84:
264:
2587:
38:
2575:
2599:
2563:
690:
Gerardo Lamas (2008) Systematics of butterflies (Lepidoptera: Hesperioidea and
Papilionoidea) in the world: current state and future perspectives (in Spanish). In: Jorge Llorente-Bousquets and AnalĂa Lanteri (eds.)
499:
Of the subfamilies of
Nymphalidae, only the Morphinae and Satyrinae are possibly paraphyletic, but these two subfamilies form a strongly supported clade with the Charaxinae as sister group.
548:
deposits dating roughly between 50 and 25 million years ago – suggest the radiation of the
Papilionoidea into the present-day families took place during that epoch.
2858:
2802:
544:, yet another prehistoric genus, also seems to belong here, but its relationships are quite obscure, indeed. However, these fossils – all found in
2750:
2776:
723:
2853:
2540:
2781:
2789:
2553:
83:
2662:
2535:
554:, from the end of the Eocene, can be quite robustly assigned to the Nymphalidae, and is quite likely a member of the
2807:
2729:
2742:
716:
503:
2843:
2615:
402:
267:
2815:
2112:
371:
19:
This article is about a group of butterflies. For the group of flowers that look like butterflies, see
2863:
2848:
2711:
982:
974:
965:
709:
2128:
1903:
1895:
301:
The members of the
Papilionoidea may be distinguished by the following combination of characters:
2653:
78:
2820:
540:
349:) are true butterflies that should be included within the Papilionoidea superfamily to reflect
2698:
2579:
1473:
673:
624:
312:
62:
2454:
1945:
1504:
663:
655:
614:
606:
263:
69:
2586:
2023:
1699:
1576:
1452:
1444:
1410:
1384:
1376:
1043:
1014:
1008:
999:
810:
804:
792:
55:
47:
37:
2737:
2724:
2603:
2591:
2567:
2244:
2031:
1709:
1634:
1600:
1558:
1483:
1468:
1111:
668:
643:
619:
594:
520:
278:
20:
595:"Cretaceous origin and repeated tertiary diversification of the redefined butterflies"
562:
fossils of
Papilionoidea are usually assignable to an extant family without problems.
2837:
2667:
2472:
2397:
2305:
2252:
2231:
2223:
1974:
1966:
1686:
1568:
1563:
1553:
1332:
1324:
1311:
1303:
1274:
1235:
1103:
1048:
1035:
1026:
951:
943:
934:
896:
890:
878:
525:
484:
158:
2574:
2491:
2459:
2441:
2389:
2321:
2281:
2081:
2073:
2039:
1924:
1916:
1737:
1727:
1714:
1668:
1660:
1647:
1639:
1626:
1584:
1545:
1535:
1512:
1491:
1460:
1423:
1415:
1402:
1282:
1230:
776:
644:"Phylogenomics provides strong evidence for relationships of butterflies and moths"
491:
379:
243:
565:
Taken together, these fossils place the origin of the
Papilionoidea in the latest
538:, is more firmly assigned to the Lycaenidae, but might belong to the Nymphalidae.
2703:
2794:
2763:
2647:
2449:
2407:
2331:
2326:
2297:
2265:
2160:
2101:
1937:
1861:
1853:
1840:
1832:
1803:
1753:
1678:
1592:
1527:
1522:
1517:
1499:
1290:
1225:
1153:
1132:
1124:
1074:
853:
847:
835:
733:
514:
478:
439:
427:
413:
406:
365:
342:
319:
238:
228:
197:
145:
135:
2598:
2515:
2423:
2415:
2368:
2352:
2339:
2313:
2289:
2273:
2194:
2181:
2173:
2152:
2144:
2060:
2052:
1995:
1987:
1953:
1811:
1745:
1732:
1704:
1691:
1613:
1240:
1182:
1087:
701:
555:
534:
472:
466:
459:
433:
397:
391:
350:
295:
271:
253:
233:
2638:
2467:
2373:
2210:
2202:
2120:
1795:
1782:
1766:
1618:
1608:
1394:
1261:
1253:
1209:
1095:
764:
752:
570:
559:
532:. It is variously placed in the Lycaenidae or Riodinidae. A similar fossil,
529:
452:
445:
346:
338:
334:
291:
212:
115:
95:
677:
659:
628:
610:
2685:
2677:
2632:
2360:
2136:
1882:
1874:
1819:
1774:
1431:
1349:
1217:
1082:
1065:
918:
770:
566:
550:
420:
385:
375:
248:
184:
171:
2562:
2381:
2013:
2005:
1722:
286:
2768:
2431:
758:
746:
593:
Heikkilä, M.; Kaila, L.; Mutanen, M.; Peña, C.; Wahlberg, N. (2012).
573:, while the extant families emerged around the early Eocene onwards.
545:
509:
337:
analyses suggest the traditionally circumscribed
Papilionoidea are a
125:
105:
2755:
2690:
2609:
323:
2613:
915:
875:
832:
789:
705:
693:
Contribuiciones taxonĂłmicas en ordens de insectos hiperdiversos
2716:
2155:(gossamer-winged butterflies: blues, coppers and relatives)
2376:(daggers, sallows, owlet moths, quakers, cutworms, darts)
504:
2363:(underwing, tiger, tussock, litter, snout, owlet moths)
648:
Proceedings of the Royal
Society B: Biological Sciences
599:
Proceedings of the Royal
Society B: Biological Sciences
361:
The six well-supported families of Papilionoidea are:
315:
are straight and clubbed or hooked as in the skippers.
2551:
2622:
2484:
2440:
2406:
2351:
2264:
2243:
2222:
2193:
2172:
2093:
2072:
2051:
2022:
2004:
1986:
1965:
1936:
1915:
1894:
1873:
1852:
1831:
1794:
1765:
1677:
1659:
1544:
1482:
1443:
1393:
1375:
1348:
1323:
1302:
1273:
1252:
1208:
1181:
1152:
1123:
1073:
1064:
1034:
1025:
1007:
998:
973:
964:
942:
933:
889:
846:
803:
1814:(castniid moths: giant butterfly-moths, sun moths)
588:
586:
390:Blues and coppers or gossamer-winged butterflies,
345:) and Neotropical moth-like butterflies (family
717:
8:
409:which contain the following 13 subfamilies:
2610:
2163:(brush-footed, or four-footed butterflies)
1372:
1368:
1205:
1178:
1070:
1031:
1004:
970:
939:
930:
926:
912:
886:
872:
843:
829:
800:
786:
724:
710:
702:
642:Kawahara, A. Y.; Breinholt, J. W. (2014).
329:The pupae are angular rather than rounded.
36:
27:
667:
618:
2342:(hawk moths, sphinx moths and hornworms)
448:(including the owls as tribe Brassolini)
290:, meaning "butterfly") contains all the
262:
2558:
2139:(whites, yellows, orangetips, sulphurs)
582:
305:The body is smaller and less moth-like.
2255:(eggars, snout moths, or lappet moths)
524:, which has long been mistaken for a
7:
2859:Taxa named by Pierre André Latreille
2743:d15819d0-778c-445c-9971-44ce6cdd650e
1777:(carpenter millers, or goat moths)
1694:(burnet, forester, or smoky moths)
14:
1098:(Amazonian primitive ghost moths)
416:(formerly the family Libytheidae)
341:group, and that skippers (family
2597:
2585:
2573:
2561:
322:do not spin cocoons in which to
82:
1114:(African primitive ghost moths)
695:. Mexico City: UNAM. Pp. 57-70.
462:(preponas and leaf butterflies)
455:(formerly the family Satyridae)
1418:(Old World spiny-winged moths)
518:is apparently a Papilionoidea
423:(formerly the family Danaidae)
1:
2316:(Kentish glory and relatives)
1135:(New Zealand primitive moths)
2462:(oriental swallowtail moths)
1740:(planthopper parasite moths)
2536:Taxonomy of the Lepidoptera
2123:(American moth-butterflies)
2084:(Old World butterfly-moths)
2063:(picture-winged leaf moths)
813:(mandibulate archaic moths)
2880:
2034:(tropical fruitworm moths)
1571:(case-bearers, case moths)
1335:(trumpet leaf miner moths)
1090:(swift moths, ghost moths)
501:
490:The tropical longwings or
18:
16:Superfamily of butterflies
2854:Lepidoptera superfamilies
2531:
2512:
2284:(American silkworm moths)
2115:(swallowtail butterflies)
1785:(dudgeon carpenter moths)
1494:(false diamondback moths)
1405:(burrowing webworm moths)
1371:
929:
925:
911:
885:
871:
842:
828:
799:
785:
741:
487:(especially the adelphas)
412:The snout butterflies or
384:Whites or yellow-whites,
357:Families of Papilionoidea
294:except for the moth-like
225:
220:
79:Scientific classification
77:
44:
35:
30:
2276:(Australian lappet moth)
1285:(pigmy, or midget moths)
444:The morphos and owls or
403:Brush-footed butterflies
1106:(miniature ghost moths)
396:Metalmark butterflies,
2485:Superfamily unassigned
1906:(bristle-legged moths)
1455:(ribbed cocoon makers)
1264:(Andean endemic moths)
1220:(fairy longhorn moths)
660:10.1098/rspb.2014.0970
611:10.1098/rspb.2011.1430
274:
268:Brush-footed butterfly
2816:Paleobiology Database
2392:(prominents, kittens)
1927:(fringe-tufted moths)
983:Acanthopteroctetidae
975:Acanthopteroctetoidea
308:The wings are larger.
266:
2738:Fauna Europaea (new)
1904:Schreckensteiniidae
1896:Schreckensteinioidea
1885:(false burnet moths)
1756:(slug, or cup moths)
1314:(Gondwanaland moths)
1293:(white eyecap moths)
1051:(archaic bell moths)
2234:(sack bearer moths)
1956:(false plume moths)
1948:(many-plumed moths)
1806:(little bear moths)
1748:(slug caterpillars)
1603:(long-horned moths)
1587:(grass-miner moths)
985:(archaic sun moths)
605:(1731): 1093–1099.
654:(1788): 20140970.
432:The glasswings or
275:
2831:
2830:
2616:Taxon identifiers
2549:
2548:
2527:
2526:
2523:
2522:
2508:
2507:
2504:
2503:
2500:
2499:
2476:
2463:
2427:
2419:
2393:
2385:
2377:
2364:
2343:
2335:
2317:
2309:
2301:
2293:
2285:
2277:
2256:
2235:
2214:
2206:
2185:
2164:
2156:
2148:
2140:
2132:
2124:
2116:
2106:
2085:
2064:
2043:
2042:(fruitworm moths)
2035:
1978:
1957:
1949:
1928:
1907:
1886:
1865:
1844:
1843:(metalmark moths)
1823:
1822:(clearwing moths)
1815:
1807:
1786:
1778:
1757:
1749:
1741:
1718:
1695:
1651:
1643:
1630:
1629:(concealer moths)
1622:
1604:
1596:
1588:
1580:
1572:
1531:
1508:
1505:Glyphipterigidae
1495:
1474:Roeslerstammiidae
1464:
1456:
1435:
1427:
1419:
1406:
1344:
1343:
1336:
1315:
1294:
1286:
1265:
1244:
1221:
1148:
1147:
1144:
1143:
1136:
1115:
1107:
1099:
1091:
1060:
1059:
1052:
1021:
1020:
994:
993:
986:
960:
959:
907:
906:
903:
902:
891:Heterobathmioidea
867:
866:
863:
862:
857:
824:
823:
820:
819:
814:
261:
260:
216:
63:Myscelia cyaniris
2871:
2824:
2823:
2811:
2810:
2798:
2797:
2795:NHMSYS0000503590
2785:
2784:
2772:
2771:
2759:
2758:
2746:
2745:
2733:
2732:
2720:
2719:
2707:
2706:
2694:
2693:
2681:
2680:
2671:
2670:
2658:
2657:
2656:
2643:
2642:
2641:
2611:
2602:
2601:
2590:
2589:
2578:
2577:
2566:
2565:
2557:
2475:(geometer moths)
2474:
2461:
2455:Pseudobistonidae
2425:
2417:
2391:
2383:
2375:
2362:
2341:
2333:
2315:
2307:
2299:
2291:
2283:
2275:
2254:
2233:
2212:
2204:
2183:
2162:
2154:
2146:
2138:
2130:
2122:
2114:
2105:
2098:
2083:
2062:
2041:
2033:
1976:
1955:
1947:
1926:
1905:
1884:
1863:
1842:
1821:
1813:
1805:
1784:
1776:
1755:
1747:
1739:
1716:
1693:
1649:
1641:
1628:
1620:
1602:
1594:
1586:
1578:
1577:Cosmopterigidae
1570:
1529:
1506:
1493:
1462:
1454:
1453:Bucculatricidae
1433:
1425:
1417:
1404:
1373:
1369:
1334:
1313:
1292:
1284:
1263:
1242:
1219:
1206:
1179:
1134:
1113:
1105:
1097:
1089:
1071:
1050:
1032:
1005:
984:
971:
940:
931:
927:
913:
897:Heterobathmiidae
887:
873:
855:
844:
830:
812:
811:Micropterigidae
805:Micropterigoidea
801:
787:
726:
719:
712:
703:
696:
688:
682:
681:
671:
639:
633:
632:
622:
590:
284:(from the genus
211:
87:
86:
70:Episcada apuleia
40:
28:
2879:
2878:
2874:
2873:
2872:
2870:
2869:
2868:
2834:
2833:
2832:
2827:
2819:
2814:
2806:
2801:
2793:
2788:
2780:
2775:
2767:
2762:
2754:
2749:
2741:
2736:
2728:
2723:
2715:
2710:
2702:
2697:
2689:
2684:
2676:
2674:
2666:
2661:
2652:
2651:
2646:
2637:
2636:
2631:
2618:
2608:
2596:
2584:
2572:
2560:
2552:
2550:
2545:
2541:Lists by region
2519:
2518:is not a clade.
2514:Note: division
2496:
2480:
2436:
2402:
2347:
2308:(Dryandra moth)
2300:(Brahmin moths)
2260:
2239:
2218:
2189:
2168:
2099:
2097:
2089:
2068:
2047:
2032:Copromorphidae
2024:Copromorphoidea
2018:
2000:
1982:
1961:
1932:
1911:
1890:
1869:
1864:(tortrix moths)
1848:
1827:
1790:
1761:
1717:(flannel moths)
1700:Himantopteridae
1673:
1655:
1601:Lecithoceridae
1595:(twirler moths)
1540:
1478:
1463:(Douglas moths)
1445:Gracillarioidea
1439:
1426:(bagworm moths)
1411:Arrhenophanidae
1389:
1377:Simaethistoidea
1363:
1361:
1359:
1357:
1355:
1353:
1351:
1340:
1319:
1298:
1269:
1248:
1200:
1198:
1196:
1194:
1192:
1190:
1188:
1186:
1184:
1173:
1171:
1169:
1167:
1165:
1163:
1161:
1159:
1157:
1155:
1140:
1133:Mnesarchaeidae
1125:Mnesarchaeoidea
1119:
1112:Prototheoridae
1056:
1044:Aenigmatineidae
1017:
1009:Lophocoronoidea
990:
956:
921:
899:
881:
879:Heterobathmiina
859:
854:Agathiphagidae
848:Agathiphagoidea
838:
816:
795:
781:
737:
730:
700:
699:
689:
685:
641:
640:
636:
592:
591:
584:
579:
506:
419:The danaids or
359:
353:relationships.
210:
81:
66:
59:
56:Calinaga buddha
51:
48:Delias eucharis
24:
17:
12:
11:
5:
2877:
2875:
2867:
2866:
2861:
2856:
2851:
2846:
2836:
2835:
2829:
2828:
2826:
2825:
2812:
2799:
2786:
2773:
2760:
2747:
2734:
2725:Fauna Europaea
2721:
2708:
2695:
2682:
2672:
2659:
2644:
2628:
2626:
2620:
2619:
2614:
2607:
2606:
2594:
2582:
2570:
2547:
2546:
2544:
2543:
2538:
2532:
2529:
2528:
2525:
2524:
2521:
2520:
2513:
2510:
2509:
2506:
2505:
2502:
2501:
2498:
2497:
2495:
2494:
2488:
2486:
2482:
2481:
2479:
2478:
2470:
2465:
2457:
2452:
2446:
2444:
2438:
2437:
2435:
2434:
2429:
2421:
2412:
2410:
2404:
2403:
2401:
2400:
2395:
2387:
2379:
2371:
2366:
2357:
2355:
2349:
2348:
2346:
2345:
2337:
2329:
2324:
2319:
2311:
2303:
2295:
2287:
2279:
2270:
2268:
2262:
2261:
2259:
2258:
2253:Lasiocampidae
2249:
2247:
2245:Lasiocampoidea
2241:
2240:
2238:
2237:
2228:
2226:
2220:
2219:
2217:
2216:
2208:
2199:
2197:
2191:
2190:
2188:
2187:
2178:
2176:
2170:
2169:
2167:
2166:
2158:
2150:
2142:
2134:
2126:
2118:
2109:
2107:
2091:
2090:
2088:
2087:
2078:
2076:
2070:
2069:
2067:
2066:
2057:
2055:
2049:
2048:
2046:
2045:
2037:
2028:
2026:
2020:
2019:
2017:
2016:
2010:
2008:
2002:
2001:
1999:
1998:
1992:
1990:
1988:Whalleyanoidea
1984:
1983:
1981:
1980:
1975:Pterophoridae
1971:
1969:
1967:Pterophoroidea
1963:
1962:
1960:
1959:
1951:
1942:
1940:
1934:
1933:
1931:
1930:
1921:
1919:
1913:
1912:
1910:
1909:
1900:
1898:
1892:
1891:
1889:
1888:
1879:
1877:
1871:
1870:
1868:
1867:
1858:
1856:
1850:
1849:
1847:
1846:
1837:
1835:
1829:
1828:
1826:
1825:
1817:
1809:
1800:
1798:
1792:
1791:
1789:
1788:
1780:
1771:
1769:
1763:
1762:
1760:
1759:
1751:
1743:
1735:
1730:
1725:
1720:
1715:Megalopygidae
1712:
1710:Somabrachyidae
1707:
1702:
1697:
1689:
1683:
1681:
1675:
1674:
1672:
1671:
1665:
1663:
1657:
1656:
1654:
1653:
1650:(timber moths)
1645:
1642:(flower moths)
1637:
1635:Pterolonchidae
1632:
1624:
1621:(mompha moths)
1616:
1611:
1606:
1598:
1590:
1582:
1579:(cosmet moths)
1574:
1569:Coleophoridae
1566:
1561:
1559:Batrachedridae
1556:
1550:
1548:
1542:
1541:
1539:
1538:
1533:
1530:(ermine moths)
1528:Yponomeutidae
1525:
1520:
1515:
1510:
1502:
1497:
1488:
1486:
1484:Yponomeutoidea
1480:
1479:
1477:
1476:
1471:
1469:Gracillariidae
1466:
1458:
1449:
1447:
1441:
1440:
1438:
1437:
1434:(fungus moths)
1429:
1421:
1413:
1408:
1399:
1397:
1391:
1390:
1388:
1387:
1385:Simaethistidae
1381:
1379:
1366:
1346:
1345:
1342:
1341:
1339:
1338:
1329:
1327:
1321:
1320:
1318:
1317:
1312:Palaephatidae
1308:
1306:
1304:Palaephatoidea
1300:
1299:
1297:
1296:
1288:
1279:
1277:
1271:
1270:
1268:
1267:
1258:
1256:
1250:
1249:
1247:
1246:
1238:
1233:
1228:
1223:
1214:
1212:
1203:
1176:
1150:
1149:
1146:
1145:
1142:
1141:
1139:
1138:
1129:
1127:
1121:
1120:
1118:
1117:
1109:
1104:Palaeosetidae
1101:
1093:
1085:
1079:
1077:
1068:
1062:
1061:
1058:
1057:
1055:
1054:
1049:Neopseustidae
1046:
1040:
1038:
1036:Neopseustoidea
1029:
1023:
1022:
1019:
1018:
1015:Lophocoronidae
1013:
1011:
1002:
996:
995:
992:
991:
989:
988:
979:
977:
968:
962:
961:
958:
957:
955:
954:
948:
946:
944:Eriocranioidea
937:
923:
922:
916:
909:
908:
905:
904:
901:
900:
895:
893:
883:
882:
876:
869:
868:
865:
864:
861:
860:
852:
850:
840:
839:
833:
826:
825:
822:
821:
818:
817:
809:
807:
797:
796:
790:
783:
782:
780:
779:
773:
767:
761:
755:
749:
742:
739:
738:
731:
729:
728:
721:
714:
706:
698:
697:
683:
634:
581:
580:
578:
575:
521:incertae sedis
497:
496:
495:
494:
488:
481:
477:The nymphs or
475:
469:
463:
456:
451:The browns or
449:
442:
436:
430:
424:
417:
400:
394:
388:
382:
369:
358:
355:
331:
330:
327:
316:
309:
306:
259:
258:
257:
256:
251:
246:
241:
236:
231:
223:
222:
218:
217:
205:
201:
200:
195:
188:
187:
182:
175:
174:
169:
162:
161:
156:
149:
148:
143:
139:
138:
133:
129:
128:
123:
119:
118:
113:
109:
108:
103:
99:
98:
93:
89:
88:
75:
74:
42:
41:
33:
32:
31:Papilionoidea
21:Papilionoideae
15:
13:
10:
9:
6:
4:
3:
2:
2876:
2865:
2862:
2860:
2857:
2855:
2852:
2850:
2847:
2845:
2844:Papilionoidea
2842:
2841:
2839:
2822:
2817:
2813:
2809:
2804:
2800:
2796:
2791:
2787:
2783:
2778:
2774:
2770:
2765:
2761:
2757:
2752:
2748:
2744:
2739:
2735:
2731:
2726:
2722:
2718:
2713:
2709:
2705:
2700:
2696:
2692:
2687:
2683:
2679:
2673:
2669:
2668:Papilionoidea
2664:
2660:
2655:
2654:Papilionoidea
2649:
2645:
2640:
2634:
2630:
2629:
2627:
2625:
2624:Papilionoidea
2621:
2617:
2612:
2605:
2600:
2595:
2593:
2588:
2583:
2581:
2576:
2571:
2569:
2564:
2559:
2555:
2542:
2539:
2537:
2534:
2533:
2530:
2517:
2511:
2493:
2490:
2489:
2487:
2483:
2477:
2471:
2469:
2466:
2464:
2460:Epicopeiidae
2458:
2456:
2453:
2451:
2448:
2447:
2445:
2443:
2439:
2433:
2430:
2428:
2422:
2420:
2414:
2413:
2411:
2409:
2405:
2399:
2398:Oenosandridae
2396:
2394:
2390:Notodontidae
2388:
2386:
2380:
2378:
2372:
2370:
2367:
2365:
2359:
2358:
2356:
2354:
2350:
2344:
2338:
2336:
2330:
2328:
2325:
2323:
2320:
2318:
2312:
2310:
2304:
2302:
2296:
2294:
2288:
2286:
2282:Apatelodidae
2280:
2278:
2272:
2271:
2269:
2267:
2263:
2257:
2251:
2250:
2248:
2246:
2242:
2236:
2232:Mimallonidae
2230:
2229:
2227:
2225:
2224:Mimallonoidea
2221:
2215:
2209:
2207:
2205:(snout moths)
2201:
2200:
2198:
2196:
2192:
2186:
2180:
2179:
2177:
2175:
2171:
2165:
2159:
2157:
2151:
2149:
2143:
2141:
2135:
2133:
2127:
2125:
2119:
2117:
2113:Papilionidae
2111:
2110:
2108:
2103:
2096:
2095:Papilionoidea
2092:
2086:
2082:Callidulidae
2080:
2079:
2077:
2075:
2074:Calliduloidea
2071:
2065:
2059:
2058:
2056:
2054:
2050:
2044:
2040:Carposinidae
2038:
2036:
2030:
2029:
2027:
2025:
2021:
2015:
2012:
2011:
2009:
2007:
2003:
1997:
1996:Whalleyanidae
1994:
1993:
1991:
1989:
1985:
1979:
1977:(plume moths)
1973:
1972:
1970:
1968:
1964:
1958:
1952:
1950:
1944:
1943:
1941:
1939:
1935:
1929:
1925:Epermeniidae
1923:
1922:
1920:
1918:
1917:Epermenioidea
1914:
1908:
1902:
1901:
1899:
1897:
1893:
1887:
1881:
1880:
1878:
1876:
1872:
1866:
1860:
1859:
1857:
1855:
1851:
1845:
1839:
1838:
1836:
1834:
1830:
1824:
1818:
1816:
1810:
1808:
1802:
1801:
1799:
1797:
1793:
1787:
1783:Dudgeoneidae
1781:
1779:
1773:
1772:
1770:
1768:
1764:
1758:
1752:
1750:
1744:
1742:
1738:Epipyropidae
1736:
1734:
1733:Cyclotornidae
1731:
1729:
1726:
1724:
1721:
1719:
1713:
1711:
1708:
1706:
1703:
1701:
1698:
1696:
1690:
1688:
1687:Heterogynidae
1685:
1684:
1682:
1680:
1676:
1670:
1667:
1666:
1664:
1662:
1661:Galacticoidea
1658:
1652:
1648:Xyloryctidae
1646:
1644:
1640:Scythrididae
1638:
1636:
1633:
1631:
1627:Oecophoridae
1625:
1623:
1617:
1615:
1614:Metachandidae
1612:
1610:
1607:
1605:
1599:
1597:
1591:
1589:
1585:Elachistidae
1583:
1581:
1575:
1573:
1567:
1565:
1564:Blastobasidae
1562:
1560:
1557:
1555:
1554:Autostichidae
1552:
1551:
1549:
1547:
1543:
1537:
1534:
1532:
1526:
1524:
1521:
1519:
1516:
1514:
1511:
1509:
1507:(sedge moths)
1503:
1501:
1498:
1496:
1492:Acrolepiidae
1490:
1489:
1487:
1485:
1481:
1475:
1472:
1470:
1467:
1465:
1461:Douglasiidae
1459:
1457:
1451:
1450:
1448:
1446:
1442:
1436:
1430:
1428:
1422:
1420:
1416:Eriocottidae
1414:
1412:
1409:
1407:
1403:Acrolophidae
1401:
1400:
1398:
1396:
1392:
1386:
1383:
1382:
1380:
1378:
1374:
1370:
1367:
1365:
1347:
1337:
1333:Tischeriidae
1331:
1330:
1328:
1326:
1325:Tischerioidea
1322:
1316:
1310:
1309:
1307:
1305:
1301:
1295:
1289:
1287:
1283:Nepticulidae
1281:
1280:
1278:
1276:
1275:Nepticuloidea
1272:
1266:
1262:Andesianidae
1260:
1259:
1257:
1255:
1254:Andesianoidea
1251:
1245:
1243:(yucca moths)
1239:
1237:
1236:Incurvariidae
1234:
1232:
1229:
1227:
1224:
1222:
1216:
1215:
1213:
1211:
1207:
1204:
1202:
1180:
1177:
1175:
1151:
1137:
1131:
1130:
1128:
1126:
1122:
1116:
1110:
1108:
1102:
1100:
1096:Neotheoridae
1094:
1092:
1086:
1084:
1081:
1080:
1078:
1076:
1072:
1069:
1067:
1063:
1053:
1047:
1045:
1042:
1041:
1039:
1037:
1033:
1030:
1028:
1024:
1016:
1012:
1010:
1006:
1003:
1001:
1000:Lophocoronina
997:
987:
981:
980:
978:
976:
972:
969:
967:
966:Acanthoctesia
963:
953:
952:Eriocraniidae
950:
949:
947:
945:
941:
938:
936:
932:
928:
924:
920:
914:
910:
898:
894:
892:
888:
884:
880:
874:
870:
858:
856:(kauri moths)
851:
849:
845:
841:
837:
831:
827:
815:
808:
806:
802:
798:
794:
788:
784:
778:
774:
772:
768:
766:
762:
760:
756:
754:
750:
748:
744:
743:
740:
735:
727:
722:
720:
715:
713:
708:
707:
704:
694:
687:
684:
679:
675:
670:
665:
661:
657:
653:
649:
645:
638:
635:
630:
626:
621:
616:
612:
608:
604:
600:
596:
589:
587:
583:
576:
574:
572:
568:
563:
561:
557:
553:
552:
547:
543:
542:
537:
536:
531:
527:
526:geometer moth
523:
522:
517:
516:
511:
505:
500:
493:
489:
486:
485:Limenitidinae
482:
480:
476:
474:
470:
468:
464:
461:
457:
454:
450:
447:
443:
441:
437:
435:
431:
429:
425:
422:
418:
415:
411:
410:
408:
404:
401:
399:
395:
393:
389:
387:
383:
381:
377:
373:
370:
367:
364:
363:
362:
356:
354:
352:
348:
344:
340:
336:
328:
325:
321:
317:
314:
310:
307:
304:
303:
302:
299:
297:
293:
289:
288:
283:
282:Papilionoidea
280:
273:
270:of subfamily
269:
265:
255:
252:
250:
247:
245:
242:
240:
237:
235:
232:
230:
227:
226:
224:
219:
214:
209:
208:Papilionoidea
206:
204:Superfamily:
203:
202:
199:
196:
193:
190:
189:
186:
183:
180:
177:
176:
173:
170:
167:
164:
163:
160:
159:Eulepidoptera
157:
154:
151:
150:
147:
144:
141:
140:
137:
134:
131:
130:
127:
124:
121:
120:
117:
114:
111:
110:
107:
104:
101:
100:
97:
94:
91:
90:
85:
80:
76:
73:
72:
71:
67:Below right:
65:
64:
58:
57:
50:
49:
43:
39:
34:
29:
26:
22:
2623:
2492:Millieriidae
2473:Geometridae
2442:Geometroidea
2426:(gold moths)
2384:(tuft moths)
2334:(saturniids)
2332:Saturniidae
2322:Eupterotidae
2306:Carthaeidae
2298:Brahmaeidae
2292:(silk moths)
2213:(grass moth)
2184:(teak moths)
2161:Nymphalidae
2147:(metalmarks)
2129:Hesperiidae
2094:
1862:Tortricidae
1854:Tortricoidea
1841:Choreutidae
1833:Choreutoidea
1804:Brachodidae
1754:Limacodidae
1728:Anomoeotidae
1669:Galacticidae
1593:Gelechiidae
1546:Gelechioidea
1536:Ypsolophidae
1513:Heliodinidae
1291:Opostegidae
1231:Heliozelidae
1083:Anomosetidae
1027:Neopseustina
777:Holometabola
775:Superorder:
769:Infraclass:
692:
686:
651:
647:
637:
602:
598:
564:
549:
539:
533:
519:
513:
507:
498:
492:Heliconiinae
380:Papilionidae
372:Swallowtails
360:
339:paraphyletic
335:phylogenetic
332:
320:caterpillars
300:
285:
281:
276:
244:Papilionidae
207:
191:
178:
165:
152:
142:Infraorder:
68:
61:
60:Below left:
54:
52:
46:
25:
2864:Obtectomera
2849:Butterflies
2764:iNaturalist
2648:Wikispecies
2450:Sematuridae
2424:Cimeliidae
2418:(hook-tips)
2416:Drepanidae
2408:Drepanoidea
2340:Sphingidae
2327:Phiditiidae
2314:Endromidae
2290:Bombycidae
2274:Anthelidae
2266:Bombycoidea
2182:Hyblaeidae
2174:Hyblaeoidea
2153:Lycaenidae
2145:Riodinidae
2102:butterflies
2061:Thyrididae
2053:Thyridoidea
1954:Tineodidae
1946:Alucitidae
1938:Alucitoidea
1812:Castniidae
1746:Dalceridae
1692:Zygaenidae
1679:Zygaenoidea
1523:Plutellidae
1518:Lyonetiidae
1500:Bedelliidae
1241:Prodoxidae
1226:Cecidosidae
1088:Hepialidae
1075:Hepialoidea
793:Zeugloptera
734:Lepidoptera
515:Lithopsyche
479:Nymphalinae
440:Calinaginae
428:Tellervinae
414:Libytheinae
407:Nymphalidae
366:Hesperiidae
343:Hesperiidae
292:butterflies
279:superfamily
239:Nymphalidae
229:Hesperiidae
198:Obtectomera
185:Apoditrysia
146:Heteroneura
136:Lepidoptera
53:Top right:
2838:Categories
2580:Arthropods
2516:Monotrysia
2374:Noctuidae
2369:Euteliidae
2353:Noctuoidea
2211:Crambidae
2203:Pyralidae
2195:Pyraloidea
2131:(skippers)
2121:Hedylidae
1705:Lacturidae
1619:Momphidae
1424:Psychidae
935:Dacnonypha
763:Subclass:
753:Arthropoda
577:References
556:Nymphalini
541:Riodinella
535:Lithodryas
502:See also:
473:Apaturinae
467:Biblidinae
460:Charaxinae
434:Ithomiinae
398:Riodinidae
392:Lycaenidae
368:(skippers)
296:Hedyloidea
272:Charaxinae
254:Riodinidae
234:Lycaenidae
116:Arthropoda
45:Top left:
2468:Uraniidae
2361:Erebidae
2137:Pieridae
1883:Urodidae
1875:Urodoidea
1820:Sesiidae
1796:Sesioidea
1775:Cossidae
1767:Cossoidea
1609:Lypusidae
1432:Tineidae
1395:Tineoidea
1218:Adelidae
1210:Adeloidea
917:Suborder
877:Suborder
836:Aglossata
834:Suborder
791:Suborder
765:Pterygota
745:Kingdom:
571:Paleogene
569:or early
560:Oligocene
530:Boarmiini
453:Satyrinae
446:Morphinae
376:birdwings
351:cladistic
347:Hedylidae
221:Families
213:Latreille
102:Kingdom:
96:Eukaryota
2686:BugGuide
2675:BioLib:
2633:Wikidata
2382:Nolidae
2006:Immoidea
1066:Exoporia
919:Glossata
771:Neoptera
751:Phylum:
747:Animalia
736:families
678:24966318
629:21920981
567:Mesozoic
551:Prodryas
421:Danainae
386:Pieridae
313:antennae
249:Pieridae
172:Ditrysia
112:Phylum:
106:Animalia
92:Domain:
2639:Q756153
2604:Biology
2592:Animals
2568:Insects
2554:Portals
2014:Immidae
1723:Aididae
759:Insecta
757:Class:
732:Extant
669:4083801
620:3267136
528:of the
333:Recent
287:Papilio
132:Order:
126:Insecta
122:Class:
2821:212998
2782:117235
2432:Doidae
676:
666:
627:
617:
546:Eocene
512:genus
510:fossil
324:pupate
215:, 1802
2808:37572
2769:47224
2678:17297
192:Clade
179:Clade
166:Clade
153:Clade
2803:NCBI
2777:ITIS
2756:1875
2751:GBIF
2730:6944
674:PMID
625:PMID
508:The
483:The
471:The
465:The
458:The
438:The
426:The
374:and
318:The
311:The
277:The
2790:NBN
2717:854
2712:EoL
2704:5G9
2699:CoL
2663:AFD
664:PMC
656:doi
652:281
615:PMC
607:doi
603:279
2840::
2818::
2805::
2792::
2779::
2766::
2753::
2740::
2727::
2714::
2701::
2691:81
2688::
2665::
2650::
2635::
672:.
662:.
650:.
646:.
623:.
613:.
601:.
597:.
585:^
558:.
405:,
378:,
298:.
194::
181::
168::
155::
2556::
2104:)
2100:(
1364:a
1362:i
1360:s
1358:y
1356:r
1354:t
1352:i
1350:D
1201:a
1199:i
1197:s
1195:y
1193:r
1191:t
1189:o
1187:n
1185:o
1183:M
1174:a
1172:r
1170:u
1168:e
1166:n
1164:o
1162:r
1160:e
1158:t
1156:e
1154:H
725:e
718:t
711:v
680:.
658::
631:.
609::
326:.
23:.
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
