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Pannoniasaurus

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528: 519:), as well as a variety of fish and crocodile teeth, were collected from the waste dump of the subterranean Ajka coal mine. The Ajka Coal Formation interdigitates with the CsehbĂĄnya Formation, the depositional environment of the latter was a floodplain, while the Ajka Coal Formation was formed in the accumulation basin of the same river system. Both of these facies were formed in the same paleogeographic area, which itself might have been part of a larger, but isolated landmass. 355: 695: 136: 451: 653: 1683: 1225: 118: 844: 495:, a fragmentary left quadrate. Though all remains (including the holotype) of Pannoniasaurus are isolated bones, the density of the specimens, the various size classes, the large number of similar elements from individual animals, and their unique characters, make it possible to link them together into a single taxon. 682:
a primitive morphology of the complete limbs in correlation with the primitive axial skeleton and pelvis is more probable, but far from certain. It is possible that the retention of a robust sacrum, pelvis and possibly non-paddle-like limbs were used to help to propel the body forward from the bottom
475:
is known from various material, including 2 isolated premaxillae, 3 maxillae, 2 postorbitofrontals, 2 quadrates, 3 dentaries, 3 splenials, 3 angulars, a coronoid, 2 surangulars, an articular, 91 isolated teeth, 20 cervical, 40 dorsal, 4 sacral, and 18 caudal vertebrae, 34 vertebral fragments, 3 ribs,
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was a medium-sized mosasauroid, estimated to grow up to a maximum of 6 m (20 ft) in length. It exhibited a combination of primitive characteristics, such as having no predental rostrum, the premaxilla-maxilla suture ends anterior to or level with the midline of the fourth maxillary tooth, a
643:
makes it the largest known predator in the waters of this paleoenvironment. Additionally, the crocodile-like flattened skull (as indicated by the premaxilla and maxilla) was a useful adaptation for water-level ambush hunting of terrestrial and shallow water prey.
678:, for example, has an anteriorly oriented, hydropelvic ilium in combination with primitive-looking proximal limb elements, and therefore a flattened, derived distal limb morphology is not entirely impossible for that taxon. For 346:, dubbed "unexpected" because it was discovered in freshwater sediments, unlike other mosasaurs, which were marine predators. It was a medium-sized mosasaur, reaching up to 6 metres (20 ft) in length. 604:
arises (two major lineages of a grade of paddle-bearing marine mosasaurs), is supported, and provides further support for the hypothesis of convergent aquatic adaptations in paddle-bearing mosasaurs.
615:, mosasauroids quickly radiated into a variety of aquatic environments, with some groups reinvading available niches in freshwater habitats, and becoming highly specialized within those ecosystems. 668:
may have looked. It had a primitive vertebral column, a posteriorly oriented ilium and an elongated humerus with a distal epiphysis, all most similar to aigialosaurs. These suggest that
910:"Geochemical study of vertebrate fossils from the Upper Cretaceous (Santonian) CsehbĂĄnya Formation (Hungary): evidence for a freshwater habitat of mosasaurs and pycnodont fish" 635:
was not a seasonal migrant from marine waters into fresh, but rather that ecologically it was a permanent member of a freshwater fauna. With the discovery and description of
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The preserved part of the maxilla bears 12 tooth sockets but the original maxillary tooth count might have been much higher. A number of isolated teeth were attributed to
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nearly straight frontoparietal suture, and a shallow quadrate alar concavity. It also had elongated stapedial pit that was at least three times longer than it was wide.
1761: 401:, sourced from a large number of individuals of differing age classes, are known from the alluvial flood-plain deposits that comprise the CsehbĂĄnya Formation. 1670: 508:, being conical and curved posterolingually, bear crowns with fine anastomosing longitudinal striae, and have a strong mesial but weaker labiodistal carina. 909: 1748: 593:, and the clades that include conventional marine mosasaur-grade mosasauroids such as halisaurs, tylosaurs and plioplatecarpines. The concept of a 1815: 965: 639:, mosasauroid evolution is now understood as also having involved important and unsuspected adaptations to freshwater ecosystems. The size of 435: 1660: 720: 135: 1825: 476:
2 humeral fragments, and 4 ilia. Since all remains are isolated bones, the basis for the referral of this material to
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Arambourg, 1952, a pachyostotic marine squamate from the latest Cretaceous phosphates of Morocco and Syria"
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had an overall aigialosaur-like postcranial morphology (including plesiopelvia and plesiopedia). However,
379: 321: 405: 325: 1787: 1224: 951: 921: 874: 805: 628: 266: 794:"The First Freshwater Mosasauroid (Upper Cretaceous, Hungary) and a New Clade of Basal Mosasauroids" 527: 1739: 700: 354: 482: 1686: 890: 130: 1792: 721:"Vertebrate remains from the Upper Cretaceous (Santonian) Ajka Coal Formation, western Hungary" 1774: 1766: 1368: 1311: 833: 789: 743: 652: 450: 1779: 480:
is based on similar methods used by other authors, such as Houssaye et al. in their study of
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Material was copied from this source, which is available under a Creative Commons License.
363: 318: 30: 925: 878: 809: 575:. Tethysaurinae was reconstructed at the base of a clade that includes the aigialosaurs 1509: 1181: 1127: 828: 793: 570: 1809: 1613: 1585: 1543: 1516: 1475: 1431: 1402: 1161: 1120: 1072: 894: 858: 563: 543: 245: 68: 1564: 1458: 1395: 1386: 1334: 1327: 1320: 1300: 1233: 1188: 1154: 1097: 1044: 1023: 1014: 601: 598: 594: 589: 551: 383: 311: 274: 739: 538:
Analysis by MakĂĄdi et al. found three equally most parsimonious trees that placed
933: 818: 1733: 1629: 1622: 1606: 1578: 1557: 1550: 1502: 1447: 1416: 1361: 1253: 1209: 1195: 1172: 1037: 674: 583: 577: 556: 378:) and referred specimens have been collected from the alluvial sediments of the 314: 43: 1724: 1482: 1468: 1269: 1202: 1106: 1086: 1058: 886: 690: 504: 203: 88: 53: 747: 1592: 1409: 1262: 1242: 1113: 1030: 612: 329: 147: 117: 93: 37: 857:
Houssaye A, Bardet N, Rage JC, Suberbiola XP, Bouya B, et al. (2011).
837: 1718: 1134: 1004: 998: 992: 974: 439: 371: 232: 216: 187: 167: 83: 78: 63: 58: 48: 664:
It is difficult to estimate how the unknown girdle and limb elements of
1753: 843: 387: 337: 333: 304: 177: 98: 73: 1065: 986: 719:Ɛsi, Attila; Bodor, Emese Réka; Makådi, Låszló; Rabi, Mårton (2016). 157: 1695: 683:
during prey-capture in shallow water, similar to extant crocodiles.
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was then named and described by Makådi, Caldwell & Ɛsi (2012).
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by Makådi, Caldwell & Ɛsi (2012), and then was classified as
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A single vertebra (MTM V.2000.21) was collected in 1999 from the
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from various exposures at the IharkĂșt open-pit bauxite mine,
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Kocsis L, Ɛsi A, Vennemann T, Trueman CN, Palmer MR (2009).
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at the waste dump of the coal mines next to the town of
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Geochemical and isotopic data are most compatible with
281: 1708: 1653: 1530: 1492: 1457: 1430: 1385: 1344: 1310: 1290: 1252: 1232: 1171: 1144: 1096: 1013: 783: 781: 779: 777: 788:MakĂĄdi, L. S.; Caldwell, M. W.; Ɛsi, A. (2012). 775: 773: 771: 769: 767: 765: 763: 761: 759: 757: 714: 712: 959: 390:since the discovery of the locality in 2000. 8: 1671:List of mosasaur-bearing stratigraphic units 611:indicates that, similar to some lineages of 397:Since then, more than one hundred bones of 1696: 1341: 1307: 1249: 1093: 966: 952: 944: 597:clade of aigialosaurs from within which a 116: 20: 827: 817: 125:Known remains and skeletal restorations 708: 631:, and evidence suggests strongly that 914:Palaeogeogr Palaeoclimatol Palaeoecol 7: 487:in 2011. The paratypes are known by 416:. It was described as belonging to 14: 1682: 1681: 1223: 842: 693: 627:having lived in a predominantly 546:. The three trees reconstructed 436:Hungarian Natural History Museum 291:MakĂĄdi, Caldwell & Ɛsi, 2012 134: 920:(3–4) (280 ed.): 532–542. 1: 1816:Fossil taxa described in 2012 1661:Timeline of mosasaur research 740:10.1016/j.cretres.2015.04.014 934:10.1016/j.palaeo.2009.07.009 861:Pachyvaranus crassispondylus 819:10.1371/journal.pone.0051781 434:are currently housed in the 265:MakĂĄdi, Caldwell & Ɛsi, 658:Pannoniasaurus inexpectatus 554:to the clade that includes 533:Pannoniasaurus inexpectatus 460:Pannoniasaurus inexpectatus 425:Pannoniasaurus inexpectatus 392:Pannoniasaurus inexpectatus 360:Pannoniasaurus inexpectatus 343:Pannoniasaurus inexpectatus 286:Pannoniasaurus inexpectatus 1842: 362:, restored with primitive 1679: 1531:Related groups and genera 1221: 981: 887:10.1017/S0016756810000580 280: 273: 131:Scientific classification 129: 124: 115: 23: 16:Extinct genus of lizards 1666:List of mosasaur genera 491:, a left quadrate, and 336:. It contains a single 661: 535: 462: 427:by Ɛsi et al. (2016). 367: 1788:Paleobiology Database 655: 531:Phylogenetic tree of 530: 511:A single vertebra of 453: 357: 629:freshwater ecosystem 542:in a new clade, the 502:that are similar to 358:Skeletal diagram of 350:Discovery and naming 1826:Mosasaurs of Europe 926:2009PPP...280..532K 879:2011GeoM..148..237H 810:2012PLoSO...751781M 728:Cretaceous Research 701:Paleontology portal 454:Holotype quadrate ( 406:Ajka Coal Formation 380:CsehbĂĄnya Formation 326:Ajka Coal Formation 322:CsehbĂĄnya Formation 662: 536: 463: 412:, 20 km from 368: 1803: 1802: 1775:Open Tree of Life 1702:Taxon identifiers 1693: 1692: 1526: 1525: 1426: 1425: 1369:Plesioplatecarpus 1312:Plioplatecarpinae 1286: 1285: 1219: 1218: 790:Butler, Richard J 660:compared to human 607:The discovery of 430:All specimens of 296: 295: 269: 1833: 1796: 1795: 1783: 1782: 1770: 1769: 1757: 1756: 1744: 1743: 1742: 1729: 1728: 1727: 1697: 1685: 1684: 1654:Related articles 1644:Vallecillosaurus 1346:Plioplatecarpini 1342: 1308: 1250: 1227: 1146:Prognathodontini 1094: 1080:Plesiotylosaurus 968: 961: 954: 945: 938: 937: 905: 899: 898: 854: 848: 846: 841: 831: 821: 785: 752: 751: 725: 716: 703: 698: 697: 696: 283: 264: 257: 244: 231: 215: 202: 139: 138: 120: 110: 40: 29:Temporal range: 21: 1841: 1840: 1836: 1835: 1834: 1832: 1831: 1830: 1821:Russellosaurins 1806: 1805: 1804: 1799: 1791: 1786: 1778: 1773: 1765: 1760: 1752: 1747: 1738: 1737: 1732: 1723: 1722: 1717: 1704: 1694: 1689: 1675: 1649: 1637:Portunatasaurus 1573:Dolichosauridae 1538:Aigialosauridae 1522: 1494:Yaguarasaurinae 1488: 1453: 1422: 1381: 1376:Plioplatecarpus 1355:Latoplatecarpus 1340: 1306: 1292:Russellosaurina 1282: 1277:Phosphorosaurus 1248: 1228: 1215: 1167: 1140: 1092: 1009: 977: 972: 942: 941: 907: 906: 902: 856: 855: 851: 787: 786: 755: 723: 718: 717: 710: 699: 694: 692: 689: 670:P. inexpectatus 666:Pannonisasaurus 650: 621: 525: 485:crassispondylus 448: 352: 319:Late Cretaceous 317:known from the 292: 289: 263: 255: 242: 233:Russellosaurina 229: 213: 200: 133: 111: 109: 108: 107: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 36:85.3–83.5  35: 34: 31:Late Cretaceous 27: 17: 12: 11: 5: 1839: 1837: 1829: 1828: 1823: 1818: 1808: 1807: 1801: 1800: 1798: 1797: 1784: 1771: 1758: 1745: 1740:Pannoniasaurus 1730: 1714: 1712: 1710:Pannoniasaurus 1706: 1705: 1700: 1691: 1690: 1680: 1677: 1676: 1674: 1673: 1668: 1663: 1657: 1655: 1651: 1650: 1648: 1647: 1640: 1633: 1626: 1619: 1618: 1617: 1610: 1603: 1596: 1589: 1582: 1570: 1569: 1568: 1561: 1554: 1547: 1534: 1532: 1528: 1527: 1524: 1523: 1521: 1520: 1513: 1510:Russellosaurus 1506: 1498: 1496: 1490: 1489: 1487: 1486: 1479: 1472: 1463: 1461: 1455: 1454: 1452: 1451: 1444: 1441:Pannoniasaurus 1436: 1434: 1428: 1427: 1424: 1423: 1421: 1420: 1413: 1406: 1399: 1391: 1389: 1383: 1382: 1380: 1379: 1372: 1365: 1358: 1350: 1348: 1339: 1338: 1331: 1324: 1316: 1314: 1305: 1304: 1296: 1294: 1288: 1287: 1284: 1283: 1281: 1280: 1273: 1266: 1258: 1256: 1247: 1246: 1238: 1236: 1230: 1229: 1222: 1220: 1217: 1216: 1214: 1213: 1206: 1199: 1192: 1185: 1182:Bentiabasaurus 1177: 1175: 1169: 1168: 1166: 1165: 1158: 1150: 1148: 1142: 1141: 1139: 1138: 1131: 1128:Igdamanosaurus 1124: 1117: 1110: 1102: 1100: 1091: 1090: 1083: 1076: 1069: 1062: 1055: 1048: 1041: 1034: 1027: 1019: 1017: 1011: 1010: 1008: 1007: 1001: 995: 989: 982: 979: 978: 973: 971: 970: 963: 956: 948: 940: 939: 900: 873:(2): 237–249. 849: 804:(12): e51781. 753: 707: 706: 705: 704: 688: 685: 680:Pannoniasaurus 649: 646: 641:Pannoniasaurus 637:Pannoniasaurus 633:Pannoniasaurus 625:Pannoniasaurus 620: 617: 609:Pannoniasaurus 571:Russellosaurus 548:Pannoniasaurus 540:Pannoniasaurus 524: 523:Classification 521: 513:Pannoniasaurus 500:Pannoniasaurus 478:Pannoniasaurus 473:Pannoniasaurus 466:Pannoniasaurus 447: 444: 432:Pannoniasaurus 421:Pannoniasaurus 399:Pannoniasaurus 351: 348: 300:Pannoniasaurus 294: 293: 290: 278: 277: 271: 270: 260:Pannoniasaurus 253: 249: 248: 240: 236: 235: 227: 220: 219: 211: 207: 206: 198: 191: 190: 185: 181: 180: 175: 171: 170: 165: 161: 160: 155: 151: 150: 145: 141: 140: 127: 126: 122: 121: 113: 112: 104: 103: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 42: 41: 28: 25:Pannoniasaurus 15: 13: 10: 9: 6: 4: 3: 2: 1838: 1827: 1824: 1822: 1819: 1817: 1814: 1813: 1811: 1794: 1789: 1785: 1781: 1776: 1772: 1768: 1763: 1759: 1755: 1750: 1746: 1741: 1735: 1731: 1726: 1720: 1716: 1715: 1713: 1711: 1707: 1703: 1698: 1688: 1678: 1672: 1669: 1667: 1664: 1662: 1659: 1658: 1656: 1652: 1646: 1645: 1641: 1639: 1638: 1634: 1632: 1631: 1627: 1625: 1624: 1620: 1616: 1615: 1614:Tetrapodophis 1611: 1609: 1608: 1604: 1602: 1601: 1597: 1595: 1594: 1590: 1588: 1587: 1586:Dolichosaurus 1583: 1581: 1580: 1576: 1575: 1574: 1571: 1567: 1566: 1562: 1560: 1559: 1555: 1553: 1552: 1548: 1546: 1545: 1544:Aigialosaurus 1541: 1540: 1539: 1536: 1535: 1533: 1529: 1519: 1518: 1517:Yaguarasaurus 1514: 1512: 1511: 1507: 1505: 1504: 1500: 1499: 1497: 1495: 1491: 1485: 1484: 1480: 1478: 1477: 1476:Taniwhasaurus 1473: 1471: 1470: 1465: 1464: 1462: 1460: 1456: 1450: 1449: 1445: 1443: 1442: 1438: 1437: 1435: 1433: 1432:Tethysaurinae 1429: 1419: 1418: 1414: 1412: 1411: 1407: 1405: 1404: 1403:Goronyosaurus 1400: 1398: 1397: 1393: 1392: 1390: 1388: 1384: 1378: 1377: 1373: 1371: 1370: 1366: 1364: 1363: 1359: 1357: 1356: 1352: 1351: 1349: 1347: 1343: 1337: 1336: 1332: 1330: 1329: 1325: 1323: 1322: 1318: 1317: 1315: 1313: 1309: 1303: 1302: 1298: 1297: 1295: 1293: 1289: 1279: 1278: 1274: 1272: 1271: 1267: 1265: 1264: 1260: 1259: 1257: 1255: 1251: 1245: 1244: 1240: 1239: 1237: 1235: 1231: 1226: 1212: 1211: 1207: 1205: 1204: 1200: 1198: 1197: 1193: 1191: 1190: 1186: 1184: 1183: 1179: 1178: 1176: 1174: 1170: 1164: 1163: 1162:Thalassotitan 1159: 1157: 1156: 1152: 1151: 1149: 1147: 1143: 1137: 1136: 1132: 1130: 1129: 1125: 1123: 1122: 1121:Harranasaurus 1118: 1116: 1115: 1111: 1109: 1108: 1104: 1103: 1101: 1099: 1095: 1089: 1088: 1084: 1082: 1081: 1077: 1075: 1074: 1073:Megapterygius 1070: 1068: 1067: 1063: 1061: 1060: 1056: 1054: 1053: 1049: 1047: 1046: 1042: 1040: 1039: 1035: 1033: 1032: 1028: 1026: 1025: 1021: 1020: 1018: 1016: 1012: 1006: 1002: 1000: 996: 994: 990: 988: 984: 983: 980: 976: 969: 964: 962: 957: 955: 950: 949: 946: 935: 931: 927: 923: 919: 915: 911: 904: 901: 896: 892: 888: 884: 880: 876: 872: 868: 864: 862: 859:"A review of 853: 850: 845: 839: 835: 830: 825: 820: 815: 811: 807: 803: 799: 795: 791: 784: 782: 780: 778: 776: 774: 772: 770: 768: 766: 764: 762: 760: 758: 754: 749: 745: 741: 737: 733: 729: 722: 715: 713: 709: 702: 691: 686: 684: 681: 677: 676: 671: 667: 659: 654: 647: 645: 642: 638: 634: 630: 626: 618: 616: 614: 610: 605: 603: 600: 596: 592: 591: 586: 585: 580: 579: 574: 572: 567: 565: 564:Yaguarasaurus 560: 558: 553: 549: 545: 544:Tethysaurinae 541: 534: 529: 522: 520: 518: 517:MTM V.2000.21 514: 509: 507: 506: 501: 496: 494: 493:MTM 2007.31.1 490: 486: 484: 479: 474: 470: 467: 461: 457: 456:MTM 2011.43.1 452: 445: 443: 441: 437: 433: 428: 426: 422: 419: 415: 411: 407: 402: 400: 395: 393: 389: 385: 381: 377: 376:MTM 2011.43.1 373: 365: 361: 356: 349: 347: 345: 344: 339: 335: 331: 327: 323: 320: 316: 313: 309: 306: 302: 301: 288: 287: 279: 276: 272: 268: 262: 261: 254: 251: 250: 247: 246:Tethysaurinae 241: 238: 237: 234: 228: 225: 222: 221: 218: 212: 209: 208: 205: 199: 196: 193: 192: 189: 186: 183: 182: 179: 176: 173: 172: 169: 166: 163: 162: 159: 156: 153: 152: 149: 146: 143: 142: 137: 132: 128: 123: 119: 114: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 39: 32: 26: 22: 19: 1709: 1642: 1635: 1628: 1621: 1612: 1605: 1598: 1591: 1584: 1577: 1565:Opetiosaurus 1563: 1556: 1549: 1542: 1515: 1508: 1501: 1481: 1474: 1467: 1459:Tylosaurinae 1446: 1440: 1439: 1415: 1408: 1401: 1396:Gavialimimus 1394: 1387:Selmasaurini 1374: 1367: 1360: 1353: 1335:Sarabosaurus 1333: 1328:Ectenosaurus 1326: 1321:Angolasaurus 1319: 1301:Haasiasaurus 1299: 1275: 1268: 1261: 1241: 1234:Halisaurinae 1208: 1201: 1194: 1189:Eremiasaurus 1187: 1180: 1160: 1155:Prognathodon 1153: 1133: 1126: 1119: 1112: 1105: 1098:Globidensini 1085: 1078: 1071: 1064: 1057: 1050: 1045:Gnathomortis 1043: 1036: 1029: 1024:Amphekepubis 1022: 1015:Mosasaurinae 917: 913: 903: 870: 866: 860: 852: 801: 797: 731: 727: 679: 673: 669: 665: 663: 657: 640: 636: 632: 624: 622: 619:Paleobiology 608: 606: 602:Mosasauridae 599:polyphyletic 595:monophyletic 590:Haasiasaurus 588: 582: 576: 569: 562: 555: 552:sister taxon 547: 539: 537: 532: 516: 512: 510: 503: 499: 497: 492: 489:MTM V.01.115 488: 483:Pachyvaranus 481: 477: 472: 471: 465: 464: 459: 455: 431: 429: 424: 420: 403: 398: 396: 391: 384:Bakony Hills 375: 369: 366:-style limbs 359: 342: 341: 312:tethysaurine 299: 298: 297: 285: 284: 275:Type species 259: 258: 223: 217:Mosasauridae 194: 24: 18: 1734:Wikispecies 1630:Judeasaurus 1623:Adriosaurus 1607:Pontosaurus 1579:Coniasaurus 1558:Komensaurus 1551:Carsosaurus 1503:Romeosaurus 1448:Tethysaurus 1417:Selmasaurus 1362:Platecarpus 1254:Halisaurini 1210:Plotosaurus 1196:Moanasaurus 1173:Mosasaurini 1052:Jormungandr 1038:Dallasaurus 734:: 228–238. 675:Dallasaurus 584:Komensaurus 578:Carsosaurus 566:columbianus 557:Tethysaurus 446:Description 442:, Hungary. 364:aigialosaur 315:mosasauroid 239:Subfamily: 1810:Categories 1600:Primitivus 1483:Tylosaurus 1469:Kaikaifilu 1270:Halisaurus 1203:Mosasaurus 1107:Carinodens 1087:Stelladens 1059:Kourisodon 687:References 505:Halisaurus 386:, Western 332:stage) of 204:Mosasauria 1593:Kaganaias 1410:Khinjaria 1263:Eonatator 1243:Pluridens 1114:Globidens 1031:Clidastes 985:Kingdom: 975:Mosasaurs 895:128706999 748:0195-6671 613:cetaceans 330:Santonian 154:Kingdom: 148:Eukaryota 1725:Q1397354 1719:Wikidata 1687:Category 1135:Xenodens 1005:Squamata 999:Reptilia 993:Chordata 991:Phylum: 987:Animalia 867:Geol Mag 838:23284766 798:PLOS ONE 656:Size of 440:Budapest 372:holotype 210:Family: 188:Squamata 178:Reptilia 168:Chordata 164:Phylum: 158:Animalia 144:Domain: 1780:6158065 1767:1483230 1754:8666962 1003:Order: 997:Class: 922:Bibcode 875:Bibcode 829:3526648 806:Bibcode 792:(ed.). 559:nopcsai 550:as the 414:IharkĂșt 388:Hungary 338:species 334:Hungary 305:extinct 252:Genus: 184:Order: 174:Class: 105:↓ 1793:253722 1066:Liodon 893:  836:  826:  746:  587:, and 573:coheni 568:, and 303:is an 1762:IRMNG 891:S2CID 724:(PDF) 648:Limbs 458:) of 308:genus 224:Clade 195:Clade 1749:GBIF 834:PMID 744:ISSN 410:Ajka 370:The 324:and 267:2012 44:PreꞒ 930:doi 918:280 883:doi 871:148 824:PMC 814:doi 736:doi 418:cf. 310:of 1812:: 1790:: 1777:: 1764:: 1751:: 1736:: 1721:: 928:. 916:. 912:. 889:. 881:. 869:. 865:. 832:. 822:. 812:. 800:. 796:. 756:^ 742:. 732:57 730:. 726:. 711:^ 581:, 561:, 438:, 340:, 226:: 197:: 94:Pg 38:Ma 33:, 1466:? 967:e 960:t 953:v 936:. 932:: 924:: 897:. 885:: 877:: 840:. 816:: 808:: 802:7 750:. 738:: 515:( 374:( 328:( 282:† 256:† 243:† 230:† 214:† 201:† 99:N 89:K 84:J 79:T 74:P 69:C 64:D 59:S 54:O 49:Ꞓ

Index

Late Cretaceous
Ma
PreꞒ
Ꞓ
O
S
D
C
P
T
J
K
Pg
N

Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Reptilia
Squamata
Mosasauria
Mosasauridae
Russellosaurina
Tethysaurinae
Pannoniasaurus
2012
Type species
extinct

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