142:
801:
3371:
3394:
3386:
2790:
452:
3379:
4254:
2175:
773:
168:
120:
787:
4240:
568:
4303:
2775:
2783:
815:
4247:
646:. Despite the broad distribution of this characteristic, its absence in some suchians makes it ambiguous whether it qualifies as a synapomorphy of Suchia, or alternatively evolved in several independent lineages within the group. Ezcurra also agreed with one of Nesbitt (2011)'s synapomorphies of the calcaneum, namely the calcaneal tuber being wider than tall.
506:. In early archosauriforms, the calcaneal tuber was taller than wide, while later archosaurs and archosaur relatives had a calcaneal tuber which was circular in cross-section. Suchians take this one step further, as early members of the group had calcaneal tubers which were much wider than tall. Although bipedal poposauroids and most loricatans (
407:. The two most comprehensive recent studies on archosaur relationships, Nesbitt (2011) and Ezcurra (2016), each came to different conclusions on what synapomorphies characterize Suchia. Nevertheless, they did agree on one trait that likely qualified as a synapomorphy of Suchia: a calcaneal tuber which was significantly wider than it was tall.
415:
As Suchia is a clade, it is defined by relationships rather than shared characteristics between its members. However, several traits are found in most suchians but not in ornithosuchids, phytosaurs, or other archosaurs and archosaur relatives, so they can be used to help determine whether a reptile
1710:
Ezcurra, Martín D.; Fiorelli, Lucas E.; Martinelli, Agustín G.; Rocher, Sebastián; Baczko, M. Belén von; Ezpeleta, Miguel; Taborda, Jeremías R. A.; Hechenleitner, E. Martín; Trotteyn, M. Jimena (11 September 2017). "Deep faunistic turnovers preceded the rise of dinosaurs in southwestern
Pangaea".
663:
united by the possession of a crurotarsal ankle, with
Pseudosuchia and Crocodilia as independent suborders within it. However, newer studies starting in the 1980s have significantly altered the traditional classification scheme of crocodilian ancestors. Rather than pseudosuchians and crocodilians
423:'s 1991 study on archosaur relationships argued that many of them were also present in ornithosuchids, phytosaurs, or avemetatarsalians, so they could not be considered traits which diagnose Suchia (under his definition of the group). Instead, Sereno listed a single synapomorphy for Suchia. The
1889:
Nesbitt, Sterling J.; Sidor, Christian A.; Angielczyk, Kenneth D.; Smith, Roger M. H.; Tsuji, Linda A. (November 2014). "A new archosaur from the Manda beds (Anisian, Middle
Triassic) of southern Tanzania and its implications for character state optimizations at Archosauria and Pseudosuchia".
539:(thigh bone), but this characteristic is also known in some archosaur relatives so it is unclear whether the presence of the trait in suchians was a synapomorphy or a plesiomorphy ("primitive" trait retained from an earlier ancestor). In addition, most
402:
There is some controversy over which traits, if any, can be used to distinguish suchians from non-suchian archosaurs. Anatomical features which evolve at the base of a group, and can thus be used to characterize the group, are known as
472:(cheek bone, below the eye) has a low, rounded longitudinal ridge in suchians, contrasting with other archosaurs which typically have no ridge, or a sharp ridge instead. As is the case for many early archosaurs, the structure of the
642:, is actually taller than the middle portion of the rear branch. This trait, formally known as the posterodorsal process of the maxilla, has been considered a synapomorphy of gracilisuchids and is also present in aetosaurs and
484:(inner ankle bone) by means of two distinct surfaces, the convex fibular condyle and the concave astragalar facet. In other archosaurs, these surfaces are continuous, rather than separate. Moreover, the fibular condyle is
658:
in 1974. His intention was to link the
Triassic pseudosuchians (which at that time were considered to have left no descendants) with the true crocodilians that would evolve later in the Mesozoic. Suchia was named as an
668:
as Krebs believed, the crocodilians are now considered to be a subset of
Pseudosuchia. This necessitated Suchia to acquire a new definition, as Kreb's original classification scheme was invalidated.
671:
Sereno (1991) assigned the name Suchia to a clade containing most pseudosuchians, which had previously been found by Benton & Clark (1988). Under this new definition, Suchia included
416:
is a suchian or not. These traits are spread out throughout the body, but different paleontologists disagree on whether they were true synapomorphies (derived distinguishing features).
435:
at the rear part of the skull, is short. This has the effect of making the lower temporal fenestra triangular in shape. However, Sereno also noted that this feature was not present in
502:. A hemicylindrical fibular condyle is also present in phytosaurs, likely convergently. Lastly, the rear edge of the calcaneum has a backwards-extending tube of bone known as a
399:(which are suchians by definition of the group). Phytosaurs are also excluded from Suchia, although it is not certain whether they qualify as pseudosuchians in the first place.
614:
which was known from significantly less fossil material than most other Suchia. These traits are the presence of large, well-rimmed pits on either side of the dorsal (back)
4403:
693:, and all descendants of their common ancestor. Nesbitt (2011) altered Sereno's definition slightly, as he defined Suchia as the least inclusive clade containing
543:
reverted to the state present in other archosaurs: a rounded medial condyle. Many of
Nesbitt (2011)'s suchian synapomorphies are not present in
2208:
535:(inner forearm bone), although the distribution of this characteristic is poorly understood. Likewise, some early suchians have a triangular
1935:"New clade of enigmatic early archosaurs yields insights into early pseudosuchian phylogeny and the biogeography of the archosaur radiation"
3393:
3370:
2789:
2161:
800:
1791:"The phylogenetic relationships of basal archosauromorphs, with an emphasis on the systematics of proterosuchian archosauriforms"
4253:
3385:
384:(the living Nile crocodile) by Nesbitt (2011). Generally the only pseudosuchian group which is omitted from Suchia is the
3378:
623:
772:
786:
4239:
2201:
2061:
Butler, R.J.; Brusatte, S.J.; Reich, M.; Nesbitt, S.J.; Schoch, R.R.; Hornung, J.J. (2011). Andrew A. Farke (ed.).
404:
141:
2314:
2774:
1933:
Butler, Richard J; Sullivan, Corwin; Ezcurra, Martín D; Liu, Jun; Lecuona, Agustina; Sookias, Roland B (2014).
167:
4398:
4306:
3677:
2124:
Hans-Dieter, Sues (10 June 1992). "A Remarkable New
Armored Archosaur from the Upper Triassic of Virginia".
536:
2067:
from the Latest Early
Triassic of Germany and the Timing and Biogeography of the Early Archosaur Radiation"
4323:
2864:
2782:
577:. Note the rounded ridge on the jugal bone beneath the eye (one of Nesbitt 's synapomorphies) visible in
516:) had circular calcaneal tubers, these features were likely reversions, since the earliest poposauroids (
4370:
2194:
2165:
730:
4171:
2133:
2078:
2009:
1946:
1899:
1860:
1720:
814:
713:
380:
3870:
461:, showing several of the calcaneal features Nesbitt (2011) considered to be synapomorphic for Suchia
451:
2820:
2807:
2741:
2673:
2287:
2180:
1210:
639:
556:
503:
419:
While Benton & Clark (1988) found numerous characteristics for the clade later renamed Suchia,
331:
468:'s major 2011 analysis of early archosaurs found several more stable syapomorphies of Suchia. The
4246:
4200:
3537:
2025:
1915:
1762:
1689:
162:
485:
4375:
4051:
4015:
3527:
2106:
1982:
1964:
1830:
1812:
1754:
1746:
1681:
630:(upper hip bone) which has a moderate length, longer than it is high, but not longer than the
581:, and the posterodorsal process of the maxilla (one of Ezcurra 's synapomorphies) visible in
4096:
3965:
3948:
3725:
2897:
2299:
2141:
2096:
2086:
2017:
1972:
1954:
1907:
1868:
1820:
1802:
1736:
1728:
1671:
1661:
1257:
840:
722:
597:
465:
424:
385:
119:
631:
4071:
3826:
3694:
3615:
3592:
3567:
3425:
3412:
3337:
3325:
3317:
2969:
2692:
2624:
2436:
1588:
1492:
970:
690:
660:
499:
489:
388:
326:
318:
291:
2137:
2082:
2013:
1950:
1903:
1864:
1724:
610:. As a result, his diagnosis for Suchia depended primarily on several traits visible in
4276:
4133:
4078:
4036:
3995:
3848:
3819:
3798:
3666:
3547:
3456:
3132:
2934:
2927:
2920:
2651:
2564:
2474:
2401:
2101:
2062:
1977:
1934:
1825:
1790:
1078:
726:
718:
593:
428:
392:
310:
128:
39:
488:(half-cylinder shaped) in contrast to the more spherical structure of ornithosuchids,
4392:
4112:
4026:
4005:
3901:
3789:
3765:
3755:
3715:
3652:
3608:
3601:
3507:
3497:
3476:
3466:
3280:
3265:
3194:
3174:
3081:
3061:
2987:
2906:
2878:
2851:
2723:
2701:
2644:
2518:
2460:
1919:
1852:
1430:
1320:
1220:
1054:
1037:
1001:
919:
754:
673:
602:
573:
532:
524:
476:(outer heel bone) is important for diagnosing Suchia. This bone connects to both the
457:
360:
and their extinct relatives). It was defined as the least inclusive clade containing
270:
77:
2029:
1766:
1693:
4361:
4286:
4126:
4064:
3924:
3910:
3812:
3775:
3745:
3638:
3629:
3557:
3295:
3287:
3251:
3235:
3227:
3218:
3204:
3164:
3143:
3101:
3051:
2960:
2941:
2913:
2730:
2637:
2603:
2596:
2589:
2550:
2453:
2445:
2380:
2367:
2326:
2217:
2145:
1872:
1477:
1454:
1382:
1197:
1129:
960:
844:
707:
686:
665:
655:
627:
626:
portion when seen from the front, and a preacetabular process (front blade) of the
551:
508:
439:, which he excluded from the group. Later studies found support for a placement of
374:
353:
301:
235:
150:
2091:
1911:
1650:"The Early Evolution of Archosaurs: Relationships and the Origin of Major Clades"
391:, although at least one analysis classifies ornithosuchids as close relatives of
4355:
4291:
4160:
4105:
3975:
3932:
3917:
3886:
3879:
3856:
3840:
3833:
3805:
3486:
3258:
3184:
3153:
3121:
3111:
3041:
3010:
3003:
2949:
2886:
2837:
2716:
2573:
2539:
2529:
2507:
2467:
2411:
1406:
1234:
1146:
984:
950:
902:
864:
545:
518:
513:
420:
281:
52:
4346:
4296:
4281:
4271:
4206:
4177:
4148:
3985:
3578:
3302:
3273:
3243:
3091:
3071:
2976:
2871:
2272:
2247:
2170:
1732:
1304:
1280:
1105:
1091:
878:
854:
701:
695:
682:
678:
494:
481:
469:
368:
362:
357:
97:
62:
1968:
1959:
1816:
1750:
1685:
4189:
4119:
3517:
2421:
2256:
473:
349:
222:
179:
102:
46:
2110:
1986:
1834:
1758:
567:
1853:"Basal Archosaurs: Phylogenetic Relationships and Functional Implications"
4340:
4085:
3735:
3645:
3446:
3025:
2662:
2610:
2241:
1368:
733:
615:
540:
531:
Some suchians have a shallow longitudinal groove on the rear edge of the
396:
199:
155:
133:
92:
87:
72:
67:
57:
3310:
2996:
2708:
2581:
2021:
1807:
1741:
635:
619:
209:
107:
82:
17:
2235:
1676:
477:
189:
4317:
2045:
A cladistic analysis of the higher systematic categories of
Diapsida
638:'s posterior process (rear branch), near the rear lower tip of the
1666:
1649:
566:
450:
432:
345:
600:
provided an alternative diagnosis for Suchia. Ezcurra found that
549:, an unusual crurotarsan sometimes considered a suchian close to
606:
was a member of the group, but also argued that Suchia excluded
443:
deep within Suchia, so Sereno's synapomorphy is likely invalid.
4321:
3410:
2805:
2365:
2228:
2190:
2186:
736:
654:
Suchia was originally formalized by German paleontologist
2047:(Ph.D. Dissertation). University of California Berkeley.
744:, Poposauroidea, Loricata, and possibly Erpetosuchidae,
634:. He also noted that in some suchians, the tip of the
4157:
4048:
4032:
4022:
4011:
4001:
3991:
3981:
3971:
3961:
3898:
3867:
3786:
3771:
3761:
3751:
3741:
3731:
3721:
3711:
3663:
3626:
3589:
3574:
3563:
3553:
3543:
3533:
3523:
3513:
3503:
3493:
3482:
3472:
3462:
3452:
3442:
3215:
3200:
3190:
3180:
3170:
3160:
3149:
3139:
3128:
3117:
3107:
3097:
3087:
3077:
3067:
3057:
3047:
3037:
2848:
2833:
2689:
2621:
2561:
2546:
2535:
2525:
2514:
2503:
2433:
2417:
2407:
2397:
323:
315:
307:
297:
288:
277:
266:
4330:
4188:
4156:
4147:
4095:
4047:
3946:
3897:
3866:
3785:
3692:
3662:
3625:
3588:
3423:
3336:
3214:
3024:
2986:
2959:
2896:
2847:
2818:
2740:
2688:
2661:
2620:
2560:
2488:
2432:
2378:
2325:
2298:
2271:
1654:Bulletin of the American Museum of Natural History
2202:
1884:
1882:
1643:
1641:
1639:
1637:
1635:
1633:
1631:
1629:
8:
2056:
2054:
835:Below is a phylogenetic cladogram by Butler
763:Taxa used by Nesbitt (2011) to define Suchia
395:(which are usually considered suchians) and
2000:Krebs, Bernard (1974). "Die Archosaurier".
4318:
4153:
4056:
3954:
3704:
3700:
3435:
3431:
3420:
3407:
3030:
2856:
2826:
2815:
2802:
2629:
2496:
2390:
2386:
2375:
2362:
2304:
2277:
2268:
2225:
2209:
2195:
2187:
140:
118:
31:
2100:
2090:
1976:
1958:
1824:
1806:
1740:
1675:
1665:
622:(upper arm bone) which has a symmetrical
431:bar, which separates the upper and lower
4404:Extant Early Triassic first appearances
1625:
768:
7:
1846:
1844:
1784:
1782:
1780:
1778:
1776:
1705:
1703:
1470:
1446:
1422:
1398:
1374:
1296:
1272:
1250:
1226:
1216:
1189:
1121:
1097:
1070:
1029:
976:
966:
956:
894:
870:
860:
850:
847:: Clade names follow Nesbitt 2011.
2126:Journal of Vertebrate Paleontology
1892:Journal of Vertebrate Paleontology
1857:Journal of Vertebrate Paleontology
839:in 2011 showing the cladistics of
528:) retained wide calcaneal tubers.
492:, and archosaur relatives such as
455:The left calcaneum (heel bone) of
25:
1789:Ezcurra, Martín D. (2016-04-28).
4302:
4301:
4252:
4245:
4238:
3392:
3384:
3377:
3369:
2788:
2781:
2773:
2173:
813:
799:
785:
771:
166:
563:Ezcurra (2016)'s synapomorphies
522:, for example) and loricatans (
447:Nesbitt (2011)'s synapomorphies
2146:10.1080/02724634.1992.10011444
1873:10.1080/02724634.1991.10011426
1713:Nature Ecology & Evolution
562:
446:
1:
2092:10.1371/journal.pone.0025693
1912:10.1080/02724634.2014.859622
674:Gracilisuchus stipanicicorum
721:). Suchia, therefore, is a
537:medial condyle of the femur
352:containing the majority of
4420:
129:Paratypothorax andressorum
4266:
4235:
4059:
3957:
3707:
3703:
3438:
3434:
3419:
3406:
3366:
3033:
2859:
2829:
2814:
2801:
2770:
2632:
2499:
2393:
2389:
2374:
2361:
2307:
2280:
2267:
2224:
2063:"The Sail-Backed Reptile
1733:10.1038/s41559-017-0305-5
1586:
1490:
1475:
1468:
1451:
1444:
1427:
1420:
1403:
1396:
1379:
1372:
1318:
1301:
1294:
1277:
1270:
1255:
1248:
1231:
1224:
1214:
1194:
1187:
1143:
1126:
1119:
1102:
1095:
1075:
1068:
1051:
1034:
1027:
998:
981:
974:
964:
954:
916:
899:
892:
875:
868:
858:
807:Prestosuchus chiniquensis
708:Prestosuchus chiniquensis
375:Prestosuchus chiniquensis
263:
258:
163:Scientific classification
161:
151:Prestosuchus chiniquensis
148:
139:
126:
117:
34:
1960:10.1186/1471-2148-14-128
1939:BMC Evolutionary Biology
1851:Sereno, Paul C. (1991).
2043:Gauthier, J.A. (1984).
2865:Benggwigwishingasuchus
1648:Nesbitt, S.J. (2011).
590:
480:(outer shin bone) and
462:
4371:Paleobiology Database
2166:Paleobiology Database
843:, focusing mostly on
793:Rauisuchus tiradentes
702:Rauisuchus tiradentes
570:
454:
369:Rauisuchus tiradentes
1859:. 11 sup 004: 1–53.
821:Crocodylus niloticus
714:Crocodylus niloticus
596:'s 2016 analysis of
381:Crocodylus niloticus
2821:Paracrocodylomorpha
2808:Paracrocodylomorpha
2742:Paracrocodylomorpha
2181:Paleontology portal
2138:1992JVPal..12..142S
2083:2011PLoSO...625693B
2014:1974NW.....61...17K
2002:Naturwissenschaften
1951:2014BMCEE..14..128B
1904:2014JVPal..34.1357N
1865:1991JVPal..11S...1S
1725:2017NatEE...1.1477E
1211:Paracrocodylomorpha
779:Aetosaurus ferratus
696:Aetosaurus ferratus
640:antorbital fenestra
557:Paracrocodylomorpha
363:Aetosaurus ferratus
332:Paracrocodylomorpha
3538:Pseudhesperosuchus
2022:10.1007/BF00602887
1808:10.7717/peerj.1778
591:
463:
433:temporal fenestrae
4386:
4385:
4324:Taxon identifiers
4315:
4314:
4262:
4261:
4233:
4232:
4229:
4228:
4225:
4224:
4221:
4220:
4217:
4216:
4143:
4142:
4052:Shartegosuchoidea
4016:Pachycheilosuchus
3942:
3941:
3688:
3687:
3528:Phyllodontosuchus
3402:
3401:
3364:
3363:
3360:
3359:
3356:
3355:
3020:
3019:
2797:
2796:
2768:
2767:
2764:
2763:
2760:
2759:
2684:
2683:
2484:
2483:
2357:
2356:
2353:
2352:
2349:
2348:
2345:
2344:
1719:(10): 1477–1483.
1617:
1616:
1608:
1607:
1599:
1598:
1575:
1574:
1566:
1565:
1557:
1556:
1548:
1547:
1539:
1538:
1530:
1529:
1521:
1520:
1512:
1511:
1503:
1502:
1358:
1357:
1349:
1348:
1340:
1339:
1331:
1330:
1176:
1175:
1167:
1166:
1158:
1157:
1013:
1012:
940:
939:
931:
930:
729:and several more
500:proterochampsians
490:avemetatarsalians
339:
338:
254:
27:Clade of reptiles
16:(Redirected from
4411:
4379:
4378:
4366:
4365:
4364:
4351:
4350:
4349:
4319:
4305:
4304:
4257:
4256:
4249:
4243:
4242:
4159:
4154:
4097:Shartegosuchidae
4057:
4050:
4034:
4024:
4013:
4003:
3993:
3983:
3973:
3966:Dianchungosaurus
3963:
3955:
3949:Mesoeucrocodylia
3900:
3869:
3788:
3773:
3763:
3753:
3743:
3733:
3726:Eopneumatosuchus
3723:
3713:
3705:
3701:
3665:
3628:
3591:
3576:
3565:
3555:
3545:
3535:
3525:
3515:
3505:
3495:
3484:
3474:
3464:
3454:
3444:
3436:
3432:
3421:
3408:
3397:
3396:
3389:
3388:
3382:
3381:
3374:
3373:
3217:
3202:
3192:
3182:
3172:
3162:
3151:
3141:
3130:
3119:
3109:
3099:
3089:
3079:
3069:
3059:
3049:
3039:
3031:
2898:Ctenosauriscidae
2857:
2850:
2835:
2827:
2816:
2803:
2792:
2786:
2785:
2778:
2777:
2691:
2630:
2623:
2563:
2548:
2537:
2527:
2516:
2505:
2497:
2435:
2419:
2409:
2399:
2391:
2387:
2376:
2363:
2315:Archosauromorpha
2305:
2300:Archosauromorpha
2278:
2269:
2262:
2261:
2226:
2211:
2204:
2197:
2188:
2183:
2178:
2177:
2176:
2150:
2149:
2121:
2115:
2114:
2104:
2094:
2058:
2049:
2048:
2040:
2034:
2033:
1997:
1991:
1990:
1980:
1962:
1930:
1924:
1923:
1898:(6): 1357–1382.
1886:
1877:
1876:
1848:
1839:
1838:
1828:
1810:
1786:
1771:
1770:
1744:
1707:
1698:
1697:
1679:
1669:
1645:
1471:
1447:
1423:
1399:
1375:
1297:
1273:
1258:Ctenosauriscidae
1251:
1227:
1217:
1190:
1122:
1098:
1071:
1030:
977:
967:
957:
895:
871:
861:
851:
841:Archosauriformes
817:
803:
789:
775:
723:node-based taxon
598:archosauromorphs
466:Sterling Nesbitt
325:
317:
309:
299:
290:
279:
268:
252:
171:
170:
144:
122:
112:
49:
38:Temporal range:
32:
21:
4419:
4418:
4414:
4413:
4412:
4410:
4409:
4408:
4389:
4388:
4387:
4382:
4374:
4369:
4360:
4359:
4354:
4345:
4344:
4339:
4326:
4316:
4311:
4258:
4251:
4237:
4213:
4184:
4139:
4091:
4072:Shantungosuchus
4043:
3951:
3938:
3893:
3862:
3827:Hemiprotosuchus
3781:
3697:
3695:Crocodyliformes
3684:
3658:
3621:
3616:Terrestrisuchus
3593:Saltoposuchidae
3584:
3568:Syntomiprosopus
3428:
3426:Crocodylomorpha
3415:
3413:Crocodylomorpha
3398:
3391:
3383:
3376:
3368:
3352:
3338:Crocodylomorpha
3332:
3326:Vytshegdosuchus
3318:Vjushkovisaurus
3210:
3016:
2982:
2970:Dolichobrachium
2955:
2892:
2843:
2823:
2810:
2793:
2780:
2772:
2756:
2736:
2693:Gracilisuchidae
2680:
2657:
2625:Aetosauriformes
2616:
2556:
2493:
2480:
2437:Ornithosuchidae
2428:
2383:
2370:
2341:
2321:
2294:
2263:
2231:
2230:
2220:
2215:
2179:
2174:
2172:
2158:
2153:
2123:
2122:
2118:
2060:
2059:
2052:
2042:
2041:
2037:
1999:
1998:
1994:
1932:
1931:
1927:
1888:
1887:
1880:
1850:
1849:
1842:
1788:
1787:
1774:
1709:
1708:
1701:
1647:
1646:
1627:
1623:
1618:
1609:
1600:
1589:Avemetatarsalia
1576:
1567:
1558:
1549:
1540:
1531:
1522:
1513:
1504:
1493:Crocodylomorpha
1359:
1350:
1341:
1332:
1177:
1168:
1159:
1014:
971:Ornithosuchidae
941:
932:
833:
828:
827:
826:
823:
818:
809:
804:
795:
790:
781:
776:
765:
764:
727:crocodylomorphs
691:Crocodylomorpha
652:
565:
504:calcaneal tuber
486:hemicylindrical
449:
413:
389:Ornithosuchidae
327:Aetosauriformes
319:Gracilisuchidae
292:Ornithosuchidae
251:
165:
113:
111:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
44:
43:
36:
28:
23:
22:
15:
12:
11:
5:
4417:
4415:
4407:
4406:
4401:
4399:Pseudosuchians
4391:
4390:
4384:
4383:
4381:
4380:
4367:
4352:
4336:
4334:
4328:
4327:
4322:
4313:
4312:
4310:
4309:
4299:
4294:
4289:
4284:
4279:
4277:Prestosuchidae
4274:
4267:
4264:
4263:
4260:
4259:
4236:
4234:
4231:
4230:
4227:
4226:
4223:
4222:
4219:
4218:
4215:
4214:
4212:
4211:
4210:
4209:
4203:
4194:
4192:
4186:
4185:
4183:
4182:
4181:
4180:
4174:
4165:
4163:
4151:
4145:
4144:
4141:
4140:
4138:
4137:
4134:Shartegosuchus
4130:
4123:
4116:
4109:
4101:
4099:
4093:
4092:
4090:
4089:
4082:
4079:Sichuanosuchus
4075:
4068:
4060:
4054:
4045:
4044:
4042:
4041:
4037:Thalattosuchia
4030:
4020:
4009:
3999:
3996:Lusitanisuchus
3989:
3979:
3969:
3958:
3952:
3947:
3944:
3943:
3940:
3939:
3937:
3936:
3929:
3921:
3914:
3906:
3904:
3895:
3894:
3892:
3891:
3883:
3875:
3873:
3871:Notochampsidae
3864:
3863:
3861:
3860:
3853:
3849:Pedeticosaurus
3845:
3837:
3830:
3823:
3820:Erythrochampsa
3816:
3809:
3802:
3799:Coloradisuchus
3794:
3792:
3783:
3782:
3780:
3779:
3769:
3759:
3749:
3739:
3729:
3719:
3708:
3698:
3693:
3690:
3689:
3686:
3685:
3683:
3682:
3681:
3680:
3678:Thalattosuchia
3671:
3669:
3667:Thalattosuchia
3660:
3659:
3657:
3656:
3649:
3642:
3634:
3632:
3623:
3622:
3620:
3619:
3612:
3605:
3597:
3595:
3586:
3585:
3583:
3582:
3572:
3561:
3551:
3548:Redondavenator
3541:
3531:
3521:
3511:
3501:
3491:
3480:
3470:
3460:
3457:Dibothrosuchus
3450:
3439:
3429:
3424:
3417:
3416:
3411:
3404:
3403:
3400:
3399:
3367:
3365:
3362:
3361:
3358:
3357:
3354:
3353:
3351:
3350:
3349:
3348:
3342:
3340:
3334:
3333:
3331:
3330:
3322:
3314:
3307:
3299:
3292:
3284:
3277:
3270:
3262:
3255:
3248:
3240:
3232:
3223:
3221:
3212:
3211:
3209:
3208:
3198:
3188:
3178:
3168:
3158:
3147:
3137:
3133:Incertovenator
3126:
3115:
3105:
3095:
3085:
3075:
3065:
3055:
3045:
3034:
3028:
3022:
3021:
3018:
3017:
3015:
3014:
3007:
3000:
2992:
2990:
2984:
2983:
2981:
2980:
2973:
2965:
2963:
2957:
2956:
2954:
2953:
2946:
2938:
2935:Hypselorhachis
2931:
2928:Ctenosauriscus
2924:
2921:Bystrowisuchus
2917:
2910:
2902:
2900:
2894:
2893:
2891:
2890:
2883:
2875:
2868:
2860:
2854:
2845:
2844:
2842:
2841:
2830:
2824:
2819:
2812:
2811:
2806:
2799:
2798:
2795:
2794:
2771:
2769:
2766:
2765:
2762:
2761:
2758:
2757:
2755:
2754:
2753:
2752:
2746:
2744:
2738:
2737:
2735:
2734:
2727:
2720:
2713:
2705:
2697:
2695:
2686:
2685:
2682:
2681:
2679:
2678:
2677:
2676:
2667:
2665:
2659:
2658:
2656:
2655:
2652:Revueltosaurus
2648:
2641:
2633:
2627:
2618:
2617:
2615:
2614:
2607:
2600:
2593:
2586:
2578:
2569:
2567:
2565:Erpetosuchidae
2558:
2557:
2555:
2554:
2544:
2533:
2523:
2512:
2500:
2494:
2489:
2486:
2485:
2482:
2481:
2479:
2478:
2475:Venaticosuchus
2471:
2464:
2457:
2450:
2441:
2439:
2430:
2429:
2427:
2426:
2415:
2405:
2402:Dasygnathoides
2394:
2384:
2379:
2372:
2371:
2366:
2359:
2358:
2355:
2354:
2351:
2350:
2347:
2346:
2343:
2342:
2340:
2339:
2338:
2337:
2331:
2329:
2323:
2322:
2320:
2319:
2318:
2317:
2308:
2302:
2296:
2295:
2293:
2292:
2291:
2290:
2281:
2275:
2265:
2264:
2260:
2259:
2250:
2244:
2238:
2229:
2222:
2221:
2216:
2214:
2213:
2206:
2199:
2191:
2185:
2184:
2169:
2168:
2157:
2156:External links
2154:
2152:
2151:
2132:(2): 142–149.
2116:
2077:(10): e25693.
2065:Ctenosauriscus
2050:
2035:
1992:
1925:
1878:
1840:
1772:
1699:
1624:
1622:
1619:
1615:
1614:
1611:
1610:
1606:
1605:
1602:
1601:
1597:
1596:
1593:
1592:
1585:
1582:
1581:
1578:
1577:
1573:
1572:
1569:
1568:
1564:
1563:
1560:
1559:
1555:
1554:
1551:
1550:
1546:
1545:
1542:
1541:
1537:
1536:
1533:
1532:
1528:
1527:
1524:
1523:
1519:
1518:
1515:
1514:
1510:
1509:
1506:
1505:
1501:
1500:
1497:
1496:
1489:
1486:
1485:
1482:
1481:
1474:
1469:
1467:
1464:
1463:
1460:
1459:
1450:
1445:
1443:
1440:
1439:
1436:
1435:
1426:
1421:
1419:
1416:
1415:
1412:
1411:
1402:
1397:
1395:
1392:
1391:
1388:
1387:
1378:
1373:
1371:
1365:
1364:
1361:
1360:
1356:
1355:
1352:
1351:
1347:
1346:
1343:
1342:
1338:
1337:
1334:
1333:
1329:
1328:
1325:
1324:
1317:
1314:
1313:
1310:
1309:
1300:
1295:
1293:
1290:
1289:
1286:
1285:
1276:
1271:
1269:
1266:
1265:
1262:
1261:
1254:
1249:
1247:
1244:
1243:
1240:
1239:
1230:
1225:
1223:
1215:
1213:
1207:
1206:
1203:
1202:
1193:
1188:
1186:
1183:
1182:
1179:
1178:
1174:
1173:
1170:
1169:
1165:
1164:
1161:
1160:
1156:
1155:
1152:
1151:
1142:
1139:
1138:
1135:
1134:
1125:
1120:
1118:
1115:
1114:
1111:
1110:
1101:
1096:
1094:
1088:
1087:
1084:
1083:
1079:Revueltosaurus
1074:
1069:
1067:
1064:
1063:
1060:
1059:
1050:
1047:
1046:
1043:
1042:
1033:
1028:
1026:
1020:
1019:
1016:
1015:
1011:
1010:
1007:
1006:
997:
994:
993:
990:
989:
980:
975:
973:
965:
963:
955:
953:
947:
946:
943:
942:
938:
937:
934:
933:
929:
928:
925:
924:
915:
912:
911:
908:
907:
898:
893:
891:
888:
887:
884:
883:
874:
869:
867:
859:
857:
849:
832:
829:
825:
824:
819:
812:
810:
805:
798:
796:
791:
784:
782:
777:
770:
767:
766:
762:
761:
760:
740:, Aetosauria,
738:Revueltosaurus
719:Nile crocodile
651:
650:Classification
648:
636:maxillary bone
632:pubic peduncle
612:Koilamasuchus,
594:Martin Ezcurra
564:
561:
448:
445:
412:
409:
405:synapomorphies
354:pseudosuchians
337:
336:
335:
334:
329:
321:
313:
311:Erpetosuchidae
305:
295:
286:
275:
261:
260:
256:
255:
246:
239:
238:
233:
226:
225:
220:
213:
212:
207:
203:
202:
197:
193:
192:
187:
183:
182:
177:
173:
172:
159:
158:
146:
145:
137:
136:
124:
123:
115:
114:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
50:
40:Early Triassic
37:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
4416:
4405:
4402:
4400:
4397:
4396:
4394:
4377:
4372:
4368:
4363:
4357:
4353:
4348:
4342:
4338:
4337:
4335:
4333:
4329:
4325:
4320:
4308:
4300:
4298:
4295:
4293:
4290:
4288:
4285:
4283:
4280:
4278:
4275:
4273:
4269:
4268:
4265:
4255:
4250:
4248:
4241:
4208:
4204:
4202:
4198:
4197:
4196:
4195:
4193:
4191:
4187:
4179:
4175:
4173:
4169:
4168:
4167:
4166:
4164:
4162:
4155:
4152:
4150:
4146:
4136:
4135:
4131:
4129:
4128:
4124:
4122:
4121:
4117:
4115:
4114:
4113:Fruitachampsa
4110:
4108:
4107:
4103:
4102:
4100:
4098:
4094:
4088:
4087:
4083:
4081:
4080:
4076:
4074:
4073:
4069:
4067:
4066:
4062:
4061:
4058:
4055:
4053:
4046:
4039:
4038:
4031:
4029:
4028:
4027:Titanochampsa
4021:
4018:
4017:
4010:
4008:
4007:
4006:Neuquensuchus
4000:
3998:
3997:
3990:
3988:
3987:
3980:
3978:
3977:
3970:
3968:
3967:
3960:
3959:
3956:
3953:
3950:
3945:
3935:
3934:
3930:
3927:
3926:
3922:
3920:
3919:
3915:
3913:
3912:
3908:
3907:
3905:
3903:
3902:Gobiosuchidae
3896:
3889:
3888:
3884:
3882:
3881:
3877:
3876:
3874:
3872:
3865:
3859:
3858:
3854:
3851:
3850:
3846:
3843:
3842:
3838:
3836:
3835:
3831:
3829:
3828:
3824:
3822:
3821:
3817:
3815:
3814:
3810:
3808:
3807:
3803:
3801:
3800:
3796:
3795:
3793:
3791:
3790:Protosuchidae
3784:
3778:
3777:
3770:
3768:
3767:
3766:Stegomosuchus
3760:
3758:
3757:
3756:Platyognathus
3750:
3748:
3747:
3740:
3738:
3737:
3730:
3728:
3727:
3720:
3718:
3717:
3716:Entradasuchus
3710:
3709:
3706:
3702:
3699:
3696:
3691:
3679:
3675:
3674:
3673:
3672:
3670:
3668:
3661:
3655:
3654:
3653:Macelognathus
3650:
3648:
3647:
3643:
3641:
3640:
3636:
3635:
3633:
3631:
3624:
3618:
3617:
3613:
3611:
3610:
3609:Saltoposuchus
3606:
3604:
3603:
3602:Litargosuchus
3599:
3598:
3596:
3594:
3587:
3581:
3580:
3573:
3570:
3569:
3562:
3560:
3559:
3552:
3550:
3549:
3542:
3540:
3539:
3532:
3530:
3529:
3522:
3520:
3519:
3512:
3510:
3509:
3508:Kayentasuchus
3502:
3500:
3499:
3498:Junggarsuchus
3492:
3489:
3488:
3481:
3479:
3478:
3477:Hesperosuchus
3471:
3469:
3468:
3467:Dromicosuchus
3461:
3459:
3458:
3451:
3449:
3448:
3441:
3440:
3437:
3433:
3430:
3427:
3422:
3418:
3414:
3409:
3405:
3395:
3390:
3387:
3380:
3372:
3346:
3345:
3344:
3343:
3341:
3339:
3335:
3328:
3327:
3323:
3320:
3319:
3315:
3313:
3312:
3308:
3305:
3304:
3300:
3298:
3297:
3293:
3290:
3289:
3285:
3283:
3282:
3281:Scolotosuchus
3278:
3276:
3275:
3271:
3268:
3267:
3266:Procerosuchus
3263:
3261:
3260:
3256:
3254:
3253:
3249:
3246:
3245:
3241:
3238:
3237:
3233:
3230:
3229:
3225:
3224:
3222:
3220:
3213:
3207:
3206:
3199:
3197:
3196:
3195:Stagonosuchus
3189:
3187:
3186:
3179:
3177:
3176:
3175:Procerosuchus
3169:
3167:
3166:
3159:
3156:
3155:
3148:
3146:
3145:
3138:
3135:
3134:
3127:
3124:
3123:
3116:
3114:
3113:
3106:
3104:
3103:
3096:
3094:
3093:
3086:
3084:
3083:
3082:Decuriasuchus
3076:
3074:
3073:
3066:
3064:
3063:
3062:Batrachotomus
3056:
3054:
3053:
3046:
3044:
3043:
3036:
3035:
3032:
3029:
3027:
3023:
3013:
3012:
3008:
3006:
3005:
3001:
2999:
2998:
2994:
2993:
2991:
2989:
2988:Shuvosauridae
2985:
2979:
2978:
2974:
2972:
2971:
2967:
2966:
2964:
2962:
2958:
2952:
2951:
2947:
2944:
2943:
2939:
2937:
2936:
2932:
2930:
2929:
2925:
2923:
2922:
2918:
2916:
2915:
2911:
2909:
2908:
2907:Arizonasaurus
2904:
2903:
2901:
2899:
2895:
2889:
2888:
2884:
2881:
2880:
2879:Schultzsuchus
2876:
2874:
2873:
2869:
2867:
2866:
2862:
2861:
2858:
2855:
2853:
2852:Poposauroidea
2846:
2840:
2839:
2832:
2831:
2828:
2825:
2822:
2817:
2813:
2809:
2804:
2800:
2791:
2787:
2784:
2776:
2750:
2749:
2748:
2747:
2745:
2743:
2739:
2733:
2732:
2728:
2726:
2725:
2724:Turfanosuchus
2721:
2719:
2718:
2714:
2711:
2710:
2706:
2704:
2703:
2702:Gracilisuchus
2699:
2698:
2696:
2694:
2687:
2675:
2671:
2670:
2669:
2668:
2666:
2664:
2660:
2654:
2653:
2649:
2647:
2646:
2645:Euscolosuchus
2642:
2640:
2639:
2635:
2634:
2631:
2628:
2626:
2619:
2613:
2612:
2608:
2606:
2605:
2601:
2599:
2598:
2594:
2592:
2591:
2587:
2584:
2583:
2579:
2576:
2575:
2571:
2570:
2568:
2566:
2559:
2553:
2552:
2545:
2542:
2541:
2534:
2532:
2531:
2524:
2521:
2520:
2519:Koilamasuchus
2513:
2510:
2509:
2502:
2501:
2498:
2495:
2492:
2487:
2477:
2476:
2472:
2470:
2469:
2465:
2463:
2462:
2461:Ornithosuchus
2458:
2456:
2455:
2451:
2448:
2447:
2443:
2442:
2440:
2438:
2431:
2424:
2423:
2416:
2414:
2413:
2406:
2404:
2403:
2396:
2395:
2392:
2388:
2385:
2382:
2377:
2373:
2369:
2364:
2360:
2335:
2334:
2333:
2332:
2330:
2328:
2324:
2316:
2312:
2311:
2310:
2309:
2306:
2303:
2301:
2297:
2289:
2285:
2284:
2283:
2282:
2279:
2276:
2274:
2270:
2266:
2258:
2254:
2251:
2249:
2245:
2243:
2239:
2237:
2233:
2232:
2227:
2223:
2219:
2212:
2207:
2205:
2200:
2198:
2193:
2192:
2189:
2182:
2171:
2167:
2163:
2160:
2159:
2155:
2147:
2143:
2139:
2135:
2131:
2127:
2120:
2117:
2112:
2108:
2103:
2098:
2093:
2088:
2084:
2080:
2076:
2072:
2068:
2066:
2057:
2055:
2051:
2046:
2039:
2036:
2031:
2027:
2023:
2019:
2015:
2011:
2007:
2003:
1996:
1993:
1988:
1984:
1979:
1974:
1970:
1966:
1961:
1956:
1952:
1948:
1944:
1940:
1936:
1929:
1926:
1921:
1917:
1913:
1909:
1905:
1901:
1897:
1893:
1885:
1883:
1879:
1874:
1870:
1866:
1862:
1858:
1854:
1847:
1845:
1841:
1836:
1832:
1827:
1822:
1818:
1814:
1809:
1804:
1800:
1796:
1792:
1785:
1783:
1781:
1779:
1777:
1773:
1768:
1764:
1760:
1756:
1752:
1748:
1743:
1738:
1734:
1730:
1726:
1722:
1718:
1714:
1706:
1704:
1700:
1695:
1691:
1687:
1683:
1678:
1673:
1668:
1667:10.1206/352.1
1663:
1659:
1655:
1651:
1644:
1642:
1640:
1638:
1636:
1634:
1632:
1630:
1626:
1620:
1613:
1612:
1604:
1603:
1595:
1594:
1591:
1590:
1584:
1583:
1580:
1579:
1571:
1570:
1562:
1561:
1553:
1552:
1544:
1543:
1535:
1534:
1526:
1525:
1517:
1516:
1508:
1507:
1499:
1498:
1495:
1494:
1488:
1487:
1484:
1483:
1480:
1479:
1473:
1472:
1466:
1465:
1462:
1461:
1458:
1457:
1456:
1449:
1448:
1442:
1441:
1438:
1437:
1434:
1433:
1432:
1431:Batrachotomus
1425:
1424:
1418:
1417:
1414:
1413:
1410:
1409:
1408:
1401:
1400:
1394:
1393:
1390:
1389:
1386:
1385:
1384:
1377:
1376:
1370:
1367:
1366:
1363:
1362:
1354:
1353:
1345:
1344:
1336:
1335:
1327:
1326:
1323:
1322:
1321:Shuvosauridae
1316:
1315:
1312:
1311:
1308:
1307:
1306:
1299:
1298:
1292:
1291:
1288:
1287:
1284:
1283:
1282:
1275:
1274:
1268:
1267:
1264:
1263:
1260:
1259:
1253:
1252:
1246:
1245:
1242:
1241:
1238:
1237:
1236:
1229:
1228:
1222:
1221:Poposauroidea
1219:
1218:
1212:
1209:
1208:
1205:
1204:
1201:
1200:
1199:
1192:
1191:
1185:
1184:
1181:
1180:
1172:
1171:
1163:
1162:
1154:
1153:
1150:
1149:
1148:
1141:
1140:
1137:
1136:
1133:
1132:
1131:
1124:
1123:
1117:
1116:
1113:
1112:
1109:
1108:
1107:
1100:
1099:
1093:
1090:
1089:
1086:
1085:
1082:
1081:
1080:
1073:
1072:
1066:
1065:
1062:
1061:
1058:
1057:
1056:
1055:Turfanosuchus
1049:
1048:
1045:
1044:
1041:
1040:
1039:
1038:Gracilisuchus
1032:
1031:
1025:
1022:
1021:
1018:
1017:
1009:
1008:
1005:
1004:
1003:
1002:Ornithosuchus
996:
995:
992:
991:
988:
987:
986:
979:
978:
972:
969:
968:
962:
959:
958:
952:
949:
948:
945:
944:
936:
935:
927:
926:
923:
922:
921:
920:Pseudopalatus
914:
913:
910:
909:
906:
905:
904:
897:
896:
890:
889:
886:
885:
882:
881:
880:
873:
872:
866:
863:
862:
856:
853:
852:
848:
846:
842:
838:
830:
822:
816:
811:
808:
802:
797:
794:
788:
783:
780:
774:
769:
759:
757:
756:
755:Euscolosuchus
751:
747:
746:Koilamasuchus
743:
739:
735:
732:
728:
724:
720:
716:
715:
710:
709:
704:
703:
698:
697:
692:
688:
684:
680:
676:
675:
669:
667:
666:sister groups
662:
657:
656:Bernard Krebs
649:
647:
645:
641:
637:
633:
629:
625:
621:
617:
613:
609:
605:
604:
603:Koilamasuchus
599:
595:
588:
584:
580:
576:
575:
574:Gracilisuchus
571:The skull of
569:
560:
558:
554:
553:
548:
547:
542:
538:
534:
529:
527:
526:
525:Batrachotomus
521:
520:
515:
511:
510:
505:
501:
497:
496:
491:
487:
483:
479:
475:
471:
467:
460:
459:
458:Batrachotomus
453:
444:
442:
438:
434:
430:
426:
422:
417:
410:
408:
406:
400:
398:
394:
393:erpetosuchids
390:
387:
383:
382:
377:
376:
371:
370:
365:
364:
359:
355:
351:
347:
343:
333:
330:
328:
322:
320:
314:
312:
306:
304:
303:
296:
293:
287:
284:
283:
276:
273:
272:
271:Koilamasuchus
265:
264:
262:
257:
250:
247:
244:
241:
240:
237:
234:
231:
228:
227:
224:
221:
218:
215:
214:
211:
208:
205:
204:
201:
198:
195:
194:
191:
188:
185:
184:
181:
178:
175:
174:
169:
164:
160:
157:
153:
152:
147:
143:
138:
135:
131:
130:
125:
121:
116:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
48:
41:
33:
30:
19:
4331:
4287:Sphenosuchia
4244:
4207:crocodilians
4178:notosuchians
4132:
4127:Nominosuchus
4125:
4118:
4111:
4104:
4084:
4077:
4070:
4065:Hadongsuchus
4063:
4035:
4025:
4014:
4004:
3994:
3984:
3974:
3964:
3931:
3925:Lisboasaurus
3923:
3916:
3911:Cassissuchus
3909:
3885:
3878:
3855:
3847:
3839:
3832:
3825:
3818:
3813:Edentosuchus
3811:
3804:
3797:
3776:Tagarosuchus
3774:
3764:
3754:
3746:Microchampsa
3744:
3734:
3724:
3714:
3651:
3644:
3639:Almadasuchus
3637:
3630:Hallopodidae
3614:
3607:
3600:
3577:
3566:
3558:Sphenosuchus
3556:
3546:
3536:
3526:
3516:
3506:
3496:
3485:
3475:
3465:
3455:
3445:
3375:
3324:
3316:
3309:
3301:
3296:Teratosaurus
3294:
3288:Scythosuchus
3286:
3279:
3272:
3264:
3257:
3252:Polonosuchus
3250:
3242:
3236:Energosuchus
3234:
3228:Arganasuchus
3226:
3219:Rauisuchidae
3205:Youngosuchus
3203:
3193:
3183:
3173:
3165:Prestosuchus
3163:
3152:
3144:Luperosuchus
3142:
3131:
3120:
3110:
3102:Fasolasuchus
3100:
3090:
3080:
3070:
3060:
3052:Arganasuchus
3050:
3040:
3009:
3002:
2995:
2975:
2968:
2961:Poposauridae
2948:
2942:Tsylmosuchus
2940:
2933:
2926:
2919:
2914:Bromsgroveia
2912:
2905:
2885:
2877:
2870:
2863:
2836:
2779:
2731:Yonghesuchus
2729:
2722:
2715:
2707:
2700:
2650:
2643:
2638:Acaenasuchus
2636:
2609:
2604:Parringtonia
2602:
2597:Pagosvenator
2595:
2590:Erpetosuchus
2588:
2580:
2572:
2551:Ticinosuchus
2549:
2538:
2528:
2517:
2506:
2490:
2473:
2466:
2459:
2454:Dynamosuchus
2452:
2446:Aenigmaspina
2444:
2420:
2410:
2400:
2381:Pseudosuchia
2368:Pseudosuchia
2327:Pseudosuchia
2252:
2218:Pseudosuchia
2129:
2125:
2119:
2074:
2070:
2064:
2044:
2038:
2008:(1): 17–24.
2005:
2001:
1995:
1942:
1938:
1928:
1895:
1891:
1856:
1798:
1794:
1716:
1712:
1657:
1653:
1587:
1491:
1478:Rauisuchidae
1476:
1455:Fasolasuchus
1453:
1452:
1429:
1428:
1405:
1404:
1383:Prestosuchus
1381:
1380:
1319:
1303:
1302:
1279:
1278:
1256:
1233:
1232:
1198:Ticinosuchus
1196:
1195:
1145:
1144:
1130:Stagonolepis
1128:
1127:
1104:
1103:
1077:
1076:
1053:
1052:
1036:
1035:
1023:
1000:
999:
983:
982:
961:Pseudosuchia
918:
917:
901:
900:
877:
876:
845:Pseudosuchia
836:
834:
820:
806:
792:
778:
753:
749:
745:
742:Ticinosuchus
741:
737:
717:(the living
712:
706:
700:
694:
687:Poposauridae
672:
670:
653:
643:
611:
607:
601:
592:
586:
582:
578:
572:
552:Ticinosuchus
550:
544:
530:
523:
517:
509:Luperosuchus
507:
493:
464:
456:
441:Prestosuchus
440:
437:Prestosuchus
436:
418:
414:
401:
379:
373:
367:
361:
358:crocodilians
341:
340:
302:Ticinosuchus
300:
280:
269:
248:
242:
236:Pseudosuchia
229:
216:
149:
127:
29:
4356:Wikispecies
4292:Protosuchia
4172:Ziphosuchia
4161:Ziphosuchia
4106:Adzhosuchus
3976:Hsisosuchus
3933:Zaraasuchus
3918:Gobiosuchus
3887:Orthosuchus
3880:Notochampsa
3857:Protosuchus
3841:Orthosuchus
3834:Hoplosuchus
3806:Dianosuchus
3487:Heteropelta
3259:Postosuchus
3185:Saurosuchus
3154:Mandasuchus
3122:Heteropelta
3112:Heptasuchus
3042:Apatosuchus
3011:Sillosuchus
3004:Shuvosaurus
2950:Xilousuchus
2887:Qianosuchus
2838:Mambawakale
2717:Parvosuchus
2574:Archeopelta
2540:Nundasuchus
2530:Mandasuchus
2508:Heteropelta
2468:Riojasuchus
2422:Phytosauria
2412:Nundasuchus
2257:Archosauria
1742:11336/41466
1407:Saurosuchus
1235:Qianosuchus
1147:Longosuchus
985:Riojasuchus
951:Archosauria
903:Smilosuchus
865:Phytosauria
750:Nundasuchus
644:Qianosuchus
608:Nundasuchus
546:Nundasuchus
519:Qianosuchus
425:postorbital
421:Paul Sereno
411:Description
282:Nundasuchus
253:Krebs, 1974
223:Archosauria
154:, an early
45:248–0
4393:Categories
4297:Mesosuchia
4282:Bathyotica
4272:Rauisuchia
4270:See also:
4149:Metasuchia
3986:Induszalim
3579:Trialestes
3347:see below↓
3303:Tikisuchus
3274:Rauisuchus
3244:Jushatyria
3092:Etjosuchus
3072:Dagasuchus
2977:Poposaurus
2872:Lotosaurus
2751:see below↓
2674:Aetosauria
2663:Aetosauria
2336:see below↓
2288:Sauropsida
2273:Sauropsida
2248:Sauropsida
1945:(1): 128.
1621:References
1305:Lotosaurus
1281:Poposaurus
1106:Aetosaurus
1092:Aetosauria
879:Parasuchus
855:Crurotarsi
725:including
683:Rauisuchia
679:Aetosauria
541:loricatans
495:Euparkeria
482:astragalus
350:archosaurs
259:Subgroups
42:- Recent,
4205:includes
4201:Neosuchia
4190:Neosuchia
4176:includes
4120:Kyasuchus
3518:Parrishia
2234:Kingdom:
1969:1471-2148
1920:129558756
1817:2167-8359
1801:: e1778.
1751:2397-334X
1686:0003-0090
1677:2246/6112
831:Phylogeny
752:, and/or
616:vertebrae
514:crownward
474:calcaneum
429:squamosal
397:aetosaurs
186:Kingdom:
180:Eukaryota
156:loricatan
4347:Q1844946
4341:Wikidata
4307:Category
4086:Zosuchus
3736:Kemkemia
3646:Hallopus
3447:Carnufex
3026:Loricata
2611:Tarjadia
2242:Chordata
2240:Phylum:
2236:Animalia
2111:22022431
2071:PLOS ONE
2030:44755441
1987:24916124
1835:27162705
1767:10007967
1759:29185518
1694:83493714
1369:Loricata
734:Triassic
624:proximal
210:Reptilia
200:Chordata
196:Phylum:
190:Animalia
176:Domain:
134:aetosaur
35:Suchians
3311:Vivaron
2997:Effigia
2709:Maehary
2582:Dyoplax
2246:Class:
2164:in the
2134:Bibcode
2102:3194824
2079:Bibcode
2010:Bibcode
1978:4061117
1947:Bibcode
1900:Bibcode
1861:Bibcode
1826:4860341
1721:Bibcode
1660:: 189.
620:humerus
206:Class:
18:Suchian
4362:Suchia
4332:Suchia
2491:Suchia
2162:Suchia
2109:
2099:
2028:
1985:
1975:
1967:
1918:
1833:
1823:
1815:
1765:
1757:
1749:
1692:
1684:
1024:Suchia
837:et al.
711:, and
664:being
533:radius
478:fibula
386:family
378:, and
342:Suchia
249:Suchia
4376:66478
2253:Clade
2026:S2CID
1916:S2CID
1795:PeerJ
1763:S2CID
1690:S2CID
731:basal
661:order
628:ilium
470:jugal
346:clade
344:is a
243:Clade
230:Clade
217:Clade
132:, an
4199:see
4170:see
3676:see
2672:see
2313:see
2286:see
2107:PMID
1983:PMID
1965:ISSN
1831:PMID
1813:ISSN
1755:PMID
1747:ISSN
1682:ISSN
618:, a
587:pdpm
555:and
512:and
498:and
53:PreꞒ
2142:doi
2097:PMC
2087:doi
2018:doi
1973:PMC
1955:doi
1908:doi
1869:doi
1821:PMC
1803:doi
1737:hdl
1729:doi
1672:hdl
1662:doi
1658:352
685:",
681:, "
348:of
4395::
4373::
4358::
4343::
2255::
2140:.
2130:12
2128:.
2105:.
2095:.
2085:.
2073:.
2069:.
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