778:
4219:
6334:
566:
4212:
2716:
3541:
3527:
119:
5786:
5007:
5772:
3520:
5779:
2722:
4226:
140:
546:
3534:
6347:
5025:
529:, an invalidly published name. He validated the name three years later in two 1926 articles by providing a diagnosis of the genus. While remaining one of the best preserved theropod skulls in Europe, and globally one of the best preserved Middle Jurassic theropod skulls, it since received little scientific attention, mainly being mentioned in studies about general aspects of theropod anatomy and evolution. The holotype skull was since
770:
5013:
5032:
6341:
5766:
422:
1337:
5020:
556:
was a small dinosaur. In restudy at 2010, total length of 2.98–3.16 m (9.8–10.4 ft) and body mass of 28–36 kg (62–79 lb) are estimated. Various books estimated it to measure 3–4 m (9.8–13.1 ft) in length and 50–100 kg (110–220 lb) in body mass.
1363:
was likely not a big game hunter, lacking the bone-crushing teeth and extremely powerful bites of the tyrannosaurids. Instead, it possessed an elongated skull, commonly found in basal coelurosaurs and basal tyrannosauroids.
482:, in honour of its discoverer. At the time it was discovered, it was one of the most complete theropod skulls known from Europe, possibly with the exception of the crushed and hard to interpret skulls of
537:, further mechanically prepared to reveal additional details of the skull, jaw, and teeth, and was re-described by the German palaeontologist Oliver W. M. Rauhut and colleagues in 2010.
651:
a junior synonym. However, no overlapping bones between the two had yet been exposed from the rock around their fossils, and future study showed that they were indeed distinct.
2045:"Palaeontology, stratigraphy and sedimentology of Woodeaton Quarry (Oxfordshire) and a new microvertebrate site from the White Limestone Formation (Bathonian, Jurassic)"
1355:
possessed a nasal crest, which may have served as a display organ but also possibly served to reduce bending stresses on the skull when biting. This may indicate
6462:
360:
was a small dinosaur, estimated to measure around 3 m (9.8 ft) in length. Its name refers to how it was originally thought to be an ancestor of
6482:
1877:
6411:
1965:"A description of Megalosaurus bucklandii (Dinosauria: Theropoda) from the Bathonian of the UK and the relationships of Middle Jurassic theropods"
4218:
6477:
4211:
6333:
1778:
1601:
1568:
2178:
666:) to be a coelurosaur, only distantly related to the ceratosaurids and allosauroids, though one opinion published in 2000 considered
3540:
5006:
1464:
3526:
5771:
1997:
2715:
495:
394:
3519:
5785:
2721:
139:
678:
among the coelurosaurs, though with only weak support, and again found an (also weakly supported) close relationship with
6212:
5024:
2151:
4225:
565:
461:
6472:
5778:
3533:
1373:
777:
6346:
5031:
6467:
6452:
2123:
697:, a member of the lineage leading to the giant tyrannosaurs of the Late Cretaceous. Furthermore, they found that
2266:
1398:
Other dinosaurs known from equivalently aged deposits from the
Bathonian of Britain include the large theropod
5012:
2084:"Machine learning confirms new records of maniraptoran theropods in Middle Jurassic UK microvertebrate faunas"
2013:
6340:
6457:
2171:
1420:
6366:
2043:
Wills, Simon; Bernard, Emma Louise; Brewer, Philippa; Underwood, Charlie J.; Ward, David J. (April 2019).
1542:
1871:
Rauhut, Oliver W M; Bakirov, Aizek A; Wings, Oliver; Fernandes, Alexandra E; Hübner, Tom R (2024-08-01).
6424:
4737:
4649:
689:
and its relationships was published in 2010 by Oliver Rauhut and colleagues. Their study concluded that
437:
287:
267:
5765:
607:
had been shown to be a much more primitive theropod and not a coelurosaur, that the classification of
6398:
6298:
4426:
4394:
1998:"The anatomy and taxonomy of Cetiosaurus (Saurischia, Sauropoda) from the Middle Jurassic of England"
398:
4750:
4537:
1617:
671:
587:
534:
510:
243:
5019:
118:
6205:
4726:
4586:
4442:
4382:
3817:
3727:
3610:
2474:
2164:
2064:
2044:
2025:
1945:
1726:
1691:
499:
457:
406:
296:
134:
1743:
von Huene, F. (1932). "Die fossile Reptil-Ordnung
Saurischia, ihre Entwicklung und Geschichte."
6429:
6087:
6004:
5978:
5817:
5804:
5682:
5240:
5192:
4024:
3374:
2453:
2105:
2017:
1937:
1896:
1853:
1774:
1770:
1763:
1709:
Milner, A. C.; Barrett, Paul M. (2016). "Smith
Woodward's contributions on fossil tetrapods".
1597:
1574:
1564:
1381:
885:
712:
545:
498:, where it is catalogued as specimen NHM R 4860. The upper part of the skull missing due to a
465:
379:
223:
781:
Skull diagram showing known material in white, with skull reconstructed after close relative
6191:
5948:
5858:
5647:
5579:
5442:
5361:
5333:
5293:
5174:
5144:
4662:
4415:
4279:
4136:
4036:
3885:
3792:
3750:
3317:
2627:
2095:
2056:
2009:
1976:
1927:
1886:
1843:
1833:
1718:
1683:
1516:
1340:
1133:
899:
795:
694:
570:
383:
2082:
Wills, Simon; Underwood, Charlie J.; Barrett, Paul M. (March 2023). Mannion, Philip (ed.).
6254:
5992:
5955:
5887:
5742:
5735:
5563:
5505:
5478:
5468:
5428:
5122:
5106:
4924:
4896:
4874:
4761:
4606:
4313:
4087:
3970:
3948:
3869:
3698:
3675:
3668:
3646:
3632:
3625:
3586:
3437:
3284:
3211:
3200:
3136:
3024:
2911:
2488:
2397:
2251:
2226:
1428:
1206:
769:
612:
453:
444:
skull discovered some time prior by F. Lewis
Bradley during excavation for a reservoir at
341:
1646:
6219:
6182:
6163:
6125:
6114:
5911:
5624:
5590:
5572:
5552:
5449:
5163:
4908:
4770:
4597:
4579:
4407:
4374:
4360:
4321:
4095:
3956:
3913:
3906:
3828:
3804:
3757:
3716:
3653:
3639:
3367:
3352:
3179:
3013:
2877:
2845:
2834:
2811:
2773:
2634:
2604:
2528:
2467:
2366:
1848:
1817:
1452:
1440:
1436:
1432:
1164:
639:
445:
410:
1687:
1672:"On a Skull of Megalosaurus from the Great Oolite of Minchinhampton (Gloucestershire)"
421:
6446:
6261:
6107:
6068:
6047:
5926:
5919:
5846:
5640:
5613:
5487:
5404:
5382:
5375:
5368:
5311:
5228:
5091:
5063:
5050:
4974:
4851:
4691:
4525:
4456:
4342:
4329:
4074:
4052:
4045:
4009:
3982:
3934:
3858:
3594:
3385:
3359:
3275:
3256:
3249:
3186:
3165:
3144:
3129:
3115:
3074:
3040:
3033:
2952:
2804:
2691:
2684:
2611:
2581:
2570:
2460:
2431:
2424:
2350:
2068:
1981:
1964:
1949:
1730:
1695:
1540:
Dinosaurs: The Most
Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages
1521:
1500:
1412:
1385:
1377:
1181:
1063:
834:
731:
634:
617:
597:
582:
490:
484:
390:
69:
2029:
6233:
6171:
6141:
6061:
6054:
6032:
6024:
5904:
5896:
5876:
5599:
5518:
5456:
5347:
5318:
5270:
5099:
5084:
4859:
4791:
4719:
4698:
4677:
4493:
4449:
4257:
4244:
4174:
4127:
4002:
3892:
3844:
3837:
3774:
3743:
3736:
3617:
3444:
3296:
3193:
3095:
3088:
2983:
2967:
2903:
2854:
2786:
2563:
2446:
2439:
2410:
2373:
2342:
1448:
1424:
1400:
1109:
964:
757:
743:
655:
592:
519:
470:
362:
306:
1794:
1336:
523:, but since the name was only used in a schematic, the name has been considered a
2060:
1891:
1872:
6287:
6268:
6198:
6148:
6134:
6076:
6017:
5962:
5839:
5633:
5525:
5494:
5411:
5397:
5129:
5077:
4945:
4867:
4830:
4712:
4684:
4637:
4557:
4435:
4292:
4143:
4116:
4016:
3963:
3899:
3878:
3851:
3602:
3415:
3408:
3394:
3220:
3067:
3058:
2960:
2931:
2861:
2826:
2819:
2619:
2597:
2590:
2549:
2541:
2517:
2502:
2481:
2389:
2381:
1416:
1406:
1312:
1086:
749:
670:
a ceratosaurid without presenting supporting evidence. Phylogenetic analyses by
586:, due to same number of premaxillary teeth (4). Later study during the 1930s by
525:
375:
44:
1873:"A new theropod dinosaur from the Callovian Balabansai Formation of Kyrgyzstan"
6226:
6155:
6100:
5721:
5536:
5354:
5340:
5325:
5277:
5211:
5137:
4959:
4952:
4938:
4844:
4777:
4705:
4669:
4622:
4564:
4550:
4517:
4509:
4502:
4483:
4306:
4271:
4059:
3660:
3451:
3401:
3338:
3309:
3264:
3242:
3235:
3227:
3151:
3047:
3006:
2999:
2991:
2975:
2895:
2766:
2642:
2495:
2358:
2217:
1039:
1015:
866:
817:
762:
725:
530:
426:
197:
89:
54:
6389:
2109:
2021:
1941:
1900:
1578:
6247:
6240:
5940:
5933:
5606:
5545:
5199:
4917:
4463:
4367:
4102:
3687:
3572:
3559:
3486:
3422:
3345:
3324:
3081:
2944:
2752:
2739:
2657:
2509:
2328:
2315:
2278:
2187:
1932:
1915:
1822:, a tyrannosauroid theropod from the Wealden Supergroup of southern England"
1621:
1444:
1390:
449:
345:
331:
210:
151:
94:
38:
30:
1857:
1594:
The
Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals
1343:
showing hypothetical complete crest-shape, similar to that of the related
6383:
5831:
5256:
5249:
5221:
5114:
4614:
3995:
3331:
3158:
2919:
2556:
2235:
2211:
1345:
923:
783:
737:
703:
441:
337:
334:
184:
171:
126:
84:
79:
64:
59:
49:
1838:
6416:
5389:
5151:
4881:
4784:
4109:
3430:
503:
349:
99:
74:
2100:
2083:
1501:"Cranial osteology and phylogenetic position of the theropod dinosaur
1419:. A diverse microvertebrate fauna is known from the equivalently aged
6040:
5728:
3172:
2205:
1722:
1558:
460:
the skull so that the left side was exposed, and submitted it to the
402:
161:
6360:
1671:
596:, and Huene thought both dinosaurs represented members of the group
4888:
4629:
4571:
1499:
Rauhut, Oliver W. M.; Milner, Angela C.; Moore-Fay, Scott (2010).
1335:
776:
768:
708:
633:
to be neither ceratosaurs nor coelurosaurs, but instead primitive
621:, again mainly due to the crest on the nose (though the idea that
564:
544:
475:
430:
420:
327:
1328:) outside Tyrannosauroidea based on their phylogenetic analysis.
6403:
6304:
4931:
6364:
5802:
5048:
4242:
3557:
2737:
2313:
2198:
2160:
625:
bore a nasal crest was later disproved). Paul considered both
569:
Mount with hypothetical crest-shape and postcranial skeleton,
1916:"The Middle Jurassic flora from Stonesfield, Oxfordshire, UK"
1505:
bradleyi(Woodward, 1910) from the Middle
Jurassic of England"
2156:
1395:
representing a probably seasonally dry coastal environment.
2014:
10.1671/0272-4634(2003)23[208:TAATOC]2.0.CO;2
1359:
used a puncture-pull strategy for hunting prey. However,
366:, due to the partially preserved portion of the crest of
637:. Furthermore, Paul considered the much larger dinosaur
701:
was most closely related to the
Chinese tyrannosauroid
502:
that had eroded the rock and was partially filled with
1745:
Monographien zur
Geologie und Palaeontologie (Serie 1)
382:, and amongst the earliest known members of the clade
517:, assuming it was the ancestor of the Jurassic genus
429:
of the holotype skull and close ups of its teeth and
356:, known from a mostly complete skull and lower jaws.
6373:
6286:
6181:
6124:
6086:
6003:
5976:
5886:
5856:
5815:
5713:
5681:
5623:
5589:
5562:
5535:
5504:
5477:
5466:
5426:
5291:
5239:
5210:
5172:
5161:
5061:
4973:
4907:
4829:
4760:
4736:
4648:
4596:
4536:
4492:
4481:
4425:
4393:
4340:
4290:
4255:
4173:
4126:
4073:
4035:
3981:
3932:
3868:
3827:
3803:
3772:
3726:
3696:
3685:
3570:
3485:
3384:
3295:
3274:
3210:
3113:
3057:
3023:
2930:
2875:
2844:
2784:
2750:
2676:
2656:
2580:
2527:
2408:
2326:
2277:
2250:
1762:
1711:Geological Society, London, Special Publications
1494:
1492:
1490:
1488:
1486:
1484:
1482:
1480:
658:studies in the early 21st century finally found
1665:
1663:
1622:"A new phylogeny of the carnivorous dinosaurs"
1552:
1550:
2172:
1372:The flora from the roughly equivalently aged
576:Arthur Smith Woodward, who initially studied
8:
370:superficially resembling the small crest of
1996:Upchurch, Paul; Martin, John (2003-04-11).
1676:Quarterly Journal of the Geological Society
1563:. Princeton University Press. p. 105.
1534:
1532:
6361:
6092:
6009:
5984:
5868:
5864:
5823:
5812:
5799:
5710:
5678:
5510:
5474:
5434:
5303:
5299:
5184:
5180:
5169:
5069:
5058:
5045:
4836:
4826:
4742:
4654:
4542:
4489:
4399:
4352:
4348:
4298:
4263:
4252:
4239:
4170:
4079:
3987:
3940:
3809:
3784:
3780:
3708:
3704:
3693:
3578:
3567:
3554:
3482:
3301:
3292:
3121:
2936:
2887:
2883:
2796:
2792:
2758:
2747:
2734:
2533:
2416:
2334:
2323:
2310:
2256:
2247:
2195:
2179:
2165:
2157:
2152:3D scan of the holotype skull on Sketchfab
2049:Proceedings of the Geologists' Association
1596:. London: Marshall Editions. p. 114.
756:Below is a cladogram from a 2022 study by
693:was in fact a coelurosaur, and moreover a
615:suggested that it was a close relative of
117:
20:
2099:
1980:
1969:Zoological Journal of the Linnean Society
1931:
1890:
1878:Zoological Journal of the Linnean Society
1847:
1837:
1769:. New York: Simon and Schuster. pp.
1520:
1509:Zoological Journal of the Linnean Society
1423:, including frogs, salamanders, turtles,
1914:Cleal, C. J.; Rees, P. M. (July 2003).
1476:
603:It was not until the late 1980s, after
494:. Currently the skull is housed at the
1560:The Princeton Field Guide to Dinosaurs
378:, specifically a member of the family
715:, defined as all theropods closer to
590:suggested a closer relationship with
7:
549:Estimated size compared to a human.
509:In 1923, the German palaeontologist
436:In 1910, the British paleontologist
6463:Middle Jurassic dinosaurs of Europe
1795:"Oldest T. rex relative identified"
1647:"Oldest T. rex relative identified"
1156:
1149:
1125:
1101:
1079:
1055:
1031:
1007:
939:
915:
891:
881:
859:
809:
802:
790:
711:containing these two dinosaurs the
513:moved the species to the new genus
374:. Now, however, it is considered a
2002:Journal of Vertebrate Paleontology
1963:Benson, Roger B. J. (April 2010).
413:while excavating for a reservoir.
14:
6483:Taxa named by Friedrich von Huene
1818:"The osteology and affinities of
1688:10.1144/GSL.JGS.1910.066.01-04.07
685:The first major re-evaluation of
6345:
6339:
6332:
5784:
5777:
5770:
5764:
5030:
5023:
5018:
5011:
5005:
4224:
4217:
4210:
3539:
3532:
3525:
3518:
2720:
2714:
1982:10.1111/j.1096-3642.2009.00569.x
1816:Naish, D.; Cau, A. (July 2022).
1765:Predatory Dinosaurs of the World
1522:10.1111/j.1096-3642.2009.00591.x
1465:Timeline of tyrannosaur research
352:. It contains a single species.
138:
1376:in Oxfordshire is dominated by
397:in London and was described in
464:. Woodward made the skull the
1:
6478:Fossil taxa described in 1926
2124:"Kirtlington 3p (Mammal Bed)"
2061:10.1016/j.pgeola.2019.02.003
1538:Holtz, Thomas R. Jr. (2008)
760:and Andrea Cau on the genus
580:, placed it as a species of
462:Geological Society of London
1374:Taynton Limestone Formation
1310:In 2024, the describers of
6499:
1892:10.1093/zoolinnean/zlae090
16:Extinct genus of dinosaurs
6329:
6095:
6012:
5987:
5871:
5867:
5826:
5811:
5798:
5762:
5709:
5677:
5513:
5437:
5306:
5302:
5187:
5183:
5072:
5057:
5044:
5003:
4839:
4825:
4745:
4657:
4545:
4402:
4355:
4351:
4301:
4266:
4251:
4238:
4207:
4169:
4082:
3990:
3943:
3812:
3787:
3783:
3711:
3707:
3581:
3566:
3553:
3515:
3481:
3304:
3124:
2939:
2890:
2886:
2799:
2795:
2761:
2746:
2733:
2711:
2536:
2419:
2337:
2322:
2309:
2259:
2246:
2194:
1557:Paul, Gregory S. (2016).
1543:Supplementary Information
1411:as well as indeterminate
1204:
1178:
1161:
1154:
1147:
1130:
1123:
1106:
1099:
1084:
1077:
1060:
1053:
1036:
1029:
1012:
1005:
961:
944:
937:
920:
913:
896:
889:
879:
864:
857:
831:
814:
807:
800:
302:
295:
273:
266:
135:Scientific classification
133:
125:
116:
23:
1670:Woodward, A. S. (1910).
1592:Palmer, D., ed. (1999).
773:Skull seen from the left
643:to be the same genus as
280:Proceratosaurus bradleyi
2088:Papers in Palaeontology
1933:10.1111/1475-4983.00319
1431:, crocodyliformes, and
1421:Forest Marble Formation
611:was again re-examined.
456:, England. Bradley had
1649:. BBC News. 2009-11-04
1349:
1320:and proceratosaurids (
787:
774:
573:
550:
496:Natural History Museum
433:
395:Natural History Museum
6425:Paleobiology Database
4738:Carcharodontosaurinae
4650:Carcharodontosauridae
2128:Paleobiology Database
1339:
780:
772:
568:
548:
438:Arthur Smith Woodward
424:
5714:Dubious coelurosaurs
4427:Metriacanthosaurinae
4395:Metriacanthosauridae
2677:Dubious neotheropods
1384:conifers as well as
674:in 2004 also placed
468:of a new species of
417:History of discovery
4751:Carcharodontosaurus
4538:Carcharodontosauria
1839:10.7717/peerj.12727
1761:Paul, G.S. (1988).
672:Thomas R. Holtz Jr.
588:Friedrich von Huene
535:University of Texas
511:Friedrich von Huene
440:reported a partial
6473:Fossils of England
6206:Archaeornithomimus
4727:Veterupristisaurus
4587:Veterupristisaurus
4443:Metriacanthosaurus
4383:Piatnitzkysauridae
3818:Eustreptospondylus
3728:Piatnitzkysauridae
3611:Chuandongocoelurus
2475:Notatesseraeraptor
1350:
788:
775:
574:
551:
434:
407:Great Oolite Group
6440:
6439:
6367:Taxon identifiers
6358:
6357:
6354:
6353:
6327:
6326:
6323:
6322:
6319:
6318:
6315:
6314:
6282:
6281:
6278:
6277:
6088:Macrocheiriformes
6005:Ornithomimosauria
5979:Maniraptoriformes
5972:
5971:
5818:Maniraptoromorpha
5805:Maniraptoromorpha
5794:
5793:
5760:
5759:
5756:
5755:
5752:
5751:
5705:
5704:
5701:
5700:
5683:Maniraptoromorpha
5673:
5672:
5669:
5668:
5665:
5664:
5661:
5660:
5657:
5656:
5422:
5421:
5287:
5286:
5241:Proceratosauridae
5193:Chingkankousaurus
5040:
5039:
5001:
5000:
4997:
4996:
4993:
4992:
4969:
4968:
4821:
4820:
4817:
4816:
4813:
4812:
4809:
4808:
4805:
4804:
4801:
4800:
4477:
4476:
4473:
4472:
4234:
4233:
4205:
4204:
4201:
4200:
4197:
4196:
4193:
4192:
4165:
4164:
4161:
4160:
4157:
4156:
4153:
4152:
4069:
4068:
4025:Vallibonavenatrix
3928:
3927:
3924:
3923:
3768:
3767:
3549:
3548:
3513:
3512:
3509:
3508:
3505:
3504:
3477:
3476:
3473:
3472:
3469:
3468:
3465:
3464:
3461:
3460:
3109:
3108:
3105:
3104:
2871:
2870:
2729:
2728:
2709:
2708:
2705:
2704:
2701:
2700:
2652:
2651:
2305:
2304:
2301:
2300:
2297:
2296:
2101:10.1002/spp2.1487
1780:978-0-671-61946-6
1620:(December 1998).
1603:978-1-84028-152-1
1570:978-1-78684-190-2
1429:rhynchocephalians
1382:cheirolepidiacean
1307:
1306:
1298:
1297:
1289:
1288:
1280:
1279:
1271:
1270:
1262:
1261:
1253:
1252:
1244:
1243:
1235:
1234:
1226:
1225:
1217:
1216:
1193:
1192:
994:
993:
985:
984:
976:
975:
886:Proceratosauridae
846:
845:
713:Proceratosauridae
707:. They named the
466:holotype specimen
405:limestone of the
380:Proceratosauridae
319:
318:
313:
247:
224:Proceratosauridae
6490:
6468:Jurassic England
6453:Proceratosaurids
6433:
6432:
6420:
6419:
6407:
6406:
6394:
6393:
6392:
6362:
6349:
6343:
6336:
6192:Aepyornithomimus
6093:
6010:
5985:
5949:Sinocalliopteryx
5869:
5865:
5859:Neocoelurosauria
5824:
5813:
5800:
5788:
5781:
5774:
5768:
5711:
5679:
5648:Zhuchengtyrannus
5580:Thanatotheristes
5511:
5475:
5443:Appalachiosaurus
5435:
5304:
5300:
5294:Pantyrannosauria
5185:
5181:
5175:Tyrannosauroidea
5170:
5145:Xinjiangovenator
5070:
5059:
5046:
5034:
5027:
5022:
5015:
5009:
4837:
4827:
4743:
4663:Acrocanthosaurus
4655:
4543:
4490:
4416:Yangchuanosaurus
4400:
4353:
4349:
4299:
4280:Lourinhanosaurus
4264:
4253:
4240:
4228:
4221:
4214:
4171:
4137:Sigilmassasaurus
4080:
4037:Ceratosuchopsini
3988:
3941:
3886:Dubreuillosaurus
3810:
3793:Streptospondylus
3785:
3781:
3751:Piatnitzkysaurus
3709:
3705:
3694:
3579:
3568:
3555:
3543:
3536:
3529:
3522:
3483:
3318:Ekrixinatosaurus
3302:
3293:
3122:
2937:
2888:
2884:
2797:
2793:
2759:
2748:
2735:
2724:
2718:
2628:Procompsognathus
2534:
2417:
2335:
2324:
2311:
2257:
2248:
2241:
2240:
2196:
2181:
2174:
2167:
2158:
2139:
2138:
2136:
2134:
2120:
2114:
2113:
2103:
2079:
2073:
2072:
2040:
2034:
2033:
1993:
1987:
1986:
1984:
1960:
1954:
1953:
1935:
1911:
1905:
1904:
1894:
1868:
1862:
1861:
1851:
1841:
1820:Eotyrannus lengi
1813:
1807:
1806:
1804:
1802:
1791:
1785:
1784:
1768:
1758:
1752:
1741:
1735:
1734:
1723:10.1144/SP430.13
1706:
1700:
1699:
1682:(1–4): 111–115.
1667:
1658:
1657:
1655:
1654:
1643:
1637:
1636:
1626:
1614:
1608:
1607:
1589:
1583:
1582:
1554:
1545:
1536:
1527:
1526:
1524:
1496:
1368:Paleoenvironment
1341:Life restoration
1157:
1150:
1134:Appalachiosaurus
1126:
1102:
1080:
1056:
1032:
1008:
940:
916:
892:
882:
860:
810:
803:
796:Tyrannosauroidea
791:
571:Worcester Museum
478:named in 1824),
448:, a town in the
384:Tyrannosauroidea
311:
282:
278:
259:P. bradleyi
242:
235:
222:
143:
142:
121:
111:
41:
29:Temporal range:
21:
6498:
6497:
6493:
6492:
6491:
6489:
6488:
6487:
6443:
6442:
6441:
6436:
6428:
6423:
6415:
6410:
6402:
6397:
6388:
6387:
6382:
6375:Proceratosaurus
6369:
6359:
6350:
6337:
6311:
6274:
6255:Sinornithomimus
6177:
6120:
6082:
5999:
5993:Compsognathidae
5981:
5968:
5956:Sinosauropteryx
5888:Compsognathidae
5882:
5861:
5852:
5820:
5807:
5790:
5789:
5775:
5748:
5743:Shanyangosaurus
5736:Phaedrolosaurus
5697:
5653:
5619:
5585:
5564:Daspletosaurini
5558:
5531:
5506:Tyrannosaurinae
5500:
5479:Albertosaurinae
5471:
5469:Tyrannosauridae
5462:
5431:
5429:Eutyrannosauria
5418:
5296:
5283:
5264:Proceratosaurus
5235:
5206:
5177:
5166:
5157:
5123:Richardoestesia
5107:Chilantaisaurus
5066:
5053:
5036:
5035:
5016:
4989:
4965:
4925:Australovenator
4903:
4897:Chilantaisaurus
4875:Phuwiangvenator
4797:
4762:Giganotosaurini
4756:
4732:
4644:
4607:Chilantaisaurus
4592:
4532:
4486:
4469:
4421:
4389:
4345:
4336:
4314:Monolophosaurus
4295:
4286:
4260:
4247:
4230:
4229:
4215:
4189:
4149:
4122:
4088:Camarillasaurus
4065:
4031:
3977:
3971:Ostafrikasaurus
3949:Camarillasaurus
3937:
3920:
3870:Afrovenatorinae
3864:
3823:
3799:
3777:
3764:
3722:
3701:
3699:Megalosauroidea
3690:
3681:
3676:Vectaerovenator
3669:Szechuanosaurus
3647:Monolophosaurus
3633:Kaijiangosaurus
3626:Cryolophosaurus
3587:Calamospondylus
3575:
3562:
3545:
3544:
3537:
3530:
3523:
3501:
3457:
3438:Pycnonemosaurus
3380:
3291:
3285:Majungasaurinae
3270:
3212:Majungasaurinae
3206:
3201:Xenotarsosaurus
3137:Dryptosauroides
3118:
3101:
3053:
3025:Elaphrosaurinae
3019:
2926:
2912:Ornithomimoides
2880:
2867:
2840:
2789:
2780:
2755:
2742:
2725:
2719:
2697:
2672:
2648:
2576:
2523:
2489:Shuangbaisaurus
2413:
2404:
2331:
2318:
2293:
2273:
2267:Avemetatarsalia
2252:Avemetatarsalia
2242:
2227:Avemetatarsalia
2201:
2200:
2190:
2185:
2148:
2143:
2142:
2132:
2130:
2122:
2121:
2117:
2081:
2080:
2076:
2042:
2041:
2037:
1995:
1994:
1990:
1962:
1961:
1957:
1913:
1912:
1908:
1870:
1869:
1865:
1815:
1814:
1810:
1800:
1798:
1793:
1792:
1788:
1781:
1760:
1759:
1755:
1742:
1738:
1708:
1707:
1703:
1669:
1668:
1661:
1652:
1650:
1645:
1644:
1640:
1624:
1616:
1615:
1611:
1604:
1591:
1590:
1586:
1571:
1556:
1555:
1548:
1537:
1530:
1503:Proceratosaurus
1498:
1497:
1478:
1473:
1461:
1404:, the sauropod
1386:bennettitaleans
1370:
1361:Proceratosaurus
1357:Proceratosaurus
1353:Proceratosaurus
1334:
1322:Proceratosaurus
1308:
1299:
1290:
1281:
1272:
1263:
1254:
1245:
1236:
1227:
1218:
1207:Tyrannosauridae
1194:
995:
986:
977:
948:Proceratosaurus
847:
717:Proceratosaurus
699:Proceratosaurus
691:Proceratosaurus
687:Proceratosaurus
676:Proceratosaurus
668:Proceratosaurus
660:Proceratosaurus
645:Proceratosaurus
627:Proceratosaurus
613:Gregory S. Paul
609:Proceratosaurus
578:Proceratosaurus
563:
554:Proceratosaurus
543:
515:Proceratosaurus
454:Gloucestershire
419:
393:is held in the
368:Proceratosaurus
358:Proceratosaurus
342:Middle Jurassic
323:Proceratosaurus
312:Woodward, 1910
291:
284:
276:
275:
262:
241:
238:Proceratosaurus
233:
220:
137:
112:
110:
109:
108:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
36:
35:
34:
27:
25:Proceratosaurus
17:
12:
11:
5:
6496:
6494:
6486:
6485:
6480:
6475:
6470:
6465:
6460:
6458:Bathonian life
6455:
6445:
6444:
6438:
6437:
6435:
6434:
6421:
6408:
6395:
6379:
6377:
6371:
6370:
6365:
6356:
6355:
6352:
6351:
6331:
6330:
6328:
6325:
6324:
6321:
6320:
6317:
6316:
6313:
6312:
6310:
6309:
6308:
6307:
6301:
6292:
6290:
6284:
6283:
6280:
6279:
6276:
6275:
6273:
6272:
6265:
6258:
6251:
6244:
6237:
6230:
6223:
6220:Dromiceiomimus
6216:
6209:
6202:
6195:
6187:
6185:
6183:Ornithomimidae
6179:
6178:
6176:
6175:
6168:
6164:Paraxenisaurus
6160:
6152:
6145:
6138:
6130:
6128:
6126:Deinocheiridae
6122:
6121:
6119:
6118:
6115:Shenzhousaurus
6111:
6104:
6096:
6090:
6084:
6083:
6081:
6080:
6073:
6065:
6058:
6051:
6044:
6037:
6029:
6021:
6013:
6007:
6001:
6000:
5998:
5997:
5988:
5982:
5977:
5974:
5973:
5970:
5969:
5967:
5966:
5959:
5952:
5945:
5937:
5930:
5923:
5916:
5912:Beipiaognathus
5908:
5901:
5892:
5890:
5884:
5883:
5881:
5880:
5872:
5862:
5857:
5854:
5853:
5851:
5850:
5843:
5836:
5827:
5821:
5816:
5809:
5808:
5803:
5796:
5795:
5792:
5791:
5783:
5769:
5763:
5761:
5758:
5757:
5754:
5753:
5750:
5749:
5747:
5746:
5739:
5732:
5725:
5717:
5715:
5707:
5706:
5703:
5702:
5699:
5698:
5696:
5695:
5694:
5693:
5687:
5685:
5675:
5674:
5671:
5670:
5667:
5666:
5663:
5662:
5659:
5658:
5655:
5654:
5652:
5651:
5644:
5637:
5629:
5627:
5625:Tyrannosaurini
5621:
5620:
5618:
5617:
5610:
5603:
5595:
5593:
5591:Teratophoneini
5587:
5586:
5584:
5583:
5576:
5573:Daspletosaurus
5568:
5566:
5560:
5559:
5557:
5556:
5553:Qianzhousaurus
5549:
5541:
5539:
5533:
5532:
5530:
5529:
5522:
5514:
5508:
5502:
5501:
5499:
5498:
5491:
5483:
5481:
5472:
5467:
5464:
5463:
5461:
5460:
5453:
5450:Bistahieversor
5446:
5438:
5432:
5427:
5424:
5423:
5420:
5419:
5417:
5416:
5408:
5401:
5394:
5386:
5379:
5372:
5365:
5358:
5351:
5344:
5337:
5330:
5322:
5315:
5307:
5297:
5292:
5289:
5288:
5285:
5284:
5282:
5281:
5274:
5267:
5260:
5253:
5245:
5243:
5237:
5236:
5234:
5233:
5225:
5217:
5215:
5208:
5207:
5205:
5204:
5196:
5188:
5178:
5173:
5167:
5164:Tyrannoraptora
5162:
5159:
5158:
5156:
5155:
5148:
5141:
5134:
5126:
5119:
5111:
5103:
5096:
5088:
5081:
5073:
5067:
5062:
5055:
5054:
5049:
5042:
5041:
5038:
5037:
5029:
5010:
5004:
5002:
4999:
4998:
4995:
4994:
4991:
4990:
4988:
4987:
4986:
4985:
4979:
4977:
4971:
4970:
4967:
4966:
4964:
4963:
4956:
4949:
4942:
4935:
4928:
4921:
4913:
4911:
4909:Megaraptoridae
4905:
4904:
4902:
4901:
4893:
4885:
4878:
4871:
4864:
4856:
4848:
4840:
4834:
4823:
4822:
4819:
4818:
4815:
4814:
4811:
4810:
4807:
4806:
4803:
4802:
4799:
4798:
4796:
4795:
4788:
4781:
4774:
4771:Giganotosaurus
4766:
4764:
4758:
4757:
4755:
4754:
4746:
4740:
4734:
4733:
4731:
4730:
4723:
4716:
4709:
4702:
4695:
4688:
4681:
4674:
4666:
4658:
4652:
4646:
4645:
4643:
4642:
4634:
4626:
4619:
4611:
4602:
4600:
4598:Neovenatoridae
4594:
4593:
4591:
4590:
4583:
4580:Ulughbegsaurus
4576:
4568:
4561:
4554:
4546:
4540:
4534:
4533:
4531:
4530:
4522:
4514:
4506:
4498:
4496:
4487:
4482:
4479:
4478:
4475:
4474:
4471:
4470:
4468:
4467:
4460:
4453:
4446:
4439:
4431:
4429:
4423:
4422:
4420:
4419:
4412:
4408:Xuanhanosaurus
4403:
4397:
4391:
4390:
4388:
4387:
4379:
4375:Xuanhanosaurus
4371:
4364:
4361:Asfaltovenator
4356:
4346:
4341:
4338:
4337:
4335:
4334:
4326:
4322:Megalosauridae
4318:
4310:
4302:
4296:
4291:
4288:
4287:
4285:
4284:
4276:
4267:
4261:
4256:
4249:
4248:
4243:
4236:
4235:
4232:
4231:
4223:
4209:
4208:
4206:
4203:
4202:
4199:
4198:
4195:
4194:
4191:
4190:
4188:
4187:
4186:
4185:
4179:
4177:
4167:
4166:
4163:
4162:
4159:
4158:
4155:
4154:
4151:
4150:
4148:
4147:
4140:
4132:
4130:
4124:
4123:
4121:
4120:
4113:
4106:
4099:
4096:Ichthyovenator
4092:
4083:
4077:
4071:
4070:
4067:
4066:
4064:
4063:
4056:
4049:
4041:
4039:
4033:
4032:
4030:
4029:
4021:
4013:
4006:
3999:
3991:
3985:
3979:
3978:
3976:
3975:
3967:
3960:
3957:Cristatusaurus
3953:
3944:
3938:
3933:
3930:
3929:
3926:
3925:
3922:
3921:
3919:
3918:
3914:Poekilopleuron
3910:
3907:Piveteausaurus
3903:
3896:
3889:
3882:
3874:
3872:
3866:
3865:
3863:
3862:
3855:
3848:
3841:
3833:
3831:
3829:Megalosaurinae
3825:
3824:
3822:
3821:
3813:
3807:
3805:Megalosauridae
3801:
3800:
3798:
3797:
3788:
3778:
3773:
3770:
3769:
3766:
3765:
3763:
3762:
3758:Xuanhanosaurus
3754:
3747:
3740:
3732:
3730:
3724:
3723:
3721:
3720:
3717:Yunyangosaurus
3712:
3702:
3697:
3691:
3686:
3683:
3682:
3680:
3679:
3672:
3665:
3657:
3654:Pandoravenator
3650:
3643:
3640:Kayentavenator
3636:
3629:
3622:
3614:
3607:
3599:
3591:
3582:
3576:
3571:
3564:
3563:
3558:
3551:
3550:
3547:
3546:
3538:
3531:
3524:
3517:
3516:
3514:
3511:
3510:
3507:
3506:
3503:
3502:
3500:
3499:
3498:
3497:
3491:
3489:
3479:
3478:
3475:
3474:
3471:
3470:
3467:
3466:
3463:
3462:
3459:
3458:
3456:
3455:
3448:
3441:
3434:
3427:
3419:
3412:
3405:
3398:
3390:
3388:
3382:
3381:
3379:
3378:
3371:
3368:Skorpiovenator
3364:
3356:
3349:
3342:
3335:
3328:
3321:
3314:
3305:
3299:
3290:
3289:
3280:
3278:
3272:
3271:
3269:
3268:
3261:
3253:
3246:
3239:
3232:
3224:
3216:
3214:
3208:
3207:
3205:
3204:
3197:
3190:
3183:
3180:Spectrovenator
3176:
3169:
3162:
3155:
3148:
3141:
3133:
3125:
3119:
3114:
3111:
3110:
3107:
3106:
3103:
3102:
3100:
3099:
3092:
3085:
3078:
3071:
3063:
3061:
3055:
3054:
3052:
3051:
3044:
3037:
3029:
3027:
3021:
3020:
3018:
3017:
3014:Spinostropheus
3010:
3003:
2996:
2988:
2980:
2972:
2964:
2957:
2949:
2940:
2934:
2928:
2927:
2925:
2924:
2916:
2908:
2900:
2891:
2881:
2878:Abelisauroidea
2876:
2873:
2872:
2869:
2868:
2866:
2865:
2858:
2850:
2848:
2846:Ceratosauridae
2842:
2841:
2839:
2838:
2835:Saltriovenator
2831:
2823:
2816:
2812:Fosterovenator
2808:
2800:
2790:
2785:
2782:
2781:
2779:
2778:
2774:Lophostropheus
2770:
2762:
2756:
2751:
2744:
2743:
2738:
2731:
2730:
2727:
2726:
2713:
2712:
2710:
2707:
2706:
2703:
2702:
2699:
2698:
2696:
2695:
2688:
2680:
2678:
2674:
2673:
2671:
2670:
2669:
2668:
2662:
2660:
2654:
2653:
2650:
2649:
2647:
2646:
2639:
2635:Pterospondylus
2631:
2624:
2616:
2608:
2605:Lucianovenator
2601:
2594:
2586:
2584:
2578:
2577:
2575:
2574:
2567:
2560:
2553:
2546:
2537:
2531:
2529:Coelophysoidea
2525:
2524:
2522:
2521:
2514:
2506:
2499:
2492:
2485:
2478:
2471:
2468:Lophostropheus
2464:
2457:
2450:
2443:
2436:
2428:
2420:
2414:
2409:
2406:
2405:
2403:
2402:
2394:
2386:
2378:
2370:
2367:Erythrovenator
2363:
2355:
2347:
2338:
2332:
2327:
2320:
2319:
2314:
2307:
2306:
2303:
2302:
2299:
2298:
2295:
2294:
2292:
2291:
2290:
2289:
2283:
2281:
2275:
2274:
2272:
2271:
2270:
2269:
2260:
2254:
2244:
2243:
2239:
2238:
2229:
2220:
2214:
2208:
2199:
2192:
2191:
2186:
2184:
2183:
2176:
2169:
2161:
2155:
2154:
2147:
2146:External links
2144:
2141:
2140:
2115:
2074:
2055:(2): 170–186.
2035:
2008:(1): 208–231.
1988:
1975:(4): 882–935.
1955:
1926:(4): 739–801.
1906:
1863:
1808:
1786:
1779:
1753:
1736:
1717:(1): 289–309.
1701:
1659:
1638:
1609:
1602:
1584:
1569:
1546:
1528:
1475:
1474:
1472:
1469:
1468:
1467:
1460:
1457:
1453:eutriconodonts
1441:morganucodonts
1437:tritylodontids
1433:mammaliamorphs
1413:ornithischians
1369:
1366:
1333:
1330:
1305:
1304:
1301:
1300:
1296:
1295:
1292:
1291:
1287:
1286:
1283:
1282:
1278:
1277:
1274:
1273:
1269:
1268:
1265:
1264:
1260:
1259:
1256:
1255:
1251:
1250:
1247:
1246:
1242:
1241:
1238:
1237:
1233:
1232:
1229:
1228:
1224:
1223:
1220:
1219:
1215:
1214:
1211:
1210:
1203:
1200:
1199:
1196:
1195:
1191:
1190:
1187:
1186:
1177:
1174:
1173:
1170:
1169:
1165:Bistahieversor
1160:
1155:
1153:
1148:
1146:
1143:
1142:
1139:
1138:
1129:
1124:
1122:
1119:
1118:
1115:
1114:
1105:
1100:
1098:
1095:
1094:
1091:
1090:
1083:
1078:
1076:
1073:
1072:
1069:
1068:
1059:
1054:
1052:
1049:
1048:
1045:
1044:
1035:
1030:
1028:
1025:
1024:
1021:
1020:
1011:
1006:
1004:
1001:
1000:
997:
996:
992:
991:
988:
987:
983:
982:
979:
978:
974:
973:
970:
969:
960:
957:
956:
953:
952:
943:
938:
936:
933:
932:
929:
928:
919:
914:
912:
909:
908:
905:
904:
895:
890:
888:
880:
878:
875:
874:
871:
870:
863:
858:
856:
853:
852:
849:
848:
844:
843:
840:
839:
830:
827:
826:
823:
822:
813:
808:
806:
801:
799:
789:
695:tyrannosauroid
649:Piveteausaurus
640:Piveteausaurus
562:
561:Classification
559:
542:
539:
446:Minchinhampton
418:
415:
411:Minchinhampton
317:
316:
315:
314:
300:
299:
293:
292:
285:
271:
270:
264:
263:
255:
253:
249:
248:
231:
227:
226:
218:
214:
213:
208:
201:
200:
195:
188:
187:
182:
175:
174:
169:
165:
164:
159:
155:
154:
149:
145:
144:
131:
130:
123:
122:
114:
113:
105:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
42:
28:
15:
13:
10:
9:
6:
4:
3:
2:
6495:
6484:
6481:
6479:
6476:
6474:
6471:
6469:
6466:
6464:
6461:
6459:
6456:
6454:
6451:
6450:
6448:
6431:
6426:
6422:
6418:
6413:
6409:
6405:
6400:
6396:
6391:
6385:
6381:
6380:
6378:
6376:
6372:
6368:
6363:
6348:
6344:
6342:
6335:
6306:
6302:
6300:
6296:
6295:
6294:
6293:
6291:
6289:
6285:
6271:
6270:
6266:
6264:
6263:
6262:Struthiomimus
6259:
6257:
6256:
6252:
6250:
6249:
6245:
6243:
6242:
6238:
6236:
6235:
6231:
6229:
6228:
6224:
6222:
6221:
6217:
6215:
6214:
6213:"Coelosaurus"
6210:
6208:
6207:
6203:
6201:
6200:
6196:
6194:
6193:
6189:
6188:
6186:
6184:
6180:
6174:
6173:
6169:
6166:
6165:
6161:
6158:
6157:
6153:
6151:
6150:
6146:
6144:
6143:
6139:
6137:
6136:
6132:
6131:
6129:
6127:
6123:
6117:
6116:
6112:
6110:
6109:
6108:Pelecanimimus
6105:
6103:
6102:
6098:
6097:
6094:
6091:
6089:
6085:
6079:
6078:
6074:
6071:
6070:
6069:Thecocoelurus
6066:
6064:
6063:
6059:
6057:
6056:
6052:
6050:
6049:
6048:Kinnareemimus
6045:
6043:
6042:
6038:
6035:
6034:
6030:
6027:
6026:
6022:
6020:
6019:
6015:
6014:
6011:
6008:
6006:
6002:
5995:
5994:
5990:
5989:
5986:
5983:
5980:
5975:
5965:
5964:
5960:
5958:
5957:
5953:
5951:
5950:
5946:
5943:
5942:
5938:
5936:
5935:
5931:
5929:
5928:
5927:Huaxiagnathus
5924:
5922:
5921:
5920:Compsognathus
5917:
5914:
5913:
5909:
5907:
5906:
5902:
5899:
5898:
5894:
5893:
5891:
5889:
5885:
5879:
5878:
5874:
5873:
5870:
5866:
5863:
5860:
5855:
5849:
5848:
5847:Ornitholestes
5844:
5842:
5841:
5837:
5834:
5833:
5829:
5828:
5825:
5822:
5819:
5814:
5810:
5806:
5801:
5797:
5787:
5782:
5780:
5773:
5767:
5745:
5744:
5740:
5738:
5737:
5733:
5731:
5730:
5726:
5724:
5723:
5719:
5718:
5716:
5712:
5708:
5691:
5690:
5689:
5688:
5686:
5684:
5680:
5676:
5650:
5649:
5645:
5643:
5642:
5641:Tyrannosaurus
5638:
5636:
5635:
5631:
5630:
5628:
5626:
5622:
5616:
5615:
5614:Teratophoneus
5611:
5609:
5608:
5604:
5602:
5601:
5597:
5596:
5594:
5592:
5588:
5582:
5581:
5577:
5575:
5574:
5570:
5569:
5567:
5565:
5561:
5555:
5554:
5550:
5548:
5547:
5543:
5542:
5540:
5538:
5534:
5528:
5527:
5523:
5521:
5520:
5516:
5515:
5512:
5509:
5507:
5503:
5497:
5496:
5492:
5490:
5489:
5488:Albertosaurus
5485:
5484:
5482:
5480:
5476:
5473:
5470:
5465:
5459:
5458:
5454:
5452:
5451:
5447:
5445:
5444:
5440:
5439:
5436:
5433:
5430:
5425:
5414:
5413:
5409:
5407:
5406:
5405:Xiongguanlong
5402:
5400:
5399:
5395:
5392:
5391:
5387:
5385:
5384:
5383:Suskityrannus
5380:
5378:
5377:
5376:Stokesosaurus
5373:
5371:
5370:
5369:Santanaraptor
5366:
5364:
5363:
5359:
5357:
5356:
5352:
5350:
5349:
5345:
5343:
5342:
5338:
5336:
5335:
5331:
5328:
5327:
5323:
5321:
5320:
5316:
5314:
5313:
5312:Alectrosaurus
5309:
5308:
5305:
5301:
5298:
5295:
5290:
5280:
5279:
5275:
5273:
5272:
5268:
5266:
5265:
5261:
5259:
5258:
5254:
5252:
5251:
5247:
5246:
5244:
5242:
5238:
5231:
5230:
5229:Tanycolagreus
5226:
5224:
5223:
5219:
5218:
5216:
5213:
5209:
5202:
5201:
5197:
5195:
5194:
5190:
5189:
5186:
5182:
5179:
5176:
5171:
5168:
5165:
5160:
5154:
5153:
5149:
5147:
5146:
5142:
5140:
5139:
5135:
5132:
5131:
5127:
5125:
5124:
5120:
5117:
5116:
5112:
5109:
5108:
5104:
5102:
5101:
5097:
5094:
5093:
5092:Bahariasaurus
5089:
5087:
5086:
5082:
5080:
5079:
5075:
5074:
5071:
5068:
5065:
5064:Coelurosauria
5060:
5056:
5052:
5051:Coelurosauria
5047:
5043:
5033:
5028:
5026:
5021:
5014:
5008:
4983:
4982:
4981:
4980:
4978:
4976:
4975:Coelurosauria
4972:
4962:
4961:
4957:
4955:
4954:
4950:
4948:
4947:
4943:
4941:
4940:
4936:
4934:
4933:
4929:
4927:
4926:
4922:
4920:
4919:
4915:
4914:
4912:
4910:
4906:
4900:
4898:
4894:
4892:
4890:
4886:
4884:
4883:
4879:
4877:
4876:
4872:
4870:
4869:
4865:
4862:
4861:
4857:
4854:
4853:
4852:Bahariasaurus
4849:
4847:
4846:
4842:
4841:
4838:
4835:
4832:
4828:
4824:
4794:
4793:
4789:
4787:
4786:
4782:
4780:
4779:
4775:
4773:
4772:
4768:
4767:
4765:
4763:
4759:
4753:
4752:
4748:
4747:
4744:
4741:
4739:
4735:
4729:
4728:
4724:
4722:
4721:
4717:
4715:
4714:
4710:
4708:
4707:
4703:
4701:
4700:
4696:
4694:
4693:
4692:Kelmayisaurus
4689:
4687:
4686:
4682:
4680:
4679:
4675:
4672:
4671:
4667:
4665:
4664:
4660:
4659:
4656:
4653:
4651:
4647:
4640:
4639:
4635:
4632:
4631:
4627:
4625:
4624:
4620:
4617:
4616:
4612:
4609:
4608:
4604:
4603:
4601:
4599:
4595:
4589:
4588:
4584:
4582:
4581:
4577:
4574:
4573:
4569:
4567:
4566:
4562:
4560:
4559:
4555:
4553:
4552:
4548:
4547:
4544:
4541:
4539:
4535:
4528:
4527:
4526:Saurophaganax
4523:
4520:
4519:
4515:
4512:
4511:
4507:
4505:
4504:
4500:
4499:
4497:
4495:
4491:
4488:
4485:
4480:
4466:
4465:
4461:
4459:
4458:
4457:Siamotyrannus
4454:
4452:
4451:
4447:
4445:
4444:
4440:
4438:
4437:
4433:
4432:
4430:
4428:
4424:
4418:
4417:
4413:
4410:
4409:
4405:
4404:
4401:
4398:
4396:
4392:
4385:
4384:
4380:
4377:
4376:
4372:
4370:
4369:
4365:
4363:
4362:
4358:
4357:
4354:
4350:
4347:
4344:
4343:Allosauroidea
4339:
4332:
4331:
4330:Spinosauridae
4327:
4324:
4323:
4319:
4316:
4315:
4311:
4309:
4308:
4304:
4303:
4300:
4297:
4294:
4289:
4282:
4281:
4277:
4274:
4273:
4269:
4268:
4265:
4262:
4259:
4254:
4250:
4246:
4241:
4237:
4227:
4222:
4220:
4213:
4183:
4182:
4181:
4180:
4178:
4176:
4172:
4168:
4146:
4145:
4141:
4139:
4138:
4134:
4133:
4131:
4129:
4125:
4119:
4118:
4114:
4112:
4111:
4107:
4105:
4104:
4100:
4098:
4097:
4093:
4090:
4089:
4085:
4084:
4081:
4078:
4076:
4075:Spinosaurinae
4072:
4062:
4061:
4057:
4055:
4054:
4053:Riparovenator
4050:
4048:
4047:
4046:Ceratosuchops
4043:
4042:
4040:
4038:
4034:
4027:
4026:
4022:
4019:
4018:
4014:
4012:
4011:
4010:Riojavenatrix
4007:
4005:
4004:
4000:
3998:
3997:
3993:
3992:
3989:
3986:
3984:
3983:Baryonychinae
3980:
3973:
3972:
3968:
3966:
3965:
3961:
3959:
3958:
3954:
3951:
3950:
3946:
3945:
3942:
3939:
3936:
3935:Spinosauridae
3931:
3916:
3915:
3911:
3909:
3908:
3904:
3902:
3901:
3897:
3895:
3894:
3890:
3888:
3887:
3883:
3881:
3880:
3876:
3875:
3873:
3871:
3867:
3861:
3860:
3859:Wiehenvenator
3856:
3854:
3853:
3849:
3847:
3846:
3842:
3840:
3839:
3835:
3834:
3832:
3830:
3826:
3820:
3819:
3815:
3814:
3811:
3808:
3806:
3802:
3795:
3794:
3790:
3789:
3786:
3782:
3779:
3776:
3771:
3760:
3759:
3755:
3753:
3752:
3748:
3746:
3745:
3741:
3739:
3738:
3734:
3733:
3731:
3729:
3725:
3719:
3718:
3714:
3713:
3710:
3706:
3703:
3700:
3695:
3692:
3689:
3684:
3678:
3677:
3673:
3671:
3670:
3666:
3663:
3662:
3658:
3656:
3655:
3651:
3649:
3648:
3644:
3642:
3641:
3637:
3635:
3634:
3630:
3628:
3627:
3623:
3620:
3619:
3615:
3613:
3612:
3608:
3605:
3604:
3600:
3597:
3596:
3595:Chienkosaurus
3592:
3589:
3588:
3584:
3583:
3580:
3577:
3574:
3569:
3565:
3561:
3556:
3552:
3542:
3535:
3528:
3521:
3495:
3494:
3493:
3492:
3490:
3488:
3484:
3480:
3454:
3453:
3449:
3447:
3446:
3442:
3440:
3439:
3435:
3433:
3432:
3428:
3425:
3424:
3420:
3418:
3417:
3413:
3411:
3410:
3406:
3404:
3403:
3399:
3397:
3396:
3392:
3391:
3389:
3387:
3386:Furileusauria
3383:
3377:
3376:
3372:
3370:
3369:
3365:
3362:
3361:
3360:Rahiolisaurus
3357:
3355:
3354:
3350:
3348:
3347:
3343:
3341:
3340:
3336:
3334:
3333:
3329:
3327:
3326:
3322:
3320:
3319:
3315:
3312:
3311:
3307:
3306:
3303:
3300:
3298:
3294:
3287:
3286:
3282:
3281:
3279:
3277:
3276:Carnotaurinae
3273:
3267:
3266:
3262:
3259:
3258:
3257:Rahiolisaurus
3254:
3252:
3251:
3250:Majungasaurus
3247:
3245:
3244:
3240:
3238:
3237:
3233:
3230:
3229:
3225:
3223:
3222:
3218:
3217:
3215:
3213:
3209:
3203:
3202:
3198:
3196:
3195:
3191:
3189:
3188:
3187:Tarascosaurus
3184:
3182:
3181:
3177:
3175:
3174:
3170:
3168:
3167:
3163:
3161:
3160:
3156:
3154:
3153:
3149:
3147:
3146:
3145:Eoabelisaurus
3142:
3139:
3138:
3134:
3132:
3131:
3130:Chenanisaurus
3127:
3126:
3123:
3120:
3117:
3116:Abelisauridae
3112:
3098:
3097:
3093:
3091:
3090:
3086:
3084:
3083:
3079:
3077:
3076:
3075:Masiakasaurus
3072:
3070:
3069:
3065:
3064:
3062:
3060:
3056:
3050:
3049:
3045:
3043:
3042:
3041:Huinculsaurus
3038:
3036:
3035:
3034:Elaphrosaurus
3031:
3030:
3028:
3026:
3022:
3016:
3015:
3011:
3009:
3008:
3004:
3002:
3001:
2997:
2994:
2993:
2989:
2986:
2985:
2981:
2978:
2977:
2973:
2970:
2969:
2965:
2963:
2962:
2958:
2955:
2954:
2953:Austrocheirus
2950:
2947:
2946:
2942:
2941:
2938:
2935:
2933:
2929:
2922:
2921:
2917:
2914:
2913:
2909:
2906:
2905:
2901:
2898:
2897:
2893:
2892:
2889:
2885:
2882:
2879:
2874:
2864:
2863:
2859:
2857:
2856:
2852:
2851:
2849:
2847:
2843:
2837:
2836:
2832:
2829:
2828:
2824:
2822:
2821:
2817:
2814:
2813:
2809:
2807:
2806:
2805:Berberosaurus
2802:
2801:
2798:
2794:
2791:
2788:
2783:
2776:
2775:
2771:
2769:
2768:
2764:
2763:
2760:
2757:
2754:
2749:
2745:
2741:
2736:
2732:
2723:
2717:
2694:
2693:
2692:Halticosaurus
2689:
2687:
2686:
2685:Teinurosaurus
2682:
2681:
2679:
2675:
2666:
2665:
2664:
2663:
2661:
2659:
2655:
2645:
2644:
2640:
2637:
2636:
2632:
2630:
2629:
2625:
2622:
2621:
2617:
2614:
2613:
2612:Megapnosaurus
2609:
2607:
2606:
2602:
2600:
2599:
2595:
2593:
2592:
2588:
2587:
2585:
2583:
2582:Coelophysidae
2579:
2573:
2572:
2571:Powellvenator
2568:
2566:
2565:
2561:
2559:
2558:
2554:
2552:
2551:
2547:
2544:
2543:
2539:
2538:
2535:
2532:
2530:
2526:
2520:
2519:
2515:
2512:
2511:
2507:
2505:
2504:
2500:
2498:
2497:
2493:
2491:
2490:
2486:
2484:
2483:
2479:
2477:
2476:
2472:
2470:
2469:
2465:
2463:
2462:
2461:Liliensternus
2458:
2456:
2455:
2451:
2449:
2448:
2444:
2442:
2441:
2437:
2434:
2433:
2432:Dolichosuchus
2429:
2427:
2426:
2425:Dilophosaurus
2422:
2421:
2418:
2415:
2412:
2407:
2400:
2399:
2398:Herrerasauria
2395:
2392:
2391:
2387:
2384:
2383:
2379:
2376:
2375:
2371:
2369:
2368:
2364:
2361:
2360:
2356:
2353:
2352:
2351:Daemonosaurus
2348:
2345:
2344:
2340:
2339:
2336:
2333:
2330:
2325:
2321:
2317:
2312:
2308:
2287:
2286:
2285:
2284:
2282:
2280:
2276:
2268:
2264:
2263:
2262:
2261:
2258:
2255:
2253:
2249:
2245:
2237:
2233:
2230:
2228:
2224:
2221:
2219:
2215:
2213:
2209:
2207:
2203:
2202:
2197:
2193:
2189:
2182:
2177:
2175:
2170:
2168:
2163:
2162:
2159:
2153:
2150:
2149:
2145:
2129:
2125:
2119:
2116:
2111:
2107:
2102:
2097:
2093:
2089:
2085:
2078:
2075:
2070:
2066:
2062:
2058:
2054:
2050:
2046:
2039:
2036:
2031:
2027:
2023:
2019:
2015:
2011:
2007:
2003:
1999:
1992:
1989:
1983:
1978:
1974:
1970:
1966:
1959:
1956:
1951:
1947:
1943:
1939:
1934:
1929:
1925:
1921:
1920:Palaeontology
1917:
1910:
1907:
1902:
1898:
1893:
1888:
1884:
1880:
1879:
1874:
1867:
1864:
1859:
1855:
1850:
1845:
1840:
1835:
1831:
1827:
1823:
1821:
1812:
1809:
1796:
1790:
1787:
1782:
1776:
1772:
1767:
1766:
1757:
1754:
1750:
1746:
1740:
1737:
1732:
1728:
1724:
1720:
1716:
1712:
1705:
1702:
1697:
1693:
1689:
1685:
1681:
1677:
1673:
1666:
1664:
1660:
1648:
1642:
1639:
1634:
1630:
1623:
1619:
1618:Holtz, Thomas
1613:
1610:
1605:
1599:
1595:
1588:
1585:
1580:
1576:
1572:
1566:
1562:
1561:
1553:
1551:
1547:
1544:
1541:
1535:
1533:
1529:
1523:
1518:
1514:
1510:
1506:
1504:
1495:
1493:
1491:
1489:
1487:
1485:
1483:
1481:
1477:
1470:
1466:
1463:
1462:
1458:
1456:
1454:
1450:
1446:
1442:
1438:
1434:
1430:
1426:
1425:choristoderes
1422:
1418:
1417:maniraptorans
1414:
1410:
1408:
1403:
1402:
1397:
1394:
1392:
1387:
1383:
1379:
1375:
1367:
1365:
1362:
1358:
1354:
1348:
1347:
1342:
1338:
1332:Palaeobiology
1331:
1329:
1327:
1323:
1319:
1315:
1314:
1303:
1302:
1294:
1293:
1285:
1284:
1276:
1275:
1267:
1266:
1258:
1257:
1249:
1248:
1240:
1239:
1231:
1230:
1222:
1221:
1213:
1212:
1209:
1208:
1202:
1201:
1198:
1197:
1189:
1188:
1185:
1184:
1183:
1182:Teratophoneus
1176:
1175:
1172:
1171:
1168:
1167:
1166:
1159:
1158:
1152:
1151:
1145:
1144:
1141:
1140:
1137:
1136:
1135:
1128:
1127:
1121:
1120:
1117:
1116:
1113:
1112:
1111:
1104:
1103:
1097:
1096:
1093:
1092:
1089:
1088:
1082:
1081:
1075:
1074:
1071:
1070:
1067:
1066:
1065:
1064:Xiongguanlong
1058:
1057:
1051:
1050:
1047:
1046:
1043:
1042:
1041:
1034:
1033:
1027:
1026:
1023:
1022:
1019:
1018:
1017:
1010:
1009:
1003:
1002:
999:
998:
990:
989:
981:
980:
972:
971:
968:
967:
966:
959:
958:
955:
954:
951:
950:
949:
942:
941:
935:
934:
931:
930:
927:
926:
925:
918:
917:
911:
910:
907:
906:
903:
902:
901:
894:
893:
887:
884:
883:
877:
876:
873:
872:
869:
868:
862:
861:
855:
854:
851:
850:
842:
841:
838:
837:
836:
835:Stokesosaurus
829:
828:
825:
824:
821:
820:
819:
812:
811:
805:
804:
798:
797:
793:
792:
786:
785:
779:
771:
767:
765:
764:
759:
754:
752:
751:
746:
745:
740:
739:
734:
733:
732:Compsognathus
728:
727:
722:
721:Tyrannosaurus
718:
714:
710:
706:
705:
700:
696:
692:
688:
683:
681:
680:Ornitholestes
677:
673:
669:
665:
664:Ornitholestes
661:
657:
652:
650:
646:
642:
641:
636:
632:
631:Ornitholestes
628:
624:
623:Ornitholestes
620:
619:
618:Ornitholestes
614:
610:
606:
601:
599:
598:Coelurosauria
595:
594:
589:
585:
584:
579:
572:
567:
560:
558:
555:
547:
540:
538:
536:
532:
528:
527:
522:
521:
516:
512:
507:
505:
501:
497:
493:
492:
491:Archaeopteryx
487:
486:
485:Compsognathus
481:
477:
473:
472:
467:
463:
459:
455:
451:
447:
443:
439:
432:
428:
423:
416:
414:
412:
408:
404:
400:
396:
392:
391:type specimen
387:
385:
381:
377:
373:
369:
365:
364:
359:
355:
351:
347:
343:
339:
336:
333:
329:
325:
324:
310:
308:
304:
303:
301:
298:
294:
289:
283:
281:
272:
269:
268:Binomial name
265:
261:
260:
254:
251:
250:
245:
240:
239:
232:
229:
228:
225:
219:
216:
215:
212:
209:
206:
203:
202:
199:
196:
193:
190:
189:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
153:
150:
147:
146:
141:
136:
132:
128:
124:
120:
115:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
40:
32:
26:
22:
19:
6374:
6338:
6267:
6260:
6253:
6246:
6239:
6234:Ornithomimus
6232:
6225:
6218:
6211:
6204:
6197:
6190:
6172:Tyrannomimus
6170:
6162:
6154:
6147:
6142:Deinocheirus
6140:
6133:
6113:
6106:
6099:
6075:
6067:
6062:Nqwebasaurus
6060:
6055:Nedcolbertia
6053:
6046:
6039:
6033:Calamosaurus
6031:
6025:Aviatyrannis
6023:
6016:
5991:
5961:
5954:
5947:
5939:
5932:
5925:
5918:
5910:
5905:Aristosuchus
5903:
5897:Aniksosaurus
5895:
5877:Aniksosaurus
5875:
5845:
5838:
5830:
5776:
5741:
5734:
5727:
5720:
5646:
5639:
5632:
5612:
5605:
5600:Dynamoterror
5598:
5578:
5571:
5551:
5544:
5524:
5519:Asiatyrannus
5517:
5493:
5486:
5457:Dryptosaurus
5455:
5448:
5441:
5410:
5403:
5396:
5388:
5381:
5374:
5367:
5360:
5353:
5348:Jinbeisaurus
5346:
5339:
5332:
5324:
5319:Aviatyrannis
5317:
5310:
5276:
5271:Sinotyrannus
5269:
5263:
5262:
5255:
5248:
5227:
5220:
5198:
5191:
5150:
5143:
5136:
5128:
5121:
5113:
5105:
5100:Bicentenaria
5098:
5090:
5085:Asiamericana
5083:
5076:
5017:
4958:
4951:
4944:
4937:
4930:
4923:
4916:
4895:
4887:
4880:
4873:
4866:
4860:Deltadromeus
4858:
4850:
4843:
4792:Tyrannotitan
4790:
4783:
4776:
4769:
4749:
4725:
4720:Taurovenator
4718:
4711:
4704:
4699:Lajasvenator
4697:
4690:
4683:
4678:Concavenator
4676:
4668:
4661:
4636:
4628:
4621:
4613:
4605:
4585:
4578:
4570:
4563:
4556:
4549:
4524:
4516:
4508:
4501:
4494:Allosauridae
4462:
4455:
4450:Shidaisaurus
4448:
4441:
4434:
4414:
4406:
4381:
4373:
4366:
4359:
4328:
4320:
4312:
4305:
4278:
4270:
4258:Avetheropoda
4245:Avetheropoda
4216:
4175:Avetheropoda
4142:
4135:
4128:Spinosaurini
4115:
4108:
4101:
4094:
4086:
4058:
4051:
4044:
4023:
4015:
4008:
4003:Protathlitis
4001:
3994:
3969:
3962:
3955:
3947:
3912:
3905:
3898:
3893:Leshansaurus
3891:
3884:
3877:
3857:
3850:
3845:Megalosaurus
3843:
3838:Duriavenator
3836:
3816:
3791:
3775:Megalosauria
3756:
3749:
3744:Marshosaurus
3742:
3737:Condorraptor
3735:
3715:
3674:
3667:
3659:
3652:
3645:
3638:
3631:
3624:
3618:Cruxicheiros
3616:
3609:
3601:
3593:
3585:
3450:
3445:Quilmesaurus
3443:
3436:
3429:
3421:
3414:
3407:
3400:
3393:
3373:
3366:
3358:
3351:
3344:
3337:
3330:
3323:
3316:
3308:
3297:Brachyrostra
3283:
3263:
3255:
3248:
3241:
3234:
3226:
3219:
3199:
3194:Tralkasaurus
3192:
3185:
3178:
3171:
3164:
3157:
3150:
3143:
3135:
3128:
3096:Vespersaurus
3094:
3089:Velocisaurus
3087:
3080:
3073:
3066:
3046:
3039:
3032:
3012:
3005:
2998:
2990:
2984:Deltadromeus
2982:
2974:
2968:Compsosuchus
2966:
2959:
2951:
2943:
2918:
2910:
2904:Lametasaurus
2902:
2894:
2860:
2855:Ceratosaurus
2853:
2833:
2825:
2818:
2810:
2803:
2787:Ceratosauria
2772:
2765:
2690:
2683:
2641:
2633:
2626:
2618:
2610:
2603:
2596:
2589:
2569:
2564:Podokesaurus
2562:
2555:
2548:
2540:
2516:
2508:
2501:
2494:
2487:
2480:
2473:
2466:
2459:
2452:
2447:Gojirasaurus
2445:
2440:Dracovenator
2438:
2430:
2423:
2411:Neotheropoda
2396:
2388:
2380:
2374:Guaibasaurus
2372:
2365:
2357:
2349:
2343:Chindesaurus
2341:
2231:
2222:
2131:. Retrieved
2127:
2118:
2094:(2): e1487.
2091:
2087:
2077:
2052:
2048:
2038:
2005:
2001:
1991:
1972:
1968:
1958:
1923:
1919:
1909:
1882:
1876:
1866:
1829:
1825:
1819:
1811:
1799:. Retrieved
1789:
1764:
1756:
1748:
1744:
1739:
1714:
1710:
1704:
1679:
1675:
1651:. Retrieved
1641:
1632:
1628:
1612:
1593:
1587:
1559:
1539:
1512:
1508:
1502:
1449:allotherians
1405:
1401:Megalosaurus
1399:
1396:
1389:
1371:
1360:
1356:
1352:
1351:
1344:
1325:
1321:
1317:
1311:
1309:
1205:
1180:
1179:
1163:
1162:
1132:
1131:
1110:Dryptosaurus
1108:
1107:
1085:
1062:
1061:
1038:
1037:
1014:
1013:
965:Sinotyrannus
963:
962:
947:
946:
945:
922:
921:
898:
897:
865:
833:
832:
816:
815:
794:
782:
761:
758:Darren Naish
755:
748:
744:Ornithomimus
742:
736:
730:
724:
720:
716:
702:
698:
690:
686:
684:
679:
675:
667:
663:
662:(as well as
659:
656:phylogenetic
653:
648:
644:
638:
635:allosauroids
630:
626:
622:
616:
608:
605:Ceratosaurus
604:
602:
593:Ceratosaurus
591:
583:Megalosaurus
581:
577:
575:
553:
552:
524:
520:Ceratosaurus
518:
514:
508:
489:
483:
479:
471:Megalosaurus
469:
435:
388:
372:Ceratosaurus
371:
367:
363:Ceratosaurus
361:
357:
353:
322:
321:
320:
307:Megalosaurus
305:
279:
274:
258:
257:
237:
236:
204:
191:
178:
24:
18:
6299:Maniraptora
6288:Maniraptora
6269:Tototlmimus
6199:Anserimimus
6149:Garudimimus
6135:Beishanlong
6077:Valdoraptor
6018:Arkansaurus
5963:Xunmenglong
5840:Juravenator
5634:Tarbosaurus
5526:Nanuqsaurus
5495:Gorgosaurus
5412:Megaraptora
5398:Timurlengia
5130:Sciurumimus
5078:Aratasaurus
4946:Murusraptor
4868:Fukuiraptor
4831:Megaraptora
4713:Shaochilong
4685:Eocarcharia
4638:Megaraptora
4558:Lusovenator
4436:Alpkarakush
4293:Carnosauria
4144:Spinosaurus
4117:Siamosaurus
4017:Suchosaurus
3964:Iberospinus
3900:Magnosaurus
3879:Afrovenator
3852:Torvosaurus
3603:Chilesaurus
3416:Carnotaurus
3409:Caletodraco
3395:Abelisaurus
3221:Arcovenator
3068:Laevisuchus
3059:Noasaurinae
2961:Berthasaura
2932:Noasauridae
2862:Genyodectes
2827:Lukousaurus
2820:Jubbulpuria
2620:Panguraptor
2598:Coelophysis
2591:Camposaurus
2550:Panguraptor
2542:Dracoraptor
2518:Zupaysaurus
2503:Tachiraptor
2482:Sarcosaurus
2382:Nhandumirim
1801:11 November
1515:: 155–195.
1427:, lizards,
1407:Cetiosaurus
1313:Alpkarakush
1087:Megaraptora
750:Deinonychus
541:Description
526:nomen nudum
480:M. bradleyi
376:coelurosaur
354:P. bradleyi
332:carnivorous
6447:Categories
6227:Gallimimus
6156:Harpymimus
6101:Harpymimus
5722:Iliosuchus
5692:see below↓
5537:Alioramini
5355:Juratyrant
5341:Eotyrannus
5326:Bagaraatan
5278:Yutyrannus
5212:Coeluridae
5138:Vayuraptor
4984:see below↓
4960:Tratayenia
4953:Orkoraptor
4939:Megaraptor
4845:Aoniraptor
4778:Mapusaurus
4706:Sauroniops
4670:Altispinax
4623:Neovenator
4565:Siamraptor
4551:Datanglong
4518:Epanterias
4510:Antrodemus
4503:Allosaurus
4484:Allosauria
4307:Altispinax
4272:Gasosaurus
4184:see below↓
4060:Suchomimus
3661:Sinosaurus
3496:see below↓
3452:Viavenator
3402:Aucasaurus
3339:Ilokelesia
3310:Dahalokely
3265:Rajasaurus
3243:Indosaurus
3236:Genusaurus
3228:Dahalokely
3152:Indosuchus
3048:Limusaurus
3007:Ligabueino
3000:Kiyacursor
2992:Genusaurus
2976:Dahalokely
2896:Betasuchus
2767:Dornraptor
2667:see below↓
2643:Segisaurus
2496:Sinosaurus
2359:Eodromaeus
2288:see below↓
2236:Dinosauria
2218:Sauropsida
1832:: e12727.
1653:2009-11-04
1471:References
1435:including
1378:araucarian
1316:recovered
1040:Eotyrannus
1016:Yutyrannus
867:Coeluridae
818:Juratyrant
763:Eotyrannus
726:Allosaurus
531:CT scanned
427:lithograph
198:Saurischia
185:Dinosauria
6303:includes
6248:Rativates
6241:Qiupalong
5941:Scipionyx
5934:Mirischia
5607:Lythronax
5546:Alioramus
5200:Labocania
4918:Aerosteon
4464:Sinraptor
4368:Erectopus
4103:Irritator
3688:Orionides
3573:Tetanurae
3560:Tetanurae
3487:Tetanurae
3423:Elemgasem
3346:Llukalkan
3325:Elemgasem
3082:Noasaurus
2945:Afromimus
2753:Averostra
2740:Averostra
2658:Averostra
2510:Velocipes
2329:Theropoda
2316:Theropoda
2279:Theropoda
2204:Kingdom:
2188:Theropoda
2133:28 August
2110:2056-2799
2069:135409990
2022:0272-4634
1950:129569932
1942:0031-0239
1901:0024-4082
1731:131347939
1696:129493139
1579:985402380
1445:docodonts
1391:Pelourdea
647:, making
450:Cotswolds
346:Bathonian
340:from the
252:Species:
244:von Huene
211:Theropoda
158:Kingdom:
152:Eukaryota
37:166
31:Bathonian
6384:Wikidata
5832:Coelurus
5257:Kileskus
5250:Guanlong
5222:Coelurus
5115:Gualicho
4615:Gualicho
3996:Baryonyx
3332:Guemesia
3159:Kryptops
2920:Ozraptor
2557:Pendraig
2212:Chordata
2210:Phylum:
2206:Animalia
2030:55360032
1858:35821895
1751:: 1–361.
1459:See also
1346:Guanlong
1326:Guanlong
924:Guanlong
784:Guanlong
738:Coelurus
719:than to
704:Guanlong
654:Several
458:prepared
442:theropod
338:dinosaur
335:theropod
309:bradleyi
297:Synonyms
290:, 1910 )
288:Woodward
217:Family:
172:Chordata
168:Phylum:
162:Animalia
148:Domain:
127:Holotype
6417:4822662
6404:4434305
6390:Q132619
5390:Timimus
5152:Zuolong
4882:Rapator
4785:Meraxes
4110:Oxalaia
3431:Koleken
2454:Lepidus
2216:Class:
1849:9271276
1771:366–369
1635:: 5–61.
533:at the
504:calcite
500:fissure
403:oolitic
350:England
277:†
256:†
230:Genus:
106:↓
6041:Hexing
5729:Kakuru
5334:Dilong
3375:Thanos
3353:Niebla
3173:Rugops
3166:Kurupi
2108:
2067:
2028:
2020:
1948:
1940:
1899:
1856:
1846:
1797:. 2009
1777:
1729:
1694:
1600:
1577:
1567:
1318:Dilong
900:Dilong
246:, 1926
129:skull
6430:38602
6305:birds
5362:Moros
4889:Siats
4630:Siats
4572:Siats
2232:Clade
2223:Clade
2065:S2CID
2026:S2CID
1946:S2CID
1885:(4).
1826:PeerJ
1727:S2CID
1692:S2CID
1625:(PDF)
747:, or
709:clade
476:genus
431:hyoid
425:1910
409:near
401:from
348:) of
328:genus
326:is a
205:Clade
192:Clade
179:Clade
6412:GBIF
6297:see
4932:Maip
2390:Tawa
2265:see
2135:2018
2106:ISSN
2018:ISSN
1938:ISSN
1897:ISSN
1854:PMID
1803:2021
1775:ISBN
1629:Gaia
1598:ISBN
1575:OCLC
1565:ISBN
1451:and
1415:and
1388:and
1380:and
1324:and
629:and
488:and
399:1910
389:The
45:PreꞒ
6399:EoL
2096:doi
2057:doi
2053:130
2010:doi
1977:doi
1973:158
1928:doi
1887:doi
1883:201
1844:PMC
1834:doi
1719:doi
1715:430
1684:doi
1517:doi
1513:158
766::
474:(a
452:in
330:of
6449::
6427::
6414::
6401::
6386::
2234::
2225::
2126:.
2104:.
2090:.
2086:.
2063:.
2051:.
2047:.
2024:.
2016:.
2006:23
2004:.
2000:.
1971:.
1967:.
1944:.
1936:.
1924:46
1922:.
1918:.
1895:.
1881:.
1875:.
1852:.
1842:.
1830:10
1828:.
1824:.
1773:.
1747:,
1725:.
1713:.
1690:.
1680:66
1678:.
1674:.
1662:^
1633:15
1631:.
1627:.
1573:.
1549:^
1531:^
1511:.
1507:.
1479:^
1455:.
1447:,
1443:,
1439:,
753:.
741:,
735:,
729:,
723:,
682:.
600:.
506:.
386:.
207::
194::
181::
95:Pg
39:Ma
33:,
6167:?
6159:?
6072:?
6036:?
6028:?
5996:?
5944:?
5915:?
5900:?
5835:?
5415:?
5393:?
5329:?
5232:?
5214:?
5203:?
5133:?
5118:?
5110:?
5095:?
4899:?
4891:?
4863:?
4855:?
4833:?
4673:?
4641:?
4633:?
4618:?
4610:?
4575:?
4529:?
4521:?
4513:?
4411:?
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4275:?
4091:?
4028:?
4020:?
3974:?
3952:?
3917:?
3796:?
3761:?
3664:?
3621:?
3606:?
3598:?
3590:?
3426:?
3363:?
3313:?
3288:?
3260:?
3231:?
3140:?
2995:?
2987:?
2979:?
2971:?
2956:?
2948:?
2923:?
2915:?
2907:?
2899:?
2830:?
2815:?
2777:?
2638:?
2623:?
2615:?
2545:?
2513:?
2435:?
2401:?
2393:?
2385:?
2377:?
2362:?
2354:?
2346:?
2180:e
2173:t
2166:v
2137:.
2112:.
2098::
2092:9
2071:.
2059::
2032:.
2012::
1985:.
1979::
1952:.
1930::
1903:.
1889::
1860:.
1836::
1805:.
1783:.
1749:4
1733:.
1721::
1698:.
1686::
1656:.
1606:.
1581:.
1525:.
1519::
1409:,
1393:,
344:(
286:(
234:†
221:†
100:N
90:K
85:J
80:T
75:P
70:C
65:D
60:S
55:O
50:Ꞓ
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