, E. (1978). "Crocodilian remains from the Eocene of Pakistan".
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930:
787:
711:
563:
173:
84:
79:
64:
59:
49:
2894:
2746:
2071:
Dyrosaurids have also been found from non-marine sediments. In northern
4604:
2092:
2076:
2012:
1363:
1297:
1246:
1184:
1073:
1061:
984:
860:
825:
593:
502:
183:
99:
74:
2675:
Hastings, A. K; Bloch, J. I.; Cadena, E. A.; Jaramillo, C. A. (2010).
664:
3058:
2271:"A large crocodyloid from the Oligocene of the Bugti Hills, Pakistan"
1385:
1221:
1160:
1113:
1048:
1027:
749:
516:
163:
109:
4533:
2201:
1302:
A large-bodied, long-snouted marine dyrosaurid; the genus spans the
2051:
Dyrosauridae had its greatest taxonomic diversity during the Early
2072:
2056:
2036:
2032:
1994:
1475:
are recovered as a pholidosaurid and a peirosaurid, respectively.
1420:
895:
631:
Dyrosaurs have a snout to skull length of about 68% and the genus
622:
605:
601:
from the fifteenth alveolus with the diameter remaining constant.
579:
540:
4591:
1398:
1285:
1234:
1172:
972:
604:
The maxilla is long (approx. two and a half times the length of
4537:
4040:
3166:
3051:
3013:
2461:
Jouve, Stephane; Bouya, Baâdi; Amaghzaz, Mbarek (March 2008).
935:
A small-bodied, short-snouted freshwater dyrosaurid from the
3009:
2095:
environments near the coast. The recently named dyrosaurids
716:
A large-bodied, long-snouted freshwater dyrosaurid from the
2809:
Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen
2575:
10.1671/0272-4634(2004)024[0764:FDTDCR]2.0.CO;2
1446:
Supratemporal fenestra anteroposteriorly strongly elongated
2024:
motion and frequency of the tail due to highly developed
2418:
Proceedings of the Royal Society B: Biological Sciences
4422:
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4343:
4276:
4209:
4193:
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4160:
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4139:
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4118:
4108:
4098:
4088:
4078:
4068:
3873:
3843:
3713:
3683:
3653:
3581:
3314:
3298:
3287:
3276:
3265:
3255:
3244:
3234:
3224:
3214:
3204:
3194:
477:
467:
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417:
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387:
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359:
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3872:
3842:
3712:
3682:
3652:
3580:
3422:
3325:
3313:
3179:
3130:
3103:
2083:as well. Some dyrosaurids, such as those from the
2167:
2165:
2163:
2161:
2159:
1078:A large-bodied, long-snouted marine dyrosaurid
792:A short-snouted freshwater dyrosaurid from the
2876:Scavezzoni, Isaure; Fischer, Valentin (2021).
3025:
8:
2064:of Sudan and Portugal and several other pre-
3000:Mikko's Phylogeny Archive Dyrosauridae page
2933:Jouve, S.; Bouya, B.; Amaghzaz, M. (2008).
2658:Berliner Geowissenschaftlische Abhandlungen
4534:
4061:
4050:
4037:
3322:
3187:
3176:
3163:
3109:
3100:
3048:
3032:
3018:
3010:
117:
27:
2958:
2903:
2893:
2700:
2521:Zoological Journal of the Linnean Society
2486:
2437:
2407:
2405:
2403:
2383:
2365:
2335:
2333:
2331:
2329:
2327:
2325:
2323:
2028:allowing a more powerful forward thrust.
2155:
679:
611:An important feature of the dyrosaurid
521:Cretaceous–Paleogene extinction event
7:
2174:"A new description of the skull of
1846:
1822:
1798:
1774:
1715:
1674:
1667:
1643:
1618:
1568:
1560:
1536:
1512:
1488:
1478:
4674:Prehistoric marine crocodylomorphs
2828:Journal of Vertebrate Paleontology
2681:Journal of Vertebrate Paleontology
2607:Journal of Vertebrate Paleontology
2563:Journal of Vertebrate Paleontology
2278:Journal of Vertebrate Paleontology
2182:Canadian Journal of Earth Sciences
1419:(2005) diagnose Dyrosauridae as a
14:
2776:Kanapaux, B. (February 2, 2010).
1368:A long-snouted marine dyrosaurid
830:A long-snouted marine dyrosaurid
4518:
4517:
4494:
4022:
4014:
4007:
4001:
3994:
2960:10.1111/j.1475-4983.2007.00747.x
2863:10.1111/j.1475-4983.2011.01092.x
2488:10.1111/j.1475-4983.2007.00747.x
1391:
1356:
1290:
1278:
1239:
1227:
1189:
1177:
1165:
1153:
1124:
1106:
1081:
1066:
1054:
1020:
995:
977:
965:
923:
888:
853:
818:
780:
742:
704:
140:
4679:Maastrichtian first appearances
4659:Late Cretaceous crocodylomorphs
3005:Paleopedia page on Dyrosauridae
1683:Arambourgisuchus khouribgaensis
1449:Symphysis about as wide as high
129:Arambourgisuchus khouribgaensis
2172:Jouve, Stéphane (March 2005).
1:
2970:Acta Palaeontologica Polonica
2298:10.1080/02724634.2019.1671427
2103:have been recovered from the
2075:, dyrosaurids are known from
2055:but it appears as though the
1423:based on the following seven
754:A marine dyrosaurid from the
2367:10.1371/journal.pone.0102189
1783:Acherontisuchus guajiraensis
1440:participates largely to the
1434:Occipital tuberosities small
566:bones separated by a single
558:followed posteriorly by two
1577:Phosphatosaurus gavialoides
1118:probable Junior synonym of
562:bones until they reach two
4700:
2782:University of Florida News
2533:10.1093/zoolinnean/zlaa081
1545:Cerrejonisuchus improcerus
18:
16:Extinct family of reptiles
4664:Paleogene crocodylomorphs
4507:
4492:
4064:
4049:
4036:
3991:
3190:
3175:
3162:
3112:
3099:
3047:
2702:10.1080/02724630903409204
1868:
1851:
1844:
1827:
1820:
1803:
1796:
1779:
1772:
1737:
1720:
1713:
1696:
1679:
1672:
1665:
1648:
1641:
1623:
1616:
1590:
1573:
1566:
1558:
1541:
1534:
1517:
1510:
1493:
1486:
276:
271:
137:Scientific classification
135:
125:
116:
30:
2830:28(2):409–421, June 2008
2244:Géologie Méditerranéenne
1872:Rhabdognathus keiniensis
1831:Rhabdognathus aslerensis
1497:Chenanisuchus lateroculi
126:Skull of the dyrosaurid
19:Not to be confused with
2256:10.3406/geolm.1978.1046
2176:Dyrosaurus phosphaticus
1807:Atlantosuchus coupatezi
1741:Dyrosaurus phosphaticus
1724:Dyrosaurus maghribensis
1594:Sokotosuchus ianwilsoni
1521:Anthracosuchus balrogus
4669:Pseudosuchian families
2430:10.1098/rspb.2008.0110
2000:
1427:or shared characters:
628:
585:
551:
4626:Paleobiology Database
4473:for extant taxa, and
2123:than they are today.
1998:
1855:Congosaurus bequaerti
1700:Luciasuchus lurusinqa
1627:Rodeosuchus machukiru
626:
583:
544:
2627:10.1671/039.029.0416
2527:(zlaa081): 475–509.
1652:Dorbignysuchus niatu
1455:4 premaxillary teeth
1120:Hyposaurus derbianus
989:The genus spans the
511:that lived from the
3733:Anteophthalmosuchus
2988:Jouve et al. (2005)
2951:2008Palgy..51..281J
2895:10.7717/peerj.11222
2855:2011Palgy..54.1095H
2747:10.1038/nature07671
2739:2009Natur.457..715H
2693:2010JVPal..30..139H
2619:2009JVPal..29.1321K
2479:2008Palgy..51..281J
2424:(1641): 1385–1391.
2358:2014PLoSO...9j2189A
2290:2019JVPal..39E1427M
2194:2005CaJES..42..323J
2085:Umm Himar Formation
4684:Eocene extinctions
4475:Extinct Crocodilia
3863:Koumpiodontosuchus
3768:Eutretauranosuchus
2993:2005-12-23 at the
2105:Cerrejón Formation
2001:
1564:Phosphatosaurinae
1328:Junior synonym of
937:Cerrejón Formation
794:Cerrejón Formation
756:Kristianstad Basin
718:Cerrejón Formation
629:
586:
552:
4641:
4640:
4540:Taxon identifiers
4531:
4530:
4513:Coelognathosuchia
4503:
4502:
4490:
4489:
4486:
4485:
4332:Musturzabalsuchus
4244:Pachycheilosuchus
4032:
4031:
3989:
3988:
3985:
3984:
3893:Brillanceausuchus
3576:
3575:
3158:
3157:
3154:
3153:
3150:
3149:
2733:(7230): 715–717.
1982:
1981:
1973:
1972:
1964:
1963:
1955:
1954:
1946:
1945:
1937:
1936:
1928:
1927:
1919:
1918:
1910:
1909:
1901:
1900:
1892:
1891:
1883:
1882:
1761:
1760:
1752:
1751:
1605:
1604:
1442:occipital condyle
1408:
1407:
652:Paleobiogeography
588:Dyrosaurs have a
492:
491:
333:Phosphatosaurinae
267:
4691:
4634:
4633:
4621:
4620:
4608:
4607:
4595:
4594:
4582:
4581:
4580:
4567:
4566:
4565:
4535:
4521:
4520:
4498:
4477:for extinct ones
4424:
4375:
4345:
4280:Allodaposuchidae
4278:
4252:Pietraroiasuchus
4213:Hylaeochampsidae
4211:
4198:Paralligatoridae
4195:
4184:
4173:
4162:
4151:
4141:
4131:
4120:
4110:
4103:Gilchristosuchus
4100:
4090:
4080:
4070:
4062:
4051:
4038:
4027:
4026:
4019:
4018:
4012:
4011:
4005:
3999:
3998:
3877:Paralligatoridae
3875:
3845:
3715:
3685:
3655:
3622:Knoetschkesuchus
3583:
3447:Arambourgisuchus
3323:
3316:
3300:
3289:
3278:
3267:
3257:
3246:
3236:
3226:
3216:
3206:
3196:
3188:
3177:
3164:
3110:
3101:
3094:
3093:
3049:
3034:
3027:
3020:
3011:
2977:
2964:
2962:
2918:
2917:
2907:
2897:
2873:
2867:
2866:
2837:
2831:
2823:
2817:
2816:
2804:
2798:
2797:
2795:
2793:
2784:. Archived from
2773:
2767:
2766:
2721:
2715:
2714:
2704:
2672:
2666:
2665:
2653:
2647:
2646:
2613:(4): 1321–1326.
2601:
2595:
2594:
2560:
2551:
2545:
2544:
2512:
2506:
2499:
2493:
2492:
2490:
2458:
2452:
2451:
2441:
2409:
2398:
2397:
2387:
2369:
2337:
2318:
2317:
2275:
2266:
2260:
2259:
2227:
2221:
2220:
2218:
2216:
2169:
2107:in northwestern
2015:as well as some
1847:
1823:
1799:
1775:
1716:
1675:
1668:
1644:
1619:
1569:
1561:
1537:
1513:
1489:
1479:
1397:
1395:
1394:
1362:
1360:
1359:
1296:
1294:
1293:
1284:
1282:
1281:
1245:
1243:
1242:
1233:
1231:
1230:
1195:
1193:
1192:
1183:
1181:
1180:
1171:
1169:
1168:
1159:
1157:
1156:
1128:
1112:
1110:
1109:
1085:
1072:
1070:
1069:
1060:
1058:
1057:
1026:
1024:
1023:
999:
983:
981:
980:
971:
969:
968:
929:
927:
926:
894:
892:
891:
859:
857:
856:
824:
822:
821:
804:Arambourgisuchus
786:
784:
783:
748:
746:
745:
710:
708:
707:
665:
645:Arambourgisuchus
479:
469:
459:
449:
439:
429:
419:
409:
399:
392:Arambourgisuchus
389:
379:
369:
361:
349:
339:
331:
321:
311:
301:
291:
281:
266:de Stefano, 1903
265:
260:
247:
145:
144:
121:
104:
41:
34:Temporal range:
28:
4699:
4698:
4694:
4693:
4692:
4690:
4689:
4688:
4644:
4643:
4642:
4637:
4629:
4624:
4616:
4611:
4603:
4598:
4590:
4585:
4576:
4575:
4570:
4561:
4560:
4555:
4542:
4532:
4527:
4499:
4482:
4454:
4417:
4405:Eothoracosaurus
4368:
4347:Aegyptosuchidae
4338:
4271:
4204:
4187:Bernissartiidae
4134:Portugalosuchus
4083:Confractosuchus
4058:
4045:
4028:
4021:
4013:
4006:
3993:
3981:
3961:
3868:
3847:Bernissartiidae
3838:
3717:Goniopholididae
3708:
3678:
3657:Stomatosuchidae
3648:
3572:
3531:Phosphatosaurus
3468:Cerrejonisuchus
3433:Acherontisuchus
3418:
3327:Pholidosauridae
3309:
3184:
3171:
3146:
3126:
3095:
3054:
3053:
3043:
3038:
2995:Wayback Machine
2984:
2967:
2932:
2922:
2921:
2875:
2874:
2870:
2849:(5): 1095–116.
2839:
2838:
2834:
2824:
2820:
2806:
2805:
2801:
2791:
2789:
2788:on June 8, 2010
2775:
2774:
2770:
2723:
2722:
2718:
2674:
2673:
2669:
2655:
2654:
2650:
2603:
2602:
2598:
2558:
2553:
2552:
2548:
2514:
2513:
2509:
2500:
2496:
2460:
2459:
2455:
2411:
2410:
2401:
2339:
2338:
2321:
2284:(4): e1671427.
2273:
2268:
2267:
2263:
2240:Phosphatosaurus
2229:
2228:
2224:
2214:
2212:
2202:10.1139/e05-008
2171:
2170:
2157:
2152:
2143:Cerrejonisuchus
2138:
2129:
2117:Acherontisuchus
2113:Cerrejonisuchus
2101:Acherontisuchus
2097:Cerrejonisuchus
2045:
1993:
1988:
1983:
1974:
1965:
1956:
1947:
1938:
1929:
1920:
1911:
1902:
1893:
1884:
1762:
1753:
1606:
1413:
1392:
1390:
1357:
1355:
1291:
1289:
1288:
1279:
1277:
1240:
1238:
1237:
1228:
1226:
1213:Phosphatosaurus
1190:
1188:
1187:
1178:
1176:
1175:
1166:
1164:
1163:
1154:
1152:
1107:
1105:
1067:
1065:
1064:
1055:
1053:
1021:
1019:
978:
976:
975:
966:
964:
924:
922:
909:Cerrejonisuchus
889:
887:
883:Late Cretaceous
854:
852:
819:
817:
781:
779:
743:
741:
705:
703:
690:Acherontisuchus
663:
654:
545:Restoration of
539:
529:Phosphatosaurus
372:Acherontisuchus
342:Phosphatosaurus
294:Cerrejonisuchus
264:
258:
245:
236:Crocodyliformes
223:Crocodylomorpha
139:
112:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
36:
35:
32:
24:
17:
12:
11:
5:
4697:
4695:
4687:
4686:
4681:
4676:
4671:
4666:
4661:
4656:
4646:
4645:
4639:
4638:
4636:
4635:
4622:
4609:
4596:
4583:
4568:
4552:
4550:
4544:
4543:
4538:
4529:
4528:
4526:
4525:
4515:
4508:
4505:
4504:
4501:
4500:
4493:
4491:
4488:
4487:
4484:
4483:
4481:
4480:
4479:
4478:
4464:
4462:
4456:
4455:
4453:
4452:
4445:
4438:
4430:
4428:
4426:Planocraniidae
4419:
4418:
4416:
4415:
4408:
4401:
4394:
4387:
4379:
4377:
4376:"Thoracosaurs"
4370:
4369:
4367:
4366:
4359:
4351:
4349:
4340:
4339:
4337:
4336:
4328:
4325:Massaliasuchus
4321:
4314:
4311:Ischyrochampsa
4307:
4299:
4296:Allodaposuchus
4292:
4284:
4282:
4273:
4272:
4270:
4269:
4262:
4255:
4248:
4240:
4237:Iharkutosuchus
4233:
4226:
4217:
4215:
4206:
4205:
4203:
4202:
4191:
4180:
4169:
4158:
4147:
4144:Pristichampsus
4137:
4127:
4116:
4106:
4096:
4093:Dolichochampsa
4086:
4076:
4065:
4059:
4054:
4047:
4046:
4041:
4034:
4033:
4030:
4029:
3992:
3990:
3987:
3986:
3983:
3982:
3980:
3979:
3978:
3977:
3971:
3969:
3963:
3962:
3960:
3959:
3952:
3945:
3938:
3931:
3924:
3917:
3910:
3903:
3896:
3889:
3881:
3879:
3870:
3869:
3867:
3866:
3859:
3851:
3849:
3840:
3839:
3837:
3836:
3828:
3821:
3814:
3806:
3799:
3796:Ophiussasuchus
3792:
3785:
3778:
3771:
3764:
3757:
3754:Denazinosuchus
3750:
3743:
3736:
3729:
3721:
3719:
3710:
3709:
3707:
3706:
3699:
3691:
3689:
3687:Paluxysuchidae
3680:
3679:
3677:
3676:
3669:
3661:
3659:
3650:
3649:
3647:
3646:
3639:
3632:
3625:
3618:
3611:
3604:
3597:
3594:Alligatorellus
3589:
3587:
3578:
3577:
3574:
3573:
3571:
3570:
3563:
3556:
3548:
3541:
3534:
3527:
3520:
3513:
3506:
3499:
3492:
3489:Dorbignysuchus
3485:
3478:
3471:
3464:
3457:
3450:
3443:
3440:Anthracosuchus
3436:
3428:
3426:
3420:
3419:
3417:
3416:
3413:Woodbinesuchus
3409:
3402:
3395:
3388:
3381:
3374:
3367:
3360:
3353:
3346:
3339:
3331:
3329:
3320:
3311:
3310:
3308:
3307:
3303:Thalattosuchia
3296:
3285:
3274:
3270:Tadzhikosuchus
3263:
3253:
3249:Stolokrosuchus
3242:
3239:Montsecosuchus
3232:
3222:
3212:
3202:
3191:
3185:
3180:
3173:
3172:
3167:
3160:
3159:
3156:
3155:
3152:
3151:
3148:
3147:
3145:
3144:
3143:
3142:
3136:
3134:
3128:
3127:
3125:
3124:
3123:
3122:
3113:
3107:
3097:
3096:
3092:
3091:
3082:
3073:
3067:
3061:
3052:
3045:
3044:
3039:
3037:
3036:
3029:
3022:
3014:
3008:
3007:
3002:
2997:
2983:
2982:External links
2980:
2979:
2978:
2965:
2945:(2): 281–294.
2930:
2920:
2919:
2868:
2832:
2818:
2799:
2768:
2716:
2687:(1): 139–162.
2667:
2648:
2596:
2569:(3): 764–768.
2546:
2507:
2494:
2473:(2): 281–294.
2453:
2399:
2352:(7): e102189.
2319:
2261:
2250:(2): 237–256.
2222:
2188:(3): 323–337.
2154:
2153:
2151:
2148:
2137:
2134:
2128:
2125:
2044:
2041:
2017:osteosclerosis
1992:
1989:
1987:
1984:
1980:
1979:
1976:
1975:
1971:
1970:
1967:
1966:
1962:
1961:
1958:
1957:
1953:
1952:
1949:
1948:
1944:
1943:
1940:
1939:
1935:
1934:
1931:
1930:
1926:
1925:
1922:
1921:
1917:
1916:
1913:
1912:
1908:
1907:
1904:
1903:
1899:
1898:
1895:
1894:
1890:
1889:
1886:
1885:
1881:
1880:
1877:
1876:
1867:
1864:
1863:
1860:
1859:
1850:
1845:
1843:
1840:
1839:
1836:
1835:
1826:
1821:
1819:
1816:
1815:
1812:
1811:
1802:
1797:
1795:
1792:
1791:
1788:
1787:
1778:
1773:
1771:
1768:
1767:
1764:
1763:
1759:
1758:
1755:
1754:
1750:
1749:
1746:
1745:
1736:
1733:
1732:
1729:
1728:
1719:
1714:
1712:
1709:
1708:
1705:
1704:
1695:
1692:
1691:
1688:
1687:
1678:
1673:
1671:
1666:
1664:
1661:
1660:
1657:
1656:
1647:
1642:
1640:
1636:
1635:
1632:
1631:
1622:
1617:
1615:
1612:
1611:
1608:
1607:
1603:
1602:
1599:
1598:
1589:
1586:
1585:
1582:
1581:
1572:
1567:
1565:
1559:
1557:
1554:
1553:
1550:
1549:
1540:
1535:
1533:
1530:
1529:
1526:
1525:
1516:
1511:
1509:
1506:
1505:
1502:
1501:
1492:
1487:
1485:
1477:
1457:
1456:
1453:
1450:
1447:
1444:
1435:
1432:
1425:synapomorphies
1412:
1409:
1406:
1405:
1403:
1401:
1388:
1383:
1380:
1372:
1371:
1369:
1366:
1353:
1348:
1345:
1337:
1336:
1334:
1326:
1324:
1322:
1320:Junior synonym
1317:
1309:
1308:
1306:
1300:
1275:
1266:
1263:
1255:
1254:
1252:
1249:
1224:
1219:
1216:
1208:
1207:
1205:
1199:
1150:
1141:
1138:
1130:
1129:
1122:
1116:
1103:
1098:
1095:
1087:
1086:
1079:
1076:
1051:
1046:
1043:
1035:
1034:
1032:
1030:
1017:
1012:
1009:
1001:
1000:
993:
987:
962:
953:
950:
942:
941:
939:
933:
920:
915:
912:
904:
903:
901:
898:
885:
880:
877:
869:
868:
866:
863:
850:
845:
842:
834:
833:
831:
828:
815:
810:
807:
799:
798:
796:
790:
777:
772:
769:
766:Anthracosuchus
761:
760:
758:
752:
739:
734:
731:
723:
722:
720:
714:
701:
696:
693:
685:
684:
681:
678:
675:
672:
669:
662:
659:
653:
650:
556:external naris
538:
535:
509:crocodyliforms
490:
489:
488:
487:
486:
485:
475:
465:
455:
445:
435:
432:Dorbignysuchus
425:
415:
405:
395:
385:
375:
357:
356:
355:
345:
327:
317:
307:
297:
287:
284:Anthracosuchus
274:
273:
269:
268:
256:
252:
251:
243:
239:
238:
233:
226:
225:
220:
213:
212:
207:
200:
199:
194:
187:
186:
181:
177:
176:
171:
167:
166:
161:
157:
156:
151:
147:
146:
133:
132:
123:
122:
114:
113:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
42:
33:
15:
13:
10:
9:
6:
4:
3:
2:
4696:
4685:
4682:
4680:
4677:
4675:
4672:
4670:
4667:
4665:
4662:
4660:
4657:
4655:
4652:
4651:
4649:
4632:
4627:
4623:
4619:
4614:
4610:
4606:
4601:
4597:
4593:
4588:
4584:
4579:
4573:
4569:
4564:
4558:
4554:
4553:
4551:
4549:
4545:
4541:
4536:
4524:
4516:
4514:
4510:
4509:
4506:
4497:
4476:
4472:
4468:
4467:
4466:
4465:
4463:
4461:
4457:
4451:
4450:
4446:
4444:
4443:
4439:
4437:
4436:
4432:
4431:
4429:
4427:
4420:
4414:
4413:
4412:Thoracosaurus
4409:
4407:
4406:
4402:
4400:
4399:
4395:
4393:
4392:
4388:
4386:
4385:
4381:
4380:
4378:
4371:
4365:
4364:
4360:
4358:
4357:
4356:Aegyptosuchus
4353:
4352:
4350:
4348:
4341:
4334:
4333:
4329:
4327:
4326:
4322:
4320:
4319:
4318:Lohuecosuchus
4315:
4313:
4312:
4308:
4305:
4304:
4300:
4298:
4297:
4293:
4291:
4290:
4286:
4285:
4283:
4281:
4274:
4268:
4267:
4263:
4261:
4260:
4256:
4254:
4253:
4249:
4246:
4245:
4241:
4239:
4238:
4234:
4232:
4231:
4230:Hylaeochampsa
4227:
4224:
4223:
4219:
4218:
4216:
4214:
4207:
4200:
4199:
4192:
4189:
4188:
4181:
4178:
4177:
4176:Atoposauridae
4170:
4167:
4166:
4165:Titanochampsa
4159:
4156:
4155:
4148:
4146:
4145:
4138:
4136:
4135:
4128:
4125:
4124:
4117:
4115:
4114:
4107:
4105:
4104:
4097:
4095:
4094:
4087:
4085:
4084:
4077:
4075:
4074:
4073:Borealosuchus
4067:
4066:
4063:
4060:
4057:
4052:
4048:
4044:
4039:
4035:
4025:
4020:
4017:
4010:
4004:
3997:
3975:
3974:
3973:
3972:
3970:
3968:
3964:
3958:
3957:
3953:
3951:
3950:
3946:
3944:
3943:
3939:
3937:
3936:
3932:
3930:
3929:
3925:
3923:
3922:
3918:
3916:
3915:
3911:
3909:
3908:
3904:
3902:
3901:
3897:
3895:
3894:
3890:
3888:
3887:
3886:Batrachomimus
3883:
3882:
3880:
3878:
3871:
3865:
3864:
3860:
3858:
3857:
3853:
3852:
3850:
3848:
3841:
3834:
3833:
3829:
3827:
3826:
3822:
3820:
3819:
3815:
3812:
3811:
3807:
3805:
3804:
3800:
3798:
3797:
3793:
3791:
3790:
3786:
3784:
3783:
3779:
3777:
3776:
3772:
3770:
3769:
3765:
3763:
3762:
3758:
3756:
3755:
3751:
3749:
3748:
3744:
3742:
3741:
3740:Calsoyasuchus
3737:
3735:
3734:
3730:
3728:
3727:
3723:
3722:
3720:
3718:
3711:
3705:
3704:
3700:
3698:
3697:
3693:
3692:
3690:
3688:
3681:
3675:
3674:
3673:Stomatosuchus
3670:
3668:
3667:
3663:
3662:
3660:
3658:
3651:
3645:
3644:
3640:
3638:
3637:
3633:
3631:
3630:
3626:
3624:
3623:
3619:
3617:
3616:
3612:
3610:
3609:
3605:
3603:
3602:
3598:
3596:
3595:
3591:
3590:
3588:
3586:
3585:Atoposauridae
3579:
3569:
3568:
3567:Tilemsisuchus
3564:
3562:
3561:
3557:
3554:
3553:
3549:
3547:
3546:
3542:
3540:
3539:
3538:Rhabdognathus
3535:
3533:
3532:
3528:
3526:
3525:
3521:
3519:
3518:
3514:
3512:
3511:
3510:Guarinisuchus
3507:
3505:
3504:
3500:
3498:
3497:
3493:
3491:
3490:
3486:
3484:
3483:
3479:
3477:
3476:
3475:Chenanisuchus
3472:
3470:
3469:
3465:
3463:
3462:
3461:Brachiosuchus
3458:
3456:
3455:
3454:Atlantosuchus
3451:
3449:
3448:
3444:
3442:
3441:
3437:
3435:
3434:
3430:
3429:
3427:
3425:
3421:
3415:
3414:
3410:
3408:
3407:
3403:
3401:
3400:
3396:
3394:
3393:
3389:
3387:
3386:
3385:Pholidosaurus
3382:
3380:
3379:
3375:
3373:
3372:
3371:Meridiosaurus
3368:
3366:
3365:
3361:
3359:
3358:
3354:
3352:
3351:
3350:Crocodilaemus
3347:
3345:
3344:
3340:
3338:
3337:
3333:
3332:
3330:
3328:
3324:
3321:
3319:
3312:
3305:
3304:
3297:
3294:
3293:
3286:
3283:
3282:
3275:
3272:
3271:
3264:
3262:
3261:
3254:
3251:
3250:
3243:
3241:
3240:
3233:
3231:
3230:
3229:Khoratosuchus
3223:
3221:
3220:
3219:Karatausuchus
3213:
3211:
3210:
3203:
3201:
3200:
3193:
3192:
3189:
3186:
3183:
3178:
3174:
3170:
3165:
3161:
3140:
3139:
3138:
3137:
3135:
3133:
3129:
3121:
3117:
3116:
3115:
3114:
3111:
3108:
3106:
3102:
3098:
3090:
3086:
3083:
3081:
3077:
3074:
3072:
3068:
3066:
3062:
3060:
3056:
3055:
3050:
3046:
3042:
3035:
3030:
3028:
3023:
3021:
3016:
3015:
3012:
3006:
3003:
3001:
2998:
2996:
2992:
2989:
2986:
2985:
2981:
2976:(3): 581–594.
2975:
2971:
2966:
2961:
2956:
2952:
2948:
2944:
2940:
2939:Palaeontology
2936:
2931:
2928:
2924:
2923:
2915:
2911:
2906:
2901:
2896:
2891:
2887:
2883:
2879:
2872:
2869:
2864:
2860:
2856:
2852:
2848:
2844:
2843:Palaeontology
2836:
2833:
2829:
2822:
2819:
2814:
2810:
2803:
2800:
2787:
2783:
2779:
2772:
2769:
2764:
2760:
2756:
2752:
2748:
2744:
2740:
2736:
2732:
2728:
2720:
2717:
2712:
2708:
2703:
2698:
2694:
2690:
2686:
2682:
2678:
2671:
2668:
2663:
2659:
2652:
2649:
2644:
2640:
2636:
2632:
2628:
2624:
2620:
2616:
2612:
2608:
2600:
2597:
2592:
2588:
2584:
2580:
2576:
2572:
2568:
2564:
2557:
2550:
2547:
2542:
2538:
2534:
2530:
2526:
2522:
2518:
2511:
2508:
2504:
2498:
2495:
2489:
2484:
2480:
2476:
2472:
2468:
2467:Palaeontology
2464:
2457:
2454:
2449:
2445:
2440:
2435:
2431:
2427:
2423:
2419:
2415:
2408:
2406:
2404:
2400:
2395:
2391:
2386:
2381:
2377:
2373:
2368:
2363:
2359:
2355:
2351:
2347:
2343:
2336:
2334:
2332:
2330:
2328:
2326:
2324:
2320:
2315:
2311:
2307:
2303:
2299:
2295:
2291:
2287:
2283:
2279:
2272:
2265:
2262:
2257:
2253:
2249:
2245:
2241:
2237:
2236:Rhabdognathus
2233:
2226:
2223:
2211:
2207:
2203:
2199:
2195:
2191:
2187:
2183:
2179:
2177:
2168:
2166:
2164:
2162:
2160:
2156:
2149:
2147:
2145:
2144:
2135:
2133:
2126:
2124:
2122:
2118:
2114:
2110:
2106:
2102:
2098:
2094:
2090:
2086:
2082:
2078:
2074:
2069:
2067:
2066:Maastrichtian
2063:
2058:
2054:
2049:
2042:
2040:
2038:
2034:
2029:
2027:
2023:
2018:
2014:
2010:
2006:
1997:
1990:
1986:Palaeobiology
1985:
1978:
1977:
1969:
1968:
1960:
1959:
1951:
1950:
1942:
1941:
1933:
1932:
1924:
1923:
1915:
1914:
1906:
1905:
1897:
1896:
1888:
1887:
1879:
1878:
1875:
1874:
1873:
1866:
1865:
1862:
1861:
1858:
1857:
1856:
1849:
1848:
1842:
1841:
1838:
1837:
1834:
1833:
1832:
1825:
1824:
1818:
1817:
1814:
1813:
1810:
1809:
1808:
1801:
1800:
1794:
1793:
1790:
1789:
1786:
1785:
1784:
1777:
1776:
1770:
1769:
1766:
1765:
1757:
1756:
1748:
1747:
1744:
1743:
1742:
1735:
1734:
1731:
1730:
1727:
1726:
1725:
1718:
1717:
1711:
1710:
1707:
1706:
1703:
1702:
1701:
1694:
1693:
1690:
1689:
1686:
1685:
1684:
1677:
1676:
1670:
1669:
1663:
1662:
1659:
1658:
1655:
1654:
1653:
1646:
1645:
1639:Hyposaurinae
1638:
1637:
1634:
1633:
1630:
1629:
1628:
1621:
1620:
1614:
1613:
1610:
1609:
1601:
1600:
1597:
1596:
1595:
1588:
1587:
1584:
1583:
1580:
1579:
1578:
1571:
1570:
1563:
1562:
1556:
1555:
1552:
1551:
1548:
1547:
1546:
1539:
1538:
1532:
1531:
1528:
1527:
1524:
1523:
1522:
1515:
1514:
1508:
1507:
1504:
1503:
1500:
1499:
1498:
1491:
1490:
1484:
1481:
1480:
1476:
1474:
1473:
1468:
1467:
1462:
1454:
1451:
1448:
1445:
1443:
1439:
1436:
1433:
1430:
1429:
1428:
1426:
1422:
1418:
1410:
1404:
1402:
1400:
1389:
1387:
1384:
1381:
1379:
1378:
1377:Tilemsisuchus
1374:
1373:
1370:
1367:
1365:
1354:
1352:
1351:Maastrichtian
1349:
1346:
1344:
1343:
1339:
1338:
1335:
1333:
1332:
1327:
1325:
1323:
1321:
1318:
1316:
1315:
1311:
1310:
1307:
1305:
1304:K–Pg boundary
1301:
1299:
1287:
1276:
1274:
1270:
1269:Maastrichtian
1267:
1264:
1262:
1261:
1260:Rhabdognathus
1257:
1256:
1253:
1250:
1248:
1236:
1225:
1223:
1220:
1217:
1215:
1214:
1210:
1209:
1206:
1204:
1203:K–Pg boundary
1200:
1198:
1186:
1174:
1162:
1151:
1149:
1145:
1144:Maastrichtian
1142:
1139:
1137:
1136:
1132:
1131:
1127:
1123:
1121:
1117:
1115:
1104:
1102:
1099:
1096:
1094:
1093:
1092:Guarinisuchus
1089:
1088:
1084:
1080:
1077:
1075:
1063:
1052:
1050:
1047:
1044:
1042:
1041:
1037:
1036:
1033:
1031:
1029:
1018:
1016:
1013:
1010:
1008:
1007:
1003:
1002:
998:
994:
992:
991:K–Pg boundary
988:
986:
974:
963:
961:
957:
956:Maastrichtian
954:
951:
949:
948:
947:Chenanisuchus
944:
943:
940:
938:
934:
932:
921:
919:
916:
913:
911:
910:
906:
905:
902:
899:
897:
886:
884:
881:
878:
876:
875:
874:Brachiosuchus
871:
870:
867:
864:
862:
851:
849:
846:
843:
841:
840:
839:Atlantosuchus
836:
835:
832:
829:
827:
816:
814:
811:
808:
806:
805:
801:
800:
797:
795:
791:
789:
778:
776:
773:
770:
768:
767:
763:
762:
759:
757:
753:
751:
740:
738:
735:
732:
730:
729:
728:Aigialosuchus
725:
724:
721:
719:
715:
713:
702:
700:
697:
694:
692:
691:
687:
686:
682:
676:
673:
670:
667:
666:
660:
658:
651:
649:
646:
642:
638:
637:Atlantosuchus
634:
633:Rhabdognathus
625:
621:
618:
614:
609:
607:
602:
600:
595:
591:
582:
578:
576:
571:
569:
565:
561:
557:
550:
549:
543:
536:
534:
531:
530:
524:
522:
518:
514:
510:
507:
504:
500:
496:
484:
483:
482:Tilemsisuchus
476:
474:
473:
472:Rhabdognathus
466:
464:
463:
456:
454:
453:
446:
444:
443:
436:
434:
433:
426:
424:
423:
416:
414:
413:
412:Brachiosuchus
406:
404:
403:
402:Atlantosuchus
396:
394:
393:
386:
384:
383:
382:Aigialosuchus
376:
374:
373:
366:
365:
364:
358:
354:
353:
346:
344:
343:
336:
335:
334:
328:
326:
325:
318:
316:
315:
314:Guarinisuchus
308:
306:
305:
304:Chenanisuchus
298:
296:
295:
288:
286:
285:
278:
277:
275:
270:
263:
257:
254:
253:
250:
244:
241:
240:
237:
234:
231:
228:
227:
224:
221:
218:
215:
214:
211:
208:
205:
202:
201:
198:
195:
192:
189:
188:
185:
182:
179:
178:
175:
172:
169:
168:
165:
162:
159:
158:
155:
152:
149:
148:
143:
138:
134:
131:
130:
124:
120:
115:
111:
107:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
40:
37:83.5–47
29:
26:
22:
4578:Dyrosauridae
4548:Dyrosauridae
4547:
4447:
4440:
4435:Boverisuchus
4433:
4410:
4403:
4396:
4389:
4382:
4361:
4354:
4330:
4323:
4316:
4309:
4301:
4294:
4287:
4264:
4257:
4250:
4242:
4235:
4228:
4220:
4196:
4185:
4174:
4163:
4152:
4142:
4132:
4121:
4113:Heterosuchus
4111:
4101:
4091:
4081:
4071:
4000:
3954:
3949:Wannchampsus
3947:
3942:Turanosuchus
3940:
3933:
3926:
3919:
3912:
3907:Paralligator
3905:
3900:Kansajsuchus
3898:
3891:
3884:
3861:
3856:Bernissartia
3854:
3832:Symptosuchus
3830:
3823:
3816:
3810:Pinacosuchus
3808:
3803:Oweniasuchus
3801:
3794:
3787:
3780:
3773:
3766:
3759:
3752:
3745:
3738:
3731:
3726:Amphicotylus
3724:
3703:Paluxysuchus
3701:
3694:
3671:
3666:Laganosuchus
3664:
3643:Varanosuchus
3641:
3636:Theriosuchus
3634:
3627:
3620:
3613:
3606:
3601:Alligatorium
3599:
3592:
3565:
3560:Sokotosuchus
3558:
3552:Sabinosuchus
3550:
3543:
3536:
3529:
3522:
3515:
3508:
3503:Fortignathus
3501:
3494:
3487:
3480:
3473:
3466:
3459:
3452:
3445:
3438:
3431:
3424:Dyrosauridae
3423:
3411:
3404:
3399:Terminonaris
3397:
3390:
3383:
3378:Oceanosuchus
3376:
3369:
3362:
3357:Dakotasuchus
3355:
3348:
3341:
3334:
3301:
3290:
3279:
3268:
3258:
3247:
3237:
3227:
3217:
3207:
3197:
3120:Pseudosuchia
3105:Pseudosuchia
3084:
3080:Pseudosuchia
3075:
2973:
2969:
2942:
2938:
2926:
2885:
2881:
2871:
2846:
2842:
2835:
2827:
2821:
2812:
2808:
2802:
2790:. Retrieved
2786:the original
2781:
2771:
2730:
2726:
2719:
2684:
2680:
2670:
2661:
2657:
2651:
2610:
2606:
2599:
2566:
2562:
2549:
2524:
2520:
2510:
2502:
2497:
2470:
2466:
2456:
2421:
2417:
2349:
2345:
2281:
2277:
2264:
2247:
2243:
2239:
2235:
2231:
2225:
2213:. Retrieved
2185:
2181:
2175:
2141:
2139:
2130:
2127:Reproduction
2116:
2112:
2100:
2096:
2091:, inhabited
2089:Saudi Arabia
2070:
2050:
2046:
2030:
2009:pachyostotic
2005:osteoporotic
2002:
1870:
1869:
1853:
1852:
1829:
1828:
1805:
1804:
1781:
1780:
1739:
1738:
1722:
1721:
1698:
1697:
1681:
1680:
1650:
1649:
1625:
1624:
1592:
1591:
1575:
1574:
1543:
1542:
1519:
1518:
1495:
1494:
1483:Dyrosauridae
1482:
1472:Fortignathus
1470:
1466:Sabinosuchus
1464:
1460:
1458:
1416:
1414:
1375:
1342:Sokotosuchus
1340:
1329:
1314:Sokotosaurus
1312:
1258:
1211:
1133:
1119:
1090:
1038:
1004:
945:
907:
872:
837:
802:
764:
726:
688:
680:Description
655:
644:
640:
636:
632:
630:
627:Jaw fragment
610:
603:
587:
572:
553:
546:
527:
525:
495:Dyrosauridae
494:
493:
480:
470:
460:
450:
440:
430:
420:
410:
400:
390:
380:
370:
363:Hyposaurinae
362:
352:Sokotosuchus
350:
340:
332:
322:
312:
302:
292:
282:
262:Dyrosauridae
261:
229:
216:
210:Pseudosuchia
203:
190:
127:
31:Dyrosauridae
25:
21:Dryosauridae
4654:Dyrosaurids
4572:Wikispecies
4449:Planocrania
4442:Duerosuchus
4384:Argochampsa
4303:Arenysuchus
4289:Agaresuchus
4259:Turcosuchus
3956:Yanjisuchus
3935:Tarsomordeo
3928:Shamosuchus
3921:Scolomastax
3818:Siamosuchus
3789:Nannosuchus
3782:Hulkepholis
3775:Goniopholis
3761:Diplosaurus
3747:Coelosuchus
3696:Deltasuchus
3629:Sabresuchus
3615:Atoposaurus
3545:Rodeosuchus
3524:Luciasuchus
3482:Congosaurus
3406:Vectisuchus
3392:Sarcosuchus
3336:Anglosuchus
3318:Tethysuchia
3292:Zhyrasuchus
3199:Burkesuchus
2792:February 3,
2140:A study on
2121:much warmer
2026:musculature
1438:Exoccipital
1006:Congosaurus
599:posteriorly
560:premaxillae
462:Luciasuchus
422:Congosaurus
324:Rodeosuchus
249:Tethysuchia
197:Archosauria
4648:Categories
4511:See also:
4471:Crocodilia
4460:Crocodilia
4391:Eogavialis
4363:Aegisuchus
4154:Susisuchus
4123:Isisfordia
3976:see below↓
3914:Rugosuchus
3825:Sunosuchus
3608:Aprosuchus
3517:Hyposaurus
3496:Dyrosaurus
3281:Wahasuchus
3260:Susisuchus
3209:Isisfordia
3141:see below↓
3089:Metasuchia
3071:Sauropsida
2888:: e11222.
2815:: 262–283.
2664:: 183–202.
2232:Dyrosaurus
2150:References
2136:Locomotion
2062:Cenomanian
2022:undulatory
1331:Hyposaurus
1135:Hyposaurus
1040:Dyrosaurus
641:Dyrosaurus
590:premaxilla
568:nasal bone
548:Dyrosaurus
506:neosuchian
452:Hyposaurus
442:Dyrosaurus
242:Suborder:
4266:Unasuchus
3364:Elosuchus
3182:Neosuchia
3169:Neosuchia
3132:Neosuchia
3057:Kingdom:
3041:Neosuchia
2643:130987967
2635:0272-4634
2583:0272-4634
2541:0024-4082
2376:1932-6203
2314:209439989
2306:0272-4634
2215:March 22,
2210:0008-4077
2093:estuarine
2053:Paleogene
2035:and left
1411:Phylogeny
1273:Paleogene
1148:Paleocene
1101:Paleocene
1015:Paleocene
960:Paleocene
918:Paleocene
848:Paleocene
813:Paleocene
775:Paleocene
737:Campanian
699:Paleocene
677:Location
615:are deep
613:dentition
513:Campanian
160:Kingdom:
154:Eukaryota
106:Campanian
4557:Wikidata
4523:Category
4398:Eosuchus
4222:Acynodon
4056:Eusuchia
4043:Eusuchia
3967:Eusuchia
3343:Chalawan
3065:Chordata
3063:Phylum:
3059:Animalia
2991:Archived
2914:34026348
2755:19194448
2711:84705605
2591:16525132
2448:18364311
2394:25032965
2346:PLOS ONE
2109:Colombia
2081:Pakistan
931:Colombia
788:Colombia
712:Colombia
617:occlusal
575:lacrimal
564:maxillae
537:Overview
255:Family:
184:Reptilia
174:Chordata
170:Phylum:
164:Animalia
150:Domain:
4605:4822244
4592:4472954
4563:Q548118
3069:Class:
2947:Bibcode
2905:8117932
2851:Bibcode
2763:4381423
2735:Bibcode
2689:Bibcode
2615:Bibcode
2475:Bibcode
2439:2602706
2385:4102515
2354:Bibcode
2286:Bibcode
2190:Bibcode
2077:fluvial
2043:Habitat
2013:ecology
1364:Nigeria
1298:Nigeria
1247:Tunisia
1185:Nigeria
1074:Tunisia
1062:Algeria
985:Morocco
861:Morocco
826:Morocco
683:Images
671:Status
661:General
594:maxilla
515:to the
503:extinct
272:Genera
180:Class:
4618:111943
2912:
2902:
2761:
2753:
2727:Nature
2709:
2641:
2633:
2589:
2581:
2539:
2505:(2008)
2503:et al.
2501:Jouve
2446:
2436:
2392:
2382:
2374:
2312:
2304:
2208:
1991:Growth
1461:et al.
1459:Jouve
1417:et al.
1415:Jouve
1396:
1386:Eocene
1382:Valid
1361:
1347:Valid
1295:
1283:
1265:Valid
1244:
1232:
1222:Eocene
1218:Valid
1194:
1182:
1170:
1161:Brazil
1158:
1140:Valid
1114:Brazil
1111:
1097:Valid
1071:
1059:
1049:Eocene
1045:Valid
1028:Angola
1025:
1011:Valid
982:
970:
952:Valid
928:
914:Valid
893:
879:Valid
858:
844:Valid
823:
809:Valid
785:
771:Valid
750:Sweden
747:
733:Valid
709:
695:Valid
668:Genus
517:Eocene
499:family
110:Eocene
4631:38370
4613:IRMNG
3085:Clade
3076:Clade
2882:PeerJ
2759:S2CID
2707:S2CID
2639:S2CID
2587:S2CID
2559:(PDF)
2310:S2CID
2274:(PDF)
2073:Sudan
2057:clade
2037:tibia
2033:femur
1999:Skull
1421:clade
896:Sudan
606:jugal
584:Tooth
497:is a
230:Clade
217:Clade
204:Clade
191:Clade
4600:GBIF
4469:see
3118:see
2910:PMID
2794:2010
2751:PMID
2631:ISSN
2579:ISSN
2537:ISSN
2444:PMID
2390:PMID
2372:ISSN
2302:ISSN
2217:2021
2206:ISSN
2115:and
2099:and
1469:and
1399:Mali
1286:Mali
1235:Mali
1173:Mali
973:Mali
674:Age
643:and
635:and
45:PreꞒ
4587:EoL
2955:doi
2927:109
2900:PMC
2890:doi
2859:doi
2813:156
2743:doi
2731:457
2697:doi
2662:120
2623:doi
2571:doi
2529:doi
2525:192
2483:doi
2434:PMC
2426:doi
2422:275
2380:PMC
2362:doi
2294:doi
2252:doi
2242:".
2198:doi
2087:in
2007:or
501:of
4650::
4628::
4615::
4602::
4589::
4574::
4559::
3087::
3078::
2974:50
2972:.
2953:.
2943:51
2941:.
2937:.
2908:.
2898:.
2884:.
2880:.
2857:.
2847:54
2845:.
2811:.
2780:.
2757:.
2749:.
2741:.
2729:.
2705:.
2695:.
2685:30
2683:.
2679:.
2660:.
2637:.
2629:.
2621:.
2611:29
2609:.
2585:.
2577:.
2567:24
2565:.
2561:.
2535:.
2523:.
2519:.
2481:.
2471:51
2469:.
2465:.
2442:.
2432:.
2420:.
2416:.
2402:^
2388:.
2378:.
2370:.
2360:.
2348:.
2344:.
2322:^
2308:.
2300:.
2292:.
2282:39
2280:.
2276:.
2246:.
2238:,
2234:,
2204:.
2196:.
2186:42
2184:.
2180:.
2158:^
1197:US
639:,
570:.
523:.
232::
219::
206::
193::
95:Pg
39:Ma
4423:†
4374:†
4344:†
4335:?
4306:?
4277:†
4247:?
4225:?
4210:†
4201:?
4194:†
4190:?
4183:†
4179:?
4172:†
4168:?
4161:†
4157:?
4150:†
4140:†
4130:†
4126:?
4119:†
4109:†
4099:†
4089:†
4079:†
4069:†
3874:†
3844:†
3835:?
3813:?
3714:†
3684:†
3654:†
3582:†
3555:?
3315:†
3306:?
3299:†
3295:?
3288:†
3284:?
3277:†
3273:?
3266:†
3256:†
3252:?
3245:†
3235:†
3225:†
3215:†
3205:†
3195:†
3033:e
3026:t
3019:v
2963:.
2957::
2949::
2929:.
2916:.
2892::
2886:9
2865:.
2861::
2853::
2796:.
2765:.
2745::
2737::
2713:.
2699::
2691::
2645:.
2625::
2617::
2593:.
2573::
2543:.
2531::
2491:.
2485::
2477::
2450:.
2428::
2396:.
2364::
2356::
2350:9
2316:.
2296::
2288::
2258:.
2254::
2248:5
2219:.
2200::
2192::
1271:–
1146:–
958:–
478:†
468:†
458:†
448:†
438:†
428:†
418:†
408:†
398:†
388:†
378:†
368:†
360:†
348:†
338:†
330:†
320:†
310:†
300:†
290:†
280:†
259:†
246:†
108:–
100:N
90:K
85:J
80:T
75:P
70:C
65:D
60:S
55:O
50:Ꞓ
23:.
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