Knowledge (XXG)

Waimanu

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551: 142: 118: 274: 1701: 1213: 288:, the order that includes modern penguins. However, although it was probably flightless like all modern penguins, with wings specialized for wing-propelled diving, its wing bones do not yet show the extreme specializations modern penguins have for an aquatic lifestyle. It may have resembled a flightless 330:
is a partial skeleton comprising almost complete right tibiotarsus, proximal half of right fibula, right tarsometatarsus, right pelvis, and synsacrum (with last thoracic vertebra attached to the synsacrum), four caudal vertebrae. It is held in the
270:(1 metre). It is one of the most important bird fossils for understanding the origin and evolution of birds because of the time period it comes from, and the position of penguins near the base of the bird family tree. 457:
Gerald Mayr; Vanesa L. De Pietri; Leigh Love; Al A. Mannering; R. Paul Scofield (2018). "A well-preserved new mid-Paleocene penguin (Aves, Sphenisciformes) from the Waipara Greensand in New Zealand".
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or extremely rapidly immediately after. DNA studies, combined with the fossil evidence, seem to indicate the latter.
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Hackett, S. J.; et al. (2008). "A Phylogenomic Study of Birds Reveals Their Evolutionary History".
1031: 503: 1212: 556: 1557: 1100: 704: 1319: 1172: 1107: 1001: 857: 764: 655: 527: 474: 136: 1811: 1640: 1588: 1411: 1384: 1374: 1305: 824: 804: 754: 1435: 1328: 1254: 1010: 1793: 1785: 1401: 1342: 1275: 1230: 1086: 947: 519: 494: 439: 348: 308: 1798: 1017: 967: 511: 466: 429: 1452: 1062: 927: 285: 267: 125: 564: 507: 117: 1671: 1654: 1531: 1289: 1072: 1047: 1828: 1758: 1121: 987: 867: 68: 478: 1630: 1511: 1486: 1442: 1351: 1298: 1139: 1093: 1079: 937: 835: 794: 531: 221: 470: 1752: 1685: 1647: 1623: 1609: 1571: 1500: 1463: 1418: 1362: 1264: 1148: 1128: 1114: 1040: 847: 784: 744: 694: 372: 320: 43: 1661: 1616: 1540: 1520: 1425: 1193: 1186: 1179: 977: 957: 774: 649: 546: 418:"Early Penguin Fossils, plus Mitochondrial Genomes, Calibrate Avian Evolution" 88: 53: 1743: 1602: 1493: 1162: 887: 618: 515: 434: 417: 393: 293: 259: 153: 93: 37: 30: 523: 443: 388:
provided evidence for debate about whether the radiation of modern birds,
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It was about the size of an 236: 213: 362:, named for Al Mannering who found and collected the 1667: 1636: 1584: 1553: 1448: 1431: 1407: 1397: 1380: 1370: 1339: 1316: 1286: 1271: 1260: 1250: 1168: 1058: 1028: 998: 983: 973: 963: 953: 943: 933: 923: 913: 903: 893: 883: 873: 863: 853: 843: 820: 810: 800: 790: 780: 770: 760: 750: 740: 730: 720: 710: 700: 690: 392:, took place before the extinction of the non-avian 1727: 1530: 1510: 1462: 1361: 1338: 1315: 1285: 1241: 1138: 1057: 1027: 997: 834: 681: 594: 411: 409: 8: 1715: 1238: 1225: 994: 678: 665: 601: 587: 579: 116: 20: 569:specimens and a phylogram of early birds. 433: 292:or diver in body shape, and possibly the 405: 574:Skull of world's oldest penguin found 264:Cretaceous–Paleogene extinction event 7: 459:Journal of Vertebrate Paleontology 416:Slack, K. E.; et al. (2006). 358:for "bird". One species is known, 14: 1699: 1211: 549: 140: 422:Molecular Biology and Evolution 284:was a very early member of the 335:in Christchurch, New Zealand. 16:Extinct genus of birds waimanu 1: 1860:Fossil taxa described in 2006 471:10.1080/02724634.2017.1398169 296:in its manner of locomotion. 1855:Extinct birds of New Zealand 366:specimen. A second species, 307:was discovered in the Basal 1876: 1697: 1237: 1224: 1209: 677: 664: 626: 227: 220: 137:Scientific classification 135: 124: 115: 23: 621:and their extinct allies 1835:Prehistoric bird genera 516:10.1126/science.1157704 277:Artist's reconstruction 258:which lived during the 278: 1807:Paleobiology Database 435:10.1093/molbev/msj124 276: 1002:Palaeospheniscinae 557:Paleontology portal 508:2008Sci...320.1763H 502:(5884): 1763–1768. 656:Austrodyptornithes 279: 231:Waimanu manneringi 1822: 1821: 1794:Open Tree of Life 1721:Taxon identifiers 1712: 1711: 1708: 1707: 1695: 1694: 1343:Tytthostonychidae 1231:Procellariiformes 1220: 1219: 1207: 1206: 1203: 1202: 1087:Archaeospheniscus 948:Palaeoapterodytes 384:The discovery of 333:Canterbury Museum 309:Waipara Greensand 262:, soon after the 243: 242: 237:Slack et al. 2006 216: 214:Slack et al. 2006 1867: 1850:Extinct penguins 1815: 1814: 1802: 1801: 1789: 1788: 1776: 1775: 1763: 1762: 1761: 1748: 1747: 1746: 1716: 1703: 1669: 1638: 1586: 1555: 1450: 1433: 1409: 1399: 1382: 1372: 1341: 1318: 1288: 1273: 1262: 1252: 1239: 1226: 1215: 1170: 1060: 1032:Paraptenodytinae 1030: 1018:Palaeospheniscus 1000: 995: 985: 975: 968:Pseudaptenodytes 965: 955: 945: 935: 925: 915: 905: 895: 885: 875: 865: 855: 845: 822: 812: 802: 792: 782: 772: 762: 752: 742: 732: 722: 712: 702: 692: 679: 666: 603: 596: 589: 580: 559: 554: 553: 552: 536: 535: 489: 483: 482: 454: 448: 447: 437: 428:(6): 1144–1155. 413: 354:for "water" and 212: 205: 145: 144: 120: 110: 40: 29:Temporal range: 21: 1875: 1874: 1870: 1869: 1868: 1866: 1865: 1864: 1845:Paleocene birds 1840:Sphenisciformes 1825: 1824: 1823: 1818: 1810: 1805: 1797: 1792: 1784: 1779: 1771: 1766: 1757: 1756: 1751: 1742: 1741: 1736: 1723: 1713: 1704: 1691: 1526: 1506: 1458: 1357: 1334: 1311: 1281: 1233: 1216: 1199: 1134: 1063:Palaeeudyptinae 1053: 1023: 993: 928:Marambiornopsis 830: 673: 671:Sphenisciformes 660: 622: 607: 555: 550: 548: 545: 540: 539: 491: 490: 486: 465:(6): e1398169. 456: 455: 451: 415: 414: 407: 402: 382: 370:, was moved to 368:Waimanu tuatahi 347:comes from the 343:The genus name 341: 302: 286:Sphenisciformes 268:emperor penguin 239: 234: 211: 203: 194:Sphenisciformes 139: 126:Tarsometatarsus 111: 109: 108: 107: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 35: 34: 27: 17: 12: 11: 5: 1873: 1871: 1863: 1862: 1857: 1852: 1847: 1842: 1837: 1827: 1826: 1820: 1819: 1817: 1816: 1803: 1790: 1777: 1764: 1749: 1733: 1731: 1725: 1724: 1719: 1710: 1709: 1706: 1705: 1698: 1696: 1693: 1692: 1690: 1689: 1682: 1675: 1672:Pterodromoides 1665: 1658: 1655:Pseudobulweria 1651: 1644: 1634: 1627: 1620: 1613: 1606: 1599: 1592: 1582: 1575: 1568: 1561: 1551: 1544: 1536: 1534: 1532:Procellariidae 1528: 1527: 1525: 1524: 1516: 1514: 1508: 1507: 1505: 1504: 1497: 1490: 1483: 1476: 1468: 1466: 1460: 1459: 1457: 1456: 1446: 1439: 1429: 1422: 1415: 1405: 1395: 1388: 1378: 1367: 1365: 1359: 1358: 1356: 1355: 1347: 1345: 1336: 1335: 1333: 1332: 1324: 1322: 1313: 1312: 1310: 1309: 1302: 1294: 1292: 1290:Diomedeoididae 1283: 1282: 1280: 1279: 1269: 1258: 1247: 1245: 1243:incertae sedis 1235: 1234: 1229: 1222: 1221: 1218: 1217: 1210: 1208: 1205: 1204: 1201: 1200: 1198: 1197: 1190: 1183: 1176: 1166: 1159: 1152: 1144: 1142: 1136: 1135: 1133: 1132: 1125: 1118: 1111: 1104: 1097: 1090: 1083: 1076: 1073:Anthropodyptes 1068: 1066: 1055: 1054: 1052: 1051: 1048:Paraptenodytes 1044: 1036: 1034: 1025: 1024: 1022: 1021: 1014: 1006: 1004: 992: 991: 981: 971: 961: 951: 941: 931: 921: 911: 901: 891: 881: 871: 861: 851: 840: 838: 832: 831: 829: 828: 818: 808: 798: 788: 778: 768: 758: 748: 738: 728: 718: 708: 698: 687: 685: 683:incertae sedis 675: 674: 669: 662: 661: 659: 658: 652: 646: 640: 634: 627: 624: 623: 608: 606: 605: 598: 591: 583: 577: 576: 571: 561: 560: 544: 543:External links 541: 538: 537: 484: 449: 404: 403: 401: 398: 381: 378: 340: 337: 301: 298: 241: 240: 235: 225: 224: 218: 217: 201: 197: 196: 191: 187: 186: 181: 177: 176: 171: 167: 166: 161: 157: 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909: 902: 900: 899: 892: 890: 889: 882: 880: 879: 872: 870: 869: 868:Duntroonornis 862: 860: 859: 852: 850: 849: 842: 841: 839: 837: 833: 827: 826: 819: 817: 816: 809: 807: 806: 799: 797: 796: 789: 787: 786: 779: 777: 776: 769: 767: 766: 759: 757: 756: 749: 747: 746: 739: 737: 736: 729: 727: 726: 719: 717: 716: 709: 707: 706: 699: 697: 696: 689: 688: 686: 684: 680: 676: 672: 667: 663: 657: 653: 651: 647: 645: 641: 639: 635: 633: 629: 628: 625: 620: 616: 612: 604: 599: 597: 592: 590: 585: 584: 581: 575: 572: 570: 568: 563: 562: 558: 547: 542: 533: 529: 525: 521: 517: 513: 509: 505: 501: 497: 496: 488: 485: 480: 476: 472: 468: 464: 460: 453: 450: 445: 441: 436: 431: 427: 423: 419: 412: 410: 406: 399: 397: 395: 391: 387: 379: 377: 375: 374: 369: 365: 361: 360:W. manneringi 357: 353: 350: 346: 338: 336: 334: 329: 324: 322: 318: 314: 313:Waipara River 310: 306: 299: 297: 295: 291: 287: 283: 275: 271: 269: 265: 261: 257: 253: 249: 248: 238: 233: 232: 226: 223: 219: 215: 210: 209: 202: 199: 198: 195: 192: 189: 188: 185: 182: 179: 178: 175: 172: 169: 168: 165: 162: 159: 158: 155: 152: 149: 148: 143: 138: 134: 131: 127: 123: 119: 114: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 39: 32: 26: 22: 19: 1728: 1684: 1677: 1670: 1660: 1653: 1646: 1639: 1631:Pelecanoides 1629: 1622: 1615: 1608: 1601: 1594: 1587: 1577: 1570: 1563: 1558:Argyrodyptes 1556: 1546: 1539: 1519: 1512:Hydrobatidae 1499: 1492: 1487:Nesofregetta 1485: 1478: 1471: 1451: 1443:Thalassarche 1441: 1434: 1424: 1417: 1410: 1400: 1390: 1383: 1373: 1352:Tytthostonyx 1350: 1327: 1304: 1299:Diomedeoides 1297: 1274: 1263: 1253: 1242: 1192: 1185: 1178: 1171: 1161: 1154: 1147: 1140:Spheniscinae 1127: 1120: 1113: 1106: 1101:Marambiornis 1099: 1094:Delphinornis 1092: 1085: 1080:Anthropornis 1078: 1071: 1046: 1039: 1016: 1009: 986: 976: 966: 956: 946: 938:Marplesornis 936: 926: 916: 906: 896: 886: 876: 866: 856: 846: 836:Spheniscidae 823: 814: 813: 803: 795:Sequiwaimanu 793: 783: 773: 763: 753: 743: 733: 723: 713: 705:Aprosdokitos 703: 693: 682: 648:Infraclass: 566: 499: 493: 487: 462: 458: 452: 425: 421: 385: 383: 380:Significance 371: 367: 359: 355: 351: 344: 342: 325: 304: 303: 281: 280: 246: 245: 244: 230: 229: 222:Type species 207: 206: 24: 18: 1753:Wikispecies 1686:Thalassoica 1648:Procellaria 1610:Macronectes 1572:Calonectris 1501:Pelagodroma 1464:Oceanitidae 1419:Phoebastria 1363:Diomedeidae 1320:Marinavidae 1265:Lonchodytes 1173:Madrynornis 1149:Aptenodytes 1129:Platydyptes 1115:Pachydyptes 1108:Mesetaornis 1041:Arthrodytes 848:Crossvallia 785:Petradyptes 765:Orthopteryx 745:Muriwaimanu 695:Arthrodytes 619:albatrosses 373:Muriwaimanu 323:, in 1980. 321:New Zealand 36:62–60  1829:Categories 1662:Pterodroma 1641:Primodroma 1617:Pachyptila 1589:Eopuffinus 1541:Aphrodroma 1521:Hydrobates 1426:Phoebetria 1412:Notoleptos 1385:Diomedavus 1375:Aldiomedes 1306:Rupelornis 1194:Spheniscus 1187:Pygoscelis 1180:Megadyptes 978:Tasidyptes 958:Perudyptes 825:Wimanornis 805:Tonniornis 775:Pakudyptes 755:Nucleornis 650:Neognathae 609:Genera of 565:Images of 400:References 317:Canterbury 1624:Pagodroma 1603:Halobaena 1494:Oceanites 1436:Plotornis 1329:Marinavis 1255:Kievornis 1163:Eudyptula 1011:Eretiscus 888:Icadyptes 630:Kingdom: 394:dinosaurs 376:in 2018. 339:Etymology 311:near the 300:Discovery 294:great auk 260:Paleocene 254:of early 160:Kingdom: 154:Eukaryota 31:Paleocene 1738:Wikidata 1679:Puffinus 1596:Fulmarus 1565:Bulweria 1480:Garrodia 1473:Fregetta 1402:Murunkus 1392:Diomedea 1276:Makahala 1156:Eudyptes 908:Inkayacu 735:Kupoupou 725:Kumimanu 638:Chordata 636:Phylum: 632:Animalia 611:penguins 524:18583609 479:89744522 444:16533822 364:holotype 328:holotype 174:Chordata 170:Phylum: 164:Animalia 150:Domain: 130:holotype 1799:3600111 1786:1025600 1773:4844859 1759:Waimanu 1744:Q138638 1729:Waimanu 1579:Daption 1548:Ardenna 918:Kairuku 815:Waimanu 654:Clade: 642:Class: 615:petrels 567:Waimanu 532:6472805 504:Bibcode 495:Science 390:Neoaves 386:Waimanu 345:Waimanu 305:Waimanu 282:Waimanu 256:penguin 247:Waimanu 208:Waimanu 200:Genus: 190:Order: 180:Class: 128:of the 105:↓ 25:Waimanu 1812:139506 898:Inguza 878:Korora 715:Kaiika 617:, and 530:  522:  477:  442:  1781:IRMNG 1453:Tydea 528:S2CID 475:S2CID 349:Māori 315:, in 252:genus 250:is a 1768:GBIF 858:Dege 644:Aves 520:PMID 440:PMID 356:manu 326:The 290:loon 184:Aves 44:PreꞒ 512:doi 500:320 467:doi 430:doi 352:wai 1831:: 1809:: 1796:: 1783:: 1770:: 1755:: 1740:: 613:, 526:. 518:. 510:. 498:. 473:. 463:37 461:. 438:. 426:23 424:. 420:. 408:^ 319:, 94:Pg 38:Ma 33:, 1668:† 1637:† 1585:† 1554:† 1449:† 1432:† 1408:† 1398:† 1381:† 1371:† 1340:† 1317:† 1287:† 1272:† 1268:? 1261:† 1251:† 1169:† 1065:" 1061:" 1059:† 1029:† 999:† 984:† 974:† 964:† 954:† 944:† 934:† 924:† 914:† 904:† 894:† 884:† 874:† 864:† 854:† 844:† 821:† 811:† 801:† 791:† 781:† 771:† 761:† 751:† 741:† 731:† 721:† 711:† 701:† 691:† 602:e 595:t 588:v 534:. 514:: 506:: 481:. 469:: 446:. 432:: 228:† 204:† 99:N 89:K 84:J 79:T 74:P 69:C 64:D 59:S 54:O 49:Ꞓ

Index

Paleocene
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Tarsometatarsus
holotype
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Aves
Sphenisciformes
Waimanu
Slack et al. 2006
Type species
Slack et al. 2006
genus
penguin

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