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Sphenophyllales

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core with secondary xylem tissue present in some species. The leaves, which can be several centimeters long, are borne on each node in whorls (called verticels) and are wedge-shaped, fan-shaped, linear, or forked. Reproductive parts are either long terminal cones (consisting of two lobes, a sterile
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Sphenophyllales are small, slender branching plants, usually growing to a height of less than 1 m (3.3 ft) tall. The long stems range from 0.5 cm (0.20 in) to 1.2 cm (0.47 in) in diameter. The stems are jointed and ribbed with weak
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or shrub-like when alive, and formed a portion of the understory in Carboniferous forests. The stem anatomy is
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of articulate land plants and a sister group to the present-day
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are shown in the cladogram below. The possible position of
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Thomas N. Taylor; Edith L. Taylor; Michael Krings (2009).
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Sphenophyllalean taxonomy is often derived from isolated
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parts of plants and classification may be based on the
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Paleobotany:the biology and evolution of fossil plants
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Grolier Concise Encyclopedia of Science and Technology
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Milan Libertín; Jiří Bek & Jana Drábková (2008).
895: 407:Sphenophyllales was first described by the British 383:lower lobe and the fertile upper lobe bearing the 855:. University of California Museum of Paleontology 720:. University of California Museum of Paleontology 8: 366:, making it probable that these plants were 454:. Sphenophyllales is typified by the genus 349:, with most of the fossils coming from the 883: 419:words σφήν ("wedge") and φύλλον ("leaf"). 109:Fossil leaves and branches of the species 100: 20: 836: 767:. Middlesex, England: The Echo Library. 684: 786: 784: 814: 812: 446:(mainly the fossilized cones and the 7: 333:). They are fossils dating from the 603: 580: 557: 537: 512: 486: 479: 378:-like), containing a solid primary 341:. They were common during the Late 468:The probable relationships within 14: 853:"Introduction to the Sphenophyta" 718:"Introduction to Sphenophyllales" 415:in 1898. The name comes from the 871: 132: 1: 825:Acta Palaeontologica Polonica 692:Sybil P. Parker, ed. (1986). 795:. Academic Press, New York. 395:, with monolete or trilete 1066: 1035:Devonian first appearances 764:Darwin and Modern Science 608: 601: 585: 578: 562: 555: 542: 535: 517: 510: 491: 484: 219: 214: 129:Scientific classification 127: 108: 99: 23: 1010:Prehistoric plant orders 112:Sphenophyllum miravallis 391:. All sphenophylls are 761:Seward, A. C. (2006). 972:Paleobiology Database 880:at Wikimedia Commons 838:10.4202/app.2008.0414 1050:Pennsylvanian plants 1045:Mississippian plants 1040:Triassic extinctions 1020:Carboniferous plants 121:Universiteit Utrecht 119:. Collection of the 566:Archaeocalamitaceae 522:Pseudobornia ursina 294:Sphenophyllostachys 438:plant parts (like 987: 986: 889:Taxon identifiers 876:Media related to 802:978-0-12-373972-8 740:"Sphenophyllales" 675: 674: 666: 665: 657: 656: 648: 647: 639: 638: 630: 629: 621: 620: 544:†Sphenophyllales 448:stratigraphically 316: 315: 210: 30:Devonian–Triassic 1057: 980: 979: 967: 966: 954: 953: 944: 943: 931: 930: 929: 916: 915: 914: 884: 875: 865: 864: 862: 860: 849: 843: 842: 840: 816: 807: 806: 788: 779: 778: 758: 752: 751: 749: 747: 742:. Access Science 736: 730: 729: 727: 725: 714: 708: 707: 689: 604: 581: 558: 538: 513: 487: 480: 476:has been added. 205: 200: 137: 136: 104: 94: 31: 27:Temporal range: 21: 1065: 1064: 1060: 1059: 1058: 1056: 1055: 1054: 1030:Triassic plants 1015:Devonian plants 990: 989: 988: 983: 975: 970: 962: 957: 949: 947: 939: 934: 927:Sphenophyllales 925: 924: 919: 910: 909: 904: 897:Sphenophyllales 891: 878:Sphenophyllales 869: 868: 858: 856: 851: 850: 846: 818: 817: 810: 803: 790: 789: 782: 775: 760: 759: 755: 745: 743: 738: 737: 733: 723: 721: 716: 715: 711: 704: 691: 690: 686: 681: 676: 667: 658: 649: 640: 631: 622: 466: 444:fructifications 405: 359: 319:Sphenophyllales 311: 304: 297: 290: 283: 276: 269: 262: 255: 248: 245:Columnisporites 241: 234: 227: 204: 202:Sphenophyllales 198: 131: 95: 93: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 42: 37: 29: 28: 25: 24:Sphenophyllales 17: 16:Order of plants 12: 11: 5: 1063: 1061: 1053: 1052: 1047: 1042: 1037: 1032: 1027: 1025:Permian plants 1022: 1017: 1012: 1007: 1002: 992: 991: 985: 984: 982: 981: 968: 955: 945: 932: 917: 901: 899: 893: 892: 887: 867: 866: 844: 808: 801: 780: 773: 753: 731: 709: 702: 683: 682: 680: 677: 673: 672: 669: 668: 664: 663: 660: 659: 655: 654: 651: 650: 646: 645: 642: 641: 637: 636: 633: 632: 628: 627: 624: 623: 619: 618: 615: 614: 607: 602: 600: 597: 596: 593: 592: 584: 579: 577: 574: 573: 570: 569: 561: 556: 554: 551: 550: 547: 546: 541: 536: 534: 531: 530: 527: 526: 516: 511: 509: 506: 505: 502: 501: 490: 485: 483: 478: 465: 462: 411:and geologist 404: 401: 358: 355: 321:is an extinct 314: 313: 252:Gondwanophyton 217: 216: 212: 211: 196: 192: 191: 186: 182: 181: 179:Polypodiopsida 176: 172: 171: 169:Polypodiophyta 166: 162: 161: 156: 149: 148: 143: 139: 138: 125: 124: 106: 105: 97: 96: 88: 83: 78: 73: 68: 63: 58: 53: 48: 43: 38: 33: 32: 26: 15: 13: 10: 9: 6: 4: 3: 2: 1062: 1051: 1048: 1046: 1043: 1041: 1038: 1036: 1033: 1031: 1028: 1026: 1023: 1021: 1018: 1016: 1013: 1011: 1008: 1006: 1003: 1001: 998: 997: 995: 978: 973: 969: 965: 960: 956: 952: 946: 942: 937: 933: 928: 922: 918: 913: 907: 903: 902: 900: 898: 894: 890: 885: 881: 879: 874: 854: 848: 845: 839: 834: 830: 826: 822: 815: 813: 809: 804: 798: 794: 787: 785: 781: 776: 774:1-4068-0480-0 770: 766: 765: 757: 754: 741: 735: 732: 719: 713: 710: 705: 703:0-7172-8525-1 699: 695: 688: 685: 678: 671: 670: 662: 661: 653: 652: 644: 643: 635: 634: 626: 625: 617: 616: 613: 612: 606: 605: 599: 598: 595: 594: 591: 590: 583: 582: 576: 575: 572: 571: 568: 567: 560: 559: 553: 552: 549: 548: 545: 540: 539: 533: 532: 529: 528: 525: 524: 523: 515: 514: 508: 507: 504: 503: 500: 498: 497: 489: 488: 482: 481: 477: 475: 471: 463: 461: 459: 458: 457:Sphenophyllum 453: 449: 445: 441: 437: 433: 429: 425: 420: 418: 414: 413:Albert Seward 410: 402: 400: 398: 394: 390: 386: 381: 377: 373: 369: 365: 356: 354: 352: 351:Carboniferous 348: 347:Early Permian 344: 343:Pennsylvanian 340: 336: 332: 328: 324: 320: 312: 310: 309: 303: 302: 301:Sphenophyllum 296: 295: 289: 288: 282: 281: 275: 274: 268: 267: 261: 260: 254: 253: 247: 246: 240: 239: 238:Cheirostrobus 233: 232: 226: 225: 218: 213: 208: 203: 197: 194: 193: 190: 187: 184: 183: 180: 177: 174: 173: 170: 167: 164: 163: 160: 159:Tracheophytes 157: 154: 151: 150: 147: 144: 141: 140: 135: 130: 126: 122: 118: 117:Carboniferous 114: 113: 107: 103: 98: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 41: 36: 22: 19: 896: 870: 857:. Retrieved 847: 828: 824: 792: 763: 756: 744:. Retrieved 734: 722:. Retrieved 712: 693: 687: 611:Equisetaceae 609: 589:Calamitaceae 586: 563: 543: 520: 518: 494: 492: 473: 467: 455: 421: 406: 360: 318: 317: 306: 299: 292: 287:Sentistrobus 285: 278: 271: 266:Peltastrobus 264: 257: 250: 243: 236: 229: 224:Hamatophyton 222: 220: 201: 152: 110: 18: 1005:Fern orders 921:Wikispecies 470:Equisetidae 393:homosporous 387:) or loose 372:protostelic 357:Description 327:Equisetales 308:Xihuphyllum 273:Rinistachya 189:Equisetidae 1000:Horsetails 994:Categories 746:27 January 724:27 January 679:References 452:synonymous 428:morphology 424:fossilized 331:horsetails 231:Bowmanites 185:Subclass: 165:Division: 464:Phylogeny 385:sporangia 280:Rotafolia 142:Kingdom: 948:BioLib: 912:Q2899203 906:Wikidata 409:botanist 403:Taxonomy 389:strobili 353:period. 339:Triassic 335:Devonian 259:Lilpopia 115:, Upper 859:31 July 436:sterile 432:anatomy 337:to the 215:Genera 195:Order: 175:Class: 146:Plantae 977:249934 941:223967 799:  771:  700:  440:leaves 397:spores 364:habits 209:, 1898 207:Seward 964:11393 959:IRMNG 951:61464 496:Ibyka 474:Ibyka 442:) or 417:Greek 380:xylem 323:order 153:Clade 936:APNI 861:2011 797:ISBN 769:ISBN 748:2011 726:2011 698:ISBN 499:(?) 430:and 376:root 368:vine 35:PreꞒ 833:doi 434:of 345:to 996:: 974:: 961:: 938:: 923:: 908:: 829:53 827:. 823:. 811:^ 783:^ 460:. 399:. 155:: 123:. 85:Pg 863:. 841:. 835:: 805:. 777:. 750:. 728:. 706:. 587:† 564:† 519:† 493:† 374:( 329:( 305:† 298:† 291:† 284:† 277:† 270:† 263:† 256:† 249:† 242:† 235:† 228:† 221:† 199:† 90:N 80:K 75:J 70:T 65:P 60:C 55:D 50:S 45:O 40:Ꞓ

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A piece of rock showing imprints of about four slender vine-like stems with small wedged-shaped leaves arranged around regularly spaced nodes.
Sphenophyllum miravallis
Carboniferous
Universiteit Utrecht
Scientific classification
Edit this classification
Plantae
Tracheophytes
Polypodiophyta
Polypodiopsida
Equisetidae
Sphenophyllales
Seward
Hamatophyton
Bowmanites
Cheirostrobus
Columnisporites
Gondwanophyton

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