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Macrogryphosaurus

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3894: 3889: 2979: 2993: 119: 3876: 1933: 132: 2986: 3900: 494: 513:, and the largest species of elasmarian. Approaching the size of many more derived iguanodontian ornithopods, it is, compared to these derived taxa, much more graceful and lightly built in form. One of the most remarked upon features of the taxon is its mineralized thoracic plates, also referred to as intercostal plates. These are a series of plates along the side of the torso, shared with a handful of other ornithischians such as 375: 1887: 3883: 1921: 1899: 504:
has been noted for its large size compared to other South American ornithopods. It has been estimated to have been around 6 metres (20 ft) in length, though the only known individual may not have been fully grown. Gregory S. Paul estimated the length of this specimen at 5 m (16 ft) in
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received a complete osteology in 2020, a study published in Cretaceous Research by Sebastián Rozadilla, Penélope Cruzado-Caballero, Jorge O. Calvo. This described the anatomy of the species in much more thoroughly and investigating its classification and biomechanics under a more modern lens. They
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specimen, MUCPv-32, consists of an essentially complete set of vertebra, a number of ribs, four mineralized thoracic plates, both sides of the pectoral girdle, and a sternum. The individual is considered to have been an adult. The preservation of the fossil was described by
1843:"New small-bodied ornithopods (Dinosauria, Neornithischia) from the Early Cretaceous Wonthaggi Formation (Strzelecki Group) of the Australian-Antarctic rift system, with revision of Qantassaurus intrepidus Rich and Vickers-Rich, 1999" 402:, an articulated, nearly complete dinosaur skeleton was discovered and excavated. It was brought to the attention of the palaeontologists by young boy Rafael Moyano, who had discovered it. Originally reported to hail from the 1719: 485:
considered the genus to be of importance, due to being the largest known elasmarian species, and for the completeness of its known remains, informing on the anatomy of the group.
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Subsequent research has not referred any other specimens to the species. However, in 2016, a study was published on MAU-Pv-PH-458, a fragmentary dorsal neural arch from the
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location, though it lived at a slightly earlier time. Other indeterminate ornithopod specimens from Argentina similarly indicate animals of similar size to the
1599: 4357: 3893: 531:. Along with a fused, birdlike sternum, its short torso would have been rather stiff in nature. This, alongside its relatively elongate neck and 2978: 4459: 3888: 1718:
Ibiricu, Lucio M.; Martínez, Rubén D.; Lamanna, Matthew C.; Casal, Gabriel A.; Luna, Marcelo; Harris, Jerald D.; Lacovara, Kenneth J. (2010).
4464: 1965: 1817: 2992: 1891: 1624:"Osteology of Ornithopod Macrogryphosaurus gondwanicus (Dinosauria, Ornithischia) from the Upper Cretaceous of Patagonia, Argentina" 432:
in 2007, in a study authored by Jorge O. Calvo, Juan D. Porfiri, and Fernando E. Novas. Its generic name was derived from the Greek
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of northern Argentina. The bone is the northernmost record of an ornithopod in South America. It was not determined to belong to a
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Cruzado-Caballero, Penélope; Filippi, Leonardo S.; Méndez, Ariel H.; Garrido, Alberto C.; Valieri, Rubén D.Juárez (2016).
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Cruzado-Caballero, Penélope; Filippi, Leonardo S.; Méndez, Ariel H.; Garrido, Alberto C.; Díaz-Martínez, Ignacio (2018).
1576:"Discovery of a new ornithopod dinosaur from the Portezuelo formation (Upper Cretaceous), Neuquen, Patagonia, Argentina" 1903: 3899: 458: 1911: 2985: 1958: 2065: 411: 341: 472:, but noted to be from an animal of similar size and anatomy, and from approximately the same geographic and 4429: 118: 4434: 4312: 2424: 2276: 1670: 4396: 4234: 2790: 2092: 1671:"New record of ornithopod dinosaur from the Plottier Formation (Upper Cretaceous), Patagonia, Argentina" 505:
length and 300 kg (660 lb) in body mass. As such, it is the largest species of South American
287: 535:-like tail, gave it a rather unique form compared to that documented in other ornithischian dinosaurs. 1758: 1623: 4424: 4344: 3654: 1951: 1854: 1770: 1682: 1635: 403: 264: 3882: 1937: 593: 4006: 3617: 2202: 1739: 1651: 465: 126: 4401: 4383: 4375: 4013: 3752: 3673: 3633: 2517: 2403: 2349: 1823: 1813: 4388: 4154: 4147: 3957: 3950: 3479: 3390: 3228: 3129: 3115: 2829: 2797: 2432: 2341: 2255: 2190: 1925: 1862: 1786: 1778: 1731: 1698: 1690: 1643: 1371: 887: 425:. Noted for bony plates on its thorax, it was identified as a large species of ornithopod. 407: 387: 345: 338: 4239: 4034: 3985: 3848: 3779: 3682: 3502: 3221: 3207: 3186: 2804: 2677: 2444: 2212: 2050: 1073: 856: 815: 422: 1858: 1774: 1686: 1639: 493: 4249: 4244: 4182: 4119: 3992: 3841: 3804: 3786: 3731: 3363: 3179: 3172: 3101: 3023: 3010: 2938: 2926: 2895: 2870: 2734: 2656: 2617: 2603: 2480: 2391: 2313: 2291: 2284: 2025: 1348: 1234: 1217: 1184: 999: 801: 654: 521: 208: 4418: 4189: 4133: 4112: 4083: 3999: 3936: 3834: 3827: 3813: 3723: 3703: 3696: 3593: 3543: 3536: 3520: 3495: 3468: 3461: 3454: 3425: 3376: 3338: 3303: 3284: 3256: 3200: 2863: 2856: 2811: 2741: 2610: 2469: 2454: 2416: 2327: 2220: 2183: 2176: 1655: 1575: 1395: 1322: 958: 925: 724: 669: 527: 510: 418: 69: 1743: 374: 4097: 4073: 4061: 4027: 4020: 3971: 3964: 3770: 3714: 3689: 3661: 3550: 3352: 3317: 3263: 3193: 3165: 3051: 3044: 2884: 2849: 2776: 2769: 2691: 2632: 2587: 2489: 2363: 2356: 2305: 2269: 2227: 2077: 2016: 1335: 1268: 1102: 839: 579: 473: 363: 192: 1782: 1647: 1694: 4286: 4175: 4140: 3978: 3820: 3793: 3745: 3738: 3571: 3557: 3529: 3509: 3488: 3432: 3411: 3404: 3397: 3331: 3324: 3277: 3242: 3214: 3150: 3143: 2912: 2762: 2748: 2720: 2698: 2640: 2503: 2334: 2298: 2262: 2158: 2145: 2104: 1974: 1308: 1087: 741: 693: 640: 532: 224: 44: 1886: 4228: 4203: 4196: 4168: 4161: 4052: 4041: 3927: 3626: 3606: 3564: 3439: 3291: 3158: 3108: 3073: 3066: 3037: 2877: 2755: 2727: 2596: 2569: 2544: 2384: 2377: 2370: 2248: 1411: 1251: 1136: 982: 908: 781: 548: 543: 515: 506: 395: 89: 54: 4335: 1827: 4210: 4126: 4105: 3943: 3640: 3578: 3087: 3059: 2919: 2783: 2713: 2684: 2510: 2496: 2239: 1122: 564: 557: 415: 391: 353: 349: 334: 327: 240: 143: 94: 38: 30: 1841:
Herne, Matthew C.; Nair, Jay P.; Evans, Alistair R.; Tait, Alan M. (2019).
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Rozadilla, Sebastián; Cruzado-Caballero, Penélope; Calvo, Jorge O. (2020).
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Satellite image of the terrain of the Neuquén Group, the geologic unit
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Poropat, S. F. The Winston Churchill Memorial Trust of Australia.
2462: 4310: 3915: 3008: 2143: 1985: 1947: 356:. It was described by Jorge Calvo and colleagues in 2007, with 1943: 1574:
Calvo, Jorge O.; Porfiri, Juan D.; Novas, Fernando E. (2007).
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of the same geologic group. This unit is dated to the
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The specific epithet 1809:The Princeton Field Guide to Dinosaurs 322:(meaning "big enigmatic lizard") is a 567:below results from analysis by Herne 7: 1387: 1363: 1341: 1314: 1209: 1159: 1152: 1128: 1118: 1094: 1065: 974: 950: 900: 893: 831: 807: 797: 773: 766: 716: 709: 685: 661: 632: 608: 586: 574: 448:refers to the ancient continent of 386:In May 1999, during field work at 14: 3898: 3892: 3887: 3881: 3874: 2991: 2984: 2977: 1931: 1919: 1897: 1885: 130: 4255:List of informal dinosaur names 394:(sixty kilometres northwest of 4265:Timeline of hadrosaur research 400:National University of Comahue 1: 4460:Fossil taxa described in 2007 1783:10.1016/j.cretres.2017.07.022 1648:10.1016/j.cretres.2019.104311 430:Macrogryphosaurus gondwanicus 300:Macrogryphosaurus gondwanicus 4465:Taxa named by Fernando Novas 1695:10.1016/j.annpal.2016.02.003 4486: 16:Extinct genus of dinosaurs 3922: 3911: 3872: 3601: 3371: 3312: 3032: 3017: 3004: 2975: 2837: 2665: 2557: 2553: 2411: 2171: 2167: 2152: 2139: 2085: 2058: 2045: 1981: 1806:Paul, Gregory S. (2016). 1724:Annals of Carnegie Museum 1409: 1392: 1385: 1368: 1361: 1346: 1339: 1319: 1312: 1265: 1248: 1231: 1214: 1207: 1181: 1164: 1157: 1150: 1133: 1126: 1116: 1099: 1092: 1070: 1063: 996: 979: 972: 955: 948: 922: 905: 898: 891: 853: 836: 829: 812: 805: 795: 778: 771: 764: 738: 721: 714: 707: 690: 683: 666: 659: 637: 630: 613: 606: 591: 584: 440:, meaning enigmatic, and 293: 286: 127:Scientific classification 125: 116: 23: 1675:Annales de PalĂ©ontologie 461:as "amazing" in nature. 412:Sierra Barrosa Formation 342:Sierra Barrosa Formation 1847:Journal of Paleontology 560:, for the two genera. 498: 459:Dr. Stephen F. Poropat 383: 4397:Paleobiology Database 4235:Basturs Poble bonebed 496: 377: 4470:Ornithischian genera 4455:Portezuelo Formation 4450:Fossils of Argentina 4445:Cretaceous Argentina 4257:(contains hadrosaur 3655:Probrachylophosaurus 1906:at Wikimedia Commons 1736:10.2992/007.079.0103 404:Portezuelo Formation 370:Discovery and naming 1868:10.1017/jpa.2018.95 1859:2019JPal...93..543H 1775:2018CrRes..83..182C 1763:Cretaceous Research 1687:2016AnPal.102..145C 1640:2020CrRes.10804311R 1628:Cretaceous Research 594:Heterodontosauridae 398:) conducted by the 279:M. gondwanicus 4007:Ornithopodichnites 3618:Brachylophosaurini 2203:Hypsilophodontidae 499: 466:Plottier Formation 384: 366:and only species. 4412: 4411: 4384:Open Tree of Life 4321:Macrogryphosaurus 4313:Taxon identifiers 4304: 4303: 4300: 4299: 4296: 4295: 4014:Shiraminesauropus 3907: 3906: 3870: 3869: 3866: 3865: 3862: 3861: 3858: 3857: 3753:Naashoibitosaurus 3674:Austrokritosauria 3634:Brachylophosaurus 3588: 3587: 3000: 2999: 2973: 2972: 2969: 2968: 2965: 2964: 2961: 2960: 2957: 2956: 2650: 2649: 2538: 2537: 2534: 2533: 2518:Transylvanosaurus 2404:Rhabdodontomorpha 2350:Notohypsilophodon 2321:Macrogryphosaurus 2135: 2134: 2131: 2130: 2127: 2126: 2123: 2122: 1904:Macrogryphosaurus 1902:Media related to 1892:Macrogryphosaurus 1819:978-1-78684-190-2 1548: 1547: 1539: 1538: 1530: 1529: 1521: 1520: 1512: 1511: 1503: 1502: 1494: 1493: 1485: 1484: 1476: 1475: 1467: 1466: 1458: 1457: 1449: 1448: 1440: 1439: 1431: 1430: 1422: 1421: 1298: 1297: 1289: 1288: 1280: 1279: 1196: 1195: 1168:Macrogryphosaurus 1047: 1046: 1038: 1037: 1029: 1028: 1020: 1019: 1011: 1010: 937: 936: 877: 876: 868: 867: 753: 752: 502:Macrogryphosaurus 470:Macrogryphosaurus 436:, meaning large, 380:Macrogryphosaurus 319:Macrogryphosaurus 315: 314: 267: 255:Macrogryphosaurus 25:Macrogryphosaurus 4477: 4405: 4404: 4392: 4391: 4379: 4378: 4366: 4365: 4353: 4352: 4340: 4339: 4338: 4308: 4281:mummy S.M. R4036 4155:Microhadrosaurus 4148:Mandschurosaurus 3958:Camptosaurichnus 3951:Bonaparteichnium 3924: 3917:Related articles 3913: 3902: 3896: 3891: 3886: 3885: 3878: 3599: 3480:Parasaurolophini 3391:Angulomastacator 3369: 3310: 3229:Shuangmiaosaurus 3130:Huehuecanauhtlus 3116:Gongpoquansaurus 3030: 3019: 3006: 2995: 2989: 2988: 2981: 2835: 2830:Hadrosauriformes 2798:Ratchasimasaurus 2663: 2555: 2551: 2433:Muttaburrasaurus 2409: 2342:Muttaburrasaurus 2256:Atlascopcosaurus 2191:Thescelosauridae 2169: 2165: 2154: 2141: 2083: 2056: 2047: 2040: 2039: 1983: 1968: 1961: 1954: 1945: 1936: 1935: 1934: 1924: 1923: 1922: 1915: 1901: 1890:Data related to 1889: 1873: 1872: 1870: 1838: 1832: 1831: 1803: 1797: 1796: 1794: 1754: 1748: 1747: 1715: 1709: 1708: 1706: 1666: 1660: 1659: 1619: 1602: 1597: 1588: 1587: 1571: 1388: 1372:Muttaburrasaurus 1364: 1342: 1315: 1210: 1160: 1153: 1129: 1119: 1095: 1066: 975: 951: 901: 894: 888:Thescelosauridae 832: 808: 798: 774: 767: 717: 710: 686: 662: 633: 609: 587: 575: 388:Mari Menuco Lake 339:Upper Cretaceous 302: 298: 259: 252: 239: 223: 207: 191: 135: 134: 121: 111: 41: 29:Temporal range: 21: 4485: 4484: 4480: 4479: 4478: 4476: 4475: 4474: 4415: 4414: 4413: 4408: 4400: 4395: 4387: 4382: 4374: 4369: 4361: 4356: 4348: 4343: 4334: 4333: 4328: 4315: 4305: 4292: 4273:mummy AMNH 5060 4240:Dakota (fossil) 4216: 4067: 4047: 4035:Tetrapodosaurus 3986:Hadrosaurichnus 3918: 3903: 3880: 3854: 3849:Shantungosaurus 3799: 3780:Augustynolophus 3765: 3709: 3683:Bonapartesaurus 3668: 3612: 3584: 3515: 3503:Parasaurolophus 3474: 3358: 3306: 3297: 3222:Qianjiangsaurus 3208:Probactrosaurus 3187:Penelopognathus 3026: 3013: 2996: 2983: 2953: 2933: 2890: 2824: 2805:Riabininohadros 2678:Bayannurosaurus 2659: 2646: 2582: 2547: 2530: 2475: 2445:Tenontosauridae 2439: 2398: 2234: 2213:Gideonmantellia 2197: 2161: 2148: 2119: 2099: 2072: 2066:Avemetatarsalia 2051:Avemetatarsalia 2041: 1988: 1987: 1977: 1972: 1942: 1932: 1930: 1920: 1918: 1910: 1882: 1877: 1876: 1840: 1839: 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3436: 3429: 3422: 3415: 3408: 3401: 3394: 3387: 3380: 3372: 3366: 3364:Lambeosaurinae 3360: 3359: 3357: 3356: 3349: 3342: 3335: 3328: 3321: 3313: 3307: 3302: 3299: 3298: 3296: 3295: 3288: 3281: 3274: 3267: 3260: 3253: 3246: 3239: 3232: 3225: 3218: 3211: 3204: 3197: 3190: 3183: 3180:Nanyangosaurus 3176: 3173:Nanningosaurus 3169: 3162: 3155: 3147: 3140: 3133: 3126: 3119: 3112: 3105: 3102:Gilmoreosaurus 3098: 3091: 3084: 3077: 3070: 3063: 3056: 3048: 3041: 3033: 3027: 3024:Hadrosauroidea 3022: 3015: 3014: 3011:Hadrosauroidea 3009: 3002: 3001: 2998: 2997: 2976: 2974: 2971: 2970: 2967: 2966: 2963: 2962: 2959: 2958: 2955: 2954: 2952: 2951: 2950: 2949: 2943: 2941: 2939:Hadrosauroidea 2935: 2934: 2932: 2931: 2927:Mantellisaurus 2923: 2916: 2909: 2900: 2898: 2896:Iguanodontidae 2892: 2891: 2889: 2888: 2881: 2874: 2871:Mantellisaurus 2867: 2860: 2853: 2846: 2838: 2832: 2826: 2825: 2823: 2822: 2815: 2808: 2801: 2794: 2787: 2780: 2773: 2766: 2759: 2752: 2745: 2738: 2735:Iguanacolossus 2731: 2724: 2717: 2710: 2702: 2695: 2688: 2681: 2674: 2666: 2660: 2657:Ankylopollexia 2655: 2652: 2651: 2648: 2647: 2645: 2644: 2637: 2629: 2621: 2618:Eousdryosaurus 2614: 2607: 2604:Dysalotosaurus 2600: 2592: 2590: 2584: 2583: 2581: 2580: 2573: 2566: 2558: 2548: 2543: 2540: 2539: 2536: 2535: 2532: 2531: 2529: 2528: 2521: 2514: 2507: 2500: 2493: 2485: 2483: 2481:Rhabdodontidae 2477: 2476: 2474: 2473: 2466: 2459: 2450: 2448: 2441: 2440: 2438: 2437: 2429: 2421: 2412: 2406: 2400: 2399: 2397: 2396: 2392:Weewarrasaurus 2388: 2381: 2374: 2367: 2360: 2353: 2346: 2338: 2331: 2324: 2317: 2314:Leaellynasaura 2310: 2302: 2295: 2292:Gasparinisaura 2288: 2285:Galleonosaurus 2281: 2273: 2266: 2259: 2252: 2244: 2242: 2236: 2235: 2233: 2232: 2224: 2217: 2208: 2206: 2199: 2198: 2196: 2195: 2187: 2180: 2172: 2162: 2157: 2150: 2149: 2144: 2137: 2136: 2133: 2132: 2129: 2128: 2125: 2124: 2121: 2120: 2118: 2117: 2116: 2115: 2109: 2107: 2101: 2100: 2098: 2097: 2096: 2095: 2086: 2080: 2074: 2073: 2071: 2070: 2069: 2068: 2059: 2053: 2043: 2042: 2038: 2037: 2028: 2026:Neornithischia 2019: 2010: 2001: 1995: 1986: 1979: 1978: 1973: 1971: 1970: 1963: 1956: 1948: 1941: 1940: 1928: 1908: 1907: 1895: 1894:at Wikispecies 1881: 1880:External links 1878: 1875: 1874: 1853:(3): 543–584. 1833: 1818: 1798: 1749: 1710: 1681:(2): 145–150. 1661: 1603: 1589: 1556: 1555: 1553: 1550: 1546: 1545: 1542: 1541: 1537: 1536: 1533: 1532: 1528: 1527: 1524: 1523: 1519: 1518: 1515: 1514: 1510: 1509: 1506: 1505: 1501: 1500: 1497: 1496: 1492: 1491: 1488: 1487: 1483: 1482: 1479: 1478: 1474: 1473: 1470: 1469: 1465: 1464: 1461: 1460: 1456: 1455: 1452: 1451: 1447: 1446: 1443: 1442: 1438: 1437: 1434: 1433: 1429: 1428: 1425: 1424: 1420: 1419: 1416: 1415: 1408: 1405: 1404: 1401: 1400: 1391: 1386: 1384: 1381: 1380: 1377: 1376: 1367: 1362: 1360: 1357: 1356: 1353: 1352: 1349:Rhabdodontidae 1345: 1340: 1338: 1332: 1331: 1328: 1327: 1318: 1313: 1311: 1305: 1304: 1301: 1300: 1296: 1295: 1292: 1291: 1287: 1286: 1283: 1282: 1278: 1277: 1274: 1273: 1264: 1261: 1260: 1257: 1256: 1247: 1244: 1243: 1240: 1239: 1235:Leaellynasaura 1230: 1227: 1226: 1223: 1222: 1218:Galleonosaurus 1213: 1208: 1206: 1203: 1202: 1199: 1198: 1194: 1193: 1190: 1189: 1185:Gasparinisaura 1180: 1177: 1176: 1173: 1172: 1163: 1158: 1156: 1151: 1149: 1146: 1145: 1142: 1141: 1132: 1127: 1125: 1117: 1115: 1112: 1111: 1108: 1107: 1098: 1093: 1091: 1083: 1082: 1079: 1078: 1069: 1064: 1062: 1054: 1053: 1050: 1049: 1045: 1044: 1041: 1040: 1036: 1035: 1032: 1031: 1027: 1026: 1023: 1022: 1018: 1017: 1014: 1013: 1009: 1008: 1005: 1004: 1000:Thescelosaurus 995: 992: 991: 988: 987: 978: 973: 971: 968: 967: 964: 963: 954: 949: 947: 944: 943: 940: 939: 935: 934: 931: 930: 921: 918: 917: 914: 913: 904: 899: 897: 892: 890: 884: 883: 880: 879: 875: 874: 871: 870: 866: 865: 862: 861: 852: 849: 848: 845: 844: 835: 830: 828: 825: 824: 821: 820: 811: 806: 804: 802:Jeholosauridae 796: 794: 791: 790: 787: 786: 777: 772: 770: 765: 763: 760: 759: 756: 755: 751: 750: 747: 746: 737: 734: 733: 730: 729: 720: 715: 713: 708: 706: 703: 702: 699: 698: 689: 684: 682: 679: 678: 675: 674: 665: 660: 658: 655:Neornithischia 650: 649: 646: 645: 636: 631: 629: 626: 625: 622: 621: 612: 607: 605: 602: 601: 598: 597: 590: 585: 583: 573: 540: 539:Classification 537: 522:Thescelosaurus 490: 487: 482:M. gondwanicus 371: 368: 359:M. gondwanicus 313: 312: 305: 291: 290: 284: 283: 275: 273: 269: 268: 248: 244: 243: 235: 228: 227: 219: 212: 211: 209:Neornithischia 203: 196: 195: 187: 180: 179: 174: 167: 166: 161: 157: 156: 151: 147: 146: 141: 137: 136: 123: 122: 114: 113: 105: 104: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 43: 42: 28: 15: 13: 10: 9: 6: 4: 3: 2: 4482: 4471: 4468: 4466: 4463: 4461: 4458: 4456: 4453: 4451: 4448: 4446: 4443: 4441: 4438: 4436: 4435:Turonian life 4433: 4431: 4428: 4426: 4423: 4422: 4420: 4403: 4398: 4394: 4390: 4385: 4381: 4377: 4372: 4368: 4364: 4359: 4355: 4351: 4346: 4342: 4337: 4331: 4327: 4326: 4324: 4322: 4318: 4314: 4309: 4289: 4288: 4284: 4282: 4280: 4279:Edmontosaurus 4276: 4274: 4272: 4271:Edmontosaurus 4268: 4266: 4263: 4260: 4256: 4253: 4251: 4248: 4246: 4243: 4241: 4238: 4236: 4233: 4231: 4230: 4226: 4225: 4223: 4219: 4213: 4212: 4208: 4206: 4205: 4201: 4199: 4198: 4194: 4192: 4191: 4190:Syngonosaurus 4187: 4185: 4184: 4180: 4178: 4177: 4173: 4171: 4170: 4166: 4164: 4163: 4159: 4157: 4156: 4152: 4150: 4149: 4145: 4143: 4142: 4138: 4136: 4135: 4134:Koutalisaurus 4131: 4129: 4128: 4124: 4122: 4121: 4117: 4115: 4114: 4113:Eucercosaurus 4110: 4108: 4107: 4103: 4100: 4099: 4095: 4093: 4092: 4088: 4086: 4085: 4084:Arstanosaurus 4081: 4080: 4078: 4076: 4075: 4070: 4064: 4063: 4059: 4058: 4056: 4054: 4050: 4044: 4043: 4039: 4037: 4036: 4032: 4030: 4029: 4025: 4023: 4022: 4018: 4016: 4015: 4011: 4009: 4008: 4004: 4002: 4001: 4000:Orcauichnites 3997: 3995: 3994: 3990: 3988: 3987: 3983: 3981: 3980: 3976: 3974: 3973: 3969: 3967: 3966: 3962: 3960: 3959: 3955: 3953: 3952: 3948: 3946: 3945: 3941: 3939: 3938: 3937:Amblydactylus 3934: 3933: 3931: 3929: 3925: 3921: 3914: 3910: 3901: 3897: 3895: 3890: 3884: 3877: 3851: 3850: 3846: 3844: 3843: 3839: 3837: 3836: 3835:Kundurosaurus 3832: 3830: 3829: 3828:Kerberosaurus 3825: 3823: 3822: 3818: 3816: 3815: 3814:Edmontosaurus 3811: 3810: 3808: 3806: 3802: 3796: 3795: 3791: 3789: 3788: 3784: 3782: 3781: 3777: 3776: 3774: 3772: 3768: 3762: 3761: 3757: 3755: 3754: 3750: 3748: 3747: 3743: 3741: 3740: 3736: 3734: 3733: 3729: 3726: 3725: 3724:Anasazisaurus 3721: 3720: 3718: 3716: 3712: 3706: 3705: 3704:Secernosaurus 3701: 3699: 3698: 3697:Kelumapusaura 3694: 3692: 3691: 3687: 3685: 3684: 3680: 3679: 3677: 3675: 3671: 3664: 3663: 3659: 3657: 3656: 3652: 3650: 3649: 3645: 3643: 3642: 3638: 3636: 3635: 3631: 3629: 3628: 3624: 3623: 3621: 3619: 3615: 3609: 3608: 3604: 3603: 3600: 3597: 3595: 3594:Saurolophinae 3591: 3581: 3580: 3576: 3574: 3573: 3569: 3567: 3566: 3562: 3560: 3559: 3555: 3553: 3552: 3548: 3546: 3545: 3544:Hypacrosaurus 3541: 3539: 3538: 3537:Corythosaurus 3534: 3532: 3531: 3527: 3526: 3524: 3522: 3521:Lambeosaurini 3518: 3512: 3511: 3507: 3505: 3504: 3500: 3498: 3497: 3496:Charonosaurus 3493: 3491: 3490: 3486: 3485: 3483: 3481: 3477: 3471: 3470: 3469:Tsintaosaurus 3466: 3464: 3463: 3462:Pararhabdodon 3459: 3457: 3456: 3455:Nipponosaurus 3452: 3450: 3449: 3445: 3442: 3441: 3437: 3435: 3434: 3430: 3428: 3427: 3426:Jaxartosaurus 3423: 3421: 3420: 3416: 3414: 3413: 3409: 3407: 3406: 3402: 3400: 3399: 3395: 3393: 3392: 3388: 3386: 3385: 3381: 3379: 3378: 3377:Adynomosaurus 3374: 3373: 3370: 3367: 3365: 3361: 3355: 3354: 3350: 3348: 3347: 3343: 3341: 3340: 3339:Lapampasaurus 3336: 3334: 3333: 3329: 3327: 3326: 3322: 3320: 3319: 3315: 3314: 3311: 3308: 3305: 3304:Hadrosauridae 3300: 3294: 3293: 3289: 3287: 3286: 3285:Zhanghenglong 3282: 3280: 3279: 3275: 3273: 3272: 3268: 3266: 3265: 3261: 3259: 3258: 3257:Telmatosaurus 3254: 3252: 3251: 3247: 3245: 3244: 3240: 3238: 3237: 3233: 3231: 3230: 3226: 3224: 3223: 3219: 3217: 3216: 3212: 3210: 3209: 3205: 3203: 3202: 3201:Portellsaurus 3198: 3196: 3195: 3191: 3189: 3188: 3184: 3182: 3181: 3177: 3175: 3174: 3170: 3168: 3167: 3163: 3161: 3160: 3156: 3153: 3152: 3148: 3146: 3145: 3141: 3139: 3138: 3134: 3132: 3131: 3127: 3125: 3124: 3120: 3118: 3117: 3113: 3111: 3110: 3106: 3104: 3103: 3099: 3097: 3096: 3092: 3090: 3089: 3085: 3083: 3082: 3078: 3076: 3075: 3071: 3069: 3068: 3064: 3062: 3061: 3057: 3054: 3053: 3049: 3047: 3046: 3042: 3040: 3039: 3035: 3034: 3031: 3028: 3025: 3020: 3016: 3012: 3007: 3003: 2994: 2990: 2987: 2980: 2947: 2946: 2945: 2944: 2942: 2940: 2936: 2929: 2928: 2924: 2922: 2921: 2917: 2915: 2914: 2910: 2907: 2906: 2902: 2901: 2899: 2897: 2893: 2887: 2886: 2882: 2880: 2879: 2875: 2873: 2872: 2868: 2866: 2865: 2864:Jinzhousaurus 2861: 2859: 2858: 2857:Hypselospinus 2854: 2852: 2851: 2847: 2845: 2844: 2840: 2839: 2836: 2833: 2831: 2827: 2821: 2820: 2816: 2814: 2813: 2812:Theiophytalia 2809: 2807: 2806: 2802: 2800: 2799: 2795: 2793: 2792: 2788: 2786: 2785: 2781: 2779: 2778: 2774: 2772: 2771: 2767: 2765: 2764: 2760: 2758: 2757: 2753: 2751: 2750: 2746: 2744: 2743: 2742:Lanzhousaurus 2739: 2737: 2736: 2732: 2730: 2729: 2725: 2723: 2722: 2718: 2716: 2715: 2711: 2708: 2707: 2703: 2701: 2700: 2696: 2694: 2693: 2689: 2687: 2686: 2682: 2680: 2679: 2675: 2673: 2672: 2668: 2667: 2664: 2661: 2658: 2653: 2643: 2642: 2638: 2635: 2634: 2630: 2627: 2626: 2622: 2620: 2619: 2615: 2613: 2612: 2611:Elrhazosaurus 2608: 2606: 2605: 2601: 2599: 2598: 2594: 2593: 2591: 2589: 2585: 2579: 2578: 2574: 2572: 2571: 2567: 2565: 2564: 2560: 2559: 2556: 2552: 2549: 2546: 2541: 2527: 2526: 2522: 2520: 2519: 2515: 2513: 2512: 2508: 2506: 2505: 2501: 2499: 2498: 2494: 2492: 2491: 2487: 2486: 2484: 2482: 2478: 2472: 2471: 2470:Tenontosaurus 2467: 2465: 2464: 2460: 2457: 2456: 2455:Convolosaurus 2452: 2451: 2449: 2446: 2442: 2435: 2434: 2430: 2427: 2426: 2422: 2419: 2418: 2417:Ampelognathus 2414: 2413: 2410: 2407: 2405: 2401: 2394: 2393: 2389: 2387: 2386: 2382: 2380: 2379: 2375: 2373: 2372: 2368: 2366: 2365: 2361: 2359: 2358: 2354: 2352: 2351: 2347: 2344: 2343: 2339: 2337: 2336: 2332: 2330: 2329: 2328:Mahuidacursor 2325: 2323: 2322: 2318: 2316: 2315: 2311: 2308: 2307: 2303: 2301: 2300: 2296: 2294: 2293: 2289: 2287: 2286: 2282: 2279: 2278: 2274: 2272: 2271: 2267: 2265: 2264: 2260: 2258: 2257: 2253: 2251: 2250: 2246: 2245: 2243: 2241: 2237: 2230: 2229: 2225: 2223: 2222: 2221:Hypsilophodon 2218: 2215: 2214: 2210: 2209: 2207: 2204: 2200: 2193: 2192: 2188: 2186: 2185: 2184:Callovosaurus 2181: 2179: 2178: 2177:Burianosaurus 2174: 2173: 2170: 2166: 2163: 2160: 2155: 2151: 2147: 2142: 2138: 2113: 2112: 2111: 2110: 2108: 2106: 2102: 2094: 2090: 2089: 2088: 2087: 2084: 2081: 2079: 2075: 2067: 2063: 2062: 2061: 2060: 2057: 2054: 2052: 2048: 2044: 2036: 2032: 2029: 2027: 2023: 2020: 2018: 2014: 2011: 2009: 2005: 2002: 2000: 1996: 1994: 1990: 1989: 1984: 1980: 1976: 1969: 1964: 1962: 1957: 1955: 1950: 1949: 1946: 1939: 1929: 1927: 1917: 1913: 1905: 1900: 1896: 1893: 1888: 1884: 1883: 1879: 1869: 1864: 1860: 1856: 1852: 1848: 1844: 1837: 1834: 1829: 1825: 1821: 1815: 1811: 1810: 1802: 1799: 1793: 1788: 1784: 1780: 1776: 1772: 1768: 1764: 1760: 1753: 1750: 1745: 1741: 1737: 1733: 1729: 1725: 1721: 1714: 1711: 1705: 1700: 1696: 1692: 1688: 1684: 1680: 1676: 1672: 1665: 1662: 1657: 1653: 1649: 1645: 1641: 1637: 1633: 1629: 1625: 1618: 1616: 1614: 1612: 1610: 1608: 1604: 1601: 1596: 1594: 1590: 1586:(4): 471–483. 1585: 1581: 1577: 1570: 1568: 1566: 1564: 1562: 1558: 1551: 1544: 1543: 1535: 1534: 1526: 1525: 1517: 1516: 1508: 1507: 1499: 1498: 1490: 1489: 1481: 1480: 1472: 1471: 1463: 1462: 1454: 1453: 1445: 1444: 1436: 1435: 1427: 1426: 1418: 1417: 1414: 1413: 1407: 1406: 1403: 1402: 1399: 1398: 1397: 1396:Tenontosaurus 1390: 1389: 1383: 1382: 1379: 1378: 1375: 1374: 1373: 1366: 1365: 1359: 1358: 1355: 1354: 1351: 1350: 1344: 1343: 1337: 1334: 1333: 1330: 1329: 1326: 1325: 1324: 1323:Hypsilophodon 1317: 1316: 1310: 1307: 1306: 1303: 1302: 1294: 1293: 1285: 1284: 1276: 1275: 1272: 1271: 1270: 1263: 1262: 1259: 1258: 1255: 1254: 1253: 1246: 1245: 1242: 1241: 1238: 1237: 1236: 1229: 1228: 1225: 1224: 1221: 1220: 1219: 1212: 1211: 1205: 1204: 1201: 1200: 1192: 1191: 1188: 1187: 1186: 1179: 1178: 1175: 1174: 1171: 1170: 1169: 1162: 1161: 1155: 1154: 1148: 1147: 1144: 1143: 1140: 1139: 1138: 1131: 1130: 1124: 1121: 1120: 1114: 1113: 1110: 1109: 1106: 1105: 1104: 1097: 1096: 1090: 1089: 1085: 1084: 1081: 1080: 1077: 1076: 1075: 1068: 1067: 1061: 1060: 1056: 1055: 1052: 1051: 1043: 1042: 1034: 1033: 1025: 1024: 1016: 1015: 1007: 1006: 1003: 1002: 1001: 994: 993: 990: 989: 986: 985: 984: 977: 976: 970: 969: 966: 965: 962: 961: 960: 959:Zephyrosaurus 953: 952: 946: 945: 942: 941: 933: 932: 929: 928: 927: 926:Koreanosaurus 920: 919: 916: 915: 912: 911: 910: 903: 902: 896: 895: 889: 886: 885: 882: 881: 873: 872: 864: 863: 860: 859: 858: 851: 850: 847: 846: 843: 842: 841: 834: 833: 827: 826: 823: 822: 819: 818: 817: 810: 809: 803: 800: 799: 793: 792: 789: 788: 785: 784: 783: 776: 775: 769: 768: 762: 761: 758: 757: 749: 748: 745: 744: 743: 736: 735: 732: 731: 728: 727: 726: 725:Hexinlusaurus 719: 718: 712: 711: 705: 704: 701: 700: 697: 696: 695: 688: 687: 681: 680: 677: 676: 673: 672: 671: 670:Lesothosaurus 664: 663: 657: 656: 652: 651: 648: 647: 644: 643: 642: 635: 634: 628: 627: 624: 623: 620: 619: 618: 611: 610: 604: 603: 600: 599: 596: 595: 589: 588: 582: 581: 577: 576: 572: 570: 566: 561: 559: 555: 551: 550: 545: 538: 536: 534: 530: 529: 528:Hypsilophodon 524: 523: 518: 517: 512: 511:Hadrosauridae 508: 503: 495: 488: 486: 483: 479: 475: 474:stratigraphic 471: 467: 462: 460: 455: 451: 447: 443: 439: 435: 431: 426: 424: 420: 417: 413: 409: 408:NeuquĂ©n Group 405: 401: 397: 393: 389: 381: 376: 369: 367: 365: 361: 360: 355: 351: 347: 346:NeuquĂ©n Group 343: 340: 336: 332: 329: 325: 321: 320: 309: 303: 301: 292: 289: 288:Binomial name 285: 281: 280: 274: 271: 270: 266: 263: 257: 256: 249: 246: 245: 242: 236: 233: 230: 229: 226: 220: 217: 214: 213: 210: 204: 201: 198: 197: 194: 188: 185: 182: 181: 178: 175: 172: 169: 168: 165: 162: 159: 158: 155: 152: 149: 148: 145: 142: 139: 138: 133: 128: 124: 120: 115: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 40: 32: 26: 22: 19: 4320: 4285: 4278: 4270: 4258: 4227: 4209: 4202: 4195: 4188: 4181: 4174: 4167: 4160: 4153: 4146: 4139: 4132: 4125: 4118: 4111: 4104: 4098:Claorhynchus 4096: 4089: 4082: 4074:Nomina dubia 4072: 4062:Guegoolithus 4060: 4040: 4033: 4028:Staurichnium 4026: 4021:Sousaichnium 4019: 4012: 4005: 3998: 3991: 3984: 3977: 3972:Dakotasaurus 3970: 3965:Caririchnium 3963: 3956: 3949: 3942: 3935: 3879: 3847: 3840: 3833: 3826: 3819: 3812: 3792: 3785: 3778: 3771:Saurolophini 3758: 3751: 3744: 3737: 3730: 3722: 3715:Kritosaurini 3702: 3695: 3690:Huallasaurus 3688: 3681: 3662:Wulagasaurus 3660: 3653: 3646: 3639: 3632: 3625: 3605: 3577: 3570: 3563: 3556: 3551:Lambeosaurus 3549: 3542: 3535: 3528: 3508: 3501: 3494: 3487: 3467: 3460: 3453: 3446: 3438: 3431: 3424: 3417: 3410: 3403: 3396: 3389: 3382: 3375: 3353:Yamatosaurus 3351: 3344: 3337: 3330: 3323: 3318:Aquilarhinus 3316: 3290: 3283: 3276: 3269: 3264:Tethyshadros 3262: 3255: 3248: 3241: 3234: 3227: 3220: 3213: 3206: 3199: 3194:Plesiohadros 3192: 3185: 3178: 3171: 3166:Lophorhothon 3164: 3157: 3149: 3142: 3135: 3128: 3121: 3114: 3107: 3100: 3093: 3086: 3079: 3072: 3065: 3058: 3052:Batyrosaurus 3050: 3045:Bactrosaurus 3043: 3036: 2982: 2925: 2918: 2911: 2903: 2885:Ouranosaurus 2883: 2876: 2869: 2862: 2855: 2850:Brighstoneus 2848: 2841: 2817: 2810: 2803: 2796: 2789: 2782: 2777:Osmakasaurus 2775: 2770:Oblitosaurus 2768: 2761: 2754: 2747: 2740: 2733: 2726: 2719: 2712: 2704: 2697: 2692:Camptosaurus 2690: 2683: 2676: 2669: 2639: 2633:Kangnasaurus 2631: 2623: 2616: 2609: 2602: 2595: 2588:Dryosauridae 2575: 2568: 2561: 2523: 2516: 2509: 2502: 2495: 2490:Matheronodon 2488: 2468: 2461: 2453: 2431: 2423: 2415: 2390: 2383: 2376: 2369: 2364:Sektensaurus 2362: 2357:Qantassaurus 2355: 2348: 2340: 2333: 2326: 2320: 2319: 2312: 2306:Kangnasaurus 2304: 2297: 2290: 2283: 2275: 2270:Diluvicursor 2268: 2261: 2254: 2247: 2228:Vectidromeus 2226: 2219: 2211: 2189: 2182: 2175: 2093:Ornithischia 2078:Ornithischia 2030: 2021: 2017:Ornithischia 2012: 2003: 1938:Paleontology 1850: 1846: 1836: 1808: 1801: 1766: 1762: 1752: 1730:(1): 39–50. 1727: 1723: 1713: 1678: 1674: 1664: 1631: 1627: 1583: 1579: 1410: 1394: 1393: 1370: 1369: 1347: 1336:Iguanodontia 1321: 1320: 1269:Diluvicursor 1267: 1266: 1250: 1249: 1233: 1232: 1216: 1215: 1183: 1182: 1167: 1166: 1165: 1135: 1134: 1103:Parksosaurus 1101: 1100: 1086: 1072: 1071: 1057: 998: 997: 981: 980: 957: 956: 924: 923: 907: 906: 855: 854: 840:Jeholosaurus 838: 837: 814: 813: 780: 779: 740: 739: 723: 722: 692: 691: 668: 667: 653: 639: 638: 615: 614: 592: 580:Ornithischia 578: 568: 562: 547: 544:Phylogenetic 542: 526: 520: 514: 501: 500: 481: 469: 463: 445: 441: 437: 433: 429: 427: 385: 379: 358: 357: 318: 317: 316: 307: 299: 294: 278: 277: 261: 254: 253: 231: 215: 199: 193:Ornithischia 183: 170: 24: 18: 4425:Elasmarians 4287:Willinakaqe 4259:nomina nuda 4176:Siluosaurus 4141:Loncosaurus 3979:Dinehichnus 3821:Kamuysaurus 3794:Saurolophus 3746:Kritosaurus 3739:Gryposaurus 3572:Sahaliyania 3558:Magnapaulia 3530:Amurosaurus 3510:Tlatolophus 3489:Adelolophus 3433:Kazaklambia 3412:Blasisaurus 3405:Arenysaurus 3398:Aralosaurus 3332:Hadrosaurus 3325:Eotrachodon 3278:Yunganglong 3243:Sirindhorna 3215:Protohadros 3151:Koshisaurus 3144:Jintasaurus 2913:Comptonatus 2763:Napaisaurus 2749:Lurdusaurus 2721:Fukuisaurus 2699:Cedrorestes 2641:Valdosaurus 2504:Pareisactus 2335:Morrosaurus 2299:Isasicursor 2263:Chakisaurus 2159:Ornithopoda 2146:Ornithopoda 2105:Ornithopoda 1975:Ornithopoda 1792:11336/71315 1769:: 182–193. 1704:11336/62352 1309:Clypeodonta 1088:Ornithopoda 742:Yandusaurus 694:Agilisaurus 641:Thyreophora 533:coelurosaur 509:outside of 489:Description 446:gondwanicus 225:Ornithopoda 37:93–89  4419:Categories 4229:Arkharavia 4204:Tichosteus 4197:Thespesius 4169:Pteropelyx 4162:Orthomerus 4042:Wintonopus 3627:Acristavus 3607:Barsboldia 3565:Olorotitan 3440:Latirhinus 3292:Zuoyunlong 3159:Levnesovia 3109:Gobihadros 3074:Datonglong 3067:Claosaurus 3038:Altirhinus 2948:see below↓ 2878:Morelladon 2756:Magnamanus 2728:Hippodraco 2597:Dryosaurus 2570:Hesperonyx 2545:Dryomorpha 2385:Trinisaura 2378:Tietasaura 2371:Talenkauen 2249:Anabisetia 2114:see below↓ 2008:Dinosauria 1634:: 104311. 1552:References 1412:Dryomorpha 1252:Anabisetia 1137:Talenkauen 983:Yueosaurus 909:Orodromeus 782:Nanosaurus 549:Talenkauen 516:Talenkauen 507:ornithopod 328:elasmarian 177:Dinosauria 4211:Trachodon 4127:Glishades 4106:Diclonius 3944:Anomoepus 3928:Ichnotaxa 3641:Maiasaura 3579:Velafrons 3088:Equijubus 3060:Choyrodon 2920:Iguanodon 2784:Planicoxa 2714:Dakotadon 2685:Calvarius 2511:Rhabdodon 2497:Mochlodon 2240:Elasmaria 1991:Kingdom: 1926:Dinosaurs 1828:985402380 1656:213679041 1123:Elasmaria 565:cladogram 558:Elasmaria 416:Coniacian 392:Argentina 354:Patagonia 350:Argentina 335:Coniacian 333:from the 272:Species: 241:Elasmaria 150:Kingdom: 144:Eukaryota 31:Coniacian 4350:47460013 4330:Wikidata 4091:Cionodon 3760:Rhinorex 3648:Ornatops 3448:Minqaria 3419:Canardia 3346:Malefica 3271:Xuwulong 3236:Siamodon 3137:Jeyawati 3123:Gonkoken 3081:Eolambia 2905:Barilium 2706:Cumnoria 2671:Barilium 2577:Owenodon 2563:Draconyx 2525:Zalmoxes 2425:Fostoria 2277:Fostoria 2035:Cerapoda 1999:Chordata 1997:Phylum: 1993:Animalia 1744:53407321 1059:Cerapoda 617:Eocursor 571:, 2019. 454:holotype 450:Gondwana 331:dinosaur 164:Chordata 160:Phylum: 154:Animalia 140:Domain: 4389:4143965 4376:1474735 4363:6141945 4336:Q138604 3384:Ajnabia 2819:Uteodon 1912:Portals 1855:Bibcode 1771:Bibcode 1683:Bibcode 1636:Bibcode 421:of the 406:of the 396:NeuquĂ©n 382:is from 362:as the 297:† 276:† 247:Genus: 106:↓ 4402:122466 4053:Ootaxa 3250:Tanius 2843:Bolong 1826:  1816:  1742:  1654:  569:et al. 525:, and 452:. The 442:saurus 438:grypho 308:et al. 306:Calvo 262:et al. 260:Calvo 4371:IRMNG 3095:Fylax 2625:Iyuku 2031:Clade 2022:Clade 2013:Clade 2004:Clade 1740:S2CID 1652:S2CID 554:clade 478:taxon 434:macro 348:) of 324:genus 232:Clade 216:Clade 200:Clade 184:Clade 171:Clade 4358:GBIF 2791:Proa 2463:Iani 2091:see 2064:see 1824:OCLC 1814:ISBN 816:Haya 563:The 364:type 337:age 310:2007 265:2007 45:Preęž’ 4345:EoL 1863:doi 1787:hdl 1779:doi 1732:doi 1699:hdl 1691:doi 1679:102 1644:doi 1632:108 419:age 352:in 326:of 4421:: 4399:: 4386:: 4373:: 4360:: 4347:: 4332:: 2033:: 2024:: 2015:: 2006:: 1861:. 1851:93 1849:. 1845:. 1822:. 1785:. 1777:. 1767:79 1765:. 1761:. 1738:. 1728:79 1726:. 1722:. 1697:. 1689:. 1677:. 1673:. 1650:. 1642:. 1630:. 1626:. 1606:^ 1592:^ 1584:65 1582:. 1578:. 1560:^ 556:, 519:, 480:. 390:, 234:: 218:: 202:: 186:: 173:: 95:Pg 39:Ma 4261:) 4101:? 3727:? 3665:? 3443:? 3154:? 3055:? 2930:? 2908:? 2709:? 2636:? 2628:? 2458:? 2447:? 2436:? 2428:? 2420:? 2395:? 2345:? 2309:? 2280:? 2231:? 2216:? 2205:? 2194:? 1967:e 1960:t 1953:v 1914:: 1871:. 1865:: 1857:: 1830:. 1795:. 1789:: 1781:: 1773:: 1746:. 1734:: 1707:. 1701:: 1693:: 1685:: 1658:. 1646:: 1638:: 344:( 251:† 238:† 222:† 206:† 190:† 100:N 90:K 85:J 80:T 75:P 70:C 65:D 60:S 55:O 50:ęž’ 34:~

Index

Coniacian
Ma
Preęž’
ęž’
O
S
D
C
P
T
J
K
Pg
N

Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Ornithischia
Neornithischia
Ornithopoda
Elasmaria
Macrogryphosaurus
2007
Binomial name
genus
elasmarian

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