4673:
6805:
1931:
4666:
3170:
3995:
133:
3981:
1904:
6257:
5461:
6243:
435:
1858:
119:
3974:
6250:
3176:
4680:
151:
3988:
1400:
2065:
6818:
5479:
1378:
5467:
1829:
5486:
6812:
1580:
6237:
1755:
1541:
1427:
581:
1711:
1633:
1512:
5474:
1483:
6925:
1733:
1689:
505:
1454:
529:
The holotypic specimen was disarticulated prior to fossilisation, with many elements of its skeleton scattered throughout the assemblage: none of the vertebral column is preserved in articulation and those vertebrae that are preserved consist of separated neural arches and centra, signifying that the
1350:
is not a megaraptoran itself according to the authors' research, with
Megaraptora representing a second wave of large-bodied tyrannosauroids that were important members of the world's ecosystem, one that may have originally slowed the evolutionary radiation of tyrannosaurids initially.
2603:
Sweetman, Steven & N. Insole, Allan. (2010). The plant debris beds of the Early
Cretaceous (Barremian) Wessex Formation of the Isle of Wight, southern England: Their genesis and palaeontological significance. Palaeogeography, Palaeoclimatology, Palaeoecology. 292. 409-424.
520:
alveolus and a dorsally-directed prong on the rostromesial margin of the notch, curving lateral furrows on the lateral surface of the dentary, a surangular with a hypertrophied gutter-like concavity near the rostrodorsal border, with the caudal end of the concavity containing
467:. Gavin Leng revealed the location of where the claw was discovered, and over the next few weeks the site was carefully excavated, and the fossils removed in a hard matrix. Over the next few years the specimen was carefully researched with scientists from the
2122:
flora was both fire and drought resistant and suggested that it was adapted to a seasonal climate with periods of marked aridity. Evidence for a wet season is provided by the frequent occurrence of fungal decay in plant material from the plant debris beds.
454:
specimen has not been revealed due to its importance and the possibility of new material to be collected as the coastline recedes. From what is mentioned in the description, the specimen was found on the southwestern coast of the Isle of Wight, between
594:
corroborates the notion that early tyrannosauroids were gracile with long forelimbs and three-fingered grasping hands, although the somewhat large size of the animal either means that early evolution for this clade was carried out at a large size or
571:
Although the specimen itself measures up to 4.5 m (15 ft) in length, it doesn't represent the size of an adult; the specimen likely belongs to a subadult due to the lack of fusion regarding the neurocentral and sacral sutures.
525:
that perforate the body of the surangular, a low coronoid process on the surangular with a concave area located caudodorsally, and an ulna and radius with a tear-drop shaped cross-section at the mid-shaft.
422:. The remains (MIWG1997.550), consisting of assorted skull, axial skeleton and appendicular skeleton elements, from a juvenile or subadult, found in a plant debris clay bed, were described by Hutt
426:
in early 2001. The etymology of the generic name refers to the animal's classification as an early tyrannosaur or "tyrant lizard", while the specific name honors the discoverer of the fossil.
533:
Due to the relative low-quality preservation of many of the skeletal elements, numerous pieces discovered have been difficult to identify: these include unidentified
6955:
463:. In 1995, local collector Gavin Leng brought a claw he had found along the coastline to Steve Hutt who worked at the old Museum of Isle of Wight Geology at
490:
fundraiser in order to release a monograph of the specimen, which received well over its goal. The monograph was eventually published in the journal
6869:
1338:
as an intermediate gracile tyrannosauroid more closely related to the true tyrannosaurids; more advanced than proceratosaurids, stokesosaurids and
599:
developed large size independently. The find of this animal in Europe puts in question to the purported Asian origin for these animals along with
482:
was given its name along with its specific epithet in honour of Mr. Leng. The material was described briefly in 2001 by Hutt et al. In July 2018
4672:
6970:
4665:
2571:
6804:
2529:
Juan D. Porfiri; Fernando E. Novas; Jorge O. Calvo; Federico L. Agnolín; Martín D. Ezcurra; Ignacio A. Cerda (2014). "Juvenile specimen of
6975:
2632:
552:
Many of the characters also presented as unique to the genus in the diagnosis of Hutt et al. (2001) are in fact widespread throughout
2118:
with a mean annual temperature of 20–25 °C with low seasonal rainfall. Watson and Alvin (1996) and Allen (1998) showed that the
3994:
5460:
2254:
998:
3980:
6242:
3169:
472:
3973:
6256:
3175:
6683:
5478:
4679:
2114:, sedimentological evidence, as well as fossils such as mud-cracks, suggests that the area experienced a warm, equable
1930:
6965:
6960:
6249:
3987:
150:
6915:
6817:
2279:"A preliminary account of a new tyrannosauroid theropod from the Wessex Formation (Cretaceous) of southern England"
5485:
2720:
2394:
and Other Lower
Cretaceous Theropod Dinosaurs From England. Unpublished Ph.D. Thesis, University of Portsmouth.
1346:' placement in the family suggests that Megaraptora are tyrannosauroids as well, even though it was found that
468:
5466:
6811:
6950:
2625:
132:
6837:
6945:
6895:
5191:
5103:
308:
1903:
6236:
2449:
556:, for example the presence of 'serrated carinae on D-shaped premaxillary teeth' is far from unique to
6769:
4880:
4848:
2483:
549:
was reconstructed with much longer tibiae, which influenced the early reconstructions of the animal.
512:
A number of characters present in the holotypic specimen are unique to the genus. These include: The
2453:
1857:
434:
5204:
4991:
2407:(1994). "The phylogenetic position of the Tyrannosauridae: implications for theropod systematics".
2404:
2131:
460:
118:
5473:
6676:
5180:
5040:
4896:
4836:
4271:
4181:
4064:
2928:
2618:
2445:
2424:
513:
337:
145:
6900:
6882:
6558:
6475:
6449:
6288:
6275:
6153:
5694:
5646:
4478:
3828:
2907:
2511:
2373:
2049:
1448:
1399:
1342:, but without the characteristics of more advanced genera. Simultaneously, the description of
1046:
641:
6887:
6929:
6662:
6419:
6329:
6118:
6034:
5896:
5815:
5787:
5747:
5628:
5598:
5116:
4869:
4733:
4590:
4490:
4339:
4246:
4204:
3771:
3081:
2583:
2550:
2542:
2501:
2491:
2461:
2457:
2416:
2363:
2353:
2293:
2127:
2119:
2087:
2079:
2064:
1851:
1358:
821:
631:
584:
553:
456:
407:
404:
263:
247:
1377:
6725:
6463:
6426:
6358:
6213:
6206:
6018:
5960:
5933:
5923:
5882:
5717:
5576:
5560:
5378:
5350:
5328:
5215:
5060:
4767:
4541:
4424:
4402:
4323:
4152:
4129:
4122:
4100:
4086:
4079:
4040:
3891:
3738:
3665:
3654:
3590:
3478:
3365:
2942:
2851:
2705:
2680:
2592:
2164:
1926:
1705:
1626:
1228:
1071:
1029:
662:
565:
2487:
6690:
6653:
6634:
6596:
6585:
6382:
6095:
6054:
6045:
6027:
6007:
5903:
5617:
5362:
5224:
5051:
5033:
4861:
4828:
4814:
4775:
4549:
4410:
4367:
4360:
4282:
4258:
4211:
4170:
4107:
4093:
3821:
3806:
3633:
3467:
3331:
3299:
3288:
3265:
3227:
3088:
3058:
2982:
2921:
2820:
2506:
2469:
2465:
2368:
2337:
2228:
2180:
1881:
1828:
1669:
1214:
419:
394:
234:
2278:
6939:
6732:
6578:
6539:
6518:
6397:
6390:
6317:
6111:
6084:
5942:
5858:
5836:
5829:
5822:
5765:
5682:
5545:
5517:
5504:
5428:
5305:
5145:
4979:
4910:
4796:
4783:
4528:
4506:
4499:
4463:
4436:
4388:
4312:
4048:
3839:
3813:
3729:
3710:
3703:
3640:
3619:
3598:
3583:
3569:
3528:
3494:
3487:
3406:
3258:
3145:
3138:
3065:
3035:
3024:
2914:
2885:
2878:
2804:
2428:
2168:
2159:
2153:
2140:
2095:
2074:
1897:
1535:
1394:
965:
948:
770:
604:
600:
439:
415:
411:
69:
1579:
6704:
6642:
6612:
6532:
6525:
6503:
6495:
6375:
6367:
6347:
6062:
5973:
5911:
5801:
5772:
5724:
5553:
5538:
5313:
5245:
5173:
5152:
5131:
4947:
4903:
4711:
4698:
4628:
4581:
4456:
4346:
4298:
4291:
4228:
4197:
4190:
4071:
3898:
3750:
3647:
3549:
3542:
3437:
3421:
3357:
3308:
3240:
3017:
2900:
2893:
2864:
2827:
2796:
2210:
2186:
2149:
2144:
2115:
2111:
1823:
1603:
917:
729:
613:
560:. Furthermore, neither the presence of a laterally flattened rostral region to the
483:
2546:
2094:
was found, was considered to have been warm and humid, similar to the present-day
1754:
1540:
1426:
2496:
6758:
6739:
6669:
6619:
6605:
6547:
6488:
6433:
6310:
6104:
5980:
5949:
5865:
5851:
5583:
5531:
5399:
5321:
5284:
5166:
5138:
5091:
5011:
4889:
4746:
4597:
4570:
4470:
4417:
4353:
4332:
4305:
4056:
3869:
3862:
3848:
3674:
3521:
3512:
3414:
3385:
3315:
3280:
3273:
3073:
3051:
3044:
3003:
2995:
2971:
2956:
2935:
2843:
2835:
2232:
2219:
2204:
2198:
2192:
2053:
1656:
1560:
1201:
1187:
1171:
486:, a colleague of Hutt who helped produce the preliminary description, created a
372:
362:
351:
44:
2098:. However, there is evidence of a phase of increasing aridity during the late
6697:
6626:
6571:
6192:
5991:
5808:
5779:
5731:
5665:
5591:
5413:
5406:
5392:
5298:
5231:
5159:
5123:
5076:
5018:
5004:
4971:
4963:
4956:
4937:
4760:
4725:
4513:
4114:
3905:
3855:
3792:
3763:
3718:
3696:
3689:
3681:
3605:
3501:
3460:
3453:
3445:
3429:
3349:
3220:
3096:
2949:
2812:
2671:
2470:"Tyrant dinosaur evolution tracks the rise and fall of Late Cretaceous oceans"
2420:
2315:
2237:
2215:
2135:
1749:
1726:
1710:
1632:
1511:
1477:
1421:
1372:
1301:
1267:
1176:
869:
753:
580:
208:
89:
54:
6860:
6718:
6711:
6411:
6404:
6077:
6069:
6000:
5653:
5371:
4917:
4821:
4556:
4141:
4026:
4013:
3940:
3876:
3799:
3778:
3535:
3398:
3206:
3193:
3111:
2963:
2782:
2769:
2732:
2641:
2342:, a tyrannosauroid theropod from the Wealden Supergroup of southern England"
2174:
2130:
possessed a wide array of fauna, including many other dinosaurs such as the
2099:
1682:
1482:
1245:
451:
221:
162:
94:
38:
30:
2515:
2377:
2297:
1732:
1688:
6854:
6302:
5710:
5703:
5675:
5568:
5068:
4449:
3785:
3612:
3373:
3010:
2689:
2665:
2587:
2474:
2241:
893:
696:
679:
487:
400:
397:
195:
182:
84:
79:
64:
59:
49:
2358:
6874:
5843:
5605:
5335:
5238:
4563:
3884:
2555:
561:
522:
504:
464:
99:
74:
2277:
Hutt, S.; Naish, D.; Martill, D.M.; Barker, M.J.; Newbery, P. (2001).
537:
elements, as well as an “ulna” which has since been recognised as the
6511:
6199:
3626:
2659:
2533:(Dinosauria, Theropoda) sheds light about tyrannosauroid radiation".
2103:
609:
538:
517:
172:
6831:
1453:
5342:
5083:
5025:
2063:
1573:
579:
542:
534:
503:
492:
433:
390:
6775:
5385:
6835:
6273:
5502:
4696:
4011:
3191:
2767:
2652:
2614:
1334:
The 2022 osteology by Naish and Cau by comparison, classifies
617:
arguing for a more complex biogeography for tyrannosauroids.
2610:
2572:"A Unified Framework for Predatory Dinosaur Macroevolution"
516:
end of dentary possessing a concave notch housing the most
6913:
545:. Before the proper identification of this fragment,
2223:. There were many contemporary mammal species which
6844:
6757:
6652:
6595:
6557:
6474:
6447:
6357:
6327:
6286:
6184:
6152:
6094:
6044:
6017:
5990:
5959:
5932:
5921:
5880:
5745:
5693:
5664:
5626:
5615:
5515:
5427:
5361:
5283:
5214:
5190:
5102:
5050:
4990:
4946:
4935:
4879:
4847:
4794:
4744:
4709:
4627:
4580:
4527:
4489:
4435:
4386:
4322:
4281:
4257:
4226:
4180:
4150:
4139:
4024:
3939:
3838:
3749:
3728:
3664:
3567:
3511:
3477:
3384:
3329:
3298:
3238:
3204:
3130:
3110:
3034:
2981:
2862:
2780:
2731:
2704:
2390:Naish, D., (2006). The Osteology and Affinities of
624:in 2013 that includes most tyrannosauroid genera.
2576:Bollettino della Società Paleontologica Italiana
2626:
8:
2272:
2270:
2331:
2329:
2327:
2325:
2323:
2316:http://www.dinosaurisle.com/eotyrannus.aspx
450:The exact location of the discovery of the
6832:
6563:
6480:
6455:
6339:
6335:
6294:
6283:
6270:
6181:
6149:
5965:
5929:
5888:
5757:
5753:
5638:
5634:
5623:
5523:
5512:
5499:
5290:
5280:
5196:
5108:
4996:
4943:
4853:
4806:
4802:
4752:
4717:
4706:
4693:
4624:
4533:
4441:
4394:
4263:
4238:
4234:
4162:
4158:
4147:
4032:
4021:
4008:
3936:
3755:
3746:
3575:
3390:
3341:
3337:
3250:
3246:
3212:
3201:
3188:
2987:
2870:
2788:
2777:
2764:
2710:
2701:
2649:
2633:
2619:
2611:
2440:
2438:
131:
117:
20:
2554:
2505:
2495:
2367:
2357:
508:Size of the holotype compared to a human
414:, located in the southwest coast of the
6920:
2266:
530:holotype was an immature individual.
7:
6956:Early Cretaceous dinosaurs of Europe
1919:
1873:
1844:
1815:
1675:
1648:
1619:
1595:
1566:
1556:
1527:
1498:
1469:
1442:
1415:
1364:
1354:
1220:
1193:
1183:
1021:
1014:
990:
940:
933:
909:
885:
861:
837:
813:
745:
721:
654:
647:
637:
627:
14:
2338:"The osteology and affinities of
6923:
6816:
6810:
6803:
6255:
6248:
6241:
6235:
5484:
5477:
5472:
5465:
5459:
4678:
4671:
4664:
3993:
3986:
3979:
3972:
3174:
3168:
2336:Naish, D.; Cau, A. (July 2022).
2255:Timeline of tyrannosaur research
2078:, with other dinosaurs from the
1929:
1902:
1856:
1827:
1753:
1731:
1709:
1687:
1631:
1578:
1539:
1510:
1481:
1452:
1425:
1398:
1376:
999:Appalachiosaurus montgomeriensis
149:
2310:Price, T. (2018, November 26).
620:Below is a cladogram by Loewen
2044:In 2024, Andrea Cau recovered
446:was discovered at Grange Chine
1:
6971:Fossil taxa described in 2001
2604:10.1016/j.palaeo.2010.03.055.
2547:10.1016/j.cretres.2014.04.007
2227:likely fed on, including the
1174:closely related to taxa like
389:(meaning "dawn tyrant") is a
2497:10.1371/journal.pone.0079420
564:nor a pronounced rim to the
2052:, distant in relation from
6992:
6976:Taxa named by Darren Naish
16:Extinct genus of dinosaurs
6800:
6566:
6483:
6458:
6342:
6338:
6297:
6282:
6269:
6233:
6180:
6148:
5968:
5891:
5760:
5756:
5641:
5637:
5526:
5511:
5498:
5457:
5293:
5279:
5199:
5111:
4999:
4856:
4809:
4805:
4755:
4720:
4705:
4692:
4661:
4623:
4536:
4444:
4397:
4266:
4241:
4237:
4165:
4161:
4035:
4020:
4007:
3969:
3935:
3758:
3578:
3393:
3344:
3340:
3253:
3249:
3215:
3200:
3187:
3165:
2990:
2873:
2791:
2776:
2763:
2713:
2700:
2648:
2421:10.1017/S0022336000026706
1924:
1917:
1895:
1878:
1871:
1849:
1842:
1820:
1813:
1746:
1724:
1702:
1680:
1673:
1653:
1646:
1624:
1617:
1600:
1593:
1571:
1564:
1554:
1532:
1525:
1503:
1496:
1474:
1467:
1447:
1440:
1420:
1413:
1391:
1369:
1362:
1298:
1281:
1264:
1242:
1225:
1218:
1198:
1191:
1069:
1043:
1026:
1019:
1012:
995:
988:
962:
945:
938:
931:
914:
907:
890:
883:
866:
859:
842:
835:
818:
811:
767:
750:
743:
726:
719:
693:
676:
659:
652:
645:
635:
568:are unique to the genus.
471:, and with help from the
343:
336:
314:
307:
146:Scientific classification
144:
139:
130:
125:
116:
23:
966:Xiongguanlong baimoensis
918:Dryptosaurus aquilunguis
771:Stokesosaurus clevelandi
663:Proceratosaurus bradleyi
469:University of Portsmouth
126:Known skeletal elements
2409:Journal of Paleontology
730:Sinotyrannus kazuoensis
2593:Supplementary Material
2298:10.1006/cres.2001.0252
2083:
1166:A 2014 analysis found
587:
509:
473:Natural History Museum
447:
6896:Paleobiology Database
5192:Carcharodontosaurinae
5104:Carcharodontosauridae
2067:
2048:as a sister taxon of
583:
507:
437:
6185:Dubious coelurosaurs
4881:Metriacanthosaurinae
4849:Metriacanthosauridae
3131:Dubious neotheropods
2588:10.4435/BSPI.2024.08
2132:carcharodontosaurian
949:Alectrosaurus olseni
894:Raptorex kriegsteini
680:Kileskus aristotocus
478:Eventually in 2001,
430:Discovery and naming
5205:Carcharodontosaurus
4992:Carcharodontosauria
2535:Cretaceous Research
2488:2013PLoSO...879420L
2359:10.7717/peerj.12727
2286:Cretaceous Research
754:Juratyrant langhami
6966:Fossils of England
6961:Cretaceous England
6677:Archaeornithomimus
5181:Veterupristisaurus
5041:Veterupristisaurus
4897:Metriacanthosaurus
4837:Piatnitzkysauridae
4272:Eustreptospondylus
4182:Piatnitzkysauridae
4065:Chuandongocoelurus
2929:Notatesseraeraptor
2172:; the ornithopods
2148:; the medium size
2084:
870:Bagaraatan ostromi
588:
510:
448:
410:beds, included in
6911:
6910:
6883:Open Tree of Life
6838:Taxon identifiers
6829:
6828:
6825:
6824:
6798:
6797:
6794:
6793:
6790:
6789:
6786:
6785:
6753:
6752:
6749:
6748:
6559:Macrocheiriformes
6476:Ornithomimosauria
6450:Maniraptoriformes
6443:
6442:
6289:Maniraptoromorpha
6276:Maniraptoromorpha
6265:
6264:
6231:
6230:
6227:
6226:
6223:
6222:
6176:
6175:
6172:
6171:
6154:Maniraptoromorpha
6144:
6143:
6140:
6139:
6136:
6135:
6132:
6131:
6128:
6127:
5876:
5875:
5741:
5740:
5695:Proceratosauridae
5647:Chingkankousaurus
5494:
5493:
5455:
5454:
5451:
5450:
5447:
5446:
5423:
5422:
5275:
5274:
5271:
5270:
5267:
5266:
5263:
5262:
5259:
5258:
5255:
5254:
4931:
4930:
4927:
4926:
4688:
4687:
4659:
4658:
4655:
4654:
4651:
4650:
4647:
4646:
4619:
4618:
4615:
4614:
4611:
4610:
4607:
4606:
4523:
4522:
4479:Vallibonavenatrix
4382:
4381:
4378:
4377:
4222:
4221:
4003:
4002:
3967:
3966:
3963:
3962:
3959:
3958:
3931:
3930:
3927:
3926:
3923:
3922:
3919:
3918:
3915:
3914:
3563:
3562:
3559:
3558:
3325:
3324:
3183:
3182:
3163:
3162:
3159:
3158:
3155:
3154:
3106:
3105:
2759:
2758:
2755:
2754:
2751:
2750:
2314:. Retrieved from
2082:in the background
2060:Palaeoenvironment
2050:Proceratosauridae
2041:
2040:
2032:
2031:
2023:
2022:
2014:
2013:
2005:
2004:
1996:
1995:
1987:
1986:
1978:
1977:
1969:
1968:
1960:
1959:
1951:
1950:
1942:
1941:
1802:
1801:
1793:
1792:
1784:
1783:
1775:
1774:
1766:
1765:
1449:Proceratosauridae
1331:
1330:
1322:
1321:
1313:
1312:
1163:
1162:
1154:
1153:
1145:
1144:
1136:
1135:
1127:
1126:
1118:
1117:
1109:
1108:
1100:
1099:
1091:
1090:
1082:
1081:
1058:
1057:
1047:Alioramus remotus
977:
976:
800:
799:
791:
790:
782:
781:
708:
707:
642:Proceratosauridae
590:The discovery of
403:hailing from the
382:
381:
288:
140:Skeletal diagram
6983:
6928:
6927:
6926:
6919:
6904:
6903:
6891:
6890:
6878:
6877:
6865:
6864:
6863:
6833:
6820:
6814:
6807:
6663:Aepyornithomimus
6564:
6481:
6456:
6420:Sinocalliopteryx
6340:
6336:
6330:Neocoelurosauria
6295:
6284:
6271:
6259:
6252:
6245:
6239:
6182:
6150:
6119:Zhuchengtyrannus
6035:Thanatotheristes
5966:
5930:
5897:Appalachiosaurus
5889:
5758:
5754:
5748:Pantyrannosauria
5639:
5635:
5629:Tyrannosauroidea
5624:
5599:Xinjiangovenator
5524:
5513:
5500:
5488:
5481:
5476:
5469:
5463:
5291:
5281:
5197:
5117:Acrocanthosaurus
5109:
4997:
4944:
4870:Yangchuanosaurus
4854:
4807:
4803:
4753:
4734:Lourinhanosaurus
4718:
4707:
4694:
4682:
4675:
4668:
4625:
4591:Sigilmassasaurus
4534:
4491:Ceratosuchopsini
4442:
4395:
4340:Dubreuillosaurus
4264:
4247:Streptospondylus
4239:
4235:
4205:Piatnitzkysaurus
4163:
4159:
4148:
4033:
4022:
4009:
3997:
3990:
3983:
3976:
3937:
3772:Ekrixinatosaurus
3756:
3747:
3576:
3391:
3342:
3338:
3251:
3247:
3213:
3202:
3189:
3178:
3172:
3082:Procompsognathus
2988:
2871:
2789:
2778:
2765:
2711:
2702:
2695:
2694:
2650:
2635:
2628:
2621:
2612:
2605:
2601:
2595:
2591:
2570:Cau, A. (2024).
2567:
2561:
2560:
2558:
2526:
2520:
2519:
2509:
2499:
2442:
2433:
2432:
2415:(5): 1100–1117.
2405:Holtz, T. R. Jr.
2401:
2395:
2392:Eotyrannus lengi
2388:
2382:
2381:
2371:
2361:
2340:Eotyrannus lengi
2333:
2318:
2312:Eotyrannus lengi
2308:
2302:
2301:
2283:
2274:
2196:; the sauropods
2128:Wessex Formation
2120:Wessex Formation
2088:Wessex Formation
2080:Wessex Formation
1933:
1920:
1906:
1874:
1860:
1852:Appalachiosaurus
1845:
1831:
1816:
1757:
1735:
1713:
1691:
1676:
1649:
1635:
1620:
1596:
1582:
1567:
1557:
1543:
1528:
1514:
1499:
1485:
1470:
1456:
1443:
1429:
1416:
1402:
1380:
1365:
1359:Tyrannosauroidea
1355:
1261:
1221:
1194:
1184:
1022:
1015:
991:
941:
934:
910:
886:
862:
846:Eotyrannus lengi
838:
822:Dilong paradoxus
814:
746:
722:
655:
648:
638:
632:Tyrannosauroidea
628:
585:Life restoration
566:antorbital fossa
554:Tyrannosauroidea
457:Atherfield Point
408:Wessex Formation
405:Early Cretaceous
356:
323:
321:Eotyrannus lengi
319:
282:
275:
264:Pantyrannosauria
262:
248:Tyrannosauroidea
246:
154:
153:
135:
121:
111:
41:
29:Temporal range:
21:
6991:
6990:
6986:
6985:
6984:
6982:
6981:
6980:
6936:
6935:
6934:
6924:
6922:
6914:
6912:
6907:
6899:
6894:
6886:
6881:
6873:
6868:
6859:
6858:
6853:
6840:
6830:
6821:
6808:
6782:
6745:
6726:Sinornithomimus
6648:
6591:
6553:
6470:
6464:Compsognathidae
6452:
6439:
6427:Sinosauropteryx
6359:Compsognathidae
6353:
6332:
6323:
6291:
6278:
6261:
6260:
6246:
6219:
6214:Shanyangosaurus
6207:Phaedrolosaurus
6168:
6124:
6090:
6040:
6019:Daspletosaurini
6013:
5986:
5961:Tyrannosaurinae
5955:
5934:Albertosaurinae
5926:
5924:Tyrannosauridae
5917:
5885:
5883:Eutyrannosauria
5872:
5750:
5737:
5718:Proceratosaurus
5689:
5660:
5631:
5620:
5611:
5577:Richardoestesia
5561:Chilantaisaurus
5520:
5507:
5490:
5489:
5470:
5443:
5419:
5379:Australovenator
5357:
5351:Chilantaisaurus
5329:Phuwiangvenator
5251:
5216:Giganotosaurini
5210:
5186:
5098:
5061:Chilantaisaurus
5046:
4986:
4940:
4923:
4875:
4843:
4799:
4790:
4768:Monolophosaurus
4749:
4740:
4714:
4701:
4684:
4683:
4669:
4643:
4603:
4576:
4542:Camarillasaurus
4519:
4485:
4431:
4425:Ostafrikasaurus
4403:Camarillasaurus
4391:
4374:
4324:Afrovenatorinae
4318:
4277:
4253:
4231:
4218:
4176:
4155:
4153:Megalosauroidea
4144:
4135:
4130:Vectaerovenator
4123:Szechuanosaurus
4101:Monolophosaurus
4087:Kaijiangosaurus
4080:Cryolophosaurus
4041:Calamospondylus
4029:
4016:
3999:
3998:
3991:
3984:
3977:
3955:
3911:
3892:Pycnonemosaurus
3834:
3745:
3739:Majungasaurinae
3724:
3666:Majungasaurinae
3660:
3655:Xenotarsosaurus
3591:Dryptosauroides
3572:
3555:
3507:
3479:Elaphrosaurinae
3473:
3380:
3366:Ornithomimoides
3334:
3321:
3294:
3243:
3234:
3209:
3196:
3179:
3173:
3151:
3126:
3102:
3030:
2977:
2943:Shuangbaisaurus
2867:
2858:
2785:
2772:
2747:
2727:
2721:Avemetatarsalia
2706:Avemetatarsalia
2696:
2681:Avemetatarsalia
2655:
2654:
2644:
2639:
2609:
2608:
2602:
2598:
2569:
2568:
2564:
2528:
2527:
2523:
2454:Sertich, J.J.W.
2444:
2443:
2436:
2403:
2402:
2398:
2389:
2385:
2335:
2334:
2321:
2309:
2305:
2281:
2276:
2275:
2268:
2263:
2251:
2165:neornithischian
2068:Restoration of
2062:
2042:
2033:
2024:
2015:
2006:
1997:
1988:
1979:
1970:
1961:
1952:
1943:
1927:Tyrannosauridae
1907:
1861:
1803:
1794:
1785:
1776:
1767:
1736:
1706:Australovenator
1692:
1636:
1627:Chilantaisaurus
1583:
1515:
1457:
1430:
1332:
1323:
1314:
1262:
1259:
1229:Australovenator
1164:
1155:
1146:
1137:
1128:
1119:
1110:
1101:
1092:
1083:
1072:Tyrannosauridae
1059:
1030:Alioramus altai
978:
801:
792:
783:
709:
697:Guanlong wucaii
578:
502:
432:
359:"Gavinosaurus"
347:
332:
325:
317:
316:
303:
281:
273:
260:
244:
148:
112:
110:
109:
108:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
36:
35:
33:
27:
17:
12:
11:
5:
6989:
6987:
6979:
6978:
6973:
6968:
6963:
6958:
6953:
6951:Barremian life
6948:
6938:
6937:
6933:
6932:
6909:
6908:
6906:
6905:
6892:
6879:
6866:
6850:
6848:
6842:
6841:
6836:
6827:
6826:
6823:
6822:
6802:
6801:
6799:
6796:
6795:
6792:
6791:
6788:
6787:
6784:
6783:
6781:
6780:
6779:
6778:
6772:
6763:
6761:
6755:
6754:
6751:
6750:
6747:
6746:
6744:
6743:
6736:
6729:
6722:
6715:
6708:
6701:
6694:
6691:Dromiceiomimus
6687:
6680:
6673:
6666:
6658:
6656:
6654:Ornithomimidae
6650:
6649:
6647:
6646:
6639:
6635:Paraxenisaurus
6631:
6623:
6616:
6609:
6601:
6599:
6597:Deinocheiridae
6593:
6592:
6590:
6589:
6586:Shenzhousaurus
6582:
6575:
6567:
6561:
6555:
6554:
6552:
6551:
6544:
6536:
6529:
6522:
6515:
6508:
6500:
6492:
6484:
6478:
6472:
6471:
6469:
6468:
6459:
6453:
6448:
6445:
6444:
6441:
6440:
6438:
6437:
6430:
6423:
6416:
6408:
6401:
6394:
6387:
6383:Beipiaognathus
6379:
6372:
6363:
6361:
6355:
6354:
6352:
6351:
6343:
6333:
6328:
6325:
6324:
6322:
6321:
6314:
6307:
6298:
6292:
6287:
6280:
6279:
6274:
6267:
6266:
6263:
6262:
6254:
6240:
6234:
6232:
6229:
6228:
6225:
6224:
6221:
6220:
6218:
6217:
6210:
6203:
6196:
6188:
6186:
6178:
6177:
6174:
6173:
6170:
6169:
6167:
6166:
6165:
6164:
6158:
6156:
6146:
6145:
6142:
6141:
6138:
6137:
6134:
6133:
6130:
6129:
6126:
6125:
6123:
6122:
6115:
6108:
6100:
6098:
6096:Tyrannosaurini
6092:
6091:
6089:
6088:
6081:
6074:
6066:
6059:
6055:Bistahieversor
6050:
6048:
6046:Teratophoneini
6042:
6041:
6039:
6038:
6031:
6028:Daspletosaurus
6023:
6021:
6015:
6014:
6012:
6011:
6008:Qianzhousaurus
6004:
5996:
5994:
5988:
5987:
5985:
5984:
5977:
5969:
5963:
5957:
5956:
5954:
5953:
5946:
5938:
5936:
5927:
5922:
5919:
5918:
5916:
5915:
5908:
5904:Bistahieversor
5900:
5892:
5886:
5881:
5878:
5877:
5874:
5873:
5871:
5870:
5862:
5855:
5848:
5840:
5833:
5826:
5819:
5812:
5805:
5798:
5791:
5784:
5776:
5769:
5761:
5751:
5746:
5743:
5742:
5739:
5738:
5736:
5735:
5728:
5721:
5714:
5707:
5699:
5697:
5691:
5690:
5688:
5687:
5679:
5671:
5669:
5662:
5661:
5659:
5658:
5650:
5642:
5632:
5627:
5621:
5618:Tyrannoraptora
5616:
5613:
5612:
5610:
5609:
5602:
5595:
5588:
5580:
5573:
5565:
5557:
5550:
5542:
5535:
5527:
5521:
5516:
5509:
5508:
5503:
5496:
5495:
5492:
5491:
5483:
5464:
5458:
5456:
5453:
5452:
5449:
5448:
5445:
5444:
5442:
5441:
5440:
5439:
5433:
5431:
5425:
5424:
5421:
5420:
5418:
5417:
5410:
5403:
5396:
5389:
5382:
5375:
5367:
5365:
5363:Megaraptoridae
5359:
5358:
5356:
5355:
5347:
5339:
5332:
5325:
5318:
5310:
5302:
5294:
5288:
5277:
5276:
5273:
5272:
5269:
5268:
5265:
5264:
5261:
5260:
5257:
5256:
5253:
5252:
5250:
5249:
5242:
5235:
5228:
5225:Giganotosaurus
5220:
5218:
5212:
5211:
5209:
5208:
5200:
5194:
5188:
5187:
5185:
5184:
5177:
5170:
5163:
5156:
5149:
5142:
5135:
5128:
5120:
5112:
5106:
5100:
5099:
5097:
5096:
5088:
5080:
5073:
5065:
5056:
5054:
5052:Neovenatoridae
5048:
5047:
5045:
5044:
5037:
5034:Ulughbegsaurus
5030:
5022:
5015:
5008:
5000:
4994:
4988:
4987:
4985:
4984:
4976:
4968:
4960:
4952:
4950:
4941:
4936:
4933:
4932:
4929:
4928:
4925:
4924:
4922:
4921:
4914:
4907:
4900:
4893:
4885:
4883:
4877:
4876:
4874:
4873:
4866:
4862:Xuanhanosaurus
4857:
4851:
4845:
4844:
4842:
4841:
4833:
4829:Xuanhanosaurus
4825:
4818:
4815:Asfaltovenator
4810:
4800:
4795:
4792:
4791:
4789:
4788:
4780:
4776:Megalosauridae
4772:
4764:
4756:
4750:
4745:
4742:
4741:
4739:
4738:
4730:
4721:
4715:
4710:
4703:
4702:
4697:
4690:
4689:
4686:
4685:
4677:
4663:
4662:
4660:
4657:
4656:
4653:
4652:
4649:
4648:
4645:
4644:
4642:
4641:
4640:
4639:
4633:
4631:
4621:
4620:
4617:
4616:
4613:
4612:
4609:
4608:
4605:
4604:
4602:
4601:
4594:
4586:
4584:
4578:
4577:
4575:
4574:
4567:
4560:
4553:
4550:Ichthyovenator
4546:
4537:
4531:
4525:
4524:
4521:
4520:
4518:
4517:
4510:
4503:
4495:
4493:
4487:
4486:
4484:
4483:
4475:
4467:
4460:
4453:
4445:
4439:
4433:
4432:
4430:
4429:
4421:
4414:
4411:Cristatusaurus
4407:
4398:
4392:
4387:
4384:
4383:
4380:
4379:
4376:
4375:
4373:
4372:
4368:Poekilopleuron
4364:
4361:Piveteausaurus
4357:
4350:
4343:
4336:
4328:
4326:
4320:
4319:
4317:
4316:
4309:
4302:
4295:
4287:
4285:
4283:Megalosaurinae
4279:
4278:
4276:
4275:
4267:
4261:
4259:Megalosauridae
4255:
4254:
4252:
4251:
4242:
4232:
4227:
4224:
4223:
4220:
4219:
4217:
4216:
4212:Xuanhanosaurus
4208:
4201:
4194:
4186:
4184:
4178:
4177:
4175:
4174:
4171:Yunyangosaurus
4166:
4156:
4151:
4145:
4140:
4137:
4136:
4134:
4133:
4126:
4119:
4111:
4108:Pandoravenator
4104:
4097:
4094:Kayentavenator
4090:
4083:
4076:
4068:
4061:
4053:
4045:
4036:
4030:
4025:
4018:
4017:
4012:
4005:
4004:
4001:
4000:
3992:
3985:
3978:
3971:
3970:
3968:
3965:
3964:
3961:
3960:
3957:
3956:
3954:
3953:
3952:
3951:
3945:
3943:
3933:
3932:
3929:
3928:
3925:
3924:
3921:
3920:
3917:
3916:
3913:
3912:
3910:
3909:
3902:
3895:
3888:
3881:
3873:
3866:
3859:
3852:
3844:
3842:
3836:
3835:
3833:
3832:
3825:
3822:Skorpiovenator
3818:
3810:
3803:
3796:
3789:
3782:
3775:
3768:
3759:
3753:
3744:
3743:
3734:
3732:
3726:
3725:
3723:
3722:
3715:
3707:
3700:
3693:
3686:
3678:
3670:
3668:
3662:
3661:
3659:
3658:
3651:
3644:
3637:
3634:Spectrovenator
3630:
3623:
3616:
3609:
3602:
3595:
3587:
3579:
3573:
3568:
3565:
3564:
3561:
3560:
3557:
3556:
3554:
3553:
3546:
3539:
3532:
3525:
3517:
3515:
3509:
3508:
3506:
3505:
3498:
3491:
3483:
3481:
3475:
3474:
3472:
3471:
3468:Spinostropheus
3464:
3457:
3450:
3442:
3434:
3426:
3418:
3411:
3403:
3394:
3388:
3382:
3381:
3379:
3378:
3370:
3362:
3354:
3345:
3335:
3332:Abelisauroidea
3330:
3327:
3326:
3323:
3322:
3320:
3319:
3312:
3304:
3302:
3300:Ceratosauridae
3296:
3295:
3293:
3292:
3289:Saltriovenator
3285:
3277:
3270:
3266:Fosterovenator
3262:
3254:
3244:
3239:
3236:
3235:
3233:
3232:
3228:Lophostropheus
3224:
3216:
3210:
3205:
3198:
3197:
3192:
3185:
3184:
3181:
3180:
3167:
3166:
3164:
3161:
3160:
3157:
3156:
3153:
3152:
3150:
3149:
3142:
3134:
3132:
3128:
3127:
3125:
3124:
3123:
3122:
3116:
3114:
3108:
3107:
3104:
3103:
3101:
3100:
3093:
3089:Pterospondylus
3085:
3078:
3070:
3062:
3059:Lucianovenator
3055:
3048:
3040:
3038:
3032:
3031:
3029:
3028:
3021:
3014:
3007:
3000:
2991:
2985:
2983:Coelophysoidea
2979:
2978:
2976:
2975:
2968:
2960:
2953:
2946:
2939:
2932:
2925:
2922:Lophostropheus
2918:
2911:
2904:
2897:
2890:
2882:
2874:
2868:
2863:
2860:
2859:
2857:
2856:
2848:
2840:
2832:
2824:
2821:Erythrovenator
2817:
2809:
2801:
2792:
2786:
2781:
2774:
2773:
2768:
2761:
2760:
2757:
2756:
2753:
2752:
2749:
2748:
2746:
2745:
2744:
2743:
2737:
2735:
2729:
2728:
2726:
2725:
2724:
2723:
2714:
2708:
2698:
2697:
2693:
2692:
2683:
2674:
2668:
2662:
2653:
2646:
2645:
2640:
2638:
2637:
2630:
2623:
2615:
2607:
2606:
2596:
2562:
2521:
2482:(11): e79420.
2466:Evans, David C
2462:Sampson, S. D.
2434:
2396:
2383:
2319:
2303:
2265:
2264:
2262:
2259:
2258:
2257:
2250:
2247:
2229:spalacotheriid
2181:Mantellisaurus
2110:lived. In the
2061:
2058:
2039:
2038:
2035:
2034:
2030:
2029:
2026:
2025:
2021:
2020:
2017:
2016:
2012:
2011:
2008:
2007:
2003:
2002:
1999:
1998:
1994:
1993:
1990:
1989:
1985:
1984:
1981:
1980:
1976:
1975:
1972:
1971:
1967:
1966:
1963:
1962:
1958:
1957:
1954:
1953:
1949:
1948:
1945:
1944:
1940:
1939:
1936:
1935:
1923:
1918:
1916:
1913:
1912:
1909:
1908:
1901:
1894:
1891:
1890:
1887:
1886:
1882:Bistahieversor
1877:
1872:
1870:
1867:
1866:
1863:
1862:
1855:
1848:
1843:
1841:
1838:
1837:
1834:
1833:
1819:
1814:
1812:
1809:
1808:
1805:
1804:
1800:
1799:
1796:
1795:
1791:
1790:
1787:
1786:
1782:
1781:
1778:
1777:
1773:
1772:
1769:
1768:
1764:
1763:
1760:
1759:
1745:
1742:
1741:
1738:
1737:
1730:
1723:
1720:
1719:
1716:
1715:
1701:
1698:
1697:
1694:
1693:
1686:
1679:
1674:
1672:
1670:Megaraptoridae
1666:
1665:
1662:
1661:
1652:
1647:
1645:
1642:
1641:
1638:
1637:
1630:
1623:
1618:
1616:
1613:
1612:
1609:
1608:
1599:
1594:
1592:
1589:
1588:
1585:
1584:
1577:
1570:
1565:
1563:
1555:
1553:
1550:
1549:
1546:
1545:
1531:
1526:
1524:
1521:
1520:
1517:
1516:
1509:
1502:
1497:
1495:
1492:
1491:
1488:
1487:
1473:
1468:
1466:
1463:
1462:
1459:
1458:
1451:
1446:
1441:
1439:
1436:
1435:
1432:
1431:
1424:
1419:
1414:
1412:
1409:
1408:
1405:
1404:
1390:
1387:
1386:
1383:
1382:
1368:
1363:
1361:
1353:
1329:
1328:
1325:
1324:
1320:
1319:
1316:
1315:
1311:
1310:
1307:
1306:
1297:
1294:
1293:
1290:
1289:
1280:
1277:
1276:
1273:
1272:
1263:
1258:
1255:
1254:
1251:
1250:
1241:
1238:
1237:
1234:
1233:
1224:
1219:
1217:
1215:Megaraptoridae
1211:
1210:
1207:
1206:
1197:
1192:
1190:
1182:
1161:
1160:
1157:
1156:
1152:
1151:
1148:
1147:
1143:
1142:
1139:
1138:
1134:
1133:
1130:
1129:
1125:
1124:
1121:
1120:
1116:
1115:
1112:
1111:
1107:
1106:
1103:
1102:
1098:
1097:
1094:
1093:
1089:
1088:
1085:
1084:
1080:
1079:
1076:
1075:
1068:
1065:
1064:
1061:
1060:
1056:
1055:
1052:
1051:
1042:
1039:
1038:
1035:
1034:
1025:
1020:
1018:
1013:
1011:
1008:
1007:
1004:
1003:
994:
989:
987:
984:
983:
980:
979:
975:
974:
971:
970:
961:
958:
957:
954:
953:
944:
939:
937:
932:
930:
927:
926:
923:
922:
913:
908:
906:
903:
902:
899:
898:
889:
884:
882:
879:
878:
875:
874:
865:
860:
858:
855:
854:
851:
850:
841:
836:
834:
831:
830:
827:
826:
817:
812:
810:
807:
806:
803:
802:
798:
797:
794:
793:
789:
788:
785:
784:
780:
779:
776:
775:
766:
763:
762:
759:
758:
749:
744:
742:
739:
738:
735:
734:
725:
720:
718:
715:
714:
711:
710:
706:
705:
702:
701:
692:
689:
688:
685:
684:
675:
672:
671:
668:
667:
658:
653:
651:
646:
644:
636:
634:
626:
601:North American
577:
576:Classification
574:
501:
498:
431:
428:
420:United Kingdom
395:tyrannosauroid
380:
379:
378:
377:
369:"Lengosaurus"
367:
357:
341:
340:
334:
333:
326:
312:
311:
305:
304:
296:
294:
290:
289:
271:
267:
266:
258:
251:
250:
242:
238:
237:
235:Tyrannoraptora
232:
225:
224:
219:
212:
211:
206:
199:
198:
193:
186:
185:
180:
176:
175:
170:
166:
165:
160:
156:
155:
142:
141:
137:
136:
128:
127:
123:
122:
114:
113:
105:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
42:
28:
15:
13:
10:
9:
6:
4:
3:
2:
6988:
6977:
6974:
6972:
6969:
6967:
6964:
6962:
6959:
6957:
6954:
6952:
6949:
6947:
6944:
6943:
6941:
6931:
6921:
6917:
6902:
6897:
6893:
6889:
6884:
6880:
6876:
6871:
6867:
6862:
6856:
6852:
6851:
6849:
6847:
6843:
6839:
6834:
6819:
6815:
6813:
6806:
6777:
6773:
6771:
6767:
6766:
6765:
6764:
6762:
6760:
6756:
6742:
6741:
6737:
6735:
6734:
6733:Struthiomimus
6730:
6728:
6727:
6723:
6721:
6720:
6716:
6714:
6713:
6709:
6707:
6706:
6702:
6700:
6699:
6695:
6693:
6692:
6688:
6686:
6685:
6684:"Coelosaurus"
6681:
6679:
6678:
6674:
6672:
6671:
6667:
6665:
6664:
6660:
6659:
6657:
6655:
6651:
6645:
6644:
6640:
6637:
6636:
6632:
6629:
6628:
6624:
6622:
6621:
6617:
6615:
6614:
6610:
6608:
6607:
6603:
6602:
6600:
6598:
6594:
6588:
6587:
6583:
6581:
6580:
6579:Pelecanimimus
6576:
6574:
6573:
6569:
6568:
6565:
6562:
6560:
6556:
6550:
6549:
6545:
6542:
6541:
6540:Thecocoelurus
6537:
6535:
6534:
6530:
6528:
6527:
6523:
6521:
6520:
6519:Kinnareemimus
6516:
6514:
6513:
6509:
6506:
6505:
6501:
6498:
6497:
6493:
6491:
6490:
6486:
6485:
6482:
6479:
6477:
6473:
6466:
6465:
6461:
6460:
6457:
6454:
6451:
6446:
6436:
6435:
6431:
6429:
6428:
6424:
6422:
6421:
6417:
6414:
6413:
6409:
6407:
6406:
6402:
6400:
6399:
6398:Huaxiagnathus
6395:
6393:
6392:
6391:Compsognathus
6388:
6385:
6384:
6380:
6378:
6377:
6373:
6370:
6369:
6365:
6364:
6362:
6360:
6356:
6350:
6349:
6345:
6344:
6341:
6337:
6334:
6331:
6326:
6320:
6319:
6318:Ornitholestes
6315:
6313:
6312:
6308:
6305:
6304:
6300:
6299:
6296:
6293:
6290:
6285:
6281:
6277:
6272:
6268:
6258:
6253:
6251:
6244:
6238:
6216:
6215:
6211:
6209:
6208:
6204:
6202:
6201:
6197:
6195:
6194:
6190:
6189:
6187:
6183:
6179:
6162:
6161:
6160:
6159:
6157:
6155:
6151:
6147:
6121:
6120:
6116:
6114:
6113:
6112:Tyrannosaurus
6109:
6107:
6106:
6102:
6101:
6099:
6097:
6093:
6087:
6086:
6085:Teratophoneus
6082:
6080:
6079:
6075:
6072:
6071:
6067:
6065:
6064:
6060:
6057:
6056:
6052:
6051:
6049:
6047:
6043:
6037:
6036:
6032:
6030:
6029:
6025:
6024:
6022:
6020:
6016:
6010:
6009:
6005:
6003:
6002:
5998:
5997:
5995:
5993:
5989:
5983:
5982:
5978:
5976:
5975:
5971:
5970:
5967:
5964:
5962:
5958:
5952:
5951:
5947:
5945:
5944:
5943:Albertosaurus
5940:
5939:
5937:
5935:
5931:
5928:
5925:
5920:
5914:
5913:
5909:
5906:
5905:
5901:
5899:
5898:
5894:
5893:
5890:
5887:
5884:
5879:
5868:
5867:
5863:
5861:
5860:
5859:Xiongguanlong
5856:
5854:
5853:
5849:
5846:
5845:
5841:
5839:
5838:
5837:Suskityrannus
5834:
5832:
5831:
5830:Stokesosaurus
5827:
5825:
5824:
5823:Santanaraptor
5820:
5818:
5817:
5813:
5811:
5810:
5806:
5804:
5803:
5799:
5797:
5796:
5792:
5790:
5789:
5785:
5782:
5781:
5777:
5775:
5774:
5770:
5768:
5767:
5766:Alectrosaurus
5763:
5762:
5759:
5755:
5752:
5749:
5744:
5734:
5733:
5729:
5727:
5726:
5722:
5720:
5719:
5715:
5713:
5712:
5708:
5706:
5705:
5701:
5700:
5698:
5696:
5692:
5685:
5684:
5683:Tanycolagreus
5680:
5678:
5677:
5673:
5672:
5670:
5667:
5663:
5656:
5655:
5651:
5649:
5648:
5644:
5643:
5640:
5636:
5633:
5630:
5625:
5622:
5619:
5614:
5608:
5607:
5603:
5601:
5600:
5596:
5594:
5593:
5589:
5586:
5585:
5581:
5579:
5578:
5574:
5571:
5570:
5566:
5563:
5562:
5558:
5556:
5555:
5551:
5548:
5547:
5546:Bahariasaurus
5543:
5541:
5540:
5536:
5534:
5533:
5529:
5528:
5525:
5522:
5519:
5518:Coelurosauria
5514:
5510:
5506:
5505:Coelurosauria
5501:
5497:
5487:
5482:
5480:
5475:
5468:
5462:
5437:
5436:
5435:
5434:
5432:
5430:
5429:Coelurosauria
5426:
5416:
5415:
5411:
5409:
5408:
5404:
5402:
5401:
5397:
5395:
5394:
5390:
5388:
5387:
5383:
5381:
5380:
5376:
5374:
5373:
5369:
5368:
5366:
5364:
5360:
5354:
5352:
5348:
5346:
5344:
5340:
5338:
5337:
5333:
5331:
5330:
5326:
5324:
5323:
5319:
5316:
5315:
5311:
5308:
5307:
5306:Bahariasaurus
5303:
5301:
5300:
5296:
5295:
5292:
5289:
5286:
5282:
5278:
5248:
5247:
5243:
5241:
5240:
5236:
5234:
5233:
5229:
5227:
5226:
5222:
5221:
5219:
5217:
5213:
5207:
5206:
5202:
5201:
5198:
5195:
5193:
5189:
5183:
5182:
5178:
5176:
5175:
5171:
5169:
5168:
5164:
5162:
5161:
5157:
5155:
5154:
5150:
5148:
5147:
5146:Kelmayisaurus
5143:
5141:
5140:
5136:
5134:
5133:
5129:
5126:
5125:
5121:
5119:
5118:
5114:
5113:
5110:
5107:
5105:
5101:
5094:
5093:
5089:
5086:
5085:
5081:
5079:
5078:
5074:
5071:
5070:
5066:
5063:
5062:
5058:
5057:
5055:
5053:
5049:
5043:
5042:
5038:
5036:
5035:
5031:
5028:
5027:
5023:
5021:
5020:
5016:
5014:
5013:
5009:
5007:
5006:
5002:
5001:
4998:
4995:
4993:
4989:
4982:
4981:
4980:Saurophaganax
4977:
4974:
4973:
4969:
4966:
4965:
4961:
4959:
4958:
4954:
4953:
4951:
4949:
4945:
4942:
4939:
4934:
4920:
4919:
4915:
4913:
4912:
4911:Siamotyrannus
4908:
4906:
4905:
4901:
4899:
4898:
4894:
4892:
4891:
4887:
4886:
4884:
4882:
4878:
4872:
4871:
4867:
4864:
4863:
4859:
4858:
4855:
4852:
4850:
4846:
4839:
4838:
4834:
4831:
4830:
4826:
4824:
4823:
4819:
4817:
4816:
4812:
4811:
4808:
4804:
4801:
4798:
4797:Allosauroidea
4793:
4786:
4785:
4784:Spinosauridae
4781:
4778:
4777:
4773:
4770:
4769:
4765:
4763:
4762:
4758:
4757:
4754:
4751:
4748:
4743:
4736:
4735:
4731:
4728:
4727:
4723:
4722:
4719:
4716:
4713:
4708:
4704:
4700:
4695:
4691:
4681:
4676:
4674:
4667:
4637:
4636:
4635:
4634:
4632:
4630:
4626:
4622:
4600:
4599:
4595:
4593:
4592:
4588:
4587:
4585:
4583:
4579:
4573:
4572:
4568:
4566:
4565:
4561:
4559:
4558:
4554:
4552:
4551:
4547:
4544:
4543:
4539:
4538:
4535:
4532:
4530:
4529:Spinosaurinae
4526:
4516:
4515:
4511:
4509:
4508:
4507:Riparovenator
4504:
4502:
4501:
4500:Ceratosuchops
4497:
4496:
4494:
4492:
4488:
4481:
4480:
4476:
4473:
4472:
4468:
4466:
4465:
4464:Riojavenatrix
4461:
4459:
4458:
4454:
4452:
4451:
4447:
4446:
4443:
4440:
4438:
4437:Baryonychinae
4434:
4427:
4426:
4422:
4420:
4419:
4415:
4413:
4412:
4408:
4405:
4404:
4400:
4399:
4396:
4393:
4390:
4389:Spinosauridae
4385:
4370:
4369:
4365:
4363:
4362:
4358:
4356:
4355:
4351:
4349:
4348:
4344:
4342:
4341:
4337:
4335:
4334:
4330:
4329:
4327:
4325:
4321:
4315:
4314:
4313:Wiehenvenator
4310:
4308:
4307:
4303:
4301:
4300:
4296:
4294:
4293:
4289:
4288:
4286:
4284:
4280:
4274:
4273:
4269:
4268:
4265:
4262:
4260:
4256:
4249:
4248:
4244:
4243:
4240:
4236:
4233:
4230:
4225:
4214:
4213:
4209:
4207:
4206:
4202:
4200:
4199:
4195:
4193:
4192:
4188:
4187:
4185:
4183:
4179:
4173:
4172:
4168:
4167:
4164:
4160:
4157:
4154:
4149:
4146:
4143:
4138:
4132:
4131:
4127:
4125:
4124:
4120:
4117:
4116:
4112:
4110:
4109:
4105:
4103:
4102:
4098:
4096:
4095:
4091:
4089:
4088:
4084:
4082:
4081:
4077:
4074:
4073:
4069:
4067:
4066:
4062:
4059:
4058:
4054:
4051:
4050:
4049:Chienkosaurus
4046:
4043:
4042:
4038:
4037:
4034:
4031:
4028:
4023:
4019:
4015:
4010:
4006:
3996:
3989:
3982:
3975:
3949:
3948:
3947:
3946:
3944:
3942:
3938:
3934:
3908:
3907:
3903:
3901:
3900:
3896:
3894:
3893:
3889:
3887:
3886:
3882:
3879:
3878:
3874:
3872:
3871:
3867:
3865:
3864:
3860:
3858:
3857:
3853:
3851:
3850:
3846:
3845:
3843:
3841:
3840:Furileusauria
3837:
3831:
3830:
3826:
3824:
3823:
3819:
3816:
3815:
3814:Rahiolisaurus
3811:
3809:
3808:
3804:
3802:
3801:
3797:
3795:
3794:
3790:
3788:
3787:
3783:
3781:
3780:
3776:
3774:
3773:
3769:
3766:
3765:
3761:
3760:
3757:
3754:
3752:
3748:
3741:
3740:
3736:
3735:
3733:
3731:
3730:Carnotaurinae
3727:
3721:
3720:
3716:
3713:
3712:
3711:Rahiolisaurus
3708:
3706:
3705:
3704:Majungasaurus
3701:
3699:
3698:
3694:
3692:
3691:
3687:
3684:
3683:
3679:
3677:
3676:
3672:
3671:
3669:
3667:
3663:
3657:
3656:
3652:
3650:
3649:
3645:
3643:
3642:
3641:Tarascosaurus
3638:
3636:
3635:
3631:
3629:
3628:
3624:
3622:
3621:
3617:
3615:
3614:
3610:
3608:
3607:
3603:
3601:
3600:
3599:Eoabelisaurus
3596:
3593:
3592:
3588:
3586:
3585:
3584:Chenanisaurus
3581:
3580:
3577:
3574:
3571:
3570:Abelisauridae
3566:
3552:
3551:
3547:
3545:
3544:
3540:
3538:
3537:
3533:
3531:
3530:
3529:Masiakasaurus
3526:
3524:
3523:
3519:
3518:
3516:
3514:
3510:
3504:
3503:
3499:
3497:
3496:
3495:Huinculsaurus
3492:
3490:
3489:
3488:Elaphrosaurus
3485:
3484:
3482:
3480:
3476:
3470:
3469:
3465:
3463:
3462:
3458:
3456:
3455:
3451:
3448:
3447:
3443:
3440:
3439:
3435:
3432:
3431:
3427:
3424:
3423:
3419:
3417:
3416:
3412:
3409:
3408:
3407:Austrocheirus
3404:
3401:
3400:
3396:
3395:
3392:
3389:
3387:
3383:
3376:
3375:
3371:
3368:
3367:
3363:
3360:
3359:
3355:
3352:
3351:
3347:
3346:
3343:
3339:
3336:
3333:
3328:
3318:
3317:
3313:
3311:
3310:
3306:
3305:
3303:
3301:
3297:
3291:
3290:
3286:
3283:
3282:
3278:
3276:
3275:
3271:
3268:
3267:
3263:
3261:
3260:
3259:Berberosaurus
3256:
3255:
3252:
3248:
3245:
3242:
3237:
3230:
3229:
3225:
3223:
3222:
3218:
3217:
3214:
3211:
3208:
3203:
3199:
3195:
3190:
3186:
3177:
3171:
3148:
3147:
3146:Halticosaurus
3143:
3141:
3140:
3139:Teinurosaurus
3136:
3135:
3133:
3129:
3120:
3119:
3118:
3117:
3115:
3113:
3109:
3099:
3098:
3094:
3091:
3090:
3086:
3084:
3083:
3079:
3076:
3075:
3071:
3068:
3067:
3066:Megapnosaurus
3063:
3061:
3060:
3056:
3054:
3053:
3049:
3047:
3046:
3042:
3041:
3039:
3037:
3036:Coelophysidae
3033:
3027:
3026:
3025:Powellvenator
3022:
3020:
3019:
3015:
3013:
3012:
3008:
3006:
3005:
3001:
2998:
2997:
2993:
2992:
2989:
2986:
2984:
2980:
2974:
2973:
2969:
2966:
2965:
2961:
2959:
2958:
2954:
2952:
2951:
2947:
2945:
2944:
2940:
2938:
2937:
2933:
2931:
2930:
2926:
2924:
2923:
2919:
2917:
2916:
2915:Liliensternus
2912:
2910:
2909:
2905:
2903:
2902:
2898:
2896:
2895:
2891:
2888:
2887:
2886:Dolichosuchus
2883:
2881:
2880:
2879:Dilophosaurus
2876:
2875:
2872:
2869:
2866:
2861:
2854:
2853:
2852:Herrerasauria
2849:
2846:
2845:
2841:
2838:
2837:
2833:
2830:
2829:
2825:
2823:
2822:
2818:
2815:
2814:
2810:
2807:
2806:
2805:Daemonosaurus
2802:
2799:
2798:
2794:
2793:
2790:
2787:
2784:
2779:
2775:
2771:
2766:
2762:
2741:
2740:
2739:
2738:
2736:
2734:
2730:
2722:
2718:
2717:
2716:
2715:
2712:
2709:
2707:
2703:
2699:
2691:
2687:
2684:
2682:
2678:
2675:
2673:
2669:
2667:
2663:
2661:
2657:
2656:
2651:
2647:
2643:
2636:
2631:
2629:
2624:
2622:
2617:
2616:
2613:
2600:
2597:
2594:
2589:
2585:
2581:
2577:
2573:
2566:
2563:
2557:
2552:
2548:
2544:
2540:
2536:
2532:
2525:
2522:
2517:
2513:
2508:
2503:
2498:
2493:
2489:
2485:
2481:
2477:
2476:
2471:
2467:
2463:
2459:
2458:Currie, P. J.
2455:
2451:
2447:
2441:
2439:
2435:
2430:
2426:
2422:
2418:
2414:
2410:
2406:
2400:
2397:
2393:
2387:
2384:
2379:
2375:
2370:
2365:
2360:
2355:
2351:
2347:
2343:
2341:
2332:
2330:
2328:
2326:
2324:
2320:
2317:
2313:
2307:
2304:
2299:
2295:
2291:
2287:
2280:
2273:
2271:
2267:
2260:
2256:
2253:
2252:
2248:
2246:
2244:
2243:
2239:
2235:
2234:
2230:
2226:
2222:
2221:
2217:
2213:
2212:
2207:
2206:
2201:
2200:
2195:
2194:
2189:
2188:
2183:
2182:
2177:
2176:
2171:
2170:
2169:Hypsilophodon
2166:
2162:
2161:
2160:Ceratosuchops
2156:
2155:
2154:Riparovenator
2151:
2147:
2146:
2142:
2141:compsognathid
2138:
2137:
2133:
2129:
2124:
2121:
2117:
2113:
2109:
2105:
2101:
2097:
2096:Mediterranean
2093:
2089:
2081:
2077:
2076:
2075:Hypsilophodon
2071:
2066:
2059:
2057:
2055:
2051:
2047:
2037:
2036:
2028:
2027:
2019:
2018:
2010:
2009:
2001:
2000:
1992:
1991:
1983:
1982:
1974:
1973:
1965:
1964:
1956:
1955:
1947:
1946:
1938:
1937:
1934:
1932:
1928:
1922:
1921:
1915:
1914:
1911:
1910:
1905:
1900:
1899:
1898:Teratophoneus
1893:
1892:
1889:
1888:
1885:
1884:
1883:
1876:
1875:
1869:
1868:
1865:
1864:
1859:
1854:
1853:
1847:
1846:
1840:
1839:
1836:
1835:
1832:
1830:
1826:
1825:
1818:
1817:
1811:
1810:
1807:
1806:
1798:
1797:
1789:
1788:
1780:
1779:
1771:
1770:
1762:
1761:
1758:
1756:
1752:
1751:
1744:
1743:
1740:
1739:
1734:
1729:
1728:
1722:
1721:
1718:
1717:
1714:
1712:
1708:
1707:
1700:
1699:
1696:
1695:
1690:
1685:
1684:
1678:
1677:
1671:
1668:
1667:
1664:
1663:
1660:
1659:
1658:
1651:
1650:
1644:
1643:
1640:
1639:
1634:
1629:
1628:
1622:
1621:
1615:
1614:
1611:
1610:
1607:
1606:
1605:
1598:
1597:
1591:
1590:
1587:
1586:
1581:
1576:
1575:
1569:
1568:
1562:
1559:
1558:
1552:
1551:
1548:
1547:
1544:
1542:
1538:
1537:
1536:Xiongguanlong
1530:
1529:
1523:
1522:
1519:
1518:
1513:
1508:
1507:
1501:
1500:
1494:
1493:
1490:
1489:
1486:
1484:
1480:
1479:
1472:
1471:
1465:
1464:
1461:
1460:
1455:
1450:
1445:
1444:
1438:
1437:
1434:
1433:
1428:
1423:
1418:
1417:
1411:
1410:
1407:
1406:
1403:
1401:
1397:
1396:
1395:Stokesosaurus
1389:
1388:
1385:
1384:
1381:
1379:
1375:
1374:
1367:
1366:
1360:
1357:
1356:
1352:
1349:
1345:
1341:
1337:
1327:
1326:
1318:
1317:
1309:
1308:
1305:
1304:
1303:
1296:
1295:
1292:
1291:
1288:
1287:
1286:
1279:
1278:
1275:
1274:
1271:
1270:
1269:
1257:
1256:
1253:
1252:
1249:
1248:
1247:
1240:
1239:
1236:
1235:
1232:
1231:
1230:
1223:
1222:
1216:
1213:
1212:
1209:
1208:
1205:
1204:
1203:
1196:
1195:
1189:
1186:
1185:
1181:
1179:
1178:
1173:
1169:
1159:
1158:
1150:
1149:
1141:
1140:
1132:
1131:
1123:
1122:
1114:
1113:
1105:
1104:
1096:
1095:
1087:
1086:
1078:
1077:
1074:
1073:
1067:
1066:
1063:
1062:
1054:
1053:
1050:
1049:
1048:
1041:
1040:
1037:
1036:
1033:
1032:
1031:
1024:
1023:
1017:
1016:
1010:
1009:
1006:
1005:
1002:
1001:
1000:
993:
992:
986:
985:
982:
981:
973:
972:
969:
968:
967:
960:
959:
956:
955:
952:
951:
950:
943:
942:
936:
935:
929:
928:
925:
924:
921:
920:
919:
912:
911:
905:
904:
901:
900:
897:
896:
895:
888:
887:
881:
880:
877:
876:
873:
872:
871:
864:
863:
857:
856:
853:
852:
849:
848:
847:
840:
839:
833:
832:
829:
828:
825:
824:
823:
816:
815:
809:
808:
805:
804:
796:
795:
787:
786:
778:
777:
774:
773:
772:
765:
764:
761:
760:
757:
756:
755:
748:
747:
741:
740:
737:
736:
733:
732:
731:
724:
723:
717:
716:
713:
712:
704:
703:
700:
699:
698:
691:
690:
687:
686:
683:
682:
681:
674:
673:
670:
669:
666:
665:
664:
657:
656:
650:
649:
643:
640:
639:
633:
630:
629:
625:
623:
618:
616:
615:
611:
607:
606:
605:Stokesosaurus
602:
598:
593:
586:
582:
575:
573:
569:
567:
563:
559:
555:
550:
548:
544:
540:
536:
531:
527:
524:
519:
515:
506:
499:
497:
495:
494:
489:
485:
481:
476:
474:
470:
466:
462:
461:Hanover Point
458:
453:
445:
441:
440:Isle of Wight
436:
429:
427:
425:
421:
417:
416:Isle of Wight
413:
412:Wealden Group
409:
406:
402:
399:
396:
392:
388:
387:
376:
374:
368:
366:
364:
358:
355:
353:
345:
344:
342:
339:
335:
330:
324:
322:
313:
310:
309:Binomial name
306:
302:
301:
300:E. lengi
295:
292:
291:
286:
280:
279:
272:
269:
268:
265:
259:
256:
253:
252:
249:
243:
241:Superfamily:
240:
239:
236:
233:
230:
227:
226:
223:
220:
217:
214:
213:
210:
207:
204:
201:
200:
197:
194:
191:
188:
187:
184:
181:
178:
177:
174:
171:
168:
167:
164:
161:
158:
157:
152:
147:
143:
138:
134:
129:
124:
120:
115:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
40:
32:
26:
22:
19:
6946:Tyrannosaurs
6845:
6809:
6738:
6731:
6724:
6717:
6710:
6705:Ornithomimus
6703:
6696:
6689:
6682:
6675:
6668:
6661:
6643:Tyrannomimus
6641:
6633:
6625:
6618:
6613:Deinocheirus
6611:
6604:
6584:
6577:
6570:
6546:
6538:
6533:Nqwebasaurus
6531:
6526:Nedcolbertia
6524:
6517:
6510:
6504:Calamosaurus
6502:
6496:Aviatyrannis
6494:
6487:
6462:
6432:
6425:
6418:
6410:
6403:
6396:
6389:
6381:
6376:Aristosuchus
6374:
6368:Aniksosaurus
6366:
6348:Aniksosaurus
6346:
6316:
6309:
6301:
6247:
6212:
6205:
6198:
6191:
6117:
6110:
6103:
6083:
6076:
6068:
6063:Dynamoterror
6061:
6053:
6033:
6026:
6006:
5999:
5979:
5974:Asiatyrannus
5972:
5948:
5941:
5912:Dryptosaurus
5910:
5902:
5895:
5864:
5857:
5850:
5842:
5835:
5828:
5821:
5814:
5807:
5802:Jinbeisaurus
5800:
5794:
5793:
5786:
5778:
5773:Aviatyrannis
5771:
5764:
5730:
5725:Sinotyrannus
5723:
5716:
5709:
5702:
5681:
5674:
5652:
5645:
5604:
5597:
5590:
5582:
5575:
5567:
5559:
5554:Bicentenaria
5552:
5544:
5539:Asiamericana
5537:
5530:
5471:
5412:
5405:
5398:
5391:
5384:
5377:
5370:
5349:
5341:
5334:
5327:
5320:
5314:Deltadromeus
5312:
5304:
5297:
5246:Tyrannotitan
5244:
5237:
5230:
5223:
5203:
5179:
5174:Taurovenator
5172:
5165:
5158:
5153:Lajasvenator
5151:
5144:
5137:
5132:Concavenator
5130:
5122:
5115:
5090:
5082:
5075:
5067:
5059:
5039:
5032:
5024:
5017:
5010:
5003:
4978:
4970:
4962:
4955:
4948:Allosauridae
4916:
4909:
4904:Shidaisaurus
4902:
4895:
4888:
4868:
4860:
4835:
4827:
4820:
4813:
4782:
4774:
4766:
4759:
4732:
4724:
4712:Avetheropoda
4699:Avetheropoda
4670:
4629:Avetheropoda
4596:
4589:
4582:Spinosaurini
4569:
4562:
4555:
4548:
4540:
4512:
4505:
4498:
4477:
4469:
4462:
4457:Protathlitis
4455:
4448:
4423:
4416:
4409:
4401:
4366:
4359:
4352:
4347:Leshansaurus
4345:
4338:
4331:
4311:
4304:
4299:Megalosaurus
4297:
4292:Duriavenator
4290:
4270:
4245:
4229:Megalosauria
4210:
4203:
4198:Marshosaurus
4196:
4191:Condorraptor
4189:
4169:
4128:
4121:
4113:
4106:
4099:
4092:
4085:
4078:
4072:Cruxicheiros
4070:
4063:
4055:
4047:
4039:
3904:
3899:Quilmesaurus
3897:
3890:
3883:
3875:
3868:
3861:
3854:
3847:
3827:
3820:
3812:
3805:
3798:
3791:
3784:
3777:
3770:
3762:
3751:Brachyrostra
3737:
3717:
3709:
3702:
3695:
3688:
3680:
3673:
3653:
3648:Tralkasaurus
3646:
3639:
3632:
3625:
3618:
3611:
3604:
3597:
3589:
3582:
3550:Vespersaurus
3548:
3543:Velocisaurus
3541:
3534:
3527:
3520:
3500:
3493:
3486:
3466:
3459:
3452:
3444:
3438:Deltadromeus
3436:
3428:
3422:Compsosuchus
3420:
3413:
3405:
3397:
3372:
3364:
3358:Lametasaurus
3356:
3348:
3314:
3309:Ceratosaurus
3307:
3287:
3279:
3272:
3264:
3257:
3241:Ceratosauria
3226:
3219:
3144:
3137:
3095:
3087:
3080:
3072:
3064:
3057:
3050:
3043:
3023:
3018:Podokesaurus
3016:
3009:
3002:
2994:
2970:
2962:
2955:
2948:
2941:
2934:
2927:
2920:
2913:
2906:
2901:Gojirasaurus
2899:
2894:Dracovenator
2892:
2884:
2877:
2865:Neotheropoda
2850:
2842:
2834:
2828:Guaibasaurus
2826:
2819:
2811:
2803:
2797:Chindesaurus
2795:
2685:
2676:
2599:
2579:
2575:
2565:
2538:
2534:
2530:
2524:
2479:
2473:
2446:Loewen, M.A.
2412:
2408:
2399:
2391:
2386:
2349:
2345:
2339:
2311:
2306:
2289:
2285:
2240:
2231:
2224:
2218:
2211:Iuticosaurus
2209:
2203:
2197:
2191:
2187:Brighstoneus
2185:
2179:
2173:
2167:
2163:; the basal
2158:
2152:
2150:spinosaurids
2145:Aristosuchus
2143:
2134:
2125:
2116:paleoclimate
2112:Wessex Basin
2107:
2091:
2085:
2073:
2069:
2045:
2043:
1925:
1896:
1880:
1879:
1850:
1824:Dryptosaurus
1822:
1821:
1748:
1747:
1725:
1704:
1703:
1681:
1655:
1654:
1625:
1604:Aniksosaurus
1602:
1601:
1572:
1534:
1533:
1505:
1504:
1476:
1475:
1393:
1392:
1371:
1370:
1347:
1343:
1339:
1335:
1333:
1300:
1299:
1284:
1283:
1282:
1266:
1265:
1244:
1243:
1227:
1226:
1200:
1199:
1175:
1172:megaraptoran
1167:
1165:
1070:
1045:
1044:
1028:
1027:
997:
996:
964:
963:
947:
946:
916:
915:
892:
891:
868:
867:
845:
844:
843:
820:
819:
769:
768:
752:
751:
728:
727:
695:
694:
678:
677:
661:
660:
621:
619:
614:Aviatyrannis
612:
603:
596:
591:
589:
570:
557:
551:
546:
541:part of the
532:
528:
511:
491:
484:Darren Naish
479:
477:
449:
443:
423:
385:
384:
383:
370:
360:
349:
346:"Fusinasus"
328:
320:
315:
299:
298:
284:
277:
276:
254:
228:
215:
202:
189:
24:
18:
6770:Maniraptora
6759:Maniraptora
6740:Tototlmimus
6670:Anserimimus
6620:Garudimimus
6606:Beishanlong
6548:Valdoraptor
6489:Arkansaurus
6434:Xunmenglong
6311:Juravenator
6105:Tarbosaurus
5981:Nanuqsaurus
5950:Gorgosaurus
5866:Megaraptora
5852:Timurlengia
5584:Sciurumimus
5532:Aratasaurus
5400:Murusraptor
5322:Fukuiraptor
5285:Megaraptora
5167:Shaochilong
5139:Eocarcharia
5092:Megaraptora
5012:Lusovenator
4890:Alpkarakush
4747:Carnosauria
4598:Spinosaurus
4571:Siamosaurus
4471:Suchosaurus
4418:Iberospinus
4354:Magnosaurus
4333:Afrovenator
4306:Torvosaurus
4057:Chilesaurus
3870:Carnotaurus
3863:Caletodraco
3849:Abelisaurus
3675:Arcovenator
3522:Laevisuchus
3513:Noasaurinae
3415:Berthasaura
3386:Noasauridae
3316:Genyodectes
3281:Lukousaurus
3274:Jubbulpuria
3074:Panguraptor
3052:Coelophysis
3045:Camposaurus
3004:Panguraptor
2996:Dracoraptor
2972:Zupaysaurus
2957:Tachiraptor
2936:Sarcosaurus
2836:Nhandumirim
2556:11336/12129
2450:Irmis, R.B.
2292:: 227–242.
2233:Yaverlestes
2220:Polacanthus
2205:Eucamerotus
2199:Ornithopsis
2193:Valdosaurus
2054:Megaraptora
1657:Fukuiraptor
1561:Megaraptora
1202:Fukuiraptor
1188:Megaraptora
500:Description
438:Map of the
373:nomen nudum
363:nomen nudum
352:nomen nudum
348:Hunt, 2002
6940:Categories
6846:Eotyrannus
6698:Gallimimus
6627:Harpymimus
6572:Harpymimus
6193:Iliosuchus
6163:see below↓
5992:Alioramini
5809:Juratyrant
5795:Eotyrannus
5780:Bagaraatan
5732:Yutyrannus
5666:Coeluridae
5592:Vayuraptor
5438:see below↓
5414:Tratayenia
5407:Orkoraptor
5393:Megaraptor
5299:Aoniraptor
5232:Mapusaurus
5160:Sauroniops
5124:Altispinax
5077:Neovenator
5019:Siamraptor
5005:Datanglong
4972:Epanterias
4964:Antrodemus
4957:Allosaurus
4938:Allosauria
4761:Altispinax
4726:Gasosaurus
4638:see below↓
4514:Suchomimus
4115:Sinosaurus
3950:see below↓
3906:Viavenator
3856:Aucasaurus
3793:Ilokelesia
3764:Dahalokely
3719:Rajasaurus
3697:Indosaurus
3690:Genusaurus
3682:Dahalokely
3606:Indosuchus
3502:Limusaurus
3461:Ligabueino
3454:Kiyacursor
3446:Genusaurus
3430:Dahalokely
3350:Betasuchus
3221:Dornraptor
3121:see below↓
3097:Segisaurus
2950:Sinosaurus
2813:Eodromaeus
2742:see below↓
2690:Dinosauria
2672:Sauropsida
2531:Megaraptor
2352:: e12727.
2261:References
2238:eobaatarid
2225:Eotyrannus
2216:ankylosaur
2214:; and the
2136:Neovenator
2108:Eotyrannus
2092:Eotyrannus
2070:Eotyrannus
2046:Eotyrannus
1750:Orkoraptor
1727:Megaraptor
1506:Eotyrannus
1478:Yutyrannus
1422:Coeluridae
1373:Juratyrant
1348:Eotyrannus
1344:Eotyrannus
1340:Yutyrannus
1336:Eotyrannus
1302:Megaraptor
1285:Eotyrannus
1268:Orkoraptor
1177:Megaraptor
1168:Eotyrannus
597:Eotyrannus
592:Eotyrannus
547:Eotyrannus
480:Eotyrannus
444:Eotyrannus
386:Eotyrannus
278:Eotyrannus
209:Saurischia
196:Dinosauria
25:Eotyrannus
6930:Dinosaurs
6774:includes
6719:Rativates
6712:Qiupalong
6412:Scipionyx
6405:Mirischia
6078:Lythronax
6070:Labocania
6001:Alioramus
5654:Labocania
5372:Aerosteon
4918:Sinraptor
4822:Erectopus
4557:Irritator
4142:Orionides
4027:Tetanurae
4014:Tetanurae
3941:Tetanurae
3877:Elemgasem
3800:Llukalkan
3779:Elemgasem
3536:Noasaurus
3399:Afromimus
3207:Averostra
3194:Averostra
3112:Averostra
2964:Velocipes
2783:Theropoda
2770:Theropoda
2733:Theropoda
2658:Kingdom:
2642:Theropoda
2541:: 35–55.
2429:129684676
2175:Iguanodon
2102:to early
2100:Barremian
1683:Aerosteon
1246:Aerosteon
496:in 2022.
293:Species:
222:Theropoda
169:Kingdom:
163:Eukaryota
37:130
31:Barremian
6855:Wikidata
6303:Coelurus
5711:Kileskus
5704:Guanlong
5676:Coelurus
5569:Gualicho
5069:Gualicho
4450:Baryonyx
3786:Guemesia
3613:Kryptops
3374:Ozraptor
3011:Pendraig
2666:Chordata
2664:Phylum:
2660:Animalia
2516:24223179
2475:PLoS ONE
2464:(2013).
2378:35821895
2249:See also
2242:Eobaatar
2236:and the
2090:, where
2072:chasing
1170:to be a
610:European
558:E. lengi
523:foramina
488:GoFundMe
452:holotype
401:dinosaur
398:theropod
338:Synonyms
183:Chordata
179:Phylum:
173:Animalia
159:Domain:
6888:4128446
6875:4822453
6861:Q131194
5844:Timimus
5606:Zuolong
5336:Rapator
5239:Meraxes
4564:Oxalaia
3885:Koleken
2908:Lepidus
2670:Class:
2507:3819173
2484:Bibcode
2468:(ed.).
2369:9271276
1260:unnamed
562:maxilla
535:cranial
514:rostral
465:Sandown
318:†
297:†
270:Genus:
106:↓
6916:Portal
6512:Hexing
6200:Kakuru
5788:Dilong
3829:Thanos
3807:Niebla
3627:Rugops
3620:Kurupi
2514:
2504:
2427:
2376:
2366:
2208:, and
2190:, and
2139:, the
2104:Aptian
622:et al.
539:distal
518:mesial
424:et al.
331:, 2001
329:et al.
287:, 2001
285:et al.
6901:68315
6776:birds
5816:Moros
5343:Siats
5084:Siats
5026:Siats
2686:Clade
2677:Clade
2582:(1).
2425:S2CID
2346:PeerJ
2282:(PDF)
2106:when
1574:Siats
543:tibia
493:PeerJ
391:genus
327:Hutt
283:Hutt
255:Clade
229:Clade
216:Clade
203:Clade
190:Clade
6870:GBIF
6768:see
5386:Maip
2844:Tawa
2719:see
2512:PMID
2374:PMID
2157:and
2126:The
2086:The
608:and
459:and
45:PreꞒ
2584:doi
2551:hdl
2543:doi
2502:PMC
2492:doi
2417:doi
2364:PMC
2354:doi
2294:doi
393:of
6942::
6898::
6885::
6872::
6857::
2688::
2679::
2580:63
2578:.
2574:.
2549:.
2539:51
2537:.
2510:.
2500:.
2490:.
2478:.
2472:.
2460:;
2456:;
2452:;
2448:;
2437:^
2423:.
2413:68
2411:.
2372:.
2362:.
2350:10
2348:.
2344:.
2322:^
2290:22
2288:.
2284:.
2269:^
2245:.
2202:,
2184:,
2178:,
2056:.
1180:.
475:.
442:;
418:,
257::
231::
218::
205::
192::
95:Pg
39:Ma
6918::
6638:?
6630:?
6543:?
6507:?
6499:?
6467:?
6415:?
6386:?
6371:?
6306:?
6073:?
6058:?
5907:?
5869:?
5847:?
5783:?
5686:?
5668:?
5657:?
5587:?
5572:?
5564:?
5549:?
5353:?
5345:?
5317:?
5309:?
5287:?
5127:?
5095:?
5087:?
5072:?
5064:?
5029:?
4983:?
4975:?
4967:?
4865:?
4840:?
4832:?
4787:?
4779:?
4771:?
4737:?
4729:?
4545:?
4482:?
4474:?
4428:?
4406:?
4371:?
4250:?
4215:?
4118:?
4075:?
4060:?
4052:?
4044:?
3880:?
3817:?
3767:?
3742:?
3714:?
3685:?
3594:?
3449:?
3441:?
3433:?
3425:?
3410:?
3402:?
3377:?
3369:?
3361:?
3353:?
3284:?
3269:?
3231:?
3092:?
3077:?
3069:?
2999:?
2967:?
2889:?
2855:?
2847:?
2839:?
2831:?
2816:?
2808:?
2800:?
2634:e
2627:t
2620:v
2590:.
2586::
2559:.
2553::
2545::
2518:.
2494::
2486::
2480:8
2431:.
2419::
2380:.
2356::
2300:.
2296::
375:)
371:(
365:)
361:(
354:)
350:(
274:†
261:†
245:†
100:N
90:K
85:J
80:T
75:P
70:C
65:D
60:S
55:O
50:Ꞓ
34:~
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