987:
1004:
1055:
970:
1021:
953:
64:
1038:
42:
284:
provides that when a subfamily is formed by reducing the rank of a family, the type of a conserved family name has precedence, making
Epacridoideae the correct name. Although Sweet and Walker-Arnott had treated the group as a subfamily, botanists preferred Brown's rank of family, as in the
259:
are also distinctive: there are fewer than twice the number of corolla lobes and their filaments are smooth. Some of these characters are individually present in other members of the family
Ericaceae. Core members of the subfamily (i.e. excluding Prionoteae) also have parallel- or somewhat
1299:
McNeill, John & Turland, Nicholas (2005), "Synopsis of
Proposals on Botanical Nomenclature - Vienna 2005. A Review of the Proposals concerning the International Code of Botanical Nomenclature Submitted to the XVII International Botanical Congress",
986:
1153:
Kron, K.A.; Judd, W.S.; Stevens, P.F.; Crayn, D.M.; Anderberg, A.A.; Gadek, P.A.; Quinn, C.J. & Luteyn, J.L. (2002), "Phylogenetic
Classification of Ericaceae: Molecular and Morphological Evidence",
281:
1279:
Article 19. International Code of
Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017
1592:
1003:
1487:
1054:
1020:
1365:
Quinn, Christopher J.; Brown, Elizabeth A.; Heslewood, Margaret M. & Crayn, Darren M. (2005), "Generic concepts in
Styphelieae (Ericaceae): the
1554:
969:
952:
305:
The subfamily is divided into tribes, whose suggested relationship is shown in the following cladogram. The number of genera is approximate.
269:
1247:
Crayn, Darren M.; Kron, Kathleen A. & Potter, Benjamin C.M. (2014), "Typification of some names in
Epacridoideae (Ericaceae)",
535:(2002), with some later modifications, genera that may be placed in the subfamily include the following (tribe in parentheses):
1474:
280:
also treated it as a subfamily in 1832 (using the name
Epacrideae). Although Sweet's name is earlier, Article 19.5 of the
1559:
1587:
1037:
235:
group within the family
Ericaceae, clearly diagnosable using a combination of morphological characters. These include a
63:
1395:
Albrecht, D.E.; Owens, C. T.; Weiller, C. M.; Quinn, C. J. (2010), "Generic concepts in
Ericaceae: Styphelioideae: the
277:
162:
1492:
1129:
1167:
273:
1010:
223:, others occurring northwards through the Pacific to Southeast Asia, with a small number in South America.
1582:
1427:
1027:
993:
1224:
1200:
1061:
372:
1465:
1317:
1179:
1171:
1044:
574:
457:
240:
219:
is also used. The subfamily contains around 35 genera and 545 species. Many species are found in
185:
58:
50:
441:
395:
322:
1541:
1500:
1080:
959:
418:
1505:
1408:
1378:
1348:
1309:
1256:
1163:
596:
286:
1277:
101:
976:
88:
1335:
Crayn, Darren M.; Brown, Elizabeth A. & Powell, Jocelyn M. (2003), "A revision of
41:
1576:
1087:, and southeastern Australia. However, some species occur in southern South America (
750:
290:
248:
1183:
1546:
849:
827:
717:
640:
1532:
1459:
1089:
1076:
926:
838:
739:
706:
629:
541:
272:
treated the group as the family Epacridaceae. It was treated as a subfamily by
220:
1282:(electronic ed.), Glashütten: International Association for Plant Taxonomy
293:
in the 1990s showed that excluding Epacridaceae from Ericaceae made the latter
1105:
904:
893:
860:
761:
684:
618:
585:
552:
345:
17:
1450:
1261:
1099:
915:
871:
816:
794:
772:
673:
607:
294:
232:
209:
147:
1526:
1444:
1084:
937:
805:
783:
695:
662:
651:
563:
137:
127:
114:
1321:
1175:
728:
1313:
1094:
882:
256:
236:
1421:
1412:
1382:
1352:
1479:
252:
244:
75:
1425:
282:
International Code of Nomenclature for algae, fungi, and plants
1225:"Indices Nominum Supragenericorum Plantarum Vascularium – S"
1201:"Indices Nominum Supragenericorum Plantarum Vascularium – E"
1168:
10.1663/0006-8101(2002)068[0335:PCOEMA]2.0.CO;2
531:
Based largely on the phylogenetic classification of Kron
1229:
Indices Nominum Supragenericorum Plantarum Vascularium
1205:
Indices Nominum Supragenericorum Plantarum Vascularium
1075:
Members of the Epacridoideae are largely confined to
260:
palmate-veined leaves and lack multicellular hairs.
1516:
1434:
1079:, being particularly diverse in southwestern
8:
1422:
1148:
1146:
1144:
1142:
40:
31:
1593:Taxa named by George Arnott Walker Arnott
1276:Turland, N.J.; et al., eds. (2018),
1260:
289:, until both morphological and molecular
231:The Epacridoideae form a well supported
1194:
1192:
1124:
1122:
1118:
948:
1249:Telopea, Journal of Plant Systematics
276:in 1828 (using the name Stypheliae).
7:
434:
411:
388:
365:
338:
315:
308:
25:
1053:
1036:
1019:
1002:
985:
968:
951:
243:, dry, membrane-like (scarious)
62:
1339:(Styphelioideae: Ericaceae)",
1128:Stevens, P.F. (2001 onwards),
1:
208:is a subfamily of the family
1401:Australian Systematic Botany
1371:Australian Systematic Botany
1341:Australian Systematic Botany
1134:Angiosperm Phylogeny Website
278:George Arnott Walker-Arnott
176:
1609:
1223:Reveal, James L. (2011),
1199:Reveal, James L. (2011),
1130:"Ericales: Epacridoideae"
455:
439:
432:
416:
409:
393:
386:
370:
363:
343:
336:
320:
313:
191:
184:
174:
169:
59:Scientific classification
57:
48:
39:
34:
1262:10.7751/telopea20147924
1103:), and Southeast Asia (
1136:, retrieved 2022-10-21
1028:Prionotes cerinthoides
1011:Oligarrhena micrantha
291:phylogenetic studies
1588:Asterid subfamilies
1062:Styphelia tubiflora
251:, and a persistent
27:Subfamily of plants
1045:Richea dracophylla
689:Brongn. & Gris
170:Tribes and genera
51:Epacris longiflora
1570:
1569:
1501:Open Tree of Life
1428:Taxon identifiers
1081:Western Australia
994:Lysinema ciliatum
960:Archeria racemosa
943:
932:
921:
910:
899:
888:
877:
866:
855:
844:
833:
822:
811:
800:
789:
778:
767:
756:
745:
734:
723:
712:
701:
690:
679:
668:
657:
646:
635:
624:
613:
602:
591:
580:
569:
558:
547:
523:
522:
514:
513:
505:
504:
496:
495:
487:
486:
478:
477:
469:
468:
218:
203:
202:
197:
165:
16:(Redirected from
1600:
1563:
1562:
1550:
1549:
1537:
1536:
1535:
1509:
1508:
1496:
1495:
1483:
1482:
1470:
1469:
1468:
1455:
1454:
1453:
1423:
1416:
1415:
1392:
1386:
1385:
1362:
1356:
1355:
1332:
1326:
1324:
1314:10.2307/25065335
1296:
1290:
1289:
1288:
1287:
1273:
1267:
1265:
1264:
1244:
1238:
1237:
1236:
1235:
1220:
1214:
1213:
1212:
1211:
1196:
1187:
1186:
1156:Botanical Review
1150:
1137:
1126:
1057:
1040:
1023:
1006:
989:
972:
955:
941:
930:
919:
908:
897:
886:
875:
864:
853:
842:
831:
820:
809:
798:
787:
776:
765:
754:
743:
732:
721:
710:
699:
688:
677:
666:
655:
644:
633:
622:
611:
600:
589:
578:
567:
556:
545:
435:
412:
389:
366:
339:
316:
309:
287:Cronquist system
216:
195:
161:
67:
66:
44:
32:
21:
1608:
1607:
1603:
1602:
1601:
1599:
1598:
1597:
1573:
1572:
1571:
1566:
1558:
1553:
1545:
1540:
1531:
1530:
1525:
1512:
1504:
1499:
1491:
1486:
1478:
1473:
1464:
1463:
1458:
1449:
1448:
1443:
1430:
1420:
1419:
1413:10.1071/SB10009
1394:
1393:
1389:
1383:10.1071/SB05005
1364:
1363:
1359:
1353:10.1071/SB03014
1334:
1333:
1329:
1298:
1297:
1293:
1285:
1283:
1275:
1274:
1270:
1246:
1245:
1241:
1233:
1231:
1222:
1221:
1217:
1209:
1207:
1198:
1197:
1190:
1152:
1151:
1140:
1127:
1120:
1115:
1073:
1066:
1058:
1049:
1041:
1032:
1024:
1015:
1014:(Oligarrheneae)
1007:
998:
990:
981:
973:
964:
956:
947:
845:(Oligarrheneae)
834:(Oligarrheneae)
713:(Oligarrheneae)
529:
524:
515:
506:
497:
488:
479:
470:
303:
266:
229:
194:Styphelioideae
160:
61:
28:
23:
22:
15:
12:
11:
5:
1606:
1604:
1596:
1595:
1590:
1585:
1575:
1574:
1568:
1567:
1565:
1564:
1551:
1538:
1522:
1520:
1518:Styphelioideae
1514:
1513:
1511:
1510:
1497:
1484:
1471:
1456:
1440:
1438:
1432:
1431:
1426:
1418:
1417:
1407:(5): 320–332,
1387:
1377:(5): 439–454,
1357:
1327:
1308:(1): 215–250,
1291:
1268:
1239:
1215:
1188:
1162:(3): 335–423,
1138:
1117:
1116:
1114:
1111:
1072:
1069:
1068:
1067:
1059:
1052:
1050:
1042:
1035:
1033:
1025:
1018:
1016:
1008:
1001:
999:
991:
984:
982:
977:Cosmelia rubra
974:
967:
965:
957:
950:
946:
945:
934:
923:
912:
901:
890:
879:
868:
857:
846:
835:
824:
813:
802:
791:
780:
769:
758:
747:
736:
725:
714:
703:
692:
681:
670:
659:
648:
637:
626:
615:
604:
593:
582:
571:
560:
549:
537:
528:
525:
521:
520:
517:
516:
512:
511:
508:
507:
503:
502:
499:
498:
494:
493:
490:
489:
485:
484:
481:
480:
476:
475:
472:
471:
467:
466:
463:
462:
454:
451:
450:
447:
446:
438:
433:
431:
428:
427:
424:
423:
415:
410:
408:
405:
404:
401:
400:
392:
387:
385:
382:
381:
378:
377:
369:
364:
362:
359:
358:
355:
354:
342:
337:
335:
332:
331:
328:
327:
319:
314:
312:
307:
302:
299:
265:
262:
228:
225:
214:Styphelioideae
201:
200:
199:
198:
189:
188:
182:
181:
172:
171:
167:
166:
155:
151:
150:
145:
141:
140:
135:
131:
130:
125:
118:
117:
112:
105:
104:
99:
92:
91:
86:
79:
78:
73:
69:
68:
55:
54:
46:
45:
37:
36:
35:Epacridoideae
26:
24:
18:Styphelioideae
14:
13:
10:
9:
6:
4:
3:
2:
1605:
1594:
1591:
1589:
1586:
1584:
1583:Epacridoideae
1581:
1580:
1578:
1561:
1556:
1552:
1548:
1543:
1539:
1534:
1528:
1524:
1523:
1521:
1519:
1515:
1507:
1502:
1498:
1494:
1489:
1485:
1481:
1476:
1472:
1467:
1466:Epacridoideae
1461:
1457:
1452:
1446:
1442:
1441:
1439:
1437:
1436:Epacridoideae
1433:
1429:
1424:
1414:
1410:
1406:
1402:
1398:
1391:
1388:
1384:
1380:
1376:
1372:
1368:
1361:
1358:
1354:
1350:
1346:
1342:
1338:
1331:
1328:
1325:, pp. 224–225
1323:
1319:
1315:
1311:
1307:
1303:
1295:
1292:
1281:
1280:
1272:
1269:
1263:
1258:
1254:
1250:
1243:
1240:
1230:
1226:
1219:
1216:
1206:
1202:
1195:
1193:
1189:
1185:
1181:
1177:
1173:
1169:
1165:
1161:
1157:
1149:
1147:
1145:
1143:
1139:
1135:
1131:
1125:
1123:
1119:
1112:
1110:
1108:
1107:
1102:
1101:
1096:
1092:
1091:
1086:
1082:
1078:
1070:
1065:(Styphelieae)
1064:
1063:
1056:
1051:
1047:
1046:
1039:
1034:
1030:
1029:
1022:
1017:
1013:
1012:
1005:
1000:
996:
995:
988:
983:
979:
978:
971:
966:
962:
961:
954:
949:
940:
939:
935:
933:(Styphelieae)
929:
928:
924:
922:(Styphelieae)
918:
917:
913:
907:
906:
902:
898:(R.Br.) Sweet
896:
895:
891:
885:
884:
880:
874:
873:
869:
867:(Styphelieae)
863:
862:
858:
856:(Styphelieae)
852:
851:
847:
841:
840:
836:
830:
829:
825:
823:(Styphelieae)
819:
818:
814:
812:(Styphelieae)
808:
807:
803:
801:(Styphelieae)
797:
796:
792:
786:
785:
781:
779:(Styphelieae)
775:
774:
770:
768:(Styphelieae)
764:
763:
759:
757:(Styphelieae)
753:
752:
751:Leptecophylla
748:
742:
741:
737:
731:
730:
726:
720:
719:
715:
709:
708:
704:
702:(Styphelieae)
698:
697:
693:
691:(Styphelieae)
687:
686:
682:
680:(Styphelieae)
676:
675:
671:
669:(Styphelieae)
665:
664:
660:
654:
653:
649:
647:(Styphelieae)
643:
642:
638:
636:(Styphelieae)
632:
631:
627:
625:(Styphelieae)
621:
620:
616:
614:(Styphelieae)
610:
609:
605:
599:
598:
594:
592:(Styphelieae)
588:
587:
583:
577:
576:
572:
570:(Styphelieae)
566:
565:
561:
559:(Styphelieae)
555:
554:
550:
548:(Styphelieae)
544:
543:
539:
538:
536:
534:
526:
519:
518:
510:
509:
501:
500:
492:
491:
483:
482:
474:
473:
465:
464:
461:
460:(16+ genera)
459:
453:
452:
449:
448:
445:
443:
437:
436:
430:
429:
426:
425:
422:
420:
414:
413:
407:
406:
403:
402:
399:
397:
391:
390:
384:
383:
380:
379:
376:
374:
373:Oligarrheneae
368:
367:
361:
360:
357:
356:
353:
351:
347:
341:
340:
334:
333:
330:
329:
326:
324:
318:
317:
311:
310:
306:
300:
298:
296:
292:
288:
283:
279:
275:
271:
263:
261:
258:
254:
250:
249:inflorescence
246:
242:
238:
234:
226:
224:
222:
215:
211:
207:
206:Epacridoideae
193:
192:
190:
187:
183:
180:
178:
173:
168:
164:
159:
158:Epacridoideae
156:
153:
152:
149:
146:
143:
142:
139:
136:
133:
132:
129:
126:
123:
120:
119:
116:
113:
110:
107:
106:
103:
100:
97:
94:
93:
90:
89:Tracheophytes
87:
84:
81:
80:
77:
74:
71:
70:
65:
60:
56:
53:
52:
47:
43:
38:
33:
30:
19:
1517:
1435:
1404:
1400:
1396:
1390:
1374:
1370:
1366:
1360:
1344:
1340:
1336:
1330:
1305:
1301:
1294:
1284:, retrieved
1278:
1271:
1252:
1248:
1242:
1232:, retrieved
1228:
1218:
1208:, retrieved
1204:
1159:
1155:
1133:
1104:
1098:
1088:
1074:
1071:Distribution
1060:
1043:
1031:(Prionoteae)
1026:
1009:
997:(Epacrideae)
992:
980:(Cosmelieae)
975:
963:(Archerieae)
958:
944:(Epacrideae)
936:
925:
914:
911:(Cosmelieae)
903:
892:
881:
878:(Prionoteae)
870:
859:
850:Pentachondra
848:
837:
828:Needhamiella
826:
815:
804:
793:
790:(Epacrideae)
782:
771:
760:
749:
746:(Prionoteae)
738:
735:(Epacrideae)
727:
718:Dracophyllum
716:
705:
694:
683:
672:
661:
658:(Cosmelieae)
650:
641:Conostephium
639:
628:
617:
606:
603:(Archerieae)
595:
584:
581:(Cosmelieae)
573:
562:
551:
540:
532:
530:
456:
440:
417:
394:
371:
349:
344:
321:
304:
295:paraphyletic
274:Robert Sweet
270:Robert Brown
267:
233:monophyletic
230:
213:
205:
204:
175:
157:
121:
108:
95:
82:
49:
29:
1460:Wikispecies
1255:: 319–321,
1090:Lebetanthus
1077:Australasia
927:Trochocarpa
865:C.M.Weiller
839:Oligarrhena
821:C.M.Weiller
755:C.M.Weiller
740:Lebetanthus
707:Dielsiodoxa
630:Coleanthera
542:Acrothamnus
458:Styphelieae
444:(5 genera)
421:(2 genera)
398:(3 genera)
375:(3 genera)
325:(2 genera)
227:Description
221:Australasia
212:. The name
154:Subfamily:
102:Angiosperms
1577:Categories
1347:(5): 600,
1286:2022-10-21
1234:2022-10-21
1210:2022-10-21
1113:References
1106:Leucopogon
1048:(Richeeae)
905:Sprengelia
900:(Richeeae)
894:Sphenotoma
889:(Richeeae)
861:Planocarpa
762:Leucopogon
724:(Richeeae)
685:Cyathopsis
667:J.M.Powell
619:Brachyloma
586:Androstoma
575:Andersonia
553:Acrotriche
442:Epacrideae
396:Cosmelieae
348:(1 genus:
346:Archerieae
323:Prionoteae
1533:Q87132837
1367:Cyathodes
1337:Lissanthe
1100:Styphelia
916:Styphelia
872:Prionotes
817:Montitega
795:Melichrus
773:Lissanthe
674:Cyathodes
634:Stschegl.
608:Astroloma
301:Phylogeny
268:In 1810,
241:epidermis
237:lignified
210:Ericaceae
148:Ericaceae
72:Kingdom:
1527:Wikidata
1480:324935-2
1445:Wikidata
1399:group",
1397:Monotoca
1369:group",
1322:25065335
1266:, p. 320
1184:35699816
1085:Tasmania
942:F.Muell.
938:Woollsia
832:L.Watson
806:Monotoca
784:Lysinema
700:F.Muell.
696:Decatoca
663:Croninia
652:Cosmelia
597:Archeria
564:Agiortia
419:Richeeae
350:Archeria
264:Taxonomy
186:Synonyms
177:See text
144:Family:
138:Ericales
128:Asterids
115:Eudicots
1451:Q781231
1176:4354425
729:Epacris
722:Labill.
678:Labill.
601:Hook.f.
590:Hook.f.
257:stamens
253:corolla
247:on the
134:Order:
76:Plantae
1506:266539
1493:217036
1320:
1182:
1174:
1095:Hawaii
883:Richea
645:Benth.
533:et al.
527:Genera
255:. The
245:bracts
1547:7NM3H
1318:JSTOR
1302:Taxon
1180:S2CID
1172:JSTOR
931:R.Br.
887:R.Br.
876:R.Br.
854:R.Br.
843:R.Br.
810:R.Br.
799:R.Br.
788:R.Br.
777:R.Br.
766:R.Br.
744:Endl.
711:Albr.
656:R.Br.
623:Sond.
612:R.Br.
579:R.Br.
568:Quinn
557:R.Br.
546:Quinn
239:leaf
217:Sweet
196:Sweet
122:Clade
109:Clade
96:Clade
83:Clade
1560:1736
1555:GRIN
1488:NCBI
1475:IPNI
733:Cav.
163:Arn.
1542:CoL
1409:doi
1379:doi
1349:doi
1310:doi
1257:doi
1164:doi
1109:).
1093:),
920:Sm.
909:Sm.
1579::
1557::
1544::
1529::
1503::
1490::
1477::
1462::
1447::
1405:23
1403:,
1375:18
1373:,
1345:16
1343:,
1316:,
1306:54
1304:,
1253:17
1251:,
1227:,
1203:,
1191:^
1178:,
1170:,
1160:68
1158:,
1141:^
1132:,
1121:^
1083:,
352:)
297:.
179:.
124::
111::
98::
85::
1411::
1381::
1351::
1312::
1259::
1166::
1097:(
20:)
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