528:
519:), as well as a variety of fish and crocodile teeth, were collected from the waste dump of the subterranean Ajka coal mine. The Ajka Coal Formation interdigitates with the CsehbĂĄnya Formation, the depositional environment of the latter was a floodplain, while the Ajka Coal Formation was formed in the accumulation basin of the same river system. Both of these facies were formed in the same paleogeographic area, which itself might have been part of a larger, but isolated landmass.
355:
695:
136:
451:
653:
1683:
1225:
118:
844:
495:, a fragmentary left quadrate. Though all remains (including the holotype) of Pannoniasaurus are isolated bones, the density of the specimens, the various size classes, the large number of similar elements from individual animals, and their unique characters, make it possible to link them together into a single taxon.
682:
a primitive morphology of the complete limbs in correlation with the primitive axial skeleton and pelvis is more probable, but far from certain. It is possible that the retention of a robust sacrum, pelvis and possibly non-paddle-like limbs were used to help to propel the body forward from the bottom
475:
is known from various material, including 2 isolated premaxillae, 3 maxillae, 2 postorbitofrontals, 2 quadrates, 3 dentaries, 3 splenials, 3 angulars, a coronoid, 2 surangulars, an articular, 91 isolated teeth, 20 cervical, 40 dorsal, 4 sacral, and 18 caudal vertebrae, 34 vertebral fragments, 3 ribs,
468:
was a medium-sized mosasauroid, estimated to grow up to a maximum of 6 m (20 ft) in length. It exhibited a combination of primitive characteristics, such as having no predental rostrum, the premaxilla-maxilla suture ends anterior to or level with the midline of the fourth maxillary tooth, a
643:
makes it the largest known predator in the waters of this paleoenvironment. Additionally, the crocodile-like flattened skull (as indicated by the premaxilla and maxilla) was a useful adaptation for water-level ambush hunting of terrestrial and shallow water prey.
678:, for example, has an anteriorly oriented, hydropelvic ilium in combination with primitive-looking proximal limb elements, and therefore a flattened, derived distal limb morphology is not entirely impossible for that taxon. For
346:, dubbed "unexpected" because it was discovered in freshwater sediments, unlike other mosasaurs, which were marine predators. It was a medium-sized mosasaur, reaching up to 6 metres (20 ft) in length.
604:
arises (two major lineages of a grade of paddle-bearing marine mosasaurs), is supported, and provides further support for the hypothesis of convergent aquatic adaptations in paddle-bearing mosasaurs.
615:, mosasauroids quickly radiated into a variety of aquatic environments, with some groups reinvading available niches in freshwater habitats, and becoming highly specialized within those ecosystems.
668:
may have looked. It had a primitive vertebral column, a posteriorly oriented ilium and an elongated humerus with a distal epiphysis, all most similar to aigialosaurs. These suggest that
910:"Geochemical study of vertebrate fossils from the Upper Cretaceous (Santonian) CsehbĂĄnya Formation (Hungary): evidence for a freshwater habitat of mosasaurs and pycnodont fish"
635:
was not a seasonal migrant from marine waters into fresh, but rather that ecologically it was a permanent member of a freshwater fauna. With the discovery and description of
498:
The preserved part of the maxilla bears 12 tooth sockets but the original maxillary tooth count might have been much higher. A number of isolated teeth were attributed to
469:
nearly straight frontoparietal suture, and a shallow quadrate alar concavity. It also had elongated stapedial pit that was at least three times longer than it was wide.
1761:
401:, sourced from a large number of individuals of differing age classes, are known from the alluvial flood-plain deposits that comprise the CsehbĂĄnya Formation.
1670:
508:, being conical and curved posterolingually, bear crowns with fine anastomosing longitudinal striae, and have a strong mesial but weaker labiodistal carina.
909:
1748:
593:, and the clades that include conventional marine mosasaur-grade mosasauroids such as halisaurs, tylosaurs and plioplatecarpines. The concept of a
1815:
965:
639:, mosasauroid evolution is now understood as also having involved important and unsuspected adaptations to freshwater ecosystems. The size of
435:
1660:
720:
135:
1825:
476:
2 humeral fragments, and 4 ilia. Since all remains are isolated bones, the basis for the referral of this material to
958:
1051:
1820:
1599:
1665:
863:
Arambourg, 1952, a pachyostotic marine squamate from the latest
Cretaceous phosphates of Morocco and Syria"
1701:
672:
had an overall aigialosaur-like postcranial morphology (including plesiopelvia and plesiopedia). However,
379:
321:
405:
325:
1787:
1224:
951:
921:
874:
805:
628:
266:
794:"The First Freshwater Mosasauroid (Upper Cretaceous, Hungary) and a New Clade of Basal Mosasauroids"
527:
1739:
700:
354:
482:
1686:
890:
130:
1792:
721:"Vertebrate remains from the Upper Cretaceous (Santonian) Ajka Coal Formation, western Hungary"
1774:
1766:
1368:
1311:
833:
789:
743:
652:
450:
1779:
480:
is based on similar methods used by other authors, such as
Houssaye et al. in their study of
413:
1643:
1345:
1145:
1079:
929:
882:
823:
813:
735:
1636:
1572:
1537:
1493:
1375:
1354:
1291:
1276:
847:
Material was copied from this source, which is available under a
Creative Commons License.
363:
318:
30:
925:
878:
809:
575:. Tethysaurinae was reconstructed at the base of a clade that includes the aigialosaurs
1509:
1181:
1127:
828:
793:
570:
1809:
1613:
1585:
1543:
1516:
1475:
1431:
1402:
1161:
1120:
1072:
894:
858:
563:
543:
245:
68:
1564:
1458:
1395:
1386:
1334:
1327:
1320:
1300:
1233:
1188:
1154:
1097:
1044:
1023:
1014:
601:
598:
594:
589:
551:
383:
311:
274:
739:
538:
Analysis by MakĂĄdi et al. found three equally most parsimonious trees that placed
933:
818:
1733:
1629:
1622:
1606:
1578:
1557:
1550:
1502:
1447:
1416:
1361:
1253:
1209:
1195:
1172:
1037:
674:
583:
577:
556:
378:) and referred specimens have been collected from the alluvial sediments of the
314:
43:
1724:
1482:
1468:
1269:
1202:
1106:
1086:
1058:
886:
690:
504:
203:
88:
53:
747:
1592:
1409:
1262:
1242:
1113:
1030:
612:
329:
147:
117:
93:
37:
857:
Houssaye A, Bardet N, Rage JC, Suberbiola XP, Bouya B, et al. (2011).
837:
1718:
1134:
1004:
998:
992:
974:
439:
371:
232:
216:
187:
167:
83:
78:
63:
58:
48:
664:
It is difficult to estimate how the unknown girdle and limb elements of
1753:
843:
387:
337:
333:
304:
177:
98:
73:
1065:
986:
719:Ćsi, Attila; Bodor, Emese RĂ©ka; MakĂĄdi, LĂĄszlĂł; Rabi, MĂĄrton (2016).
157:
1695:
683:
during prey-capture in shallow water, similar to extant crocodiles.
394:
was then named and described by MakĂĄdi, Caldwell & Ćsi (2012).
651:
526:
449:
353:
307:
423:
by MakĂĄdi, Caldwell & Ćsi (2012), and then was classified as
404:
A single vertebra (MTM V.2000.21) was collected in 1999 from the
409:
1699:
947:
417:
382:
from various exposures at the IharkĂșt open-pit bauxite mine,
943:
908:
Kocsis L, Ćsi A, Vennemann T, Trueman CN, Palmer MR (2009).
408:
at the waste dump of the coal mines next to the town of
623:
Geochemical and isotopic data are most compatible with
281:
1708:
1653:
1530:
1492:
1457:
1430:
1385:
1344:
1310:
1290:
1252:
1232:
1171:
1144:
1096:
1013:
783:
781:
779:
777:
788:MakĂĄdi, L. S.; Caldwell, M. W.; Ćsi, A. (2012).
775:
773:
771:
769:
767:
765:
763:
761:
759:
757:
714:
712:
959:
390:since the discovery of the locality in 2000.
8:
1671:List of mosasaur-bearing stratigraphic units
611:indicates that, similar to some lineages of
397:Since then, more than one hundred bones of
1696:
1341:
1307:
1249:
1093:
966:
952:
944:
597:clade of aigialosaurs from within which a
116:
20:
827:
817:
125:Known remains and skeletal restorations
708:
631:, and evidence suggests strongly that
914:Palaeogeogr Palaeoclimatol Palaeoecol
7:
487:in 2011. The paratypes are known by
416:. It was described as belonging to
14:
1682:
1681:
1223:
842:
693:
627:having lived in a predominantly
546:. The three trees reconstructed
436:Hungarian Natural History Museum
291:MakĂĄdi, Caldwell & Ćsi, 2012
134:
920:(3â4) (280 ed.): 532â542.
1:
1816:Fossil taxa described in 2012
1661:Timeline of mosasaur research
740:10.1016/j.cretres.2015.04.014
934:10.1016/j.palaeo.2009.07.009
861:Pachyvaranus crassispondylus
819:10.1371/journal.pone.0051781
434:are currently housed in the
265:MakĂĄdi, Caldwell & Ćsi,
658:Pannoniasaurus inexpectatus
554:to the clade that includes
533:Pannoniasaurus inexpectatus
460:Pannoniasaurus inexpectatus
425:Pannoniasaurus inexpectatus
392:Pannoniasaurus inexpectatus
360:Pannoniasaurus inexpectatus
343:Pannoniasaurus inexpectatus
286:Pannoniasaurus inexpectatus
1842:
362:, restored with primitive
1679:
1531:Related groups and genera
1221:
981:
887:10.1017/S0016756810000580
280:
273:
131:Scientific classification
129:
124:
115:
23:
16:Extinct genus of lizards
1666:List of mosasaur genera
491:, a left quadrate, and
336:. It contains a single
661:
535:
462:
427:by Ćsi et al. (2016).
367:
1788:Paleobiology Database
655:
531:Phylogenetic tree of
530:
511:A single vertebra of
453:
357:
629:freshwater ecosystem
542:in a new clade, the
502:that are similar to
358:Skeletal diagram of
350:Discovery and naming
1826:Mosasaurs of Europe
926:2009PPP...280..532K
879:2011GeoM..148..237H
810:2012PLoSO...751781M
728:Cretaceous Research
701:Paleontology portal
454:Holotype quadrate (
406:Ajka Coal Formation
380:CsehbĂĄnya Formation
326:Ajka Coal Formation
322:CsehbĂĄnya Formation
662:
536:
463:
412:, 20 km from
368:
1803:
1802:
1775:Open Tree of Life
1702:Taxon identifiers
1693:
1692:
1526:
1525:
1426:
1425:
1369:Plesioplatecarpus
1312:Plioplatecarpinae
1286:
1285:
1219:
1218:
790:Butler, Richard J
660:compared to human
607:The discovery of
430:All specimens of
296:
295:
269:
1833:
1796:
1795:
1783:
1782:
1770:
1769:
1757:
1756:
1744:
1743:
1742:
1729:
1728:
1727:
1697:
1685:
1684:
1654:Related articles
1644:Vallecillosaurus
1346:Plioplatecarpini
1342:
1308:
1250:
1227:
1146:Prognathodontini
1094:
1080:Plesiotylosaurus
968:
961:
954:
945:
938:
937:
905:
899:
898:
854:
848:
846:
841:
831:
821:
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698:
697:
696:
283:
264:
257:
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202:
139:
138:
120:
110:
40:
29:Temporal range:
21:
1841:
1840:
1836:
1835:
1834:
1832:
1831:
1830:
1821:Russellosaurins
1806:
1805:
1804:
1799:
1791:
1786:
1778:
1773:
1765:
1760:
1752:
1747:
1738:
1737:
1732:
1723:
1722:
1717:
1704:
1694:
1689:
1675:
1649:
1637:Portunatasaurus
1573:Dolichosauridae
1538:Aigialosauridae
1522:
1494:Yaguarasaurinae
1488:
1453:
1422:
1381:
1376:Plioplatecarpus
1355:Latoplatecarpus
1340:
1306:
1292:Russellosaurina
1282:
1277:Phosphorosaurus
1248:
1228:
1215:
1167:
1140:
1092:
1009:
977:
972:
942:
941:
907:
906:
902:
856:
855:
851:
787:
786:
755:
723:
718:
717:
710:
699:
694:
692:
689:
670:P. inexpectatus
666:Pannonisasaurus
650:
621:
525:
485:crassispondylus
448:
352:
319:Late Cretaceous
317:known from the
292:
289:
263:
255:
242:
233:Russellosaurina
229:
213:
200:
133:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
36:85.3â83.5
35:
34:
31:Late Cretaceous
27:
17:
12:
11:
5:
1839:
1837:
1829:
1828:
1823:
1818:
1808:
1807:
1801:
1800:
1798:
1797:
1784:
1771:
1758:
1745:
1740:Pannoniasaurus
1730:
1714:
1712:
1710:Pannoniasaurus
1706:
1705:
1700:
1691:
1690:
1680:
1677:
1676:
1674:
1673:
1668:
1663:
1657:
1655:
1651:
1650:
1648:
1647:
1640:
1633:
1626:
1619:
1618:
1617:
1610:
1603:
1596:
1589:
1582:
1570:
1569:
1568:
1561:
1554:
1547:
1534:
1532:
1528:
1527:
1524:
1523:
1521:
1520:
1513:
1510:Russellosaurus
1506:
1498:
1496:
1490:
1489:
1487:
1486:
1479:
1472:
1463:
1461:
1455:
1454:
1452:
1451:
1444:
1441:Pannoniasaurus
1436:
1434:
1428:
1427:
1424:
1423:
1421:
1420:
1413:
1406:
1399:
1391:
1389:
1383:
1382:
1380:
1379:
1372:
1365:
1358:
1350:
1348:
1339:
1338:
1331:
1324:
1316:
1314:
1305:
1304:
1296:
1294:
1288:
1287:
1284:
1283:
1281:
1280:
1273:
1266:
1258:
1256:
1247:
1246:
1238:
1236:
1230:
1229:
1222:
1220:
1217:
1216:
1214:
1213:
1206:
1199:
1192:
1185:
1182:Bentiabasaurus
1177:
1175:
1169:
1168:
1166:
1165:
1158:
1150:
1148:
1142:
1141:
1139:
1138:
1131:
1128:Igdamanosaurus
1124:
1117:
1110:
1102:
1100:
1091:
1090:
1083:
1076:
1069:
1062:
1055:
1048:
1041:
1034:
1027:
1019:
1017:
1011:
1010:
1008:
1007:
1001:
995:
989:
982:
979:
978:
973:
971:
970:
963:
956:
948:
940:
939:
900:
873:(2): 237â249.
849:
804:(12): e51781.
753:
707:
706:
705:
704:
688:
685:
680:Pannoniasaurus
649:
646:
641:Pannoniasaurus
637:Pannoniasaurus
633:Pannoniasaurus
625:Pannoniasaurus
620:
617:
609:Pannoniasaurus
571:Russellosaurus
548:Pannoniasaurus
540:Pannoniasaurus
524:
523:Classification
521:
513:Pannoniasaurus
500:Pannoniasaurus
478:Pannoniasaurus
473:Pannoniasaurus
466:Pannoniasaurus
447:
444:
432:Pannoniasaurus
421:Pannoniasaurus
399:Pannoniasaurus
351:
348:
300:Pannoniasaurus
294:
293:
290:
278:
277:
271:
270:
260:Pannoniasaurus
253:
249:
248:
240:
236:
235:
227:
220:
219:
211:
207:
206:
198:
191:
190:
185:
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160:
155:
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150:
145:
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140:
127:
126:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
25:Pannoniasaurus
15:
13:
10:
9:
6:
4:
3:
2:
1838:
1827:
1824:
1822:
1819:
1817:
1814:
1813:
1811:
1794:
1789:
1785:
1781:
1776:
1772:
1768:
1763:
1759:
1755:
1750:
1746:
1741:
1735:
1731:
1726:
1720:
1716:
1715:
1713:
1711:
1707:
1703:
1698:
1688:
1678:
1672:
1669:
1667:
1664:
1662:
1659:
1658:
1656:
1652:
1646:
1645:
1641:
1639:
1638:
1634:
1632:
1631:
1627:
1625:
1624:
1620:
1616:
1615:
1614:Tetrapodophis
1611:
1609:
1608:
1604:
1602:
1601:
1597:
1595:
1594:
1590:
1588:
1587:
1586:Dolichosaurus
1583:
1581:
1580:
1576:
1575:
1574:
1571:
1567:
1566:
1562:
1560:
1559:
1555:
1553:
1552:
1548:
1546:
1545:
1544:Aigialosaurus
1541:
1540:
1539:
1536:
1535:
1533:
1529:
1519:
1518:
1517:Yaguarasaurus
1514:
1512:
1511:
1507:
1505:
1504:
1500:
1499:
1497:
1495:
1491:
1485:
1484:
1480:
1478:
1477:
1476:Taniwhasaurus
1473:
1471:
1470:
1465:
1464:
1462:
1460:
1456:
1450:
1449:
1445:
1443:
1442:
1438:
1437:
1435:
1433:
1432:Tethysaurinae
1429:
1419:
1418:
1414:
1412:
1411:
1407:
1405:
1404:
1403:Goronyosaurus
1400:
1398:
1397:
1393:
1392:
1390:
1388:
1384:
1378:
1377:
1373:
1371:
1370:
1366:
1364:
1363:
1359:
1357:
1356:
1352:
1351:
1349:
1347:
1343:
1337:
1336:
1332:
1330:
1329:
1325:
1323:
1322:
1318:
1317:
1315:
1313:
1309:
1303:
1302:
1298:
1297:
1295:
1293:
1289:
1279:
1278:
1274:
1272:
1271:
1267:
1265:
1264:
1260:
1259:
1257:
1255:
1251:
1245:
1244:
1240:
1239:
1237:
1235:
1231:
1226:
1212:
1211:
1207:
1205:
1204:
1200:
1198:
1197:
1193:
1191:
1190:
1186:
1184:
1183:
1179:
1178:
1176:
1174:
1170:
1164:
1163:
1162:Thalassotitan
1159:
1157:
1156:
1152:
1151:
1149:
1147:
1143:
1137:
1136:
1132:
1130:
1129:
1125:
1123:
1122:
1121:Harranasaurus
1118:
1116:
1115:
1111:
1109:
1108:
1104:
1103:
1101:
1099:
1095:
1089:
1088:
1084:
1082:
1081:
1077:
1075:
1074:
1073:Megapterygius
1070:
1068:
1067:
1063:
1061:
1060:
1056:
1054:
1053:
1049:
1047:
1046:
1042:
1040:
1039:
1035:
1033:
1032:
1028:
1026:
1025:
1021:
1020:
1018:
1016:
1012:
1006:
1002:
1000:
996:
994:
990:
988:
984:
983:
980:
976:
969:
964:
962:
957:
955:
950:
949:
946:
935:
931:
927:
923:
919:
915:
911:
904:
901:
896:
892:
888:
884:
880:
876:
872:
868:
864:
862:
859:"A review of
853:
850:
845:
839:
835:
830:
825:
820:
815:
811:
807:
803:
799:
795:
791:
784:
782:
780:
778:
776:
774:
772:
770:
768:
766:
764:
762:
760:
758:
754:
749:
745:
741:
737:
733:
729:
722:
715:
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709:
702:
691:
686:
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677:
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667:
659:
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630:
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614:
610:
605:
603:
600:
596:
592:
591:
586:
585:
580:
579:
574:
572:
567:
565:
564:Yaguarasaurus
560:
558:
553:
549:
545:
544:Tethysaurinae
541:
534:
529:
522:
520:
518:
517:MTM V.2000.21
514:
509:
507:
506:
501:
496:
494:
493:MTM 2007.31.1
490:
486:
484:
479:
474:
470:
467:
461:
457:
456:MTM 2011.43.1
452:
445:
443:
441:
437:
433:
428:
426:
422:
419:
415:
411:
407:
402:
400:
395:
393:
389:
385:
381:
377:
376:MTM 2011.43.1
373:
365:
361:
356:
349:
347:
345:
344:
339:
335:
331:
327:
323:
320:
316:
313:
309:
306:
302:
301:
288:
287:
279:
276:
272:
268:
262:
261:
254:
251:
250:
247:
246:Tethysaurinae
241:
238:
237:
234:
228:
225:
222:
221:
218:
212:
209:
208:
205:
199:
196:
193:
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179:
176:
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159:
156:
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137:
132:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
1709:
1642:
1635:
1628:
1621:
1612:
1605:
1598:
1591:
1584:
1577:
1565:Opetiosaurus
1563:
1556:
1549:
1542:
1515:
1508:
1501:
1481:
1474:
1467:
1459:Tylosaurinae
1446:
1440:
1439:
1415:
1408:
1401:
1396:Gavialimimus
1394:
1387:Selmasaurini
1374:
1367:
1360:
1353:
1335:Sarabosaurus
1333:
1328:Ectenosaurus
1326:
1321:Angolasaurus
1319:
1301:Haasiasaurus
1299:
1275:
1268:
1261:
1241:
1234:Halisaurinae
1208:
1201:
1194:
1189:Eremiasaurus
1187:
1180:
1160:
1155:Prognathodon
1153:
1133:
1126:
1119:
1112:
1105:
1098:Globidensini
1085:
1078:
1071:
1064:
1057:
1050:
1045:Gnathomortis
1043:
1036:
1029:
1024:Amphekepubis
1022:
1015:Mosasaurinae
917:
913:
903:
870:
866:
860:
852:
801:
797:
731:
727:
679:
673:
669:
665:
663:
657:
640:
636:
632:
624:
622:
619:Paleobiology
608:
606:
602:Mosasauridae
599:polyphyletic
595:monophyletic
590:Haasiasaurus
588:
582:
576:
569:
562:
555:
552:sister taxon
547:
539:
537:
532:
516:
512:
510:
503:
499:
497:
492:
489:MTM V.01.115
488:
483:Pachyvaranus
481:
477:
472:
471:
465:
464:
459:
455:
431:
429:
424:
420:
403:
398:
396:
391:
384:Bakony Hills
375:
369:
366:-style limbs
359:
342:
341:
312:tethysaurine
299:
298:
297:
285:
284:
275:Type species
259:
258:
223:
217:Mosasauridae
194:
24:
18:
1734:Wikispecies
1630:Judeasaurus
1623:Adriosaurus
1607:Pontosaurus
1579:Coniasaurus
1558:Komensaurus
1551:Carsosaurus
1503:Romeosaurus
1448:Tethysaurus
1417:Selmasaurus
1362:Platecarpus
1254:Halisaurini
1210:Plotosaurus
1196:Moanasaurus
1173:Mosasaurini
1052:Jormungandr
1038:Dallasaurus
734:: 228â238.
675:Dallasaurus
584:Komensaurus
578:Carsosaurus
566:columbianus
557:Tethysaurus
446:Description
442:, Hungary.
364:aigialosaur
315:mosasauroid
239:Subfamily:
1810:Categories
1600:Primitivus
1483:Tylosaurus
1469:Kaikaifilu
1270:Halisaurus
1203:Mosasaurus
1107:Carinodens
1087:Stelladens
1059:Kourisodon
687:References
505:Halisaurus
386:, Western
332:stage) of
204:Mosasauria
1593:Kaganaias
1410:Khinjaria
1263:Eonatator
1243:Pluridens
1114:Globidens
1031:Clidastes
985:Kingdom:
975:Mosasaurs
895:128706999
748:0195-6671
613:cetaceans
330:Santonian
154:Kingdom:
148:Eukaryota
1725:Q1397354
1719:Wikidata
1687:Category
1135:Xenodens
1005:Squamata
999:Reptilia
993:Chordata
991:Phylum:
987:Animalia
867:Geol Mag
838:23284766
798:PLOS ONE
656:Size of
440:Budapest
372:holotype
210:Family:
188:Squamata
178:Reptilia
168:Chordata
164:Phylum:
158:Animalia
144:Domain:
1780:6158065
1767:1483230
1754:8666962
1003:Order:
997:Class:
922:Bibcode
875:Bibcode
829:3526648
806:Bibcode
792:(ed.).
559:nopcsai
550:as the
414:IharkĂșt
388:Hungary
338:species
334:Hungary
305:extinct
252:Genus:
184:Order:
174:Class:
105:↓
1793:253722
1066:Liodon
893:
836:
826:
746:
587:, and
573:coheni
568:, and
303:is an
1762:IRMNG
891:S2CID
724:(PDF)
648:Limbs
458:) of
308:genus
224:Clade
195:Clade
1749:GBIF
834:PMID
744:ISSN
410:Ajka
370:The
324:and
267:2012
44:Preê
930:doi
918:280
883:doi
871:148
824:PMC
814:doi
736:doi
418:cf.
310:of
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1751::
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889:.
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812:.
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796:.
756:^
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732:57
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711:^
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561:,
438:,
340:,
226::
197::
94:Pg
38:Ma
33:,
1466:?
967:e
960:t
953:v
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