3346:
3489:
3341:
440:
2501:
866:
140:
451:
4449:
2506:
3541:
3523:
119:
4461:
3505:
3496:
3514:
659:
3550:
3532:
507:, though offering many advantages, may also have led to its vulnerability. The constantly changing environment on a large geological scale may have reduced its long-term survival, favoring smaller species more resilient to ecological shifts. In other words, it was over-specialised and couldn't survive when its habitat changed, unlike smaller related species of caiman.
3568:
3561:
481:
reached about 10.3 metres (34 ft) in length, weighing about 5.16 metric tons (5.69 short tons). Another estimate gave a larger size of 12.5 metres (41 ft) in length, ranging from 9.9–15.8 metres (32–52 ft), and 8.4 metric tons (9.3 short tons) in weight, ranging from 5.6–12.6 metric
486:
reached only 10.9 metres (36 ft) long and 5.6 metric tons (6.2 short tons). A 2022 study estimated a length of 7.6–9.2 metres (25–30 ft) and a mass of 2–6.2 metric tons (2.2–6.8 short tons) using a phylogenetic approach; and a length of 9.2–10 metres (30–33 ft) and mass of 3.9–4.9
541:
hunted large vertebrates, as these ridges are used for puncturing and holding on to flesh. The teeth are slightly flattened at the top and are roughly conical, which means that they would have been unlikely to break on impact with a thick bone. Teeth at the anterior are taller and more pointed,
491:-force), with upper estimates indicating that Purussaurus was capable of generating 69,000 N (around 7 metric tons-force). The large size and estimated strength of this animal appears to have allowed it to include a wide range of prey in its diet, making it an
487:
metric tons (4.3–5.4 short tons) using a non-phylogenetic approach. As only skulls have been found, the actual length is not certain. Bite force has been estimated to be around 52,500 N (around 5.3
723:, and river dolphins. Rivers, floodplains, and lake environments were present. Marine and freshwater fish, turtles, crocodilians, and terrestrial and aquatic mammals are associated with Venezuelan
1178:
Blanco, R. E.; Jones, W. W.; Villamil, J. N. (April 16, 2014). "The 'death roll' of giant fossil crocodyliforms (Crocodylomorpha: Neosuchia): Allometric and skull strength analysis".
3578:
3400:
4855:
4534:
1127:
Scheyer, Torsten M.; Hutchinson, John R.; Strauss, Olivier; Delfino, Massimo; Carrillo-Briceño, Jorge D.; Sánchez, Rodolfo; Sánchez-Villagra, Marcelo R. (2019).
3805:
3560:
1414:
4755:
4750:
533:, but are always around 50 mm (2 in) long and curve slightly backwards. They have small ridges along two of the edges which resemble those in
903:"Phylogenetic analysis of a new morphological dataset elucidates the evolutionary history of Crocodylia and resolves the long-standing gharial problem"
4740:
514:
shows some adaptations for more upright limb orientation or weight support. Unlike all other members of the crown
Crocodylia, which have two sacrals,
4681:
4614:
4521:
1370:
4850:
3393:
3488:
4452:
3386:
4745:
3340:
1407:
3345:
2673:
4830:
2500:
389:
2719:
2588:
4845:
3796:
3591:
4825:
3701:
503:
present, with no real competition from sympatric, smaller, carnivores. Researchers have proposed that the large size of
999:
Aureliano, Tito; Ghilardi, Aline M.; Guilherme, Edson; Souza-Filho, Jonas P.; Cavalcanti, Mauro; Riff, Douglas (2015).
3671:
1400:
2580:
1747:
1340:
Langston, W (1965). "Fossil crocodilians from
Colombia and the Cenozoic history of the Crocodilia in South America".
139:
598:
as it had a much broader, shorter snout and this would require a thicker, stronger neck to support the larger head.
4835:
4820:
4815:
3762:
3739:
2801:
482:
tons (6.2–13.9 short tons), with a mean daily food intake of 40.6 kilograms (90 lb). It is also likely that
4800:
3621:
2773:
270:
4840:
2419:
1304:
1740:
2505:
4860:
4805:
4795:
2794:
1070:
Paiva, Ana Laura S.; Godoy, Pedro L.; Souza, Ray B. B.; Klein, Wilfried; Hsiou, Annie S. (August 13, 2022).
3280:
3054:
4760:
4735:
4476:
4264:
4149:
3471:
3304:
3078:
2787:
4810:
4707:
4627:
4560:
3895:
3823:
3709:
3639:
3634:
3435:
3264:
3257:
3070:
3062:
2946:
1583:
1503:
808:
804:
404:
4456:
4105:
3684:
3504:
3288:
847:
419:
1359:
826:
4785:
4765:
4668:
4601:
4508:
4357:
4349:
4205:
3752:
3531:
2780:
2469:
2462:
2412:
2210:
1263:
1083:
1016:
495:
in its ecosystem. As an adult, it would have preyed upon large to very large vertebrates such as the
4085:
3874:
3679:
3644:
3629:
3495:
2448:
2175:
1380:
871:
1219:
sp. nov. (Pleurodira, Podocnemididae) from the
Miocene–Pliocene of the Solimões Formation, Brazil"
439:
4770:
4432:
4424:
4383:
4339:
4331:
4320:
4049:
3902:
3775:
3747:
3513:
3445:
3312:
3221:
3213:
2939:
2217:
2196:
2189:
1850:
1726:
1530:
1287:
1195:
1109:
737:
719:
616:. During the summer of 2005, a Franco-Peruvian expedition (the Fitzcarrald expedition) found new
331:
134:
4712:
4632:
4565:
3611:
3450:
534:
4790:
4780:
4694:
4547:
4539:
4231:
4223:
4196:
4056:
4035:
4014:
3888:
3839:
3719:
3714:
3549:
3522:
3228:
2727:
2694:
2455:
2182:
1987:
1279:
1160:
1101:
1052:
1034:
936:
839:
817:
743:
415:
411:
4699:
4552:
4460:
959:"Size and Palaeoecology of Giant Miocene South American Crocodiles (Archosauria: Crocodylia)"
3599:
3206:
2441:
2304:
2267:
2203:
2098:
1964:
1271:
1230:
1187:
1150:
1140:
1091:
1042:
1024:
970:
926:
916:
713:
207:
194:
4775:
4367:
4257:
4142:
4114:
4042:
3770:
3656:
3651:
3296:
3272:
3235:
3155:
2903:
2630:
2389:
2374:
2339:
2282:
2152:
2008:
1953:
1770:
1754:
830:
646:
All sense organs (eyes, ears, nostrils) were at the very top of the head, indicating that
625:
549:
397:
1267:
1087:
1020:
4306:
4241:
4176:
4121:
4074:
3916:
3729:
3724:
3661:
3540:
3466:
3440:
3430:
3169:
3147:
2889:
2881:
2839:
2637:
2572:
2558:
2431:
2144:
2128:
2057:
2015:
1973:
1945:
1923:
1880:
1815:
1802:
1777:
1762:
1733:
1664:
1620:
1605:
1254:(Crocodyliformes, Alligatoridae) from the Upper Miocene Urumaco Formation, Venezuela".
1155:
1128:
1047:
1000:
931:
902:
681:
1327:
Rodrigues, J.B. (1892). ""Les reptiles fossils de la Vallée de L'Amazone". Vellosia".
1250:
Aguilera, Orangel A.; Riff, Douglas; Bocquentin-Villanueva, Jean (2006). "A new giant
450:
4729:
4376:
4313:
4299:
4212:
3972:
3832:
3367:
3199:
3192:
3177:
3162:
3125:
3111:
3023:
3008:
3000:
2917:
2896:
2867:
2860:
2853:
2743:
2657:
2645:
2595:
2565:
2550:
2536:
2318:
2289:
2259:
2121:
2090:
2077:
2065:
2049:
2031:
1994:
1894:
1718:
1696:
1129:"Giant extinct caiman breaks constraint on the axial skeleton of extant crocodylians"
1113:
958:
732:
600:
566:
492:
373:
227:
76:
1291:
1199:
4399:
4391:
4292:
4250:
4135:
3979:
3937:
3930:
3860:
3362:
3185:
3118:
3031:
3016:
2983:
2972:
2924:
2706:
2616:
2603:
2523:
2402:
2353:
2346:
2274:
2136:
2113:
1938:
1930:
1901:
1843:
1836:
1711:
1596:
1488:
1463:
974:
500:
470:
294:
4659:
1191:
1096:
1071:
1029:
4285:
4278:
4271:
4021:
4000:
3993:
3923:
3909:
3867:
3853:
3846:
3812:
3689:
3140:
3133:
2931:
2910:
2874:
2846:
2825:
2360:
2332:
2325:
2311:
2252:
2105:
2041:
2001:
1980:
1908:
1887:
1829:
1703:
1671:
1628:
1310:
1072:"Body size estimation of Caimaninae specimens from the miocene of South America"
880:
774:
693:
687:
675:
560:
554:
385:
118:
51:
38:
4592:
4413:
4406:
4128:
4007:
3986:
3951:
3944:
3094:
2958:
2763:
2687:
2680:
2666:
2245:
1862:
1642:
1542:
1454:
1424:
1275:
861:
757:
727:. Its environment is described as tropical and coastal. The earlier Colombian
640:
583:
579:
488:
341:
217:
96:
61:
4499:
1283:
1105:
1038:
4185:
4164:
4156:
3881:
3567:
3044:
2382:
2165:
1649:
1515:
851:
658:
643:" maneuver used by extant crocodilians to subdue and dismember their prey.
496:
423:
151:
101:
45:
3604:
3378:
1164:
1056:
940:
1235:
1214:
4653:
4586:
4493:
3958:
3247:
2735:
2296:
1916:
1656:
1472:
1448:
812:
762:
708:
401:
381:
171:
91:
86:
71:
66:
56:
34:
1145:
921:
4686:
4619:
4526:
3965:
1635:
1392:
843:
699:
609:
526:
377:
362:
250:
181:
106:
81:
30:
1001:"Morphometry, Bite-Force, and Paleobiology of the Late Miocene Caiman
4094:
2232:
1442:
821:
799:
617:
408:
393:
369:
237:
161:
4470:
1678:
907:
704:
657:
613:
522:
365:
4673:
4606:
4513:
2751:
1613:
834:
629:
477:
is 1,453 millimetres (57.2 in). It has been estimated that
4474:
3382:
2521:
1800:
1581:
1435:
1396:
392:
classification. It is known from skull material found in the
3566:
3559:
1342:
University of
California Publications in Geological Sciences
717:) weighing up to 700 kilograms (1540 lb), the primate
552:
ever to have existed. Three other extinct crocodyliformes,
542:
whereas those at the posterior are lower and more rounded.
901:
Rio, Jonathan P.; Mannion, Philip D. (September 6, 2021).
469:
The skull length of the largest known individual of the
731:
lived alongside a massive variety of fauna, including
761:. This fauna dates from 13 million years ago, in the
4643:
4576:
4483:
4423:
4366:
4348:
4330:
4240:
4222:
4195:
4175:
4104:
4084:
4066:
3822:
3795:
3788:
3761:
3738:
3700:
3670:
3620:
3590:
3577:
3459:
3423:
3414:
3245:
3093:
3042:
2982:
2956:
2823:
2761:
2705:
2656:
2614:
2534:
2429:
2400:
2231:
2163:
2076:
2029:
1963:
1860:
1813:
1688:
1595:
1541:
1514:
1487:
635:Analysis of a biomechanical model of the skull of
650:was an ambush predator like many modern caimans.
639:indicated that it was capable of performing the "
994:
992:
990:
988:
986:
984:
952:
950:
586:, respectively. One study also indicates that
3394:
1408:
8:
1329:Contribuições do Museu Botânico do Amazonas
582:much older, dating from the Early and Late
4471:
3792:
3587:
3420:
3401:
3387:
3379:
3103:
2992:
2964:
2831:
2711:
2622:
2542:
2531:
2518:
2237:
2082:
2073:
1872:
1868:
1821:
1810:
1797:
1592:
1578:
1520:
1493:
1484:
1432:
1415:
1401:
1393:
117:
20:
1234:
1154:
1144:
1095:
1046:
1028:
930:
920:
1076:Journal of South American Earth Sciences
893:
608:, but was slightly smaller, had a more
578:had similar proportions, but both were
707:, bats, rodents related to the modern
7:
4856:Taxa named by JoĂŁo Barbosa Rodrigues
777:where its fossils were first found.
755:, and the terrestrial crocodyliform
1256:Journal of Systematic Palaeontology
1223:Revista Brasileira de Paleontologia
673:likely included the extinct turtle
963:Journal of Vertebrate Paleontology
590:may have been heavier than either
14:
4756:Miocene reptiles of North America
4751:Miocene reptiles of South America
459:Size comparisons of estimates of
4741:Prehistoric pseudosuchian genera
4459:
4448:
4447:
3548:
3539:
3530:
3521:
3512:
3503:
3494:
3487:
3344:
3339:
2504:
2499:
1213:Bocqentin, J.; Melo, J. (2006).
864:
449:
438:
138:
975:10.1080/02724634.2007.10010458
703:birds, and mammals including:
604:lived around the same time as
1:
4851:Fossil taxa described in 1892
1192:10.1080/08912963.2014.893300
1097:10.1016/j.jsames.2022.103970
1030:10.1371/journal.pone.0117944
957:Jorge Moreno-Bernal (2007).
773:The genus was named for the
548:is one of the largest known
765:stage of the Late Miocene.
4877:
4445:
3482:
3357:
3337:
3106:
2995:
2967:
2834:
2714:
2625:
2545:
2530:
2517:
2497:
2240:
2085:
1875:
1871:
1824:
1809:
1796:
1591:
1577:
1523:
1496:
1483:
1431:
1360:Tetrapod Zoology post on
1276:10.1017/S147720190600188X
679:, crocodilians including
337:
330:
297:) Barbosa-Rodrigues, 1892
283:
278:
135:Scientific classification
133:
125:
116:
23:
16:Extinct genus of reptiles
4578:Purussaurus brasiliensis
3409:Paleontology in Colombia
2674:"Crocodylus" gariepensis
1003:Purussaurus brasiliensis
664:Purussaurus brasiliensis
570:had similar body sizes.
510:The skeletal anatomy of
4746:Miocene crocodylomorphs
2947:"Tomistoma" lusitanicum
2720:"Crocodylus" megarhinus
2589:"Crocodylus" megarhinus
747:, the early species of
525:vary between the three
4150:Montrichardia aquatica
3571:
3564:
666:
537:. This indicates that
4831:Honda Group, Colombia
4708:Paleobiology Database
4628:Paleobiology Database
4561:Paleobiology Database
3896:Galbula hylochoreutes
3570:
3563:
1236:10.4072/rbp.2006.2.02
809:Castilletes Formation
669:The probable diet of
661:
126:Skull restoration of
4846:Fossils of Venezuela
4645:Purussaurus mirandai
4206:Pachyrhizodus etayoi
3222:"Tomistoma" coppensi
3214:"Tomistoma" cairense
2940:"Tomistoma" cairense
2581:"Crocodylus" affinis
1748:"Crocodylus" affinis
798:Solimões Formation,
789:have been found in:
405:Villavieja Formation
4826:Fossils of Colombia
3875:Dracaena colombiana
3583:stratigraphic units
3229:"Tomistoma" dowsoni
1268:2006JSPal...4..221A
1146:10.7554/eLife.49972
1088:2022JSAES.11803970P
1021:2015PLoSO..1017944A
922:10.7717/peerj.12094
872:Paleontology portal
612:snout and lived in
4321:Stenorhynchosaurus
3903:Granastrapotherium
3572:
3565:
2802:C. thorbjarnarsoni
1851:Listrognathosuchus
1727:Brachyuranochampsa
1217:Stupendemys souzai
1180:Historical Biology
738:Granastrapotherium
667:
628:(600 km from
420:Socorro Formations
4836:Fossils of Panama
4821:Fossils of Brazil
4816:Neogene Venezuela
4723:
4722:
4695:Open Tree of Life
4548:Open Tree of Life
4477:Taxon identifiers
4468:
4467:
4441:
4440:
4057:Xenastrapotherium
4036:Saimiri annectens
3889:Eunectes stirtoni
3840:Aramus paludigrus
3784:
3783:
3480:
3479:
3376:
3375:
3353:
3352:
3335:
3334:
3331:
3330:
3327:
3326:
3323:
3322:
3089:
3088:
2819:
2818:
2815:
2814:
2811:
2810:
2774:C. anthropophagus
2513:
2512:
2495:
2494:
2491:
2490:
2487:
2486:
2483:
2482:
2479:
2478:
2227:
2226:
2025:
2024:
1988:Eurycephalosuchus
1792:
1791:
1788:
1787:
1741:"Crocodylus" acer
1573:
1572:
1569:
1568:
1565:
1564:
1561:
1560:
848:Socorro Formation
840:Urumaco Formation
818:Culebra Formation
744:Xenastrapotherium
412:Culebra Formation
354:
353:
274:
4868:
4801:Neogene Colombia
4716:
4715:
4703:
4702:
4690:
4689:
4677:
4676:
4664:
4663:
4662:
4636:
4635:
4623:
4622:
4610:
4609:
4597:
4596:
4595:
4569:
4568:
4556:
4555:
4543:
4542:
4530:
4529:
4517:
4516:
4504:
4503:
4502:
4472:
4463:
4451:
4450:
3793:
3588:
3552:
3543:
3534:
3525:
3516:
3507:
3498:
3491:
3421:
3403:
3396:
3389:
3380:
3348:
3343:
3305:G. pachyrhynchus
3207:Siquisiquesuchus
3104:
2993:
2965:
2832:
2712:
2623:
2543:
2532:
2519:
2508:
2503:
2470:C. wannlangstoni
2463:C. venezuelensis
2305:Globidentosuchus
2268:Centenariosuchus
2238:
2099:Allognathosuchus
2083:
2074:
1965:Orientalosuchina
1873:
1869:
1822:
1811:
1798:
1593:
1579:
1521:
1494:
1485:
1478:
1477:
1433:
1417:
1410:
1403:
1394:
1379:
1369:
1349:
1336:
1314:
1302:
1296:
1295:
1247:
1241:
1240:
1238:
1210:
1204:
1203:
1175:
1169:
1168:
1158:
1148:
1124:
1118:
1117:
1099:
1067:
1061:
1060:
1050:
1032:
996:
979:
978:
954:
945:
944:
934:
924:
898:
874:
869:
868:
867:
827:Fitzcarrald Arch
714:Josephoartigasia
453:
442:
380:epoch, from the
348:
324:
309:
298:
269:
262:
208:Archosauriformes
195:Archosauromorpha
143:
142:
121:
111:
48:
29:Temporal range:
21:
4876:
4875:
4871:
4870:
4869:
4867:
4866:
4865:
4841:Fossils of Peru
4726:
4725:
4724:
4719:
4711:
4706:
4698:
4693:
4685:
4680:
4672:
4667:
4658:
4657:
4652:
4639:
4631:
4626:
4618:
4613:
4605:
4600:
4591:
4590:
4585:
4572:
4564:
4559:
4551:
4546:
4538:
4533:
4525:
4520:
4512:
4507:
4498:
4497:
4492:
4479:
4469:
4464:
4437:
4419:
4362:
4344:
4340:Dinosaur tracks
4326:
4258:Callawayasaurus
4236:
4218:
4191:
4171:
4143:Cerrejonisuchus
4115:Acherontisuchus
4100:
4080:
4062:
4043:Saimiri fieldsi
3818:
3789:Notable fossils
3780:
3757:
3734:
3710:La Frontera Fm.
3696:
3666:
3635:Castilletes Fm.
3616:
3582:
3580:
3573:
3558:
3557:
3556:
3555:
3554:
3553:
3545:
3544:
3536:
3535:
3527:
3526:
3518:
3517:
3509:
3508:
3500:
3499:
3476:
3455:
3416:
3410:
3407:
3377:
3372:
3349:
3319:
3281:G. curvirostris
3258:G. bengawanicus
3241:
3236:Toyotamaphimeia
3156:Hesperogavialis
3097:
3085:
3038:
2986:
2978:
2961:
2952:
2904:Megadontosuchus
2828:
2807:
2795:C. palaeindicus
2757:
2701:
2652:
2631:Aldabrachampsus
2619:
2610:
2539:
2526:
2509:
2475:
2449:C. brevirostris
2425:
2396:
2390:Wannaganosuchus
2375:Stangerochampsa
2340:Orthogenysuchus
2283:Chinatichampsus
2223:
2159:
2153:Wannaganosuchus
2072:
2034:
2021:
2009:Orientalosuchus
1959:
1954:Stangerochampsa
1865:
1856:
1818:
1805:
1784:
1771:Prodiplocynodon
1755:Portugalosuchus
1684:
1587:
1557:
1537:
1510:
1479:
1438:
1437:
1427:
1421:
1377:
1368:(in Portuguese)
1367:
1356:
1339:
1326:
1323:
1321:Further reading
1318:
1317:
1303:
1299:
1249:
1248:
1244:
1212:
1211:
1207:
1177:
1176:
1172:
1126:
1125:
1121:
1069:
1068:
1064:
1015:(2): e0117944.
998:
997:
982:
956:
955:
948:
900:
899:
895:
890:
870:
865:
863:
860:
831:Pebas Formation
783:
771:
662:Restoration of
656:
626:Peruvian Amazon
550:crocodyliformes
479:P. brasiliensis
475:P. brasiliensis
467:
466:
465:
464:
461:P. brasiliensis
456:
455:
454:
445:
444:
443:
432:
398:Peruvian Amazon
346:
318:
307:
292:
289:P. brasiliensis
268:
260:
137:
128:P. brasiliensis
112:
110:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
43:
42:
27:
17:
12:
11:
5:
4874:
4872:
4864:
4863:
4861:Apex predators
4858:
4853:
4848:
4843:
4838:
4833:
4828:
4823:
4818:
4813:
4808:
4806:Neogene Panama
4803:
4798:
4796:Neogene Brazil
4793:
4788:
4783:
4778:
4773:
4768:
4763:
4758:
4753:
4748:
4743:
4738:
4728:
4727:
4721:
4720:
4718:
4717:
4704:
4691:
4678:
4665:
4649:
4647:
4641:
4640:
4638:
4637:
4624:
4611:
4598:
4582:
4580:
4574:
4573:
4571:
4570:
4557:
4544:
4531:
4518:
4505:
4489:
4487:
4481:
4480:
4475:
4466:
4465:
4446:
4443:
4442:
4439:
4438:
4436:
4435:
4429:
4427:
4421:
4420:
4418:
4417:
4410:
4403:
4396:
4387:
4380:
4372:
4370:
4364:
4363:
4361:
4360:
4354:
4352:
4350:Valle Alto Fm.
4346:
4345:
4343:
4342:
4336:
4334:
4328:
4327:
4325:
4324:
4317:
4310:
4307:Platypterygius
4303:
4296:
4289:
4282:
4275:
4268:
4261:
4254:
4246:
4244:
4238:
4237:
4235:
4234:
4228:
4226:
4220:
4219:
4217:
4216:
4209:
4201:
4199:
4193:
4192:
4190:
4189:
4181:
4179:
4173:
4172:
4170:
4169:
4160:
4153:
4146:
4139:
4132:
4125:
4122:Anthracosuchus
4118:
4110:
4108:
4102:
4101:
4099:
4098:
4090:
4088:
4082:
4081:
4079:
4078:
4075:Colombitherium
4070:
4068:
4064:
4063:
4061:
4060:
4053:
4046:
4039:
4032:
4025:
4018:
4011:
4004:
3997:
3990:
3983:
3976:
3969:
3962:
3955:
3948:
3941:
3934:
3927:
3920:
3917:Hilarcotherium
3913:
3906:
3899:
3892:
3885:
3878:
3871:
3864:
3857:
3850:
3843:
3836:
3828:
3826:
3820:
3819:
3817:
3816:
3809:
3801:
3799:
3790:
3786:
3785:
3782:
3781:
3779:
3778:
3773:
3767:
3765:
3759:
3758:
3756:
3755:
3753:Valle Alto Fm.
3750:
3744:
3742:
3736:
3735:
3733:
3732:
3727:
3722:
3717:
3712:
3706:
3704:
3698:
3697:
3695:
3694:
3693:
3692:
3682:
3676:
3674:
3668:
3667:
3665:
3664:
3659:
3654:
3649:
3648:
3647:
3637:
3632:
3626:
3624:
3618:
3617:
3615:
3614:
3609:
3608:
3607:
3596:
3594:
3585:
3575:
3574:
3547:
3546:
3538:
3537:
3529:
3528:
3520:
3519:
3511:
3510:
3502:
3501:
3493:
3492:
3486:
3485:
3484:
3483:
3481:
3478:
3477:
3475:
3474:
3469:
3463:
3461:
3457:
3456:
3454:
3453:
3448:
3443:
3438:
3433:
3427:
3425:
3418:
3412:
3411:
3408:
3406:
3405:
3398:
3391:
3383:
3374:
3373:
3371:
3370:
3365:
3358:
3355:
3354:
3351:
3350:
3338:
3336:
3333:
3332:
3329:
3328:
3325:
3324:
3321:
3320:
3318:
3317:
3309:
3301:
3293:
3285:
3277:
3269:
3261:
3253:
3251:
3243:
3242:
3240:
3239:
3232:
3225:
3218:
3210:
3203:
3196:
3189:
3182:
3174:
3170:Maomingosuchus
3166:
3159:
3152:
3148:Harpacochampsa
3144:
3137:
3130:
3122:
3115:
3107:
3101:
3091:
3090:
3087:
3086:
3084:
3083:
3075:
3071:T. lusitanicum
3067:
3059:
3055:T. calaritanum
3050:
3048:
3040:
3039:
3037:
3036:
3028:
3020:
3013:
3005:
2996:
2990:
2980:
2979:
2977:
2976:
2968:
2962:
2957:
2954:
2953:
2951:
2950:
2943:
2936:
2928:
2921:
2914:
2907:
2900:
2893:
2890:Maomingosuchus
2886:
2882:Leptorrhamphus
2878:
2871:
2864:
2857:
2850:
2843:
2840:Dollosuchoides
2835:
2829:
2824:
2821:
2820:
2817:
2816:
2813:
2812:
2809:
2808:
2806:
2805:
2798:
2791:
2788:C. falconensis
2784:
2777:
2769:
2767:
2759:
2758:
2756:
2755:
2748:
2740:
2732:
2724:
2715:
2709:
2703:
2702:
2700:
2699:
2691:
2684:
2677:
2670:
2662:
2660:
2654:
2653:
2651:
2650:
2642:
2638:Dzungarisuchus
2634:
2626:
2620:
2615:
2612:
2611:
2609:
2608:
2600:
2592:
2585:
2577:
2573:Australosuchus
2569:
2562:
2559:Antecrocodylus
2555:
2546:
2540:
2535:
2528:
2527:
2522:
2515:
2514:
2511:
2510:
2498:
2496:
2493:
2492:
2489:
2488:
2485:
2484:
2481:
2480:
2477:
2476:
2474:
2473:
2466:
2459:
2452:
2445:
2437:
2435:
2427:
2426:
2424:
2423:
2420:M. latrubessei
2416:
2408:
2406:
2398:
2397:
2395:
2394:
2386:
2379:
2371:
2364:
2357:
2350:
2343:
2336:
2329:
2322:
2315:
2308:
2301:
2293:
2286:
2279:
2271:
2264:
2256:
2249:
2241:
2235:
2229:
2228:
2225:
2224:
2222:
2221:
2214:
2207:
2200:
2193:
2186:
2179:
2171:
2169:
2161:
2160:
2158:
2157:
2149:
2145:Procaimanoidea
2141:
2133:
2129:Hassiacosuchus
2125:
2118:
2110:
2102:
2095:
2086:
2080:
2071:
2070:
2062:
2058:Menatalligator
2054:
2046:
2037:
2035:
2030:
2027:
2026:
2023:
2022:
2020:
2019:
2016:Protoalligator
2012:
2005:
1998:
1991:
1984:
1977:
1974:Dongnanosuchus
1969:
1967:
1961:
1960:
1958:
1957:
1950:
1946:Procaimanoidea
1942:
1935:
1927:
1924:Hassiacosuchus
1920:
1913:
1905:
1898:
1891:
1884:
1881:Albertochampsa
1876:
1866:
1861:
1858:
1857:
1855:
1854:
1847:
1840:
1833:
1825:
1819:
1816:Alligatoroidea
1814:
1807:
1806:
1803:Alligatoroidea
1801:
1794:
1793:
1790:
1789:
1786:
1785:
1783:
1782:
1778:Planocraniidae
1774:
1767:
1763:Pristichampsus
1759:
1751:
1744:
1737:
1734:Charactosuchus
1730:
1723:
1715:
1708:
1700:
1692:
1690:
1686:
1685:
1683:
1682:
1675:
1668:
1665:Trilophosuchus
1661:
1653:
1646:
1639:
1632:
1625:
1621:Harpacochampsa
1617:
1610:
1606:Australosuchus
1601:
1599:
1589:
1588:
1582:
1575:
1574:
1571:
1570:
1567:
1566:
1563:
1562:
1559:
1558:
1556:
1555:
1554:
1553:
1547:
1545:
1539:
1538:
1536:
1535:
1534:
1533:
1524:
1518:
1512:
1511:
1509:
1508:
1507:
1506:
1497:
1491:
1481:
1480:
1476:
1475:
1466:
1457:
1451:
1445:
1436:
1429:
1428:
1422:
1420:
1419:
1412:
1405:
1397:
1391:
1390:
1387:DinosaurusBlog
1381:Article about
1375:
1365:
1355:
1354:External links
1352:
1351:
1350:
1337:
1322:
1319:
1316:
1315:
1297:
1262:(3): 221–232.
1242:
1229:(2): 187–192.
1205:
1186:(5): 514–524.
1170:
1119:
1062:
980:
946:
892:
891:
889:
886:
885:
884:
876:
875:
859:
856:
855:
854:
837:
824:
815:
802:
795:
794:
782:
779:
770:
767:
682:Charactosuchus
655:
652:
458:
457:
448:
447:
446:
437:
436:
435:
434:
433:
431:
428:
372:that lived in
352:
351:
350:
349:
347:Langston, 1965
335:
334:
328:
327:
326:
325:
310:
299:
281:
280:
276:
275:
258:
254:
253:
248:
241:
240:
235:
231:
230:
225:
221:
220:
215:
211:
210:
205:
198:
197:
192:
185:
184:
179:
175:
174:
169:
165:
164:
159:
155:
154:
149:
145:
144:
131:
130:
123:
122:
114:
113:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
49:
28:
15:
13:
10:
9:
6:
4:
3:
2:
4873:
4862:
4859:
4857:
4854:
4852:
4849:
4847:
4844:
4842:
4839:
4837:
4834:
4832:
4829:
4827:
4824:
4822:
4819:
4817:
4814:
4812:
4809:
4807:
4804:
4802:
4799:
4797:
4794:
4792:
4789:
4787:
4784:
4782:
4779:
4777:
4774:
4772:
4769:
4767:
4764:
4762:
4761:Montehermosan
4759:
4757:
4754:
4752:
4749:
4747:
4744:
4742:
4739:
4737:
4736:Alligatoridae
4734:
4733:
4731:
4714:
4709:
4705:
4701:
4696:
4692:
4688:
4683:
4679:
4675:
4670:
4666:
4661:
4655:
4651:
4650:
4648:
4646:
4642:
4634:
4629:
4625:
4621:
4616:
4612:
4608:
4603:
4599:
4594:
4588:
4584:
4583:
4581:
4579:
4575:
4567:
4562:
4558:
4554:
4549:
4545:
4541:
4536:
4532:
4528:
4523:
4519:
4515:
4510:
4506:
4501:
4495:
4491:
4490:
4488:
4486:
4482:
4478:
4473:
4462:
4458:
4454:
4444:
4434:
4431:
4430:
4428:
4426:
4422:
4416:
4415:
4411:
4409:
4408:
4404:
4402:
4401:
4397:
4395:
4394:
4393:
4388:
4386:
4385:
4381:
4379:
4378:
4377:Antarctilamna
4374:
4373:
4371:
4369:
4365:
4359:
4356:
4355:
4353:
4351:
4347:
4341:
4338:
4337:
4335:
4333:
4329:
4323:
4322:
4318:
4316:
4315:
4314:Sachicasaurus
4311:
4309:
4308:
4304:
4302:
4301:
4300:Padillasaurus
4297:
4295:
4294:
4290:
4288:
4287:
4283:
4281:
4280:
4276:
4274:
4273:
4269:
4267:
4266:
4265:Desmatochelys
4262:
4260:
4259:
4255:
4253:
4252:
4248:
4247:
4245:
4243:
4239:
4233:
4230:
4229:
4227:
4225:
4221:
4215:
4214:
4213:Yaguarasaurus
4210:
4208:
4207:
4203:
4202:
4200:
4198:
4194:
4188:
4187:
4183:
4182:
4180:
4178:
4174:
4168:
4167:
4166:
4161:
4159:
4158:
4154:
4152:
4151:
4147:
4145:
4144:
4140:
4138:
4137:
4133:
4131:
4130:
4126:
4124:
4123:
4119:
4117:
4116:
4112:
4111:
4109:
4107:
4103:
4097:
4096:
4092:
4091:
4089:
4087:
4083:
4077:
4076:
4072:
4071:
4069:
4067:Gualanday Gp.
4065:
4059:
4058:
4054:
4052:
4051:
4047:
4045:
4044:
4040:
4038:
4037:
4033:
4031:
4030:
4026:
4024:
4023:
4019:
4017:
4016:
4012:
4010:
4009:
4005:
4003:
4002:
3998:
3996:
3995:
3991:
3989:
3988:
3984:
3982:
3981:
3977:
3975:
3974:
3973:Miocallicebus
3970:
3968:
3967:
3963:
3961:
3960:
3956:
3954:
3953:
3949:
3947:
3946:
3942:
3940:
3939:
3935:
3933:
3932:
3928:
3926:
3925:
3921:
3919:
3918:
3914:
3912:
3911:
3907:
3905:
3904:
3900:
3898:
3897:
3893:
3891:
3890:
3886:
3884:
3883:
3879:
3877:
3876:
3872:
3870:
3869:
3865:
3863:
3862:
3858:
3856:
3855:
3851:
3849:
3848:
3844:
3842:
3841:
3837:
3835:
3834:
3833:Anachlysictis
3830:
3829:
3827:
3825:
3821:
3815:
3814:
3810:
3808:
3807:
3803:
3802:
3800:
3798:
3794:
3791:
3787:
3777:
3774:
3772:
3769:
3768:
3766:
3764:
3760:
3754:
3751:
3749:
3746:
3745:
3743:
3741:
3737:
3731:
3728:
3726:
3723:
3721:
3718:
3716:
3713:
3711:
3708:
3707:
3705:
3703:
3699:
3691:
3688:
3687:
3686:
3683:
3681:
3678:
3677:
3675:
3673:
3669:
3663:
3660:
3658:
3655:
3653:
3650:
3646:
3643:
3642:
3641:
3638:
3636:
3633:
3631:
3630:Barzalosa Fm.
3628:
3627:
3625:
3623:
3619:
3613:
3610:
3606:
3603:
3602:
3601:
3598:
3597:
3595:
3593:
3589:
3586:
3584:
3581:fossiliferous
3576:
3569:
3562:
3551:
3542:
3533:
3524:
3515:
3506:
3497:
3490:
3473:
3470:
3468:
3465:
3464:
3462:
3458:
3452:
3449:
3447:
3444:
3442:
3439:
3437:
3434:
3432:
3429:
3428:
3426:
3422:
3419:
3413:
3404:
3399:
3397:
3392:
3390:
3385:
3384:
3381:
3369:
3368:Gryposuchinae
3366:
3364:
3360:
3359:
3356:
3347:
3342:
3315:
3314:
3310:
3307:
3306:
3302:
3299:
3298:
3294:
3291:
3290:
3286:
3283:
3282:
3278:
3275:
3274:
3270:
3267:
3266:
3262:
3260:
3259:
3255:
3254:
3252:
3250:
3249:
3244:
3238:
3237:
3233:
3231:
3230:
3226:
3224:
3223:
3219:
3216:
3215:
3211:
3209:
3208:
3204:
3202:
3201:
3200:Rhamphosuchus
3197:
3195:
3194:
3193:Piscogavialis
3190:
3188:
3187:
3183:
3180:
3179:
3178:Paratomistoma
3175:
3172:
3171:
3167:
3165:
3164:
3163:Ikanogavialis
3160:
3158:
3157:
3153:
3150:
3149:
3145:
3143:
3142:
3138:
3136:
3135:
3131:
3128:
3127:
3126:Gavialosuchus
3123:
3121:
3120:
3116:
3114:
3113:
3112:Aktiogavialis
3109:
3108:
3105:
3102:
3100:
3096:
3092:
3081:
3080:
3079:T. taiwanicum
3076:
3073:
3072:
3068:
3065:
3064:
3060:
3057:
3056:
3052:
3051:
3049:
3047:
3046:
3041:
3034:
3033:
3029:
3026:
3025:
3024:Paratomistoma
3021:
3019:
3018:
3014:
3011:
3010:
3009:Gavialosuchus
3006:
3003:
3002:
3001:Brasilosuchus
2998:
2997:
2994:
2991:
2989:
2988:sensu stricto
2985:
2981:
2975:
2974:
2970:
2969:
2966:
2963:
2960:
2955:
2949:
2948:
2944:
2942:
2941:
2937:
2934:
2933:
2929:
2927:
2926:
2922:
2920:
2919:
2918:Paratomistoma
2915:
2913:
2912:
2908:
2906:
2905:
2901:
2899:
2898:
2897:Maroccosuchus
2894:
2892:
2891:
2887:
2884:
2883:
2879:
2877:
2876:
2872:
2870:
2869:
2868:Gunggamarandu
2865:
2863:
2862:
2861:Gavialosuchus
2858:
2856:
2855:
2854:Ferganosuchus
2851:
2849:
2848:
2844:
2842:
2841:
2837:
2836:
2833:
2830:
2827:
2822:
2804:
2803:
2799:
2797:
2796:
2792:
2790:
2789:
2785:
2783:
2782:
2778:
2776:
2775:
2771:
2770:
2768:
2766:
2765:
2760:
2754:
2753:
2749:
2746:
2745:
2744:Tzaganosuchus
2741:
2738:
2737:
2733:
2730:
2729:
2725:
2722:
2721:
2717:
2716:
2713:
2710:
2708:
2704:
2697:
2696:
2692:
2690:
2689:
2685:
2683:
2682:
2678:
2676:
2675:
2671:
2669:
2668:
2664:
2663:
2661:
2659:
2658:Osteolaeminae
2655:
2648:
2647:
2646:Oxysdonsaurus
2643:
2640:
2639:
2635:
2633:
2632:
2628:
2627:
2624:
2621:
2618:
2613:
2606:
2605:
2601:
2598:
2597:
2596:Jiangxisuchus
2593:
2591:
2590:
2586:
2583:
2582:
2578:
2575:
2574:
2570:
2568:
2567:
2566:Astorgosuchus
2563:
2561:
2560:
2556:
2553:
2552:
2551:Albertosuchus
2548:
2547:
2544:
2541:
2538:
2537:Crocodyloidea
2533:
2529:
2525:
2520:
2516:
2507:
2502:
2472:
2471:
2467:
2465:
2464:
2460:
2458:
2457:
2456:C. gasparinae
2453:
2451:
2450:
2446:
2444:
2443:
2439:
2438:
2436:
2434:
2433:
2428:
2422:
2421:
2417:
2415:
2414:
2410:
2409:
2407:
2405:
2404:
2399:
2392:
2391:
2387:
2385:
2384:
2380:
2377:
2376:
2372:
2370:
2369:
2365:
2363:
2362:
2358:
2356:
2355:
2351:
2349:
2348:
2344:
2342:
2341:
2337:
2335:
2334:
2330:
2328:
2327:
2323:
2321:
2320:
2319:Kuttanacaiman
2316:
2314:
2313:
2309:
2307:
2306:
2302:
2299:
2298:
2294:
2292:
2291:
2290:Culebrasuchus
2287:
2285:
2284:
2280:
2277:
2276:
2272:
2270:
2269:
2265:
2262:
2261:
2260:Brachychampsa
2257:
2255:
2254:
2250:
2248:
2247:
2243:
2242:
2239:
2236:
2234:
2230:
2220:
2219:
2215:
2213:
2212:
2211:A. prenasalis
2208:
2206:
2205:
2201:
2199:
2198:
2194:
2192:
2191:
2187:
2185:
2184:
2180:
2178:
2177:
2173:
2172:
2170:
2168:
2167:
2162:
2155:
2154:
2150:
2147:
2146:
2142:
2139:
2138:
2134:
2131:
2130:
2126:
2124:
2123:
2122:Chrysochampsa
2119:
2116:
2115:
2111:
2108:
2107:
2103:
2101:
2100:
2096:
2093:
2092:
2091:Akanthosuchus
2088:
2087:
2084:
2081:
2079:
2078:Alligatorinae
2075:
2068:
2067:
2066:Sajkanosuchus
2063:
2060:
2059:
2055:
2052:
2051:
2050:Lianghusuchus
2047:
2044:
2043:
2039:
2038:
2036:
2033:
2032:Alligatoridae
2028:
2018:
2017:
2013:
2011:
2010:
2006:
2004:
2003:
1999:
1997:
1996:
1995:Jiangxisuchus
1992:
1990:
1989:
1985:
1983:
1982:
1978:
1976:
1975:
1971:
1970:
1968:
1966:
1962:
1956:
1955:
1951:
1948:
1947:
1943:
1941:
1940:
1936:
1933:
1932:
1928:
1926:
1925:
1921:
1919:
1918:
1914:
1911:
1910:
1906:
1904:
1903:
1899:
1897:
1896:
1895:Brachychampsa
1892:
1890:
1889:
1885:
1883:
1882:
1878:
1877:
1874:
1870:
1867:
1864:
1859:
1853:
1852:
1848:
1846:
1845:
1841:
1839:
1838:
1834:
1832:
1831:
1827:
1826:
1823:
1820:
1817:
1812:
1808:
1804:
1799:
1795:
1780:
1779:
1775:
1773:
1772:
1768:
1765:
1764:
1760:
1757:
1756:
1752:
1750:
1749:
1745:
1743:
1742:
1738:
1736:
1735:
1731:
1729:
1728:
1724:
1721:
1720:
1719:Borealosuchus
1716:
1714:
1713:
1709:
1706:
1705:
1701:
1699:
1698:
1697:Albertosuchus
1694:
1693:
1691:
1687:
1681:
1680:
1676:
1674:
1673:
1669:
1667:
1666:
1662:
1659:
1658:
1654:
1652:
1651:
1647:
1645:
1644:
1640:
1638:
1637:
1633:
1631:
1630:
1626:
1623:
1622:
1618:
1616:
1615:
1611:
1608:
1607:
1603:
1602:
1600:
1598:
1594:
1590:
1585:
1580:
1576:
1551:
1550:
1549:
1548:
1546:
1544:
1540:
1532:
1528:
1527:
1526:
1525:
1522:
1519:
1517:
1513:
1505:
1501:
1500:
1499:
1498:
1495:
1492:
1490:
1486:
1482:
1474:
1470:
1467:
1465:
1461:
1458:
1456:
1452:
1450:
1446:
1444:
1440:
1439:
1434:
1430:
1426:
1418:
1413:
1411:
1406:
1404:
1399:
1398:
1395:
1389:
1388:
1384:
1376:
1374:
1373:
1366:
1364:
1363:
1358:
1357:
1353:
1347:
1343:
1338:
1334:
1330:
1325:
1324:
1320:
1312:
1308:
1307:
1301:
1298:
1293:
1289:
1285:
1281:
1277:
1273:
1269:
1265:
1261:
1257:
1253:
1246:
1243:
1237:
1232:
1228:
1224:
1220:
1218:
1209:
1206:
1201:
1197:
1193:
1189:
1185:
1181:
1174:
1171:
1166:
1162:
1157:
1152:
1147:
1142:
1138:
1134:
1130:
1123:
1120:
1115:
1111:
1107:
1103:
1098:
1093:
1089:
1085:
1081:
1077:
1073:
1066:
1063:
1058:
1054:
1049:
1044:
1040:
1036:
1031:
1026:
1022:
1018:
1014:
1010:
1006:
1004:
995:
993:
991:
989:
987:
985:
981:
976:
972:
968:
964:
960:
953:
951:
947:
942:
938:
933:
928:
923:
918:
914:
910:
909:
904:
897:
894:
887:
883:
882:
878:
877:
873:
862:
857:
853:
849:
845:
841:
838:
836:
832:
828:
825:
823:
819:
816:
814:
810:
806:
803:
801:
797:
796:
792:
791:
790:
788:
780:
778:
776:
768:
766:
764:
760:
759:
754:
750:
746:
745:
740:
739:
734:
733:astrapotheres
730:
726:
722:
721:
716:
715:
710:
706:
702:
701:
696:
695:
690:
689:
684:
683:
678:
677:
672:
665:
660:
653:
651:
649:
644:
642:
638:
633:
631:
627:
623:
619:
615:
611:
607:
603:
602:
601:Rhamphosuchus
597:
593:
589:
585:
581:
577:
573:
569:
568:
567:Rhamphosuchus
563:
562:
557:
556:
551:
547:
543:
540:
536:
532:
528:
524:
519:
517:
513:
508:
506:
502:
501:notoungulates
498:
494:
493:apex predator
490:
485:
480:
476:
472:
462:
452:
441:
429:
427:
425:
421:
417:
413:
410:
406:
403:
399:
395:
391:
387:
383:
379:
375:
374:South America
371:
367:
364:
360:
359:
345:
343:
339:
338:
336:
333:
329:
322:
317:
316:
311:
306:
305:
300:
296:
291:
290:
285:
284:
282:
277:
272:
267:
266:
259:
256:
255:
252:
249:
246:
243:
242:
239:
236:
233:
232:
229:
228:Alligatoridae
226:
223:
222:
219:
216:
213:
212:
209:
206:
203:
200:
199:
196:
193:
190:
187:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
153:
150:
147:
146:
141:
136:
132:
129:
124:
120:
115:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
47:
40:
36:
32:
26:
22:
19:
4811:Neogene Peru
4644:
4577:
4484:
4425:Floresta Fm.
4412:
4405:
4400:Holoptychius
4398:
4392:Bothriolepis
4390:
4389:
4382:
4375:
4332:Arcabuco Fm.
4319:
4312:
4305:
4298:
4293:Muiscasaurus
4291:
4284:
4277:
4270:
4263:
4256:
4251:Acostasaurus
4249:
4211:
4204:
4184:
4163:
4162:
4155:
4148:
4141:
4136:Cerrejonemys
4134:
4127:
4120:
4113:
4106:CerrejĂłn Fm.
4093:
4073:
4055:
4048:
4041:
4034:
4028:
4027:
4020:
4013:
4006:
3999:
3992:
3985:
3980:Miocochilius
3978:
3971:
3964:
3957:
3950:
3943:
3938:Huilatherium
3936:
3931:Hondadelphys
3929:
3922:
3915:
3908:
3901:
3894:
3887:
3880:
3873:
3866:
3861:Cebupithecia
3859:
3852:
3845:
3838:
3831:
3811:
3806:Gomphotheres
3804:
3776:Floresta Fm.
3748:Arcabuco Fm.
3685:CerrejĂłn Fm.
3460:19th century
3363:Brevirostres
3313:G. papuensis
3311:
3303:
3295:
3287:
3279:
3271:
3265:G. breviceps
3263:
3256:
3246:
3234:
3227:
3220:
3212:
3205:
3198:
3191:
3186:Penghusuchus
3184:
3176:
3168:
3161:
3154:
3146:
3139:
3132:
3124:
3119:Dadagavialis
3117:
3110:
3098:
3077:
3069:
3061:
3053:
3043:
3032:Thecachampsa
3030:
3022:
3017:Melitosaurus
3015:
3007:
2999:
2987:
2984:Tomistominae
2973:Sacacosuchus
2971:
2945:
2938:
2930:
2925:Thecachampsa
2923:
2916:
2909:
2902:
2895:
2888:
2880:
2873:
2866:
2859:
2852:
2845:
2838:
2800:
2793:
2786:
2781:C. checchiai
2779:
2772:
2762:
2750:
2742:
2734:
2726:
2718:
2707:Crocodylinae
2693:
2686:
2679:
2672:
2665:
2644:
2636:
2629:
2617:Crocodylidae
2604:Mekosuchinae
2602:
2594:
2587:
2579:
2571:
2564:
2557:
2549:
2524:Longirostres
2468:
2461:
2454:
2447:
2442:C. australis
2440:
2430:
2418:
2411:
2403:Melanosuchus
2401:
2388:
2381:
2373:
2367:
2366:
2359:
2354:Paranasuchus
2352:
2347:Paranacaiman
2345:
2338:
2331:
2324:
2317:
2310:
2303:
2295:
2288:
2281:
2275:Ceratosuchus
2273:
2266:
2258:
2251:
2244:
2216:
2209:
2202:
2195:
2188:
2181:
2176:A. hailensis
2174:
2164:
2151:
2143:
2137:Navajosuchus
2135:
2127:
2120:
2114:Ceratosuchus
2112:
2104:
2097:
2089:
2064:
2056:
2048:
2040:
2014:
2007:
2000:
1993:
1986:
1979:
1972:
1952:
1944:
1939:Navajosuchus
1937:
1931:Leidyosuchus
1929:
1922:
1915:
1907:
1902:Ceratosuchus
1900:
1893:
1886:
1879:
1849:
1844:Leidyosuchus
1842:
1837:Diplocynodon
1835:
1828:
1776:
1769:
1761:
1753:
1746:
1739:
1732:
1725:
1717:
1712:Asiatosuchus
1710:
1702:
1695:
1677:
1670:
1663:
1655:
1648:
1641:
1634:
1627:
1619:
1612:
1604:
1597:Mekosuchinae
1586:crocodilians
1504:Pseudosuchia
1489:Pseudosuchia
1468:
1464:Pseudosuchia
1459:
1425:crocodilians
1386:
1382:
1371:
1361:
1345:
1341:
1332:
1328:
1305:
1300:
1259:
1255:
1251:
1245:
1226:
1222:
1216:
1208:
1183:
1179:
1173:
1136:
1132:
1122:
1079:
1075:
1065:
1012:
1008:
1002:
969:(3 ): A120.
966:
962:
912:
906:
896:
879:
786:
784:
781:Distribution
772:
756:
752:
748:
742:
736:
729:P. neivensis
728:
724:
718:
712:
698:
692:
686:
680:
674:
670:
668:
663:
654:Paleoecology
647:
645:
636:
634:
621:
610:gharial-like
605:
599:
596:Deinosuchus,
595:
591:
587:
580:geologically
575:
571:
565:
559:
553:
545:
544:
538:
530:
520:
518:have three.
515:
511:
509:
504:
483:
478:
474:
471:type species
468:
460:
422:of northern
357:
356:
355:
340:
320:
314:
313:
304:P. neivensis
303:
302:
295:type species
288:
287:
264:
263:
244:
201:
188:
127:
44:16–5.3
24:
18:
4786:Colloncuran
4766:Huayquerian
4485:Purussaurus
4384:Asterolepis
4286:Leyvachelys
4279:Leivanectes
4272:Kronosaurus
4197:Hondita Fm.
4029:Purussaurus
4022:Potamosiren
4001:Neotamandua
3994:Mourasuchus
3924:Hoazinoides
3910:Gryposuchus
3868:Colombophis
3854:Boreostemma
3847:Balanerodus
3813:Megatherium
3797:Pleistocene
3720:Hondita Fm.
3640:Honda Group
3592:Pleistocene
3446:Hoffstetter
3417:researchers
3289:G. leptodus
3141:Hanyusuchus
3134:Gryposuchus
3063:T. gaudense
2932:Tienosuchus
2911:Ocepesuchus
2875:Kentisuchus
2847:Dollosuchus
2826:Gavialoidea
2368:Purussaurus
2361:Protocaiman
2333:Necrosuchus
2326:Mourasuchus
2312:Gnatusuchus
2253:Bottosaurus
2218:A. thomsoni
2197:A. munensis
2190:A. mefferdi
2106:Arambourgia
2042:Balanerodus
2002:Krabisuchus
1981:Eoalligator
1909:Deinosuchus
1888:Arambourgia
1830:Deinosuchus
1704:Arenysuchus
1672:Ultrastenos
1629:Kalthifrons
1383:Purussaurus
1362:Purussaurus
1311:Fossilworks
1306:Purussaurus
1252:Purussaurus
881:Mourasuchus
805:Honda Group
787:Purussaurus
785:Fossils of
775:Purus River
753:Gryposuchus
749:Mourasuchus
725:P. mirandai
694:Mourasuchus
688:Gryposuchus
676:Stupendemys
671:Purussaurus
648:Purussaurus
637:Purussaurus
622:Purussaurus
606:Purussaurus
592:Sarcosuchus
588:Purussaurus
576:Deinosuchus
572:Sarcosuchus
561:Deinosuchus
555:Sarcosuchus
546:Purussaurus
539:Purussaurus
531:Purussaurus
516:P. mirandai
512:P. mirandai
505:Purussaurus
497:xenarthrans
489:metric tons
484:Purussaurus
430:Description
386:Huayquerian
376:during the
358:Purussaurus
315:P. mirandai
265:Purussaurus
234:Subfamily:
39:Huayquerian
25:Purussaurus
4730:Categories
4433:Trilobites
4414:Strepsodus
4407:Pterygotus
4129:Carbonemys
4086:Bogotá Fm.
4008:Nuciruptor
3987:Mohanamico
3952:Langstonia
3945:Lagonimico
3702:Cretaceous
3680:Bogotá Fm.
3600:Sabana Fm.
3424:After 1900
3361:See also:
3099:sensu lato
3095:Gavialinae
2959:Gavialidae
2764:Crocodylus
2688:Rimasuchus
2681:Euthecodon
2667:Brochuchus
2413:M. fisheri
2246:Acresuchus
2233:Caimaninae
2183:A. mcgrewi
1863:Globidonta
1643:Mekosuchus
1552:see below↓
1543:Crocodilia
1455:Sauropsida
1378:(in Czech)
1139:: e49972.
1082:: 103970.
915:: e12094.
888:References
758:Langstonia
641:death roll
584:Cretaceous
535:ziphodonts
409:Panamanian
342:Dinosuchus
308:Mook, 1941
238:Caimaninae
218:Crocodilia
4771:Chasicoan
4660:Q48789986
4368:Cuche Fm.
4232:Ammonites
4186:Eonatator
4177:OlinĂ Gp.
4165:Titanoboa
4157:Puentemys
4050:Stirtonia
3882:Dukecynus
3824:Honda Gp.
3771:Cuche Fm.
3725:OlinĂ Gp.
3672:Paleogene
3657:Uitpa Fm.
3652:Jimol Fm.
3612:Soatá Fm.
3297:G. lewisi
3273:G. browni
3045:Tomistoma
2383:Tsoabichi
2204:A. olseni
2166:Alligator
1650:Paludirex
1531:Neosuchia
1516:Neosuchia
1441:Kingdom:
1284:1477-2019
1114:251560425
1106:0895-9811
1039:1932-6203
852:Venezuela
769:Etymology
720:Stirtonia
424:Venezuela
402:Colombian
394:Brazilian
368:of giant
344:neivensis
319:Aguilera
271:Rodrigues
158:Kingdom:
152:Eukaryota
4791:Friasian
4781:Laventan
4674:42335147
4654:Wikidata
4607:42335146
4593:Q1241636
4587:Wikidata
4494:Wikidata
4453:Category
4242:Paja Fm.
4224:HilĂł Fm.
4015:Patasola
3959:Lycopsis
3763:Devonian
3740:Jurassic
3730:Paja Fm.
3715:HilĂł Fm.
3690:CerrejĂłn
3662:Ware Fm.
3645:La Venta
3472:Humboldt
3248:Gavialis
2736:Quinkana
2297:Eocaiman
1917:Eocaiman
1657:Quinkana
1473:Eusuchia
1449:Chordata
1447:Phylum:
1443:Animalia
1423:Extinct
1348:: 1–169.
1335:: 41–60.
1292:85950121
1200:84880200
1165:31843051
1057:25689140
1009:PLOS ONE
941:34567843
858:See also
813:Colombia
763:Laventan
709:capybara
382:Friasian
332:Synonyms
279:Species
224:Family:
182:Reptilia
172:Chordata
168:Phylum:
162:Animalia
148:Domain:
35:Friasian
4700:6149899
4687:8632836
4620:8565800
4553:4129092
4540:1129895
4527:4821898
4514:4472819
4500:Q133296
3966:Micodon
3622:Neogene
3441:Hettner
3415:Notable
2728:Kinyang
2695:Kinyang
1636:Kambara
1453:Class:
1264:Bibcode
1156:6917493
1084:Bibcode
1048:4331287
1017:Bibcode
932:8428266
844:Urumaco
793:Miocene
700:Anhinga
624:in the
618:fossils
527:species
416:Urumaco
388:in the
384:to the
378:Miocene
363:extinct
323:., 2006
257:Genus:
251:Jacarea
214:Order:
178:Class:
31:Miocene
4776:Mayoan
4713:265905
4633:265904
4566:194770
4457:Images
4095:Etayoa
3605:TibitĂł
3467:Cuvier
3451:Páramo
3436:Hammen
2432:Caiman
1689:Others
1290:
1282:
1198:
1163:
1153:
1112:
1104:
1055:
1045:
1037:
939:
929:
822:Panama
800:Brazil
705:sloths
691:, and
564:, and
370:caiman
361:is an
273:, 1892
4535:IRMNG
4358:Flora
3579:Major
3431:Etayo
1679:Volia
1584:Basal
1469:Clade
1460:Clade
1288:S2CID
1196:S2CID
1133:eLife
1110:S2CID
908:PeerJ
735:like
614:India
523:teeth
390:SALMA
366:genus
321:et al
245:Clade
202:Clade
189:Clade
4682:GBIF
4615:GBIF
4522:GBIF
2752:Voay
1614:Baru
1529:see
1502:see
1372:UFAC
1313:.org
1280:ISSN
1161:PMID
1102:ISSN
1053:PMID
1035:ISSN
937:PMID
846:and
835:Peru
829:and
807:and
751:and
741:and
630:Lima
574:and
521:The
499:and
418:and
396:and
52:Preęž’
4669:EoL
4602:EoL
4509:EoL
1385:on
1309:at
1272:doi
1231:doi
1188:doi
1151:PMC
1141:doi
1092:doi
1080:118
1043:PMC
1025:doi
971:doi
927:PMC
917:doi
632:).
620:of
594:or
529:of
41:),
4732::
4710::
4697::
4684::
4671::
4656::
4630::
4617::
4604::
4589::
4563::
4550::
4537::
4524::
4511::
4496::
4455:•
1471::
1462::
1346:52
1344:.
1331:.
1286:.
1278:.
1270:.
1258:.
1225:.
1221:.
1194:.
1184:27
1182:.
1159:.
1149:.
1135:.
1131:.
1108:.
1100:.
1090:.
1078:.
1074:.
1051:.
1041:.
1033:.
1023:.
1013:10
1011:.
1007:.
983:^
967:27
965:.
961:.
949:^
935:.
925:.
911:.
905:.
850:,
842:,
833:,
820:,
811:,
697:,
685:,
558:,
473:,
426:.
414:,
407:,
400:,
247::
204::
191::
102:Pg
46:Ma
3402:e
3395:t
3388:v
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3308:?
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3292:?
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3276:?
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3027:?
3012:?
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1934:?
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1707:?
1660:?
1624:?
1609:?
1416:e
1409:t
1402:v
1333:2
1294:.
1274::
1266::
1260:4
1239:.
1233::
1227:9
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1202:.
1190::
1167:.
1143::
1137:8
1116:.
1094::
1086::
1059:.
1027::
1019::
1005:"
977:.
973::
943:.
919::
913:9
711:(
463:.
312:â€
301:â€
293:(
286:â€
261:â€
107:N
97:K
92:J
87:T
82:P
77:C
72:D
67:S
62:O
57:ęž’
37:-
33:(
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